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<mods:title id="732044FC4890409E88A142921100B8DE">Fossils reveal a high diversity of the staghorn coral genera Acropora and Isopora (Scleractinia: Acroporidae) in the Neogene of Indonesia</mods:title>
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<subSubSection id="C318C3F2FFC3F635FF38FF167EF8FF61" box="[216,686,198,219]" pageId="44" pageNumber="721" type="nomenclature">
<paragraph id="8BBD9079FFC3F635FF38FF167EF8FF61" blockId="44.[216,686,198,219]" box="[216,686,198,219]" pageId="44" pageNumber="721">
<heading id="D0F52715FFC3F635FF38FF167EF8FF61" box="[216,686,198,219]" centered="true" fontSize="8" level="2" pageId="44" pageNumber="721" reason="8">
<taxonomicName id="4C02EBFAFFC3F635FF38FF167EF8FF61" ID-CoL="9NXF" authority="WALLACE, 2008" authorityName="WALLACE" authorityYear="2008" box="[216,686,198,219]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="44" pageNumber="721" phylum="Cnidaria" rank="species" species="bartonensis">
<emphasis id="B9764C6BFFC3F635FF38FF167DA4FF60" box="[216,498,198,219]" italics="true" pageId="44" pageNumber="721">ACROPORA BARTONENSIS</emphasis>
<bibRefCitation id="EF93ED88FFC3F635FE1AFF167EF8FF61" author="Wallace CC" box="[506,686,198,219]" pageId="44" pageNumber="721" pagination="313 - 328" refId="ref53922" refString="Wallace CC. 2008. New species and records from the Eocene of England and France support early diversification of the coral genus Acropora. Journal of Paleontology 82: 313 - 328." type="journal article" year="2008">WALLACE, 2008</bibRefCitation>
</taxonomicName>
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<paragraph id="8BBD9079FFC3F635FE67FF3D7DA9FEB8" blockId="44.[391,511,237,258]" box="[391,511,237,258]" pageId="44" pageNumber="721">
<figureCitation id="13398CFCFFC3F635FE67FF3D7DA9FEB8" box="[391,511,237,258]" captionStart="Figure 21" captionStartId="44.[144,221,1765,1784]" captionTargetBox="[161,1404,901,1736]" captionTargetId="figure-277@44.[572,885,900,1333]" captionTargetPageId="44" captionText="Figure 21. Acropora bartonensis. NHMUK PI AZ 6879, TF79, Batu Cermen, Samarinda, East Kalimantan, Middle Miocene (14.815.3 Ma). A, whole specimen showing short branchlets indicative of hispidose form. B, detail of branchlet showing axial corallite and radial corallites. C, electron micrograph showing incipient branchlet and radial corallites. D, electron micrograph of incipient branchlet with radial corallite. E, electron micrograph of coenosteum formed of aligned simple flattened spinules." figureDoi="http://doi.org/10.5281/zenodo.5339608" httpUri="https://zenodo.org/record/5339608/files/figure.png" pageId="44" pageNumber="721">FIGURE 21</figureCitation>
</paragraph>
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<subSubSection id="C318C3F2FFC3F635FF70FEC37E12FEFD" pageId="44" pageNumber="721" type="reference_group">
<paragraph id="8BBD9079FFC3F635FF70FEC37E12FEFD" blockId="44.[144,759,275,328]" pageId="44" pageNumber="721">
<treatmentCitationGroup id="AB12B757FFC3F635FF70FEC37E12FEFD" pageId="44" pageNumber="721">
