treatments-xml/data/03/C5/E0/03C5E0447145FFCAFED4E61AFEA8D95E.xml
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<document id="E7C037044C1EBDE68E0724A8000D1677" ID-CLB-Dataset="6338" ID-DOI="10.5281/zenodo.4689932" ID-GBIF-Dataset="34c0b6e4-12a3-4470-8aa2-e632c4e8854f" ID-ISSN="1638-9387" ID-Zenodo-Dep="4689932" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618438023714" checkinUser="felipe" docAuthor="Salazar-Vallejo, Sergio I., Bailey-Brock, Julie H. &amp; Dreyer, Jennifer C." docDate="2007" docId="03C5E0447145FFCAFED4E61AFEA8D95E" docLanguage="en" docName="Zoosystema.29.3.535-553.pdf" docOrigin="Zoosystema 29 (3)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Pseudexogone Augener 1922" docType="treatment" docVersion="5" lastPageNumber="539" masterDocId="FFFC983C7146FFCEFE6AE376FF92DD74" masterDocTitle="Revision of Pseudexogone Augener, 1922 (Annelida, Polychaeta, Syllidae), and its transfer to Pilargidae" masterLastPageNumber="553" masterPageNumber="535" pageNumber="538" updateTime="1698936629142" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5DD5A7008CC0B80D1033277A9C1F18DB">Revision of Pseudexogone Augener, 1922 (Annelida, Polychaeta, Syllidae), and its transfer to Pilargidae</mods:title>
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<mods:namePart id="9806C4C391E90FB547ED54B2E0CCC614">Salazar-Vallejo, Sergio I.</mods:namePart>
<mods:affiliation id="B862D332E0519A2283CD3D0EDCBCAE71">El Colegio de la Frontera Sur, Departamento Ecología Acuática, Apartado Postal 424, Chetumal, Quintana Roo, 77000 (Mexico) salazar @ ecosur-qroo. mx</mods:affiliation>
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<mods:namePart id="4FD627B22ADEFE31475AB0CCB1E9D35F">Bailey-Brock, Julie H.</mods:namePart>
<mods:affiliation id="D804C920EF69B6DD6D35C73FC7E5794A">University of Hawaii at Manoa, Department of Zoology, Honolulu, HI 96822 (USA) jbrock @ hawaii. edu</mods:affiliation>
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<mods:namePart id="0556E8C84D78CCF7EFE698BEF412F598">Dreyer, Jennifer C.</mods:namePart>
<mods:affiliation id="4966040BE706754F0B54472318780360">University of Hawaii at Manoa, Department of Zoology, Honolulu, HI 96822 (USA) Present address: Virginia Institute of Marine Science, Gloucester Point, VA 23062 (USA) jcdrey @ vims. edu</mods:affiliation>
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<mods:title id="94F48A8AA088D9B5866F6605AC1314E9">Zoosystema</mods:title>
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<mods:date id="9A11096527F6A9523D958A46C1816BA4">2007</mods:date>
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<treatment id="03C5E0447145FFCAFED4E61AFEA8D95E" ID-DOI="http://doi.org/10.5281/zenodo.4893367" ID-GBIF-Taxon="182484811" ID-Zenodo-Dep="4893367" LSID="urn:lsid:plazi:treatment:03C5E0447145FFCAFED4E61AFEA8D95E" httpUri="http://treatment.plazi.org/id/03C5E0447145FFCAFED4E61AFEA8D95E" lastPageId="4" lastPageNumber="539" pageId="3" pageNumber="538">
<subSubSection id="C37602D97145FFCDFED4E61AFDA8D8F2" box="[190,570,1388,1414]" pageId="3" pageNumber="538" type="nomenclature">
<paragraph id="8BD351527145FFCDFED4E61AFDA8D8F2" blockId="3.[190,570,1388,1414]" box="[190,570,1388,1414]" pageId="3" pageNumber="538">
<heading id="D09BE63E7145FFCDFED4E61AFDA8D8F2" box="[190,570,1388,1414]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="538" reason="2">
Genus
<taxonomicName id="4C6C2AD17145FFCDFF61E61AFDA8D8F2" ID-CoL="84SVY" authority="Augener, 1922" authorityName="Augener" authorityYear="1922" box="[267,570,1388,1414]" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="genus">
<emphasis id="B9188D407145FFCDFF61E61AFE06D8F2" box="[267,404,1388,1414]" italics="true" pageId="3" pageNumber="538">Pseudexogone</emphasis>
