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<document id="CFB8A95C5A7CE72697C93F20962E9CC2" ID-CLB-Dataset="45370" ID-DOI="10.5281/zenodo.215126" ID-GBIF-Dataset="7b3ca294-d1e3-48a3-973f-5f002e9d20e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="215126" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460444712012" checkinUser="plazi" docAuthor="Hunger, Holger &amp; Schiel, Franz-Josef" docDate="2012" docId="039BA549FFBBFFF7FF18FEC7FB12FA17" docLanguage="en" docName="zt03202p050.pdf" docOrigin="Zootaxa 3202" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Protallagma hoffmanni Hunger &amp; Schiel, 2012, sp. nov." docType="treatment" docVersion="11" lastPageNumber="33" masterDocId="FFA2DD31FFBAFFF2FF8FFA6BFFDEFFF3" masterDocTitle="Description of Protallagma hoffmanni sp. nov. from the Peruvian Andes (Odonata: Coenagrionidae), including description of its larva" masterLastPageNumber="50" masterPageNumber="28" pageNumber="29" updateTime="1698333227736" updateUser="ExternalLinkService">
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<mods:title id="BDD2D942154E587DE3784499B890713C">Description of Protallagma hoffmanni sp. nov. from the Peruvian Andes (Odonata: Coenagrionidae), including description of its larva</mods:title>
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<mods:namePart id="A4D018F8B19BE5FE6DC3070EB60793C6">Hunger, Holger</mods:namePart>
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<mods:namePart id="C52574ECC6CDCA4F73B668129E8EA741">Schiel, Franz-Josef</mods:namePart>
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<mods:date id="61C9F08A80553F4E57136F5BCC370960">2012</mods:date>
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<treatment id="039BA549FFBBFFF7FF18FEC7FB12FA17" ID-DOI="http://doi.org/10.5281/zenodo.5625800" ID-GBIF-Taxon="119509201" ID-Zenodo-Dep="5625800" LSID="urn:lsid:plazi:treatment:039BA549FFBBFFF7FF18FEC7FB12FA17" httpUri="http://treatment.plazi.org/id/039BA549FFBBFFF7FF18FEC7FB12FA17" lastPageId="5" lastPageNumber="33" pageId="1" pageNumber="29">
<subSubSection id="C32847D4FFBBFFF3FF18FEC7FDC6FB36" box="[151,536,1195,1222]" pageId="1" pageNumber="29" type="nomenclature">
<paragraph id="8B8D145FFFBBFFF3FF18FEC7FDC6FB36" blockId="1.[151,536,1195,1222]" box="[151,536,1195,1222]" pageId="1" pageNumber="29">
<heading id="D0C5A333FFBBFFF3FF18FEC7FDC6FB36" bold="true" box="[151,536,1195,1222]" fontSize="11" level="1" pageId="1" pageNumber="29" reason="1">
<emphasis id="B946C84DFFBBFFF3FF18FEC7FDC6FB36" bold="true" box="[151,536,1195,1222]" pageId="1" pageNumber="29">
<taxonomicName id="4C326FDCFFBBFFF3FF18FEC7FE6DFB35" ID-CoL="9X45B" box="[151,435,1196,1222]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="29" phylum="Arthropoda" rank="species" species="hoffmanni" status="sp. nov.">
<emphasis id="B946C84DFFBBFFF3FF18FEC7FE6DFB35" bold="true" box="[151,435,1196,1222]" italics="true" pageId="1" pageNumber="29">Protallagma hoffmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2757536FFBBFFF3FE34FEC0FDC6FB36" box="[443,536,1195,1221]" pageId="1" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32847D4FFBBFFF3FF18FE98FDD3FAA1" pageId="1" pageNumber="29" type="etymology">
<paragraph id="8B8D145FFFBBFFF3FF18FE98FDD3FAA1" blockId="1.[151,1437,1266,1938]" pageId="1" pageNumber="29">
<emphasis id="B946C84DFFBBFFF3FF18FE98FEC0FAF8" bold="true" box="[151,286,1267,1291]" pageId="1" pageNumber="29">Etymology.</emphasis>
This species is named
<taxonomicName id="4C326FDCFFBBFFF3FDACFE98FD4BFAF8" box="[547,661,1267,1291]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="29" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="B946C84DFFBBFFF3FDACFE98FD4BFAF8" box="[547,661,1267,1291]" italics="true" pageId="1" pageNumber="29">hoffmanni</emphasis>
</taxonomicName>
after our good friend and colleague Joachim Hoffmann who worked at the Museo de Historia Natural in Lima for many years and is an expert on
<taxonomicName id="4C326FDCFFBBFFF3FC61FF7DFB93FADD" box="[1006,1101,1302,1326]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="29" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
from
<collectingCountry id="F32554CFFFBBFFF3FB1FFF7CFB19FADD" box="[1168,1223,1303,1326]" name="Peru" pageId="1" pageNumber="29">Peru</collectingCountry>
. We found the new species on a field trip led by him.
</paragraph>
</subSubSection>
<subSubSection id="C32847D4FFBBFFF3FF46FF34FC4AFA11" pageId="1" pageNumber="29" type="materials_examined">
<paragraph id="8B8D145FFFBBFFF3FF46FF34FC4AFA11" blockId="1.[151,1437,1266,1938]" pageId="1" pageNumber="29">
<emphasis id="B946C84DFFBBFFF3FF46FF34FE24FA84" bold="true" box="[201,506,1374,1399]" pageId="1" pageNumber="29">
<typeStatus id="5489AAFDFFBBFFF3FF46FF34FEDAFA84" box="[201,260,1375,1399]" pageId="1" pageNumber="29">Type</typeStatus>
material.
<typeStatus id="5489AAFDFFBBFFF3FE0BFF34FE2BFA84" box="[388,501,1375,1399]" pageId="1" pageNumber="29" type="holotype">Holotype</typeStatus>
:
</emphasis>
3 from Laguna Querococha, Ancash region,
<collectingCountry id="F32554CFFFBBFFF3FB9CFF34FB97FA85" box="[1043,1097,1375,1398]" name="Peru" pageId="1" pageNumber="29">Peru</collectingCountry>
,
<geoCoordinate id="EE067298FFBBFFF3FBD7FF35FAD2FA85" box="[1112,1292,1374,1398]" direction="south" orientation="latitude" pageId="1" pageNumber="29" precision="1" value="-9.726">S 09° 43' 33.6&quot;</geoCoordinate>
<geoCoordinate id="EE067298FFBBFFF3FA99FF34FF08FA69" direction="west" orientation="longitude" pageId="1" pageNumber="29" precision="1" value="-77.331055">W 077° 19' 51.8&quot;</geoCoordinate>
,
<quantity id="4CCAB9BAFFBBFFF3FF6CFFE9FE9FFA69" box="[227,321,1410,1434]" metricMagnitude="3" metricUnit="m" metricValue="3.9799999999999995" pageId="1" pageNumber="29" unit="m" value="3980.0">3,980 m</quantity>
a.s.l.,
<date id="FF8C329FFFBBFFF3FE01FFE9FD85FA69" box="[398,603,1410,1434]" pageId="1" pageNumber="29" value="2007-02-18">18 February 2007</date>
, leg. Holger Hunger and Franz-Josef Schiel (deposited at
<collectionCode id="ED238C9AFFBBFFF3FB7AFFE9FA93FA69" box="[1269,1357,1410,1434]" httpUri="http://grbio.org/cool/4jmm-iazx" name="Musee Guimet d'Histoire Naturelle de Lyon" pageId="1" pageNumber="29">MHNL</collectionCode>
).