<treatmentCitation id="0AA3B668FFC3F635FF70FEC37ECBFE93" author="Wallace CC" box="[144,669,275,297]" page="327" pageId="44" pageNumber="721" year="2008">
<taxonomicName id="4C02EBFAFFC3F635FF70FEC37ECBFE93" ID-CoL="9NXF" authority="Wallace, 2008: 326 - 327" authorityName="WALLACE" authorityPageNumber="326 - 327" authorityYear="2008" box="[144,669,275,297]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="44" pageNumber="721" phylum="Cnidaria" rank="species" species="bartonensis">
<emphasis id="B9764C6BFFC3F635FF70FEC37DD5FE92" box="[144,387,275,296]" italics="true" pageId="44" pageNumber="721">Acropora bartonensis</emphasis>
<bibRefCitation id="EF93ED88FFC3F635FE6CFEC37ECBFE93" author="Wallace CC" box="[396,669,275,297]" pageId="44" pageNumber="721" pagination="313 - 328" refId="ref53922" refString="Wallace CC. 2008. New species and records from the Eocene of England and France support early diversification of the coral genus Acropora. Journal of Paleontology 82: 313 - 328." type="journal article" year="2008">Wallace, 2008: 326327</bibRefCitation>
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, fig. 10;
<treatmentCitation id="0AA3B668FFC3F635FF70FEE27DBFFEFD" author="Wallace CC &amp; Bosellini FR" box="[144,489,306,328]" page="14" pageId="44" pageNumber="721" year="2014">
<bibRefCitation id="EF93ED88FFC3F635FF70FEE27DBFFEFD" author="Wallace CC &amp; Bosellini FR" box="[144,489,306,328]" pageId="44" pageNumber="721" pagination="447 - 469" refId="ref54012" refString="Wallace CC, Bosellini FR. 2014. Acropora (Scleractinia) from the Oligocene and Miocene of Europe: species longevity, origination and turnover following the Eocene - Oligocene transition. Journal of Systematic Palaeontology 13: 447 - 469." type="journal article" year="2014">Wallace &amp; Bosellini, 2014: 14</bibRefCitation>
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, fig. 10.
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</paragraph>
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<subSubSection id="C318C3F2FFC3F634FF70FEBB7E86FB57" lastPageId="45" lastPageNumber="722" pageId="44" pageNumber="721" type="diagnosis">
<paragraph id="8BBD9079FFC3F635FF70FEBB7D56FE3A" blockId="44.[144,759,363,537]" box="[144,256,363,384]" pageId="44" pageNumber="721">
<emphasis id="B9764C6BFFC3F635FF70FEBB7D56FE3A" box="[144,256,363,384]" italics="true" pageId="44" pageNumber="721">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BBD9079FFC3F635FF70FE587EC1FDA3" blockId="44.[144,759,363,537]" pageId="44" pageNumber="721">
Colony hispidose, slender axis with short branches; smaller radial corallites mainly clustered around the branchlets, or irregularly sparse along main branches, evenly sized. Coenosteum with lateral flattened simple spinules aligned throughout (
<bibRefCitation id="EF93ED88FFC3F635FE07FDD37EDAFDA3" author="Wallace CC" box="[487,652,515,537]" pageId="44" pageNumber="721" pagination="313 - 328" refId="ref53922" refString="Wallace CC. 2008. New species and records from the Eocene of England and France support early diversification of the coral genus Acropora. Journal of Paleontology 82: 313 - 328." type="journal article" year="2008">Wallace, 2008</bibRefCitation>
).