<bibRefCitation id="EFFD2CA37145FFCDFFF1E61AFDA8D8F2" author="AUGENER H." box="[411,570,1388,1414]" pageId="3" pageNumber="538" pagination="161 - 218" refId="ref9805" refString="AUGENER H. 1922. - Litorale Polychaeten von Juan Fernandez, in SKOTTSBERG C. (ed.), The Natural History of Juan Fernandez and Easter Island, volume 3, Zoology, Part 2. Almqvist &amp; Wiksells, Uppsala: 161 - 218." type="book chapter" year="1922">Augener, 1922</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37602D97145FFCDFE1BE6D6FD15D8CC" box="[113,647,1440,1465]" pageId="3" pageNumber="538" type="reference_group">
<paragraph id="8BD351527145FFCDFE1BE6D6FD15D8CC" blockId="3.[113,647,1440,1699]" box="[113,647,1440,1465]" pageId="3" pageNumber="538">
<treatmentCitationGroup id="AB7C767C7145FFCDFE1BE6D6FD15D8CC" box="[113,647,1440,1465]" pageId="3" pageNumber="538">
<treatmentCitation id="0ACD77437145FFCDFE1BE6D6FE3BD8CC" author="AUGENER H." box="[113,425,1440,1465]" page="191" pageId="3" pageNumber="538" year="1922">
<taxonomicName id="4C6C2AD17145FFCDFE1BE6D6FE3BD8CC" ID-CoL="84SVY" authority="Augener, 1922: 191" authorityName="Augener" authorityPageNumber="191" authorityYear="1922" box="[113,425,1440,1465]" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="genus">
<emphasis id="B9188D407145FFCDFE1BE6D6FF75D8CC" box="[113,231,1440,1464]" italics="true" pageId="3" pageNumber="538">Pseudexogone</emphasis>
<bibRefCitation id="EFFD2CA37145FFCDFE84E6D7FE3BD8CC" author="AUGENER H." box="[238,425,1440,1465]" pageId="3" pageNumber="538" pagination="161 - 218" refId="ref9805" refString="AUGENER H. 1922. - Litorale Polychaeten von Juan Fernandez, in SKOTTSBERG C. (ed.), The Natural History of Juan Fernandez and Easter Island, volume 3, Zoology, Part 2. Almqvist &amp; Wiksells, Uppsala: 161 - 218." type="book chapter" year="1922">Augener, 1922: 191</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. —
<treatmentCitation id="0ACD77437145FFCDFFBBE6D6FD11D8CC" author="FAUCHALD K." box="[465,643,1440,1464]" page="84" pageId="3" pageNumber="538" year="1977">
<bibRefCitation id="EFFD2CA37145FFCDFFBBE6D6FD11D8CC" author="FAUCHALD K." box="[465,643,1440,1464]" pageId="3" pageNumber="538" pagination="1 - 188" refId="ref10002" refString="FAUCHALD K. 1977. - The polychaete worms: definitions and keys to the orders, families and genera. Natural History Museum of Los Angeles County, Science Series 28: 1 - 188." type="book chapter" year="1977">Fauchald 1977: 84</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37602D97145FFCDFE1BE6BAFEB8DB37" pageId="3" pageNumber="538" type="materials_examined">
<paragraph id="8BD351527145FFCDFE1BE6BAFECAD88A" blockId="3.[113,647,1440,1699]" pageId="3" pageNumber="538">
<typeStatus id="54D7EFF07145FFCDFE1BE6BAFF33D895" box="[113,161,1484,1507]" pageId="3" pageNumber="538">TYPE</typeStatus>
SPECIES. —
<taxonomicName id="4C6C2AD17145FFCDFF7EE6BDFD11D897" authority="Augener, 1922" authorityName="Augener" authorityYear="1922" box="[276,643,1483,1507]" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="species" species="backstromi">
<emphasis id="B9188D407145FFCDFF7EE6BDFE60D897" box="[276,498,1483,1507]" italics="true" pageId="3" pageNumber="538">Pseudexogone backstromi</emphasis>
<bibRefCitation id="EFFD2CA37145FFCDFF9DE6BDFD11D897" author="AUGENER H." box="[503,643,1483,1507]" pageId="3" pageNumber="538" pagination="161 - 218" refId="ref9805" refString="AUGENER H. 1922. - Litorale Polychaeten von Juan Fernandez, in SKOTTSBERG C. (ed.), The Natural History of Juan Fernandez and Easter Island, volume 3, Zoology, Part 2. Almqvist &amp; Wiksells, Uppsala: 161 - 218." type="book chapter" year="1922">Augener, 1922</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