<emphasis id="B946C84DFFBBFFF3FAEEFFE9FF0DFA4C" bold="true" pageId="1" pageNumber="29">Allotype:</emphasis>
Ƥ same data (
<collectionCode id="ED238C9AFFBBFFF3FEFCFFCDFE12FA4D" box="[371,460,1446,1470]" httpUri="http://grbio.org/cool/4jmm-iazx" name="Musee Guimet d'Histoire Naturelle de Lyon" pageId="1" pageNumber="29">MHNL</collectionCode>
).
<emphasis id="B946C84DFFBBFFF3FE51FFCCFD81FA4C" bold="true" box="[478,607,1447,1471]" pageId="1" pageNumber="29">
<typeStatus id="5489AAFDFFBBFFF3FE51FFCCFD84FA4C" box="[478,602,1447,1471]" pageId="1" pageNumber="29" type="paratype">Paratypes</typeStatus>
:
</emphasis>
7 3, 8 Ƥ, same data (2 3 and 2 Ƥ
<typeStatus id="5489AAFDFFBBFFF3FC6CFFCCFB90FA4C" box="[995,1102,1447,1471]" pageId="1" pageNumber="29" type="paratype">paratypes</typeStatus>
at FNS, remainder in
<collectionCode id="ED238C9AFFBBFFF3FACAFFCCFA42FA4C" box="[1349,1436,1447,1471]" pageId="1" pageNumber="29">INULA</collectionCode>
collection).
<emphasis id="B946C84DFFBBFFF3FE93FFA0FEA5FA10" bold="true" box="[284,379,1483,1507]" pageId="1" pageNumber="29">Larvae:</emphasis>
<specimenCount id="9D34DFD6FFBBFFF3FE0DFFA1FE32FA11" box="[386,492,1482,1506]" pageId="1" pageNumber="29" type="larva">19 larvae</specimenCount>
, same data (all in
<collectionCode id="ED238C9AFFBBFFF3FD38FFA0FCD0FA10" box="[695,782,1483,1507]" pageId="1" pageNumber="29">INULA</collectionCode>
collection).
</paragraph>
</subSubSection>
<subSubSection id="C32847D4FFBBFFF7FF46FF85FB98FCCB" lastPageId="5" lastPageNumber="33" pageId="1" pageNumber="29" type="description">
<paragraph id="8B8D145FFFBBFFF3FF46FF85FDD5F9F5" blockId="1.[151,1437,1266,1938]" box="[201,523,1518,1543]" pageId="1" pageNumber="29">
<emphasis id="B946C84DFFBBFFF3FF46FF85FE84F9F4" bold="true" box="[201,346,1518,1543]" pageId="1" pageNumber="29">Description.</emphasis>
Male
<typeStatus id="5489AAFDFFBBFFF3FE2DFF85FDD6F9F5" box="[418,520,1518,1542]" pageId="1" pageNumber="29" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B8D145FFFBBFFF3FF46FC78FF06F965" blockId="1.[151,1437,1266,1938]" pageId="1" pageNumber="29">
<emphasis id="B946C84DFFBBFFF3FF46FC78FECEF9D8" bold="true" box="[201,272,1555,1579]" pageId="1" pageNumber="29">Head.</emphasis>
Head and labium yellowish brown. Frontal suture black, as well as sutures on both sides of median ocellus and suture behind left and right ocelli and occipital bar, which is pale. Whole head, especially antefrons to labium, with very long (up to ca.
<quantity id="4CCAB9BAFFBBFFF3FD9FFC31FDB6F981" box="[528,616,1626,1650]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="29" unit="mm" value="0.6">0.6 mm</quantity>
) setae. Distinct row of setae also on occipital bar (
<figureCitation id="130908DAFFBBFFF3FB2CFC31FB3BF981" box="[1187,1253,1626,1650]" captionStart="FIGURE 1. P" captionStartId="2.[151,250,1073,1096]" captionTargetBox="[153,1418,198,1054]" captionTargetId="figure@2.[151,1436,187,1060]" captionTargetPageId="2" captionText="FIGURE 1. P. hoffmanni, head of male holotype, dorsal view." httpUri="https://zenodo.org/record/215127/files/figure.png" pageId="1" pageNumber="29">Fig. 1</figureCitation>
). No postocular spots.
</paragraph>
<paragraph id="8B8D145FFFBBFFF3FF46FCC8FF08F861" blockId="1.[151,1437,1266,1938]" pageId="1" pageNumber="29">
<emphasis id="B946C84DFFBBFFF3FF46FCC8FEF7F948" bold="true" box="[201,297,1699,1723]" pageId="1" pageNumber="29">Thorax.</emphasis>
Pronotal hindlobe undulate with nozzle-like structure facing toward head (
<figureCitation id="130908DAFFBBFFF3FB00FCC9FB09F949" box="[1167,1239,1698,1722]" captionStart="FIGURE 2. P" captionStartId="2.[151,250,1758,1781]" captionTargetBox="[175,1406,1147,1743]" captionTargetId="figure@2.[163,1423,1139,1746]" captionTargetPageId="2" captionText="FIGURE 2. P. hoffmanni, pronotum and mesostigmal plate of male holotype, dorsal view." httpUri="https://zenodo.org/record/215128/files/figure.png" pageId="1" pageNumber="29">Fig. 2</figureCitation>
). Legs yellowish brown with long black spines. Supplementary pretarsal claws distinct, forming an acute angle with claws. Wings hyaline with very light brownish tinge; pterostigma light brown, very narrow. Number of postnodal crossveins: FW 11 (left), 10 (right); HW 9 (left), 10 (right). Thorax yellowish brown, densely covered with fine, long setae. Darker band along both sides of middorsal carina. Humeral, metapleural, and interpleural stripes very fine and diffuse. Broad, light greenish antehumeral stripe. Venter of thorax bears a metasternal tubercle covered with long pale setae.
</paragraph>
<caption id="DF4D44D7FFB8FFF0FF18FE5AFCD0FBB4" box="[151,782,1073,1096]" httpUri="https://zenodo.org/record/215127/files/figure.png" pageId="2" pageNumber="30" targetBox="[153,1418,198,1054]" targetPageId="2">
<paragraph id="8B8D145FFFB8FFF0FF18FE5AFCD0FBB4" blockId="2.[151,782,1073,1096]" box="[151,782,1073,1096]" pageId="2" pageNumber="30">
<emphasis id="B946C84DFFB8FFF0FF18FE5AFECDFBB5" bold="true" box="[151,275,1073,1096]" pageId="2" pageNumber="30">FIGURE 1.</emphasis>
<taxonomicName id="4C326FDCFFB8FFF0FE96FE59FE47FBB4" box="[281,409,1073,1095]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="30" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="B946C84DFFB8FFF0FE96FE59FE47FBB4" box="[281,409,1073,1095]" italics="true" pageId="2" pageNumber="30">P. hoffmanni</emphasis>
</taxonomicName>
, head of male holotype, dorsal view.