</paragraph>
<paragraph id="8BBD9079FFC3F635FF70FDEB7D18FDEA" blockId="44.[144,759,571,746]" box="[144,334,571,592]" pageId="44" pageNumber="721">
<emphasis id="B9764C6BFFC3F635FF70FDEB7D18FDEA" box="[144,334,571,592]" italics="true" pageId="44" pageNumber="721">Material studied</emphasis>
</paragraph>
<paragraph id="8BBD9079FFC3F635FF70FD8A7D7EFD50" blockId="44.[144,759,571,746]" pageId="44" pageNumber="721">
<materialsCitation id="3B6A9A24FFC3F635FF70FD8A7D7EFD50" accessionNumber="R14571, R19125" collectionCode="NHMUK, PI, R" county="Fossil" location="Eocene" municipality="Barton" pageId="44" pageNumber="721" specimenCode="AZ 6879" specimenCount="1" stateProvince="Kalimantan Timur" typeStatus="holotype">
<collectingRegion id="49C65E9BFFC3F635FF70FD8A7D0DFDCA" box="[144,347,602,624]" country="Indonesia" name="Kalimantan Timur" pageId="44" pageNumber="721">East Kalimantan</collectingRegion>
:
<collectionCode id="ED1308BCFFC3F635FE88FD8A7D99FDD5" box="[360,463,602,623]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="44" pageNumber="721" type="Museum">NHMUK</collectionCode>
<collectionCode id="ED1308BCFFC3F635FE39FD8A7DA5FDD5" box="[473,499,602,623]" country="Russia" name="Paleontological Institute" pageId="44" pageNumber="721">PI</collectionCode>
<specimenCode id="DBA43802FFC3F635FE1BFD8A7E09FDCA" box="[507,607,602,624]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="44" pageNumber="721">AZ 6879</specimenCode>
,
<specimenCount id="9D045BF0FFC3F635FD8EFD8A7EA4FDCA" box="[622,754,602,624]" pageId="44" pageNumber="721" type="generic">1 specimen</specimenCount>
.
<emphasis id="B9764C6BFFC3F635FF48FDA87D8DFD37" box="[168,475,632,654]" italics="true" pageId="44" pageNumber="721">
<collectingCounty id="62DCE8F5FFC3F635FF48FDA87CBFFD37" box="[168,233,632,653]" pageId="44" pageNumber="721">Fossil</collectingCounty>
comparative material
</emphasis>
:
<collectionCode id="ED1308BCFFC3F635FE09FDA97E18FD34" box="[489,590,633,654]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="44" pageNumber="721" type="Museum">NHMUK</collectionCode>
<collectionCode id="ED1308BCFFC3F635FDB5FDA97E39FD34" box="[597,623,633,654]" country="Russia" name="Paleontological Institute" pageId="44" pageNumber="721">PI</collectionCode>
AZ
<collectionCode id="ED1308BCFFC3F635FD7DFDA87EE6FD34" box="[669,688,632,654]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="44" pageNumber="721">R</collectionCode>
14571,
<typeStatus id="54B92EDBFFC3F635FF70FD477CAFFD17" box="[144,249,663,685]" pageId="44" pageNumber="721" type="holotype">Holotype</typeStatus>
,
<collectingMunicipality id="6BD90A03FFC3F635FEE8FD477D0CFD16" box="[264,346,663,684]" pageId="44" pageNumber="721">Barton</collectingMunicipality>
,
<collectingRegion id="49C65E9BFFC3F635FE89FD477DBDFD17" box="[361,491,663,685]" country="United Kingdom" name="Hampshire" pageId="44" pageNumber="721">Hampshire</collectingRegion>
,
<location id="8EDDC6A2FFC3F635FE1BFD477E18FD16" LSID="urn:lsid:plazi:treatment:03AB216FFFC3F634FF38FF167E86FB57:8EDDC6A2FFC3F635FE1BFD477E18FD16" box="[507,590,663,684]" county="Fossil" municipality="Barton" name="Eocene" pageId="44" pageNumber="721" stateProvince="Kalimantan Timur">Eocene</location>
,
<location id="8EDDC6A2FFC3F635FDBDFD477EA4FD17" LSID="urn:lsid:plazi:treatment:03AB216FFFC3F634FF38FF167E86FB57:8EDDC6A2FFC3F635FDBDFD477EA4FD17" box="[605,754,663,685]" county="Fossil" municipality="Barton" name="Barton Beds" pageId="44" pageNumber="721" stateProvince="Kalimantan Timur">Barton Beds</location>
;
<collectionCode id="ED1308BCFFC3F635FF70FD667CA1FD71" box="[144,247,694,715]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="44" pageNumber="721" type="Museum">NHMUK</collectionCode>