<paragraph id="8BD351527145FFCDFE1BE567FEB8DB37" blockId="3.[113,647,1440,1699]" pageId="3" pageNumber="538">POTENTIAL APOMORPHY BASED DEFINITION. — Bidentate curved notospines.</paragraph>
</subSubSection>
<subSubSection id="C37602D97145FFCDFE1BE52EFD15DBD7" pageId="3" pageNumber="538" type="description">
<paragraph id="8BD351527145FFCDFE1BE52EFD15DBD7" blockId="3.[113,647,1440,1699]" pageId="3" pageNumber="538">
SPECIES INCLUDED. —
<taxonomicName id="4C6C2AD17145FFCDFF26E520FF32DBFD" authority="Augener, 1922" authorityName="Augener" authorityYear="1922" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="species" species="backstromi">
<emphasis id="B9188D407145FFCDFF26E520FDBEDB1A" box="[332,556,1622,1646]" italics="true" pageId="3" pageNumber="538">Pseudexogone backstromi</emphasis>
<bibRefCitation id="EFFD2CA37145FFCDFC59E520FF32DBFD" author="AUGENER H." pageId="3" pageNumber="538" pagination="161 - 218" refId="ref9805" refString="AUGENER H. 1922. - Litorale Polychaeten von Juan Fernandez, in SKOTTSBERG C. (ed.), The Natural History of Juan Fernandez and Easter Island, volume 3, Zoology, Part 2. Almqvist &amp; Wiksells, Uppsala: 161 - 218." type="book chapter" year="1922">Augener, 1922</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6C2AD17145FFCDFEC2E507FDBCDBFD" authority="(Katzmann, Laubier &amp; Ramos, 1974)" authorityName="Salazar-Vallejo &amp; Bailey-Brock &amp; Dreyer" authorityYear="2007" baseAuthorityName="Katzmann, Laubier &amp; Ramos" baseAuthorityYear="1974" box="[168,558,1649,1673]" class="Polychaeta" family="Pilargidae" genus="Synelmis" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="species" species="dineti" status="comb. nov.">
<emphasis id="B9188D407145FFCDFEC2E507FF7BDBFD" box="[168,233,1649,1673]" italics="true" pageId="3" pageNumber="538">P.dineti</emphasis>
(
<bibRefCitation id="EFFD2CA37145FFCDFE99E507FDBADBFD" author="KATZMANN W. &amp; LAUBIER L. &amp; RAMOS J." box="[243,552,1649,1673]" pageId="3" pageNumber="538" pagination="1 - 40" refId="ref10108" refString="KATZMANN W., LAUBIER L. &amp; RAMOS J. 1974. - Pilargidae (annelides polychetes errantes) de Mediterranee. Bulletin de l'Institut oceanographique, Monaco 71: 1 - 40." type="journal article" year="1974">Katzmann, Laubier &amp; Ramos, 1974</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22B303B7145FFCDFC59E507FD16DBFD" box="[563,644,1649,1673]" pageId="3" pageNumber="538" rank="species">n. comb.</taxonomicNameLabel>
,
<taxonomicName id="4C6C2AD17145FFCDFE1BE5FAFF58DBD7" authority="Salazar-Vallejo &amp; Bailey-Brock &amp; Dreyer, 2007" authorityName="Salazar-Vallejo &amp; Bailey-Brock &amp; Dreyer" authorityYear="2007" box="[113,202,1675,1699]" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="species" species="helmuti" status="sp. nov.">
<emphasis id="B9188D407145FFCDFE1BE5FAFF58DBD7" box="[113,202,1675,1699]" italics="true" pageId="3" pageNumber="538">P. helmuti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B303B7145FFCDFEBAE5FAFE97DBD7" box="[208,261,1676,1699]" pageId="3" pageNumber="538" rank="species">n. sp.</taxonomicNameLabel>
,
<taxonomicName id="4C6C2AD17145FFCDFF64E5FAFEE7DBD7" authority="Salazar-Vallejo &amp; Bailey-Brock &amp; Dreyer, 2007" authorityName="Salazar-Vallejo &amp; Bailey-Brock &amp; Dreyer" authorityYear="2007" box="[270,373,1676,1699]" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="species" species="imajimai" status="sp. nov.">