</paragraph>
</caption>
<caption id="DF4D44D7FFB8FFF0FF18FCB5FBF4F907" box="[151,1066,1758,1781]" httpUri="https://zenodo.org/record/215128/files/figure.png" pageId="2" pageNumber="30" targetBox="[175,1406,1147,1743]" targetPageId="2">
<paragraph id="8B8D145FFFB8FFF0FF18FCB5FBF4F907" blockId="2.[151,1066,1758,1781]" box="[151,1066,1758,1781]" pageId="2" pageNumber="30">
<emphasis id="B946C84DFFB8FFF0FF18FCB5FECDF900" bold="true" box="[151,275,1758,1781]" pageId="2" pageNumber="30">FIGURE 2.</emphasis>
<taxonomicName id="4C326FDCFFB8FFF0FE96FCB4FE47F907" box="[281,409,1758,1780]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="30" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="B946C84DFFB8FFF0FE96FCB4FE47F907" box="[281,409,1758,1780]" italics="true" pageId="2" pageNumber="30">P. hoffmanni</emphasis>
</taxonomicName>
, pronotum and mesostigmal plate of male holotype, dorsal view.
</paragraph>
</caption>
<paragraph id="8B8D145FFFB8FFF1FF46FD71FEAAFE92" blockId="2.[151,1436,1818,2019]" lastBlockId="3.[151,1437,151,389]" lastPageId="3" lastPageNumber="31" pageId="2" pageNumber="30">
<emphasis id="B946C84DFFB8FFF0FF46FD71FE9BF8C0" bold="true" box="[201,325,1818,1843]" pageId="2" pageNumber="30">Abdomen.</emphasis>
Completely covered with fine, short pale setae; ventral side of S13 including area around secondary genitalia with long black setae. Abdomen yellowish brown ventrally; S14 completely coral red dorsally, with fine black transverse rings at intersegmental articulations (present on S26). S56 red with a black dot at distal third, washed on S5, more conspicuous on S6. S710 with azure markings forming a characteristic indented band, caused by a black lateral band on S710 with flame-like dorsad projections. Posterior margin of S79 with a few black denticles. Posterior margin of S10 with a wedge-shaped mid-dorsal emargination. Cercus dorsally brown, in lateral view whitish, mitten-shaped, “thumb“ very pronounced and long, with pointed black tip, overlapping paraproct (Fig. 3). Paraproct conical, wide base gradually narrowing toward blackish tip pointing upwards. Genital opercula pointed posteriorly, reaching close to distal margin of S9 (
<figureCitation id="130908DAFFB9FFF1FC1CFAB4FC00FF04" box="[915,990,223,247]" captionStart="FIGURE 4. P" captionStartId="4.[151,250,1327,1350]" captionTargetBox="[529,1060,186,1281]" captionTargetId="figure@4.[525,1062,185,1299]" captionTargetPageId="4" captionText="FIGURE 4. P. hoffmanni, tip of abdomen of male holotype with genital opercula and appendages, ventral view." httpUri="https://zenodo.org/record/215129/files/figure.png" pageId="3" pageNumber="31">Fig. 4</figureCitation>
). Genital ligula (taken from
<typeStatus id="5489AAFDFFB9FFF1FAB3FA8BFA42FF0B" box="[1340,1436,224,248]" pageId="3" pageNumber="31" type="paratype">paratype</typeStatus>
specimen): distal segment of genital ligula with an inner fold proximal to the flexure and with two lateral lobes on each side. On the outer side with a paired ledge-shaped structure (
<figureCitation id="130908DAFFB9FFF1FC1FFB4DFC21FECD" box="[912,1023,294,318]" captionStart-0="FIGURE 5. P" captionStart-1="FIGURE 6. P" captionStartId-0="4.[151,250,1989,2012]" captionStartId-1="6.[151,250,1571,1594]" captionTargetBox-0="[234,1332,1414,1957]" captionTargetId-0="figure@4.[233,1354,1401,1965]" captionTargetPageId-0="4" captionText-0="FIGURE 5. P. hoffmanni, genital ligula, taken from paratype, lateral view." captionText-1="FIGURE 6. P. hoffmanni, genital ligula, taken from paratype, ectal view." httpUri-0="https://zenodo.org/record/215130/files/figure.png" pageId="3" pageNumber="31">Figs. 56</figureCitation>
), the two ledges, seen from caudad, converging distally.
</paragraph>
<paragraph id="8B8D145FFFB9FFF1FF46FB06FCB8FE77" blockId="3.[151,1437,151,389]" box="[201,870,364,389]" pageId="3" pageNumber="31">
<emphasis id="B946C84DFFB9FFF1FF46FB06FEA1FE76" bold="true" box="[201,383,365,389]" pageId="3" pageNumber="31">Measurements.</emphasis>
HW 17.5, abdomen 19.2, total length 26.3.
</paragraph>
<paragraph id="8B8D145FFFB9FFF1FF18FE24FBC5FB97" blockId="3.[151,1051,1102,1124]" box="[151,1051,1102,1124]" pageId="3" pageNumber="31">
<emphasis id="B946C84DFFB9FFF1FF18FE24FED8FB90" bold="true" box="[151,262,1103,1123]" pageId="3" pageNumber="31">FIGURE 3.</emphasis>
<taxonomicName id="4C326FDCFFB9FFF1FE83FE24FE52FB97" box="[268,396,1102,1124]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="31" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="B946C84DFFB9FFF1FE83FE24FE52FB97" box="[268,396,1102,1124]" italics="true" pageId="3" pageNumber="31">P. hoffmanni</emphasis>
</taxonomicName>
, tip of abdomen with appendages of male
<typeStatus id="5489AAFDFFB9FFF1FCB5FE24FC4AFB97" box="[826,916,1103,1124]" pageId="3" pageNumber="31" type="paratype">paratype</typeStatus>
, lateral view.
</paragraph>
<paragraph id="8B8D145FFFB9FFF1FF46FEE6FE01FAFC" blockId="3.[151,1436,1164,1969]" pageId="3" pageNumber="31">
<emphasis id="B946C84DFFB9FFF1FF46FEE6FE3CFB56" bold="true" box="[201,482,1165,1189]" pageId="3" pageNumber="31">Female allotype. Head.</emphasis>
Labium to postfrons yellowish brown, frontal suture and sutures right and left of median ocellus dark brown. Vertex light olive, occiput yellowish brown. Whole head, especially antefrons to labium, with very long (up to ca.