<collectionCode id="ED1308BCFFC3F635FF1FFD667D4FFD71" box="[255,281,694,715]" country="Russia" name="Paleontological Institute" pageId="44" pageNumber="721">PI</collectionCode>
AZ
<collectionCode id="ED1308BCFFC3F635FEABFD657D08FD71" box="[331,350,693,715]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="44" pageNumber="721">R</collectionCode>
19125,
<location id="8EDDC6A2FFC3F635FE51FD667E55FD71" LSID="urn:lsid:plazi:treatment:03AB216FFFC3F634FF38FF167E86FB57:8EDDC6A2FFC3F635FE51FD667E55FD71" box="[433,515,694,715]" county="Fossil" municipality="Barton" name="Barton" pageId="44" pageNumber="721" stateProvince="Kalimantan Timur">Barton</location>
,
<collectingRegion id="49C65E9BFFC3F635FDF0FD657EC4FD71" box="[528,658,693,715]" country="United Kingdom" name="Hampshire" pageId="44" pageNumber="721">Hampshire</collectingRegion>
,
<location id="8EDDC6A2FFC3F635FD7FFD667EA4FD71" LSID="urn:lsid:plazi:treatment:03AB216FFFC3F634FF38FF167E86FB57:8EDDC6A2FFC3F635FD7FFD667EA4FD71" box="[671,754,694,715]" county="Fossil" municipality="Barton" name="Eocene" pageId="44" pageNumber="721" stateProvince="Kalimantan Timur">Eocene</location>
, Barton Beds.
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</paragraph>
<paragraph id="8BBD9079FFC3F635FF70FCDC7DC1FC9B" blockId="44.[144,759,780,864]" box="[144,407,780,801]" pageId="44" pageNumber="721">
<emphasis id="B9764C6BFFC3F635FF70FCDC7DC1FC9B" box="[144,407,780,801]" italics="true" pageId="44" pageNumber="721">Skeletal characteristics</emphasis>
</paragraph>
<paragraph id="8BBD9079FFC3F635FF70FCFB79D8FE08" blockId="44.[144,759,780,864]" lastBlockId="44.[806,1422,198,862]" pageId="44" pageNumber="721">
<taxonomicName id="4C02EBFAFFC3F635FF70FCFB7CAAFCFA" authorityName="Ehrenberg" authorityYear="1834" box="[144,252,811,832]" genus="Corallum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="721" phylum="Cnidaria" rank="genus">
<emphasis id="B9764C6BFFC3F635FF70FCFB7CAAFCFA" box="[144,252,811,832]" italics="true" pageId="44" pageNumber="721">Corallum</emphasis>
</taxonomicName>
. Overall length of specimen
<quantity id="4CFA3D9CFFC3F635FDB6FCFB7EEDFCFA" box="[598,699,811,833]" metricMagnitude="-2" metricUnit="m" metricValue="5.01" pageId="44" pageNumber="721" unit="mm" value="50.1">50.1 mm</quantity>
, mid branch diameter 7.108.23
<quantity id="4CFA3D9CFFC3F635FE33FC9A7E13FCE5" box="[467,581,842,863]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="44" pageNumber="721" unit="mm" value="10.1">10.10 mm</quantity>
, multiple short branchlets extending from main branch indicating hispidose form; branchlets arising regularly from main branch in angles of 49.6863.5388.45°, distance between branchlets 8.209.42
<quantity id="4CFA3D9CFFC3F635FBD7FEF278F9FE8D" box="[1079,1199,290,312]" metricMagnitude="-2" metricUnit="m" metricValue="1.046" pageId="44" pageNumber="721" unit="mm" value="10.46">10.46 mm</quantity>
, with tips mostly abraded, tapering, branchlet length 3.25.3
<quantity id="4CFA3D9CFFC3F635FAD1FE9079DEFEEC" box="[1329,1416,320,342]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="44" pageNumber="721" unit="mm" value="7.2">7.2 mm</quantity>
, basal branchlet diameter 4.905.36
<quantity id="4CFA3D9CFFC3F635FB2DFE8F7967FECE" box="[1229,1329,351,373]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="44" pageNumber="721" unit="mm" value="6.2">6.20 mm</quantity>
, middle branch diameter 3.904.58
<quantity id="4CFA3D9CFFC3F635FB84FEAD789EFE29" box="[1124,1224,381,403]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="44" pageNumber="721" unit="mm" value="5.4">5.40 mm</quantity>
, surfaces extending between branches with a few sparse radial corallites.