<emphasis id="B9188D407145FFCDFF64E5FAFEE7DBD7" box="[270,373,1676,1699]" italics="true" pageId="3" pageNumber="538">P. imajimai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B303B7145FFCDFF11E5FAFE3CDBD7" box="[379,430,1676,1699]" pageId="3" pageNumber="538" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="4C6C2AD17145FFCDFFB1E5FAFDDCDBD7" authority="Salazar-Vallejo &amp; Bailey-Brock &amp; Dreyer, 2007" authorityName="Salazar-Vallejo &amp; Bailey-Brock &amp; Dreyer" authorityYear="2007" box="[475,590,1675,1699]" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="species" species="williamsae" status="sp. nov.">
<emphasis id="B9188D407145FFCDFFB1E5FAFDDCDBD7" box="[475,590,1675,1699]" italics="true" pageId="3" pageNumber="538">P. williamsae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B303B7145FFCDFC3EE5FAFD15DBD7" box="[596,647,1676,1699]" pageId="3" pageNumber="538" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C37602D97145FFCDFCCFE3AEFD66DCD7" pageId="3" pageNumber="538" type="distribution">
<paragraph id="8BD351527145FFCDFCCFE3AEFC63DD9B" blockId="3.[677,1009,215,239]" box="[677,1009,215,239]" pageId="3" pageNumber="538">DISTRIBUTION. — Cosmopolitan.</paragraph>
<paragraph id="8BD351527145FFCDFCCFE271FD66DCD7" blockId="3.[677,1210,263,419]" pageId="3" pageNumber="538">
EMENDED DIAGNOSIS. —
<taxonomicName id="4C6C2AD17145FFCDFDF0E271FB9BDC6B" authorityName="Salazar-Vallejo" authorityYear="1987" box="[922,1033,263,287]" class="Polychaeta" family="Pilargidae" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="subFamily" subFamily="Synelminae">Synelminae</taxonomicName>
with palps simple, each with a ventrolateral papilla. Three antennae. Eyes present or absent. Two pairs of tentacular cirri. Parapodia with bidentate curved notospines, neurochaetae include pectinate and denticulate capillaries and furcate chaetae.
</paragraph>
</subSubSection>
<subSubSection id="C37602D97145FFCAFCCFE2A0FEA8D95E" lastPageId="4" lastPageNumber="539" pageId="3" pageNumber="538" type="discussion">
<paragraph id="8BD351527145FFCDFCCFE2A0FC9BDC9A" blockId="3.[677,1210,470,1700]" box="[677,777,470,496]" pageId="3" pageNumber="538">REMARKS</paragraph>
<paragraph id="8BD351527145FFCDFCCFE283FCF3D8F2" blockId="3.[677,1210,470,1700]" pageId="3" pageNumber="538">
Although the anterior end of
<taxonomicName id="4C6C2AD17145FFCDFD89E280FBF9DF7B" authorityName="Augener" authorityYear="1922" box="[995,1131,501,527]" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="species" species="backstromi">
<emphasis id="B9188D407145FFCDFD89E280FBF9DF7B" box="[995,1131,501,527]" italics="true" pageId="3" pageNumber="538">P. backstromi</emphasis>
</taxonomicName>
resembles species of
<taxonomicName id="4C6C2AD17145FFCDFD22E163FBAEDF5B" authority="Orsted, 1845" authorityName="Orsted" authorityYear="1845" box="[840,1084,531,559]" class="Polychaeta" family="Syllidae" genus="Exogone" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="genus">
<emphasis id="B9188D407145FFCDFD22E163FC0FDF5A" box="[840,925,533,558]" italics="true" pageId="3" pageNumber="538">Exogone</emphasis>
Ørsted, 1845
</taxonomicName>
, its general appearance is very similar to some pilargids that have been included in
<taxonomicName id="4C6C2AD17145FFCDFDF9E122FB2BDF1A" authority="Chamberlin, 1919" authorityName="Chamberlin" authorityYear="1919" box="[915,1209,596,622]" class="Polychaeta" family="Pilargidae" genus="Synelmis" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="genus">
<emphasis id="B9188D407145FFCDFDF9E122FC7EDF1A" box="[915,1004,596,622]" italics="true" pageId="3" pageNumber="538">Synelmis</emphasis>
Chamberlin, 1919
</taxonomicName>
(Salazar-Vallejo 2003); besides, the differences in the structure of the alimentary canal, especially the presence of a proventricle in syllids, are so important that Augener had some doubts about its placement among syllids.