<quantity id="4CCAB9BAFFB9FFF1FD95FEB8FDAAFB18" box="[538,628,1235,1259]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="31" unit="mm" value="0.6">0.6 mm</quantity>
) setae. Row of setae also on occipital bar which is pale as in the male (
<figureCitation id="130908DAFFB9FFF1FF10FE9CFF3AFAFC" box="[159,228,1271,1295]" captionStart="FIGURE 7. P" captionStartId="7.[151,250,960,983]" captionTargetBox="[247,1349,206,931]" captionTargetId="figure@7.[235,1351,195,944]" captionTargetPageId="7" captionText="FIGURE 7. P. hoffmanni, head of female paratype, dorsal view." httpUri="https://zenodo.org/record/215131/files/figure.png" pageId="3" pageNumber="31">Fig. 7</figureCitation>
). No postocular spots.
</paragraph>
<paragraph id="8B8D145FFFB9FFF1FF46FF70FC01FA6E" blockId="3.[151,1436,1164,1969]" pageId="3" pageNumber="31">
<emphasis id="B946C84DFFB9FFF1FF46FF70FEF7FAC0" bold="true" box="[201,297,1307,1331]" pageId="3" pageNumber="31">Thorax.</emphasis>
Pronotum yellowish brown with darker areas, pronotal hindlobe similar to that of
<typeStatus id="5489AAFDFFB9FFF1FB44FF71FAEEFAC1" box="[1227,1328,1306,1330]" pageId="3" pageNumber="31" type="holotype">holotype</typeStatus>
, undulate with nozzle facing toward head. Legs and supplementary pretarsal claws as in
<typeStatus id="5489AAFDFFB9FFF1FB9BFF55FBA4FAA5" box="[1044,1146,1342,1366]" pageId="3" pageNumber="31" type="holotype">holotype</typeStatus>
. Wings hyaline with very light brown tinge, pterostigma light brown and small. Postnodal crossveins: FW 11, HW 9. Thorax similar to
<typeStatus id="5489AAFDFFB9FFF1FAEEFF0AFF15FA6D" pageId="3" pageNumber="31" type="holotype">holotype</typeStatus>
; as well as abundant occurrence of setae, metasternal tubercle present.
</paragraph>
<paragraph id="8B8D145FFFB9FFF1FF46FFC3FBC1F94B" blockId="3.[151,1436,1164,1969]" pageId="3" pageNumber="31">
<emphasis id="B946C84DFFB9FFF1FF46FFC3FE9BFA32" bold="true" box="[201,325,1448,1473]" pageId="3" pageNumber="31">Abdomen.</emphasis>
Venter of S110 yellowish brown/light ochre. S26 with a fine black line at intersegmental articulation. S2 with a black dorsal marking not reaching black line at dorsal half of intersegmental articulation. S35 with sword-like markings, formed by bands tapering at both sides and dorsally connected to a circular “hilt” which is connected to the transversal ring at the intersegmental articulations. Sword shape is clearest on S34 and black markings become wider from S3 to S5. S6 with a broad black band tapering anteriorly. S710 with a broad black band with a conspicuous, narrow azure middorsal band (
<figureCitation id="130908DAFFB9FFF1FCAEFC32FCA9F982" box="[801,887,1625,1649]" captionStart="FIGURE 10. P" captionStartId="8.[151,250,2006,2029]" captionTargetBox="[571,1002,843,1973]" captionTargetId="figure@8.[569,1017,828,1987]" captionTargetPageId="8" captionText="FIGURE 10. P. hoffmanni, abdomen (S 7 10) with appendages of female allotype, dorsal view." httpUri="https://zenodo.org/record/215134/files/figure.png" pageId="3" pageNumber="31">Fig. 10</figureCitation>
). As in
<typeStatus id="5489AAFDFFB9FFF1FC5AFC32FBE4F982" box="[981,1082,1625,1649]" pageId="3" pageNumber="31" type="holotype">holotype</typeStatus>
, posterior margin of S10 has a wedge-shaped mid-dorsal emargination. Cercus white, cone-shaped; paraproct white. S8 with vulvar spine, tip of ovipositor valve reaching distal margin of S10, stylus curved downward (
<figureCitation id="130908DAFFB9FFF1FC42FCCBFBCFF94B" box="[973,1041,1696,1720]" captionStart="FIGURE 9. P" captionStartId="8.[151,250,766,789]" captionTargetBox="[194,1420,195,722]" captionTargetId="figure@8.[151,1436,195,742]" captionTargetPageId="8" captionText="FIGURE 9. P. hoffmanni, tip of abdomen with appendages of female allotype, lateral view." httpUri="https://zenodo.org/record/215133/files/figure.png" pageId="3" pageNumber="31">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8B8D145FFFB9FFF1FF46FCAEFC23F92F" blockId="3.[151,1436,1164,1969]" box="[201,1021,1732,1757]" pageId="3" pageNumber="31">
<emphasis id="B946C84DFFB9FFF1FF46FCAEFEA1F92E" bold="true" box="[201,383,1733,1757]" pageId="3" pageNumber="31">Measurements.</emphasis>
HW
<quantity id="4CCAB9BAFFB9FFF1FE33FCAFFDFCF92F" box="[444,546,1732,1756]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="3" pageNumber="31" unit="mm" value="17.7">17.7 mm</quantity>
, abdomen
<quantity id="4CCAB9BAFFB9FFF1FD14FCAFFCDFF92F" box="[667,769,1732,1756]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="3" pageNumber="31" unit="mm" value="17.8">17.8 mm</quantity>
, total length
<quantity id="4CCAB9BAFFB9FFF1FC1DFCAFFC27F92F" box="[914,1017,1732,1756]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" pageId="3" pageNumber="31" unit="mm" value="24.5">24.5 mm</quantity>
.
</paragraph>
<paragraph id="8B8D145FFFB9FFF1FF46FC8CFD2FF8D0" blockId="3.[151,1436,1164,1969]" pageId="3" pageNumber="31">
<emphasis id="B946C84DFFB9FFF1FF46FC8CFE0DF8F3" bold="true" box="[201,467,1767,1792]" pageId="3" pageNumber="31">
Variation in
<typeStatus id="5489AAFDFFB9FFF1FED2FC83FE0DF8F3" box="[349,467,1768,1792]" pageId="3" pageNumber="31" type="paratype">paratypes</typeStatus>
</emphasis>
: As
<typeStatus id="5489AAFDFFB9FFF1FD88FC83FDB3F8F3" box="[519,621,1768,1792]" pageId="3" pageNumber="31" type="holotype">holotype</typeStatus>
; for variations in dimensions, see above and tables 1 and 2; for variations in number of postnodal crossveins, see tables 1 and 2.