</paragraph>
<paragraph id="8BBD9079FFC3F635FCDFFE6B78D9FCE4" blockId="44.[806,1422,198,862]" pageId="44" pageNumber="721">
<emphasis id="B9764C6BFFC3F635FCDFFE6B7FFBFE6A" box="[831,941,443,464]" italics="true" pageId="44" pageNumber="721">Corallites</emphasis>
. Axial corallites visible at some branchlet tips, round calice, outer diameter 1.511.93
<quantity id="4CFA3D9CFFC3F635FCC6FE287FD8FDB7" box="[806,910,504,525]" metricMagnitude="-3" metricUnit="m" metricValue="2.19" pageId="44" pageNumber="721" unit="mm" value="2.19">2.19 mm</quantity>
, inner diameter 0.961.24
<quantity id="4CFA3D9CFFC3F635FB04FE28791CFDB7" box="[1252,1354,504,526]" metricMagnitude="-3" metricUnit="m" metricValue="1.43" pageId="44" pageNumber="721" unit="mm" value="1.43">1.43 mm</quantity>
, wall thickness 0.290.31
<quantity id="4CFA3D9CFFC3F635FBEBFDC7783AFD96" box="[1035,1132,535,556]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="44" pageNumber="721" unit="mm" value="0.39">0.39 mm</quantity>
, primary septa visible but worn, secondary septa may be present; radial corallites mostly worn and crowded in short branchlets, three to six radials per branchlet, mostly not touching, probably short tubular to labellate with lower wall slightly thickened, profile length 0.290.42
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, angle 43.0453.5562.10°, outer diameter 0.820.86
<quantity id="4CFA3D9CFFC3F635FAC9FD1E79DFFD59" box="[1321,1417,718,740]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="44" pageNumber="721" unit="mm" value="0.94">0.94 mm</quantity>
, inner diameter 0.470.60
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, wall thickness 0.12 0.14
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, septa worn out, evidence for primary septa larger than secondary septa. Corallite arrangement sequence 1[34]-610?.
</paragraph>
<caption id="DF7DC0F1FFC3F635FF70F9357D02F8D4" ID-DOI="http://doi.org/10.5281/zenodo.5339608" ID-Zenodo-Dep="5339608" httpUri="https://zenodo.org/record/5339608/files/figure.png" pageId="44" pageNumber="721" startId="44.[144,221,1765,1784]" targetBox="[161,1404,901,1736]" targetPageId="44">
<paragraph id="8BBD9079FFC3F635FF70F9357D02F8D4" blockId="44.[144,1421,1765,1902]" pageId="44" pageNumber="721">
<emphasis id="B9764C6BFFC3F635FF70F9357D54F942" bold="true" box="[144,258,1765,1784]" pageId="44" pageNumber="721">Figure 21.</emphasis>
<emphasis id="B9764C6BFFC3F635FEECF9367DBDF942" box="[268,491,1765,1785]" italics="true" pageId="44" pageNumber="721">
<taxonomicName id="4C02EBFAFFC3F635FEECF9367DB1F942" authorityName="WALLACE" authorityYear="2008" box="[268,487,1765,1785]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="44" pageNumber="721" phylum="Cnidaria" rank="species" species="bartonensis">Acropora bartonensis</taxonomicName>
.
</emphasis>
NHMUK PI AZ 6879, TF79, Batu Cermen, Samarinda, East Kalimantan, Middle Miocene (14.815.3 Ma). A, whole specimen showing short branchlets indicative of hispidose form. B, detail of branchlet showing axial corallite and radial corallites. C, electron micrograph showing incipient branchlet and radial corallites. D, electron micrograph of incipient branchlet with radial corallite. E, electron micrograph of coenosteum formed of aligned simple flattened spinules.