<taxonomicName id="4C6C2AD17145FFCDFDC4E184FB96DE78" authorityName="Chamberlin" authorityYear="1919" box="[942,1028,754,780]" class="Polychaeta" family="Pilargidae" genus="Synelmis" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="genus">
<emphasis id="B9188D407145FFCDFDC4E184FB96DE78" box="[942,1028,754,780]" italics="true" pageId="3" pageNumber="538">Synelmis</emphasis>
</taxonomicName>
is usually defined by having biarticulate palps and straight notospines, but its perceived similarity to syllids is depicted by the generic placement that some species received in the past, e.g.,
<taxonomicName id="4C6C2AD17145FFCDFD0BE007FD4FDEDF" authority="Langerhans, 1881" authorityName="Langerhans" authorityYear="1881" class="Polychaeta" family="Pilargidae" genus="Ancistrosyllis" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="species" species="albini">
<emphasis id="B9188D407145FFCDFD0BE007FBBEDEFF" box="[865,1068,881,907]" italics="true" pageId="3" pageNumber="538">Ancistrosyllis albini</emphasis>
Langerhans, 1881
</taxonomicName>
and
<taxonomicName id="4C6C2AD17145FFCDFD7DE0E7FB99DEDF" authority="Fauvel, 1919" authorityName="Fauvel" authorityYear="1919" box="[791,1035,913,939]" class="Polychaeta" family="Pilargidae" genus="Ancistrosyllis" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="species" species="rigida">
<emphasis id="B9188D407145FFCDFD7DE0E7FCE7DEDF" box="[791,885,913,939]" italics="true" pageId="3" pageNumber="538">A. rigida</emphasis>
Fauvel, 1919
</taxonomicName>
.
<taxonomicName id="4C6C2AD17145FFCDFA76E0E7FBE4DEDF" authorityName="Chamberlin" authorityYear="1919" box="[1052,1142,913,939]" class="Polychaeta" family="Pilargidae" genus="Synelmis" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="genus">
<emphasis id="B9188D407145FFCDFA76E0E7FBE4DEDF" box="[1052,1142,913,939]" italics="true" pageId="3" pageNumber="538">Synelmis</emphasis>
</taxonomicName>
is the
<typeStatus id="54D7EFF07145FFCDFCCFE0C7FD41DEBF" box="[677,723,945,971]" pageId="3" pageNumber="538">type</typeStatus>
genus for
<taxonomicName id="4C6C2AD17145FFCDFD57E0C6FC29DEBE" authorityName="Salazar-Vallejo" authorityYear="1987" box="[829,955,944,970]" class="Polychaeta" family="Pilargidae" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="subFamily" subFamily="Synelminae">Synelminae</taxonomicName>
, which has been defined by having a smooth integument, body cylindrical, and straight or curved notospines (Salazar-Vallejo 2003).
<taxonomicName id="4C6C2AD17145FFCDFC98E779FCE9D95D" authorityName="Augener" authorityYear="1922" box="[754,891,1039,1065]" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="genus">
<emphasis id="B9188D407145FFCDFC98E779FCE9D95D" box="[754,891,1039,1065]" italics="true" pageId="3" pageNumber="538">Pseudexogone</emphasis>
</taxonomicName>
belongs to this subfamily and differs from
<taxonomicName id="4C6C2AD17145FFCDFD44E759FC1AD93D" authorityName="Chamberlin" authorityYear="1919" box="[814,904,1071,1097]" class="Polychaeta" family="Pilargidae" genus="Synelmis" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="genus">
<emphasis id="B9188D407145FFCDFD44E759FC1AD93D" box="[814,904,1071,1097]" italics="true" pageId="3" pageNumber="538">Synelmis</emphasis>
</taxonomicName>
especially by having simple palps, and bidentate sigmoid notospines. These two pilargid synelmin genera are apparently closely allied. Further,
<taxonomicName id="4C6C2AD17145FFCDFD6EE7F8FC1FD9DC" authorityName="Augener" authorityYear="1922" box="[772,909,1166,1192]" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="genus">
<emphasis id="B9188D407145FFCDFD6EE7F8FC1FD9DC" box="[772,909,1166,1192]" italics="true" pageId="3" pageNumber="538">Pseudexogone</emphasis>
</taxonomicName>
is closely allied to
<taxonomicName id="4C6C2AD17145FFCDFA37E7F9FCD2D9BC" authority="Pearson, 1970" authorityName="Pearson" authorityYear="1970" class="Polychaeta" family="Pilargidae" genus="Litocorsa" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="genus">