</paragraph>
<paragraph id="8B8D145FFFB9FFF1FF46FD44FE2FF842" blockId="3.[151,1436,1164,1969]" pageId="3" pageNumber="31">
<emphasis id="B946C84DFFB9FFF1FF46FD44FEA1F8B4" bold="true" box="[201,383,1839,1863]" pageId="3" pageNumber="31">Measurements.</emphasis>
Males (n = 8, including
<typeStatus id="5489AAFDFFB9FFF1FD2DFD45FCD4F8B5" box="[674,778,1838,1862]" pageId="3" pageNumber="31" type="holotype">holotype</typeStatus>
): HW 15.018.3 (mean 16.6); abdomen 17.020.0 (mean 18.1); total length: 23.028.3 (mean 24.8). Females (n = 8; including allotype; one of the
<typeStatus id="5489AAFDFFB9FFF1FB0BFD38FB3AF898" box="[1156,1252,1875,1899]" pageId="3" pageNumber="31" type="paratype">paratype</typeStatus>
females lacking wings and therefore not taken into account): HW 15.519.2 (mean 17.7); abdomen 17.820.0 (mean 18.5); total length: 23.028.3 (mean 25.1).
</paragraph>
<caption id="DF4D44D7FFBEFFF6FF18FF44FADAFAB6" box="[151,1284,1327,1350]" httpUri="https://zenodo.org/record/215129/files/figure.png" pageId="4" pageNumber="32" targetBox="[529,1060,186,1281]" targetPageId="4">
<paragraph id="8B8D145FFFBEFFF6FF18FF44FADAFAB6" blockId="4.[151,1284,1327,1350]" box="[151,1284,1327,1350]" pageId="4" pageNumber="32">
<emphasis id="B946C84DFFBEFFF6FF18FF44FECCFAB7" bold="true" box="[151,274,1327,1350]" pageId="4" pageNumber="32">FIGURE 4.</emphasis>
<taxonomicName id="4C326FDCFFBEFFF6FE96FF5BFE47FAB6" box="[281,409,1327,1349]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="32" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="B946C84DFFBEFFF6FE96FF5BFE47FAB6" box="[281,409,1327,1349]" italics="true" pageId="4" pageNumber="32">P. hoffmanni</emphasis>
</taxonomicName>
, tip of abdomen of male holotype with genital opercula and appendages, ventral view.
</paragraph>
</caption>
<caption id="DF4D44D7FFBEFFF6FF18FDAEFC52F82F" box="[151,908,1989,2012]" httpUri="https://zenodo.org/record/215130/files/figure.png" pageId="4" pageNumber="32" targetBox="[234,1332,1414,1957]" targetPageId="4">
<paragraph id="8B8D145FFFBEFFF6FF18FDAEFC52F82F" blockId="4.[151,908,1989,2012]" box="[151,908,1989,2012]" pageId="4" pageNumber="32">
<emphasis id="B946C84DFFBEFFF6FF18FDAEFECCF828" bold="true" box="[151,274,1989,2012]" pageId="4" pageNumber="32">FIGURE 5.</emphasis>
<taxonomicName id="4C326FDCFFBEFFF6FE96FDACFE47F82F" box="[281,409,1990,2012]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="32" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="B946C84DFFBEFFF6FE96FDACFE47F82F" box="[281,409,1990,2012]" italics="true" pageId="4" pageNumber="32">P. hoffmanni</emphasis>
</taxonomicName>
, genital ligula, taken from paratype, lateral view.
</paragraph>
</caption>
<paragraph id="8B8D145FFFBFFFF7FF46FAF3FE3FFF5C" blockId="5.[151,1437,151,1508]" box="[201,481,151,176]" pageId="5" pageNumber="33">
<emphasis id="B946C84DFFBFFFF7FF46FAF3FEA6FF43" bold="true" box="[201,376,151,176]" pageId="5" pageNumber="33">Male F0 larva.</emphasis>
(
<figureCitation id="130908DAFFBFFFF7FE08FAFCFE06FF5C" box="[391,472,151,175]" captionStart="FIGURE 17. P" captionStartId="11.[151,250,1577,1600]" captionTargetBox="[168,1416,1156,1540]" captionTargetId="figure@11.[151,1436,1143,1556]" captionTargetPageId="11" captionText="FIGURE 17. P. hoffmanni, male FO-larva, lateral view." httpUri="https://zenodo.org/record/215141/files/figure.png" pageId="5" pageNumber="33">Fig. 17</figureCitation>
)
</paragraph>
<paragraph id="8B8D145FFFBFFFF7FF46FAD7FE3CFE97" blockId="5.[151,1437,151,1508]" pageId="5" pageNumber="33">
<emphasis id="B946C84DFFBFFFF7FF46FAD7FECEFF27" bold="true" box="[201,272,188,212]" pageId="5" pageNumber="33">Head.</emphasis>
Antennae with seven segments (
<figureCitation id="130908DAFFBFFFF7FD0FFAD7FD0DFF27" box="[640,723,188,212]" captionStart="FIGURE 18. P" captionStartId="12.[151,250,878,901]" captionText="FIGURE 18. P. hoffmanni, head of larva, dorsal view." pageId="5" pageNumber="33">Fig. 18</figureCitation>
). Prementum short and broad, articulation of pre- and postmentum reaching backwards behind procoxae; two medium-sized mental setae on each side of prementum ental surface; ligula entire, smoothly convex, and devoid of denticles (
<figureCitation id="130908DAFFBFFFF7FCDBFB6FFC76FEEF" box="[852,936,260,284]" captionStart="FIGURE 19. P" captionStartId="12.[151,250,1922,1945]" captionText="FIGURE 19. P. hoffmanni, prementum of larva, dorsal view." pageId="5" pageNumber="33">Fig. 19</figureCitation>
). Labial palp with two lobes and three large setae distal to movable hook and five small setae along outer border (
<figureCitation id="130908DAFFBFFFF7FC12FB4CFBF9FECC" box="[925,1063,295,319]" captionStart-0="FIGURE 19. P" captionStart-1="FIGURE 20. P" captionStartId-0="12.[151,250,1922,1945]" captionStartId-1="13.[151,250,972,995]" captionText-0="FIGURE 19. P. hoffmanni, prementum of larva, dorsal view." captionText-1="FIGURE 20. P. hoffmanni, prementum of larva, ventral view." pageId="5" pageNumber="33">Figs. 1920</figureCitation>
). Mandibular formula L 1234 y x a b / R 1234 y 0 a 0 (
<figureCitation id="130908DAFFBFFFF7FE0EFB27FE0BFE97" box="[385,469,332,356]" captionStart="FIGURE 21. P" captionStartId="13.[151,250,1974,1997]" captionTargetBox="[193,1395,1044,1957]" captionTargetId="figure@13.[178,1410,1044,1959]" captionTargetPageId="13" captionText="FIGURE 21. P. hoffmanni, mandibles of larva, ventrointernal view." httpUri="https://zenodo.org/record/215142/files/figure.png" pageId="5" pageNumber="33">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="8B8D145FFFBFFFF7FF46FB1BFC1FFE74" blockId="5.[151,1437,151,1508]" box="[201,961,367,392]" pageId="5" pageNumber="33">
<emphasis id="B946C84DFFBFFFF7FF46FB1BFEF7FE7B" bold="true" box="[201,297,368,392]" pageId="5" pageNumber="33">Thorax.</emphasis>
Wing sheaths reaching from middle to distal margin of S4.