</paragraph>
</caption>
<paragraph id="8BBD9079FFC2F634FF5CFF167D58FE8D" blockId="45.[164,779,198,311]" pageId="45" pageNumber="722">
<emphasis id="B9764C6BFFC2F634FF5CFF167D12FF61" box="[188,324,198,219]" italics="true" pageId="45" pageNumber="722">Coenosteum</emphasis>
. Simple flattened spinules evenly distributed and aligned, both on corallite walls and between radial corallites. Coenosteum amount 1.111.85
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.
</paragraph>
<paragraph id="8BBD9079FFC2F634FF44FE8F7D74FECE" blockId="45.[164,780,351,679]" box="[164,290,351,372]" pageId="45" pageNumber="722">
<emphasis id="B9764C6BFFC2F634FF44FE8F7D74FECE" box="[164,290,351,372]" italics="true" pageId="45" pageNumber="722">Occurrence</emphasis>
</paragraph>
<paragraph id="8BBD9079FFC2F634FF44FEAE7D02FD1D" blockId="45.[164,780,351,679]" pageId="45" pageNumber="722">
Eocene to Middle Miocene. The earliest occurrence of the species is from its
<typeStatus id="54B92EDBFFC2F634FE59FE4D7DBCFE08" box="[441,490,413,434]" pageId="45" pageNumber="722">type</typeStatus>
locality, Barton,
<collectingRegion id="49C65E9BFFC2F634FD5CFE4D7C88FE6B" country="United Kingdom" name="Hampshire" pageId="45" pageNumber="722">Hampshire</collectingRegion>
,
<collectingCountry id="F315D0E9FFC2F634FF09FE6B7D44FE6A" box="[233,274,443,464]" name="United Kingdom" pageId="45" pageNumber="722">UK</collectingCountry>
, of Bartonian age, 3841 Ma (
<bibRefCitation id="EF93ED88FFC2F634FD83FE6B7F56FE6B" author="Wallace CC" box="[611,768,443,465]" pageId="45" pageNumber="722" pagination="313 - 328" refId="ref53922" refString="Wallace CC. 2008. New species and records from the Eocene of England and France support early diversification of the coral genus Acropora. Journal of Paleontology 82: 313 - 328." type="journal article" year="2008">Wallace, 2008</bibRefCitation>
). Additional records include Miocene specimens from
<collectingCountry id="F315D0E9FFC2F634FF44FE287CA5FDB7" box="[164,243,504,525]" name="France" pageId="45" pageNumber="722">France</collectingCountry>
and
<collectingCountry id="F315D0E9FFC2F634FED0FE287D30FDB4" box="[304,358,504,526]" name="Italy" pageId="45" pageNumber="722">Italy</collectingCountry>
of Aquitanian and Burdigalian age, 1623 Ma (
<bibRefCitation id="EF93ED88FFC2F634FEC7FDC77E01FD97" author="Wallace CC &amp; Bosellini FR" box="[295,599,535,557]" pageId="45" pageNumber="722" pagination="447 - 469" refId="ref54012" refString="Wallace CC, Bosellini FR. 2014. Acropora (Scleractinia) from the Oligocene and Miocene of Europe: species longevity, origination and turnover following the Eocene - Oligocene transition. Journal of Systematic Palaeontology 13: 447 - 469." type="journal article" year="2014">Wallace &amp; Bosellini, 2014</bibRefCitation>
). The specimen studied was recovered from the locality TF79, Samarinda, East Kalimantan, of Langhian age, 14.8 15.3 Ma. This record is the youngest occurrence of this extinct species.