<emphasis id="B9188D407145FFCDFA37E7F9FB2BD9DC" box="[1117,1209,1167,1192]" italics="true" pageId="3" pageNumber="538">Litocorsa</emphasis>
Pearson, 1970
</taxonomicName>
; they share a thin transparent body and simple palps. However, they differ because
<taxonomicName id="4C6C2AD17145FFCDFCCFE798FC96D873" authorityName="Pearson" authorityYear="1970" box="[677,772,1262,1287]" class="Polychaeta" family="Pilargidae" genus="Litocorsa" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="genus">
<emphasis id="B9188D407145FFCDFCCFE798FC96D873" box="[677,772,1262,1287]" italics="true" pageId="3" pageNumber="538">Litocorsa</emphasis>
</taxonomicName>
has straight emergent notospines with tapering tips, instead of having notospines with bidentate tips. Further,
<taxonomicName id="4C6C2AD17145FFCDFDC9E65BFC6DD832" authorityName="Pearson" authorityYear="1970" box="[931,1023,1325,1350]" class="Polychaeta" family="Pilargidae" genus="Litocorsa" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="genus">
<emphasis id="B9188D407145FFCDFDC9E65BFC6DD832" box="[931,1023,1325,1350]" italics="true" pageId="3" pageNumber="538">Litocorsa</emphasis>
</taxonomicName>
has the potential autopomorphy of having neurospines (
<bibRefCitation id="EFFD2CA37145FFCDFA2AE63AFCC7D8F2" author="DARBYSHIRE T. &amp; MACKIE A. S. Y." pageId="3" pageNumber="538" pagination="63 - 73" refId="ref9883" refString="DARBYSHIRE T. &amp; MACKIE A. S. Y. 2003. - Species of Litocorsa (Polychaeta: Pilargidae) from the Indian Ocean and South China Sea, in SIGVALDADOTTIR E., MACKIE A. S. Y., HELGASON G. V., REISH D. J., SVA- VARSSON J., STEINGRIMSSON S. A. &amp; GUDMUNDSSON G. (eds), Advances in polychaete research, Proceedings of the 7 th International Polychaete Conference held in Reykjavik, Iceland, 2 - 6 July 2001. Developments in Hydrobiology 170, series editor H. J. Dumont, Hydrobiologia 496: 63 - 73." type="journal article" year="2003">Darbyshire &amp; Mackie 2003</bibRefCitation>
).
</paragraph>
<paragraph id="8BD351527145FFCAFCD1E6FAFF5EDF1A" blockId="3.[677,1210,470,1700]" lastBlockId="4.[113,647,216,1066]" lastPageId="4" lastPageNumber="539" pageId="3" pageNumber="538">
In the original diagnosis,
<bibRefCitation id="EFFD2CA37145FFCDFDBEE6FAFB2BD8D2" author="AUGENER H." box="[980,1209,1420,1446]" pageId="3" pageNumber="538" pagination="161 - 218" refId="ref9805" refString="AUGENER H. 1922. - Litorale Polychaeten von Juan Fernandez, in SKOTTSBERG C. (ed.), The Natural History of Juan Fernandez and Easter Island, volume 3, Zoology, Part 2. Almqvist &amp; Wiksells, Uppsala: 161 - 218." type="book chapter" year="1922">Augener (1922: 191)</bibRefCitation>
indicated that
<taxonomicName id="4C6C2AD17145FFCDFD2EE6DDFC5FD8B1" authorityName="Augener" authorityYear="1922" box="[836,973,1451,1477]" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="genus">
<emphasis id="B9188D407145FFCDFD2EE6DDFC5FD8B1" box="[836,973,1451,1477]" italics="true" pageId="3" pageNumber="538">Pseudexogone</emphasis>
</taxonomicName>
was syllid-like, had a nematoid body with simple palps, simple hooks in median and posterior chaetigers, and an unarmed pharynx. After the description of
<taxonomicName id="4C6C2AD17145FFCDFA61E57DFB01DB50" authorityName="Augener" authorityYear="1922" box="[1035,1171,1546,1572]" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="538" phylum="Annelida" rank="species" species="backstromi">
<emphasis id="B9188D407145FFCDFA61E57DFB01DB50" box="[1035,1171,1546,1572]" italics="true" pageId="3" pageNumber="538">P. backstromi</emphasis>
</taxonomicName>
, he tentatively placed the animal close to syllids (
<bibRefCitation id="EFFD2CA37145FFCDFAFAE55CFCC9DB10" author="AUGENER H." pageId="3" pageNumber="538" pagination="161 - 218" refId="ref9805" refString="AUGENER H. 1922. - Litorale Polychaeten von Juan Fernandez, in SKOTTSBERG C. (ed.), The Natural History of Juan Fernandez and Easter Island, volume 3, Zoology, Part 2. Almqvist &amp; Wiksells, Uppsala: 161 - 218." type="book chapter" year="1922">Augener 1922: 194</bibRefCitation>
).