</paragraph>
<paragraph id="8B8D145FFFBFFFF7FF46FBF8FA59FE07" blockId="5.[151,1437,151,1508]" pageId="5" pageNumber="33">
<emphasis id="B946C84DFFBFFFF7FF46FBF8FE9BFE5F" bold="true" box="[201,325,403,428]" pageId="5" pageNumber="33">Abdomen.</emphasis>
Lacking lateral spines; integument completely covered with small bristles. Caudal lamellae ovalshaped with tip forming a small point (
<figureCitation id="130908DAFFBFFFF7FDC1FBDCFD7DFE3C" box="[590,675,439,463]" captionStart="FIGURE 22. P" captionStartId="14.[151,250,788,811]" captionText="FIGURE 22. P. hoffmannii, median caudal lamella of larva." pageId="5" pageNumber="33">Fig. 22</figureCitation>
). Lateral and median lamellae shaped identically. Larvae uniformly brown without any definite pattern. Male gonapophyses as in
<figureCitation id="130908DAFFBFFFF7FCC4FBB7FC18FE07" box="[843,966,476,500]" captionStart-0="FIGURE 23. P" captionStart-1="FIGURE 24. P" captionStartId-0="14.[151,250,1305,1328]" captionStartId-1="14.[151,250,1982,2005]" captionText-0="FIGURE 23. P. hoffmanni, male gonapophyses of larva, lateral view." captionText-1="FIGURE 24. P. hoffmanni, male gonapophyses of larva, ventral view." pageId="5" pageNumber="33">Fig. 2324</figureCitation>
, female gonapophyses as in
<figureCitation id="130908DAFFBFFFF7FA89FBB7FA5CFE07" box="[1286,1410,476,500]" captionStart-0="FIGURE 25. P" captionStart-1="FIGURE 26. P" captionStartId-0="15.[151,250,652,675]" captionStartId-1="15.[151,250,1363,1386]" captionText-0="FIGURE 25. P. hoffmanni, female gonapophyses of larva, lateral view." captionText-1="FIGURE 26. P. hoffmanni, female gonapophyses of larva, ventral view." pageId="5" pageNumber="33">Fig. 2526</figureCitation>
.
</paragraph>
<paragraph id="8B8D145FFFBFFFF7FF46F86BFDD3FDAC" blockId="5.[151,1437,151,1508]" pageId="5" pageNumber="33">
<emphasis id="B946C84DFFBFFFF7FF46F86BFEA1FDEB" bold="true" box="[201,383,512,536]" pageId="5" pageNumber="33">Measurements.</emphasis>
Total length including caudal lamellae 18.4; head width 3.75; abdomen length 8.3; lateral lamellae length 3.5, width 2.1; median lamella length 3.25, width 2.25; hind femur length 3; hind tibia length 2.5; prementum length 2.6, width 2.3.
</paragraph>
<paragraph id="8B8D145FFFBFFFF7FF46F800FB98FCCB" blockId="5.[151,1437,151,1508]" pageId="5" pageNumber="33">
<emphasis id="B946C84DFFBFFFF7FF46F800FDCCFD77" bold="true" box="[201,530,619,644]" pageId="5" pageNumber="33">Variation in measurements.</emphasis>
The
<specimenCount id="9D34DFD6FFBFFFF7FDC4F807FD6FFD77" box="[587,689,620,644]" pageId="5" pageNumber="33" type="larva">19 larvae</specimenCount>
collected belonged to four different stadia: (1) Four larvae were in the F0 stadium with completely developed wing sheaths surpassing middle to end of S4, head width 3.43.75 (mean 3.56), total length incl. caudal lamellae 16.018.5 (mean 17.35); (2) five larvae with wing sheaths reaching end of S2, head width 2.62.8 (mean 2.66), total length 12.513.1 (mean 12.75); (3) nine larvae with wing sheaths reaching end of S1, head width 2.12.4 (mean 2.27) and total length of 9.612.4 (mean 11.0), and (4) one small larva with wing sheaths just reaching S1, head width of 1.8 and a total length of 7.7.
</paragraph>
</subSubSection>
<subSubSection id="C32847D4FFBFFFF7FF46F928FB12FA17" pageId="5" pageNumber="33" type="diagnosis">
<paragraph id="8B8D145FFFBFFFF7FF46F928FE5BFB53" blockId="5.[151,1437,151,1508]" pageId="5" pageNumber="33">
<emphasis id="B946C84DFFBFFFF7FF46F928FE9DFCAF" bold="true" box="[201,323,835,860]" pageId="5" pageNumber="33">Diagnosis.</emphasis>
<taxonomicName id="4C326FDCFFBFFFF7FEC3F92FFD8CFCAF" box="[332,594,836,860]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="B946C84DFFBFFFF7FEC3F92FFD8CFCAF" box="[332,594,836,860]" italics="true" pageId="5" pageNumber="33">Protallagma hoffmanni</emphasis>
</taxonomicName>
is distinguished from the only other species in this genus,
<taxonomicName id="4C326FDCFFBFFFF7FB7CF92EFAAEFCAF" box="[1267,1392,837,860]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="titicacae">
<emphasis id="B946C84DFFBFFFF7FB7CF92EFAAEFCAF" box="[1267,1392,837,860]" italics="true" pageId="5" pageNumber="33">P. titicacae</emphasis>
</taxonomicName>
, by several characters. While
<taxonomicName id="4C326FDCFFBFFFF7FE36F902FDEAFC73" box="[441,564,873,896]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="titicacae">
<emphasis id="B946C84DFFBFFFF7FE36F902FDEAFC73" box="[441,564,873,896]" italics="true" pageId="5" pageNumber="33">P. titicacae</emphasis>
</taxonomicName>
has extended black areas around the ocelli, in
<taxonomicName id="4C326FDCFFBFFFF7FBCEF902FB11FC73" box="[1089,1231,872,896]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="B946C84DFFBFFFF7FBCEF902FB11FC73" box="[1089,1231,872,896]" italics="true" pageId="5" pageNumber="33">P. hoffmanni</emphasis>
</taxonomicName>
black is restricted to the frontal suture and other sutures around the ocelli. Pale occipital spots are present in
<taxonomicName id="4C326FDCFFBFFFF7FBF2F9E6FB26FC57" box="[1149,1272,909,932]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="titicacae">
<emphasis id="B946C84DFFBFFFF7FBF2F9E6FB26FC57" box="[1149,1272,909,932]" italics="true" pageId="5" pageNumber="33">P. titicacae</emphasis>
</taxonomicName>
but missing in
<taxonomicName id="4C326FDCFFBFFFF7FF18F9DAFEFAFC3B" box="[151,292,944,968]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="B946C84DFFBFFFF7FF18F9DAFEFAFC3B" box="[151,292,944,968]" italics="true" pageId="5" pageNumber="33">P. hoffmanni</emphasis>
</taxonomicName>
(
<figureCitation id="130908DAFFBFFFF7FEBCF9C4FEAAFC34" box="[307,372,943,967]" captionStart="FIGURE 1. P" captionStartId="2.[151,250,1073,1096]" captionTargetBox="[153,1418,198,1054]" captionTargetId="figure@2.[151,1436,187,1060]" captionTargetPageId="2" captionText="FIGURE 1. P. hoffmanni, head of male holotype, dorsal view." httpUri="https://zenodo.org/record/215127/files/figure.png" pageId="5" pageNumber="33">Fig. 1</figureCitation>
). In both sexes, the pronotal hind lobe is undulate with a nozzle facing toward the head (