</paragraph>
<paragraph id="8BBD9079FFC2F634FF44FD1E7D28FD59" blockId="45.[164,779,718,832]" box="[164,382,718,739]" pageId="45" pageNumber="722">
<emphasis id="B9764C6BFFC2F634FF44FD1E7D28FD59" box="[164,382,718,739]" italics="true" pageId="45" pageNumber="722">Palaeoenvironment</emphasis>
</paragraph>
<paragraph id="8BBD9079FFC2F634FF44FD3D7D64FCFA" blockId="45.[164,779,718,832]" pageId="45" pageNumber="722">
The mixed siliciclasticcarbonate limestones of locality TF79 have been interpreted as patch reefs (
<bibRefCitation id="EF93ED88FFC2F634FD50FCDC7D71FCFA" author="Renema W &amp; Warter V &amp; Novak V &amp; Young JR &amp; Marshall N &amp; Hasibuan F" pageId="45" pageNumber="722" pagination="26 - 39" refId="ref52535" refString="Renema W, Warter V, Novak V, Young JR, Marshall N, Hasibuan F. 2015. Ages of Miocene fossil localities in the northern Kutai Basin (East Kalimantan, Indonesia). Palaios 30: 26 - 39. doi: 10.2110 / palo. 2013.127." type="journal article" year="2015">
Renema
<emphasis id="B9764C6BFFC2F634FF44FCFB7C82FC85" box="[164,212,810,832]" italics="true" pageId="45" pageNumber="722">et al</emphasis>
., 2015
</bibRefCitation>
).
</paragraph>
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<emphasis id="B9764C6BFFC2F634FF44FCB87D5FFCC7" box="[164,265,872,893]" italics="true" pageId="45" pageNumber="722">Remarks</emphasis>
</paragraph>
<paragraph id="8BBD9079FFC2F634FF44FC567E86FB57" blockId="45.[164,779,872,1261]" pageId="45" pageNumber="722">
This specimen compares well with the
<typeStatus id="54B92EDBFFC2F634FD66FC567EBFFC26" box="[646,745,902,924]" pageId="45" pageNumber="722" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C02EBFAFFC2F634FF44FC757D01FC00" authorityName="WALLACE" authorityYear="2008" box="[164,343,933,954]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="45" pageNumber="722" phylum="Cnidaria" rank="species" species="bartonensis">
<emphasis id="B9764C6BFFC2F634FF44FC757D01FC00" box="[164,343,933,954]" italics="true" pageId="45" pageNumber="722">A. bartonensis</emphasis>
</taxonomicName>
by sharing regularly distributed branchlets indicative of hispidose form and with radial corallites arranged throughout the main axis and also clustered around the branchlets. Although the coenosteum of the Indonesian fossil specimen is not as costate as in the
<typeStatus id="54B92EDBFFC2F634FE6AFBEE7DBCFBEE" box="[394,490,1086,1108]" pageId="45" pageNumber="722" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C02EBFAFFC2F634FDEFFBEE7EE7FBE9" authorityName="WALLACE" authorityYear="2008" box="[527,689,1086,1107]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="45" pageNumber="722" phylum="Cnidaria" rank="species" species="bartonensis">
<emphasis id="B9764C6BFFC2F634FDEFFBEE7EE7FBE9" box="[527,689,1086,1107]" italics="true" pageId="45" pageNumber="722">A. bartonensis</emphasis>
</taxonomicName>
, simple flattened spinules are aligned and it is probable that the costae have not been preserved in this highly recrystallized specimen. Together with
<taxonomicName id="4C02EBFAFFC2F634FD7CFB4A7D66FB77" baseAuthorityName="Michelin" baseAuthorityYear="1840" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="45" pageNumber="722" phylum="Cnidaria" rank="species" species="lavandulina">
<emphasis id="B9764C6BFFC2F634FD7CFB4A7D66FB77" italics="true" pageId="45" pageNumber="722">Acropora lavandulina</emphasis>
</taxonomicName>
, these are the only Tethyan species also recorded in the fossil record of the Indo-Pacific.
</paragraph>
</subSubSection>
</treatment>
</document>