<bibRefCitation id="EFFD2CA37145FFCDFD04E53CFBC3DB10" author="AUGENER H." box="[878,1105,1610,1636]" pageId="3" pageNumber="538" pagination="161 - 218" refId="ref9805" refString="AUGENER H. 1922. - Litorale Polychaeten von Juan Fernandez, in SKOTTSBERG C. (ed.), The Natural History of Juan Fernandez and Easter Island, volume 3, Zoology, Part 2. Almqvist &amp; Wiksells, Uppsala: 161 - 218." type="book chapter" year="1922">Augener (1922: 193)</bibRefCitation>
confused the position of the bidentate notospines, since he regarded them as ventral (neuropodial). This could be explained because of either the polychaete body was too transparent, or because he confused the typical morphology of the pygidium; i.e. anal cirri are generally ventral or ventrolateral, which is shown in his original drawings, but if the specimen was twisted, then there would be some confusion about which surface was dorsal or ventral in the posterior end. An important missing detail in the original description is the start of notospines; since this is a rather conservative useful feature (see below), this species could be separated by using additional materials or by employing other morphological features.
</paragraph>
<paragraph id="8BD351527142FFCAFEE2E102FEA8D95E" blockId="4.[113,647,216,1066]" pageId="4" pageNumber="539">
Further delay in recognizing this genus as a member of the
<taxonomicName id="4C6C2AD17142FFCAFED9E1E5FE8EDFD9" authorityName="de Saint-Joseph" authorityYear="1899" box="[179,284,659,685]" class="Polychaeta" family="Pilargidae" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="539" phylum="Annelida" rank="family">Pilargidae</taxonomicName>
resulted because
<taxonomicName id="4C6C2AD17142FFCAFFB8E1E5FDC8DFD9" authorityName="Augener" authorityYear="1922" box="[466,602,659,685]" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="539" phylum="Annelida" rank="genus">
<emphasis id="B9188D407142FFCAFFB8E1E5FDC8DFD9" box="[466,602,659,685]" italics="true" pageId="4" pageNumber="539">Pseudexogone</emphasis>
</taxonomicName>
was regarded as a syllid in two major reviews (
<bibRefCitation id="EFFD2CA37142FFCAFC48E1C5FF72DF99" author="HARTMAN O." pageId="4" pageNumber="539" pagination="1 - 353" refId="ref10039" refString="HARTMAN O. 1959. - Catalogue of the polychaetous annelids of the world, part 1. Allan Hancock Foundation Publications, Occasional Paper 23: 1 - 353." type="journal article" year="1959">Hartman 1959: 220</bibRefCitation>
;
<bibRefCitation id="EFFD2CA37142FFCAFE83E1A5FE22DF99" author="FAUCHALD K." box="[233,432,723,749]" pageId="4" pageNumber="539" pagination="1 - 188" refId="ref10002" refString="FAUCHALD K. 1977. - The polychaete worms: definitions and keys to the orders, families and genera. Natural History Museum of Los Angeles County, Science Series 28: 1 - 188." type="book chapter" year="1977">Fauchald 1977: 84</bibRefCitation>
). However, the same morphological features such as body pattern and structures (nematoid body, simple palps, bidentate emergent spines) were recognized in a pilargid by
<bibRefCitation id="EFFD2CA37142FFCAFEFEE024FE0ADE18" author="KATZMANN W. &amp; LAUBIER L. &amp; RAMOS J." box="[148,408,849,876]" pageId="4" pageNumber="539" pagination="1 - 40" refId="ref10108" refString="KATZMANN W., LAUBIER L. &amp; RAMOS J. 1974. - Pilargidae (annelides polychetes errantes) de Mediterranee. Bulletin de l'Institut oceanographique, Monaco 71: 1 - 40." type="journal article" year="1974">
Katzmann
<emphasis id="B9188D407142FFCAFF64E024FED1DE1F" box="[270,323,849,875]" italics="true" pageId="4" pageNumber="539">et al.</emphasis>
(1974)
</bibRefCitation>
, when they described
<taxonomicName id="4C6C2AD17142FFCAFE1BE007FE8BDEFF" authorityName="Salazar-Vallejo &amp; Bailey-Brock &amp; Dreyer" authorityYear="2007" baseAuthorityName="Katzmann, Laubier &amp; Ramos" baseAuthorityYear="1974" box="[113,281,881,907]" class="Polychaeta" family="Pilargidae" genus="Synelmis" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="539" phylum="Annelida" rank="species" species="dineti">
<emphasis id="B9188D407142FFCAFE1BE007FE8BDEFF" box="[113,281,881,907]" italics="true" pageId="4" pageNumber="539">Synelmis dineti</emphasis>
</taxonomicName>
(see below). As herein defined,
<taxonomicName id="4C6C2AD17142FFCAFE1BE0E7FE92DEDF" authorityName="Augener" authorityYear="1922" box="[113,256,913,939]" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="539" phylum="Annelida" rank="genus">
<emphasis id="B9188D407142FFCAFE1BE0E7FE92DEDF" box="[113,256,913,939]" italics="true" pageId="4" pageNumber="539">Pseudexogone</emphasis>
</taxonomicName>
includes species living in subtidal depths and rarely from deeper waters (
<emphasis id="B9188D407142FFCAFC7DE0C7FDB2DEBE" box="[535,544,945,970]" italics="true" pageId="4" pageNumber="539">c</emphasis>
.