<figureCitation id="130908DAFFBFFFF7FADCF9C4FA49FC34" box="[1363,1431,943,967]" captionStart="FIGURE 2. P" captionStartId="2.[151,250,1758,1781]" captionTargetBox="[175,1406,1147,1743]" captionTargetId="figure@2.[163,1423,1139,1746]" captionTargetPageId="2" captionText="FIGURE 2. P. hoffmanni, pronotum and mesostigmal plate of male holotype, dorsal view." httpUri="https://zenodo.org/record/215128/files/figure.png" pageId="5" pageNumber="33">Fig. 2</figureCitation>
,
<figureCitation id="130908DAFFBFFFF7FF18F9BFFF07FC1F" box="[151,217,980,1004]" captionStart="FIGURE 8. P" captionStartId="7.[151,250,1736,1759]" captionTargetBox="[152,1419,1050,1708]" captionTargetId="figure@7.[151,1436,1030,1713]" captionTargetPageId="7" captionText="FIGURE 8. P. hoffmanni, pronotum and mesostigmal plate of female paratype." httpUri="https://zenodo.org/record/215132/files/figure.png" pageId="5" pageNumber="33">Fig. 8</figureCitation>
), whereas in
<taxonomicName id="4C326FDCFFBFFFF7FEE1F9BEFE34FC1F" box="[366,490,981,1004]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="titicacae">
<emphasis id="B946C84DFFBFFFF7FEE1F9BEFE34FC1F" box="[366,490,981,1004]" italics="true" pageId="5" pageNumber="33">P. titicacae</emphasis>
</taxonomicName>
it is bi- or trilobed. In lateral view, the male cercus of
<emphasis id="B946C84DFFBFFFF7FBC6F9BEFB0CFC1F" box="[1097,1234,980,1004]" italics="true" pageId="5" pageNumber="33">P. h o f f m a n n i</emphasis>
is mitten-shaped, with the thumb of the mitten very pronounced and long, overlapping the paraproct (Fig. 3). In comparison, the thumb of the mitten in males of
<taxonomicName id="4C326FDCFFBFFFF7FD88FE76FD5AFBC7" box="[519,644,1053,1076]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="titicacae">
<emphasis id="B946C84DFFBFFFF7FD88FE76FD5AFBC7" box="[519,644,1053,1076]" italics="true" pageId="5" pageNumber="33">P. titicacae</emphasis>
</taxonomicName>
is very small and not overlapping the paraproct. The number of postnodal crossveins as well as the overall number of veins and cells in both FW and HW of both sexes of
<taxonomicName id="4C326FDCFFBFFFF7FA80FE2AFA42FBAB" box="[1295,1436,1088,1112]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="B946C84DFFBFFFF7FA80FE2AFA42FBAB" box="[1295,1436,1088,1112]" italics="true" pageId="5" pageNumber="33">P. hoffmanni</emphasis>
</taxonomicName>
is significantly higher than in
<taxonomicName id="4C326FDCFFBFFFF7FE64FE0EFDB9FB8F" box="[491,615,1125,1148]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="titicacae">
<emphasis id="B946C84DFFBFFFF7FE64FE0EFDB9FB8F" box="[491,615,1125,1148]" italics="true" pageId="5" pageNumber="33">P. titicacae</emphasis>
</taxonomicName>
. More details and further characteristics are given in the discussion section (
<tableCitation id="C6B021E4FFBFFFF7FF5EFEE3FEEEFB53" box="[209,304,1160,1184]" captionStart="TABLE 1" captionStartId="19.[151,235,152,173]" captionTargetBox="[159,1422,283,1296]" captionTargetPageId="19" captionText="TABLE 1. Differences between males of Protallagma titicacace and P. hoffmanni." httpUri="http://table.plazi.org/id/DF4D44D7FFA9FFE1FF18FAF3FC02FF5D" pageId="5" pageNumber="33" tableUuid="DF4D44D7FFA9FFE1FF18FAF3FC02FF5D">Tables 1</tableCitation>
and
<tableCitation id="C6B021E4FFBFFFF7FEE8FEE3FEA6FB53" box="[359,376,1160,1184]" captionStart="TABLE 2" captionStartId="20.[151,235,332,353]" captionTargetBox="[159,1424,464,1079]" captionTargetPageId="20" captionText="TABLE 2. Differences between females of Protallagma titicacae and P. hoffmanni." httpUri="http://table.plazi.org/id/DF4D44D7FFAEFFE6FF18FB27FC3BFE91" pageId="5" pageNumber="33" tableUuid="DF4D44D7FFAEFFE6FF18FB27FC3BFE91">2</tableCitation>
).
</paragraph>
<paragraph id="8B8D145FFFBFFFF7FF46FEC7FE9EFAFF" blockId="5.[151,1437,151,1508]" pageId="5" pageNumber="33">
Common trait of the larvae of
<taxonomicName id="4C326FDCFFBFFFF7FDA4FEC7FCFFFB37" box="[555,801,1196,1220]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="titicacae">
<emphasis id="B946C84DFFBFFFF7FDA4FEC7FCFFFB37" box="[555,801,1196,1220]" italics="true" pageId="5" pageNumber="33">Protallagma titicacae</emphasis>
</taxonomicName>
described by
<bibRefCitation id="EFA369AEFFBFFFF7FC4BFEC7FB89FB37" author="Bulla" box="[964,1111,1196,1220]" pageId="5" pageNumber="33" refString="Bulla, L. A. (1972) La ninfa de Protallagma titicacae (Calvert) (Odonata, Coenagriidae). Neotropica, 18 (57), 129 - 133." type="journal article" year="1972">Bulla (1972)</bibRefCitation>
, drawings of which are also depicted in
<bibRefCitation id="EFA369AEFFBFFFF7FE94FEBBFE02FB1B" author="Heckman" box="[283,476,1232,1256]" pageId="5" pageNumber="33" refString="Heckman, C. W. (2008) Encyclopaedia of South American Aquatic Insects: Odonata - Zygoptera. Illustrated keys to known families, genera, and species in South America. Springer Science + Business Media B. V., VIII + 692 pp." type="book" year="2008">Heckman (2008)</bibRefCitation>
, and
<taxonomicName id="4C326FDCFFBFFFF7FD96FEBAFD71FB1B" box="[537,687,1232,1256]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="hotffmanni">
<emphasis id="B946C84DFFBFFFF7FD96FEBAFD71FB1B" box="[537,687,1232,1256]" italics="true" pageId="5" pageNumber="33">P. hotffmanni</emphasis>
</taxonomicName>
: a distinct median notch at the occipital margin (
<bibRefCitation id="EFA369AEFFBFFFF7FB52FEBBFEEDFAFF" author="Bulla" pageId="5" pageNumber="33" refString="Bulla, L. A. (1972) La ninfa de Protallagma titicacae (Calvert) (Odonata, Coenagriidae). Neotropica, 18 (57), 129 - 133." type="journal article" year="1972">
Bulla 1972: 130,
<figureCitation id="130908DAFFBFFFF7FF18FE9FFF05FAFF" box="[151,219,1268,1292]" captionStart="FIGURE 2. P" captionStartId="2.[151,250,1758,1781]" captionTargetBox="[175,1406,1147,1743]" captionTargetId="figure@2.[163,1423,1139,1746]" captionTargetPageId="2" captionText="FIGURE 2. P. hoffmanni, pronotum and mesostigmal plate of male holotype, dorsal view." httpUri="https://zenodo.org/record/215128/files/figure.png" pageId="5" pageNumber="33">Fig. 2</figureCitation>
;
<figureCitation id="130908DAFFBFFFF7FF68FE9FFEEDFAFF" box="[231,307,1268,1292]" captionStart="FIGURE 16. P" captionStartId="11.[151,250,1071,1094]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@11.[151,1436,193,1050]" captionTargetPageId="11" captionText="FIGURE 16. P. hoffmanni, female, dorsal view. Laguna Querococha, Feb. 18, 2007." httpUri="https://zenodo.org/record/215140/files/figure.png" pageId="5" pageNumber="33">Fig.16</figureCitation>
</bibRefCitation>
).