<quantity id="4C94FCB77142FFCAFC5AE0C7FD14DEBF" box="[560,646,945,971]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="539" unit="m" value="1000.0">1000 m</quantity>
depth). Besides the
<typeStatus id="54D7EFF07142FFCAFF57E0A7FEF9DE9F" box="[317,363,977,1003]" pageId="4" pageNumber="539">type</typeStatus>
species, it includes
<taxonomicName id="4C6C2AD17142FFCAFC5EE0A7FD15DE9E" authorityName="Salazar-Vallejo &amp; Bailey-Brock &amp; Dreyer" authorityYear="2007" baseAuthorityName="Katzmann, Laubier &amp; Ramos" baseAuthorityYear="1974" box="[564,647,976,1002]" class="Polychaeta" family="Pilargidae" genus="Synelmis" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="539" phylum="Annelida" rank="species" species="dineti" status="comb. nov.">
<emphasis id="B9188D407142FFCAFC5EE0A7FD15DE9E" box="[564,647,976,1002]" italics="true" pageId="4" pageNumber="539">P. dineti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B303B7142FFCAFE1BE086FF47D97E" box="[113,213,1008,1034]" pageId="4" pageNumber="539" rank="species">n. comb.</taxonomicNameLabel>
,
<taxonomicName id="4C6C2AD17142FFCAFEB5E087FEDAD97E" authority="Salazar-Vallejo &amp; Bailey-Brock &amp; Dreyer, 2007" authorityName="Salazar-Vallejo &amp; Bailey-Brock &amp; Dreyer" authorityYear="2007" box="[223,328,1008,1034]" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="539" phylum="Annelida" rank="species" species="helmuti" status="sp. nov.">
<emphasis id="B9188D407142FFCAFEB5E087FEDAD97E" box="[223,328,1008,1034]" italics="true" pageId="4" pageNumber="539">P. helmuti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B303B7142FFCAFF24E086FE19D97E" box="[334,395,1008,1034]" pageId="4" pageNumber="539" rank="species">n. sp.</taxonomicNameLabel>
,
<taxonomicName id="4C6C2AD17142FFCAFFFCE087FD9DD97E" authority="Salazar-Vallejo &amp; Bailey-Brock &amp; Dreyer, 2007" authorityName="Salazar-Vallejo &amp; Bailey-Brock &amp; Dreyer" authorityYear="2007" box="[406,527,1009,1034]" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="539" phylum="Annelida" rank="species" species="imajimai" status="sp. nov.">
<emphasis id="B9188D407142FFCAFFFCE087FD9DD97E" box="[406,527,1009,1034]" italics="true" pageId="4" pageNumber="539">P. imajimai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B303B7142FFCAFC7CE086FDC0D97E" box="[534,594,1008,1034]" pageId="4" pageNumber="539" rank="species">n. sp.</taxonomicNameLabel>
, and
<taxonomicName id="4C6C2AD17142FFCAFE1BE766FF6AD95D" authority="Salazar-Vallejo &amp; Bailey-Brock &amp; Dreyer, 2007" authorityName="Salazar-Vallejo &amp; Bailey-Brock &amp; Dreyer" authorityYear="2007" box="[113,248,1039,1065]" class="Polychaeta" family="Pilargidae" genus="Pseudexogone" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="539" phylum="Annelida" rank="species" species="williamsae" status="sp. nov.">
<emphasis id="B9188D407142FFCAFE1BE766FF6AD95D" box="[113,248,1039,1065]" italics="true" pageId="4" pageNumber="539">P. williamsae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B303B7142FFCAFE95E766FEA8D95E" box="[255,314,1040,1066]" pageId="4" pageNumber="539" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>