</paragraph>
<paragraph id="8B8D145FFFBFFFF7FF46FF73FB12FA17" blockId="5.[151,1437,151,1508]" pageId="5" pageNumber="33">
Differences between the larvae of the two species: Total length without caudal lamellae higher in
<taxonomicName id="4C326FDCFFBFFFF7FA88FF72FA4BFAC3" box="[1287,1429,1304,1328]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="B946C84DFFBFFFF7FA88FF72FA4BFAC3" box="[1287,1429,1304,1328]" italics="true" pageId="5" pageNumber="33">P. hoffmanni</emphasis>
</taxonomicName>
:
<quantity id="4CCAB9BAFFBFFFF7FF18FF57FEFAFAA7" box="[151,292,1340,1364]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.683" metricValueMin="2.667" pageId="5" pageNumber="33" unit="in" value="12.5" valueMax="14.5" valueMin="10.5">10.514.5 in</quantity>
<taxonomicName id="4C326FDCFFBFFFF7FEA5FF56FE7BFAA7" box="[298,421,1341,1364]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="titicacae">
<emphasis id="B946C84DFFBFFFF7FEA5FF56FE7BFAA7" box="[298,421,1341,1364]" italics="true" pageId="5" pageNumber="33">P. titicacae</emphasis>
</taxonomicName>
compared to 13.0
<quantity id="4CCAB9BAFFBFFFF7FD04FF57FD07FAA7" box="[651,729,1340,1364]" metricMagnitude="-1" metricUnit="m" metricValue="3.8608" pageId="5" pageNumber="33" unit="in" value="15.2">15.2 in</quantity>
<taxonomicName id="4C326FDCFFBFFFF7FD50FF56FCB3FAA7" box="[735,877,1340,1364]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="B946C84DFFBFFFF7FD50FF56FCB3FAA7" box="[735,877,1340,1364]" italics="true" pageId="5" pageNumber="33">P. hoffmanni</emphasis>
</taxonomicName>
. Length of caudal lamellae lower in
<emphasis id="B946C84DFFBFFFF7FA88FF56FA4EFAA7" box="[1287,1424,1340,1364]" italics="true" pageId="5" pageNumber="33">P. h o f f m a n n i</emphasis>
:
<quantity id="4CCAB9BAFFBFFFF7FF18FF0BFED7FA8B" box="[151,265,1376,1400]" metricMagnitude="-1" metricUnit="m" metricValue="1.0922000000000003" metricValueMax="1.1176000000000001" metricValueMin="1.0668000000000002" pageId="5" pageNumber="33" unit="in" value="4.3" valueMax="4.4" valueMin="4.2">4.24.4 in</quantity>
<taxonomicName id="4C326FDCFFBFFFF7FE9DFF0AFE50FA8B" box="[274,398,1377,1400]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="titicacae">
<emphasis id="B946C84DFFBFFFF7FE9DFF0AFE50FA8B" box="[274,398,1377,1400]" italics="true" pageId="5" pageNumber="33">P. titicacae</emphasis>
</taxonomicName>
(not specified whether median or lateral) compared to 3.5 (lateral) and 3.25 (median) in
<taxonomicName id="4C326FDCFFBFFFF7FA08FF0AFED7FA6F" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="B946C84DFFBFFFF7FA08FF0AFED7FA6F" italics="true" pageId="5" pageNumber="33">P. hoffmanni</emphasis>
</taxonomicName>
. Whereas the caudal lamellae in
<taxonomicName id="4C326FDCFFBFFFF7FDF6FFEEFCD8FA6F" box="[633,774,1412,1436]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="B946C84DFFBFFFF7FDF6FFEEFCD8FA6F" box="[633,774,1412,1436]" italics="true" pageId="5" pageNumber="33">P. hoffmanni</emphasis>
</taxonomicName>
(
<figureCitation id="130908DAFFBFFFF7FC9BFFEFFCBBFA6F" box="[788,869,1412,1436]" captionStart="FIGURE 22. P" captionStartId="14.[151,250,788,811]" captionText="FIGURE 22. P. hoffmannii, median caudal lamella of larva." pageId="5" pageNumber="33">Fig. 22</figureCitation>
) are short (3.5), oval-shaped and bearing a distinct small point, the caudal lamellae of
<taxonomicName id="4C326FDCFFBFFFF7FDAAFFC2FD7CFA33" box="[549,674,1449,1472]" class="Insecta" family="Coenagrionidae" genus="Protallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="titicacae">
<emphasis id="B946C84DFFBFFFF7FDAAFFC2FD7CFA33" box="[549,674,1449,1472]" italics="true" pageId="5" pageNumber="33">P. titicacae</emphasis>
</taxonomicName>
(no indication given whether lateral or median is depicted in
<bibRefCitation id="EFA369AEFFBFFFF7FAD0FFC3FF0DFA17" author="Bulla" pageId="5" pageNumber="33" refString="Bulla, L. A. (1972) La ninfa de Protallagma titicacae (Calvert) (Odonata, Coenagriidae). Neotropica, 18 (57), 129 - 133." type="journal article" year="1972">Bulla 1972</bibRefCitation>
: 130,
<figureCitation id="130908DAFFBFFFF7FE9AFFA7FE86FA17" box="[277,344,1484,1508]" captionStart="FIGURE 6. P" captionStartId="6.[151,250,1571,1594]" captionText="FIGURE 6. P. hoffmanni, genital ligula, taken from paratype, ectal view." pageId="5" pageNumber="33">Fig. 6</figureCitation>
) are longer (4.24.4), lanceolate, tapering distally and forming an elongate tip.
</paragraph>
</subSubSection>
</treatment>
</document>