692 lines
76 KiB
XML
692 lines
76 KiB
XML
<document ID-CLB-Dataset="7319" ID-DOI="http://dx.doi.org/10.3897/zookeys.1031.62181" ID-GBIF-Dataset="7cf05e72-a279-4e29-82eb-7be8499a4eb3" ID-PMC="PMC8062429" ID-Pensoft-Pub="1313-2970-1031-143" ID-Pensoft-UUID="9FB8610C17435068844C1BEBF3EE409F" ID-PubMed="33958911" ID-ZooBank="85A6CB62A3504F16954BF3E1D657CC02" ModsDocID="1313-2970-1031-143" checkinTime="1618544357638" checkinUser="pensoft" docAuthor="Brescovit, Antonio D., Zampaulo, Robson de A. & Cizauskas, Igor" docDate="2021" docId="4FE7FB7679865ABAB75606F44AE07506" docLanguage="en" docName="ZooKeys 1031: 143-159" docOrigin="ZooKeys 1031" docPubDate="2021-04-15" docSource="http://dx.doi.org/10.3897/zookeys.1031.62181" docTitle="Ochyrocera ritxoco Brescovit & Zampaulo & Cizauskas 2021, sp. nov." docType="treatment" docUuid="3A17CF00-CFE6-482A-89B7-2CF243171883" docUuidSource="ZooBank" docVersion="6" id="9FB8610C17435068844C1BEBF3EE409F" lastPageNumber="143" masterDocId="9FB8610C17435068844C1BEBF3EE409F" masterDocTitle="The first two blind troglobitic spiders of the genus Ochyrocera from caves in Floresta Nacional de Carajas, state of Para, Brazil (Araneae, Ochyroceratidae)" masterLastPageNumber="159" masterPageNumber="143" pageNumber="143" updateTime="1732846385162" updateUser="ExternalLinkService">
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<mods:mods id="D6F959A9F008D6EE699126734B9A68C3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F6373C103AC1C436DADF06596DE99190">
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<mods:title id="70DC92D01C3C962242CF640A41C836D5">The first two blind troglobitic spiders of the genus Ochyrocera from caves in Floresta Nacional de Carajas, state of Para, Brazil (Araneae, Ochyroceratidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="E2FA50D30553B6122ECBDDD63FD9EF1F" type="personal">
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<mods:role id="A300A856465ABD99B63E062243226B08">
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<mods:roleTerm id="3AE01947D3D2FFEF4BEB98C9A0081E81">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C94749B7EF249C666C6D18CD4D49BC2C">Brescovit, Antonio D.</mods:namePart>
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<mods:nameIdentifier id="D5106BA055E4173FE529443C279DBBB4" type="ORCID">https://orcid.org/0000-0002-1511-5324</mods:nameIdentifier>
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<mods:affiliation id="717B529AA865077CC746DF7C0AD98966">Laboratorio de Colecoes Zoologicas, Instituto Butantan, Av. Vital Brasil, 1500, Butanta, Sao Paulo, 05503 - 900, Brazil</mods:affiliation>
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<mods:nameIdentifier id="2DC263354E99E17C3A84945F35BDCA84" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
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</mods:name>
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<mods:name id="2F0A35104080478E4BF404B02FC95706" type="personal">
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<mods:role id="FFE217F5BDB092C984F7BB1A32BF26CE">
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<mods:roleTerm id="14C0AB7F2DA577C08F0858A1EBB07579">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C12D601418F7C3F66924380741BDCD59">Zampaulo, Robson de A.</mods:namePart>
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<mods:affiliation id="5D92074A699F3228BBEAC3D6B2E77F47">Gerencia de Licenciamento Ambiental e Espeleologia, Vale SA, Avenida Doutor Marco Paulo Simon Jardim, 34006 - 200 Nova Lima, Minas Gerais, Brazil</mods:affiliation>
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</mods:name>
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<mods:name id="A9DB90E8E6D73CC5A9BFE6E9C1924A43" type="personal">
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<mods:roleTerm id="30C373F94AFC2F9392C4D1913E32D9FB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="92C1B0354AF17CAAE6417EF8EBB7AF41">Cizauskas, Igor</mods:namePart>
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<mods:affiliation id="250FE54B5626E34FD66F8A3BB36E09DE">Organizacao de Apoio a Pesquisa da Biodiversidade (OAPBio), Rua Frei Inacio da 238, 05362 - 040, Sao Paulo, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="B2D64CA7ED8411FD21361C3B44B8F237">text</mods:typeOfResource>
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<mods:title id="BCAA7D3842D6382787F299E0B896FC9A">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="9D3F3EB8DCF11AC07BD6209CC96A1499">
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<mods:date id="302A6D8F759054338AD786F48FB4D201">2021</mods:date>
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<mods:detail id="6D000B0ABDAFD7EC4DE5CD7B42D778B1" type="pubDate">
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<mods:number id="43662C9A7752CF47F8C0AE1A3D137ADC">2021-04-15</mods:number>
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</mods:detail>
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<mods:detail id="D00839961E31822FA43E947311811A0E" type="volume">
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<mods:number id="3F4BD8F9FB641012195B9CD6FBD180FB">1031</mods:number>
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<mods:start id="C2FD52B5BAF47281156E50BD6C9E51AC">143</mods:start>
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<mods:end id="25A0A7C09DA1D4051C309F079457B88F">159</mods:end>
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</mods:extent>
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<mods:location id="A81B45EBF6EAA2022C29D8FEAE44C763">
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<mods:url id="F44123C095515DA3980D896D06050AB3">http://dx.doi.org/10.3897/zookeys.1031.62181</mods:url>
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</mods:location>
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<mods:classification id="CC9486B644CD4A172241A85ECF7C9AED">journal article</mods:classification>
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<mods:identifier id="7C2B9FC75C6948DE692A80B433F729C6" type="DOI">http://dx.doi.org/10.3897/zookeys.1031.62181</mods:identifier>
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<mods:identifier id="2AF47346B7E45A46E3C69CA4BB3EC933" type="Pensoft-Pub">1313-2970-1031-143</mods:identifier>
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<mods:identifier id="9C53800B81A2BE2A21AC23C136DFCFA8" type="ZooBank">85A6CB62A3504F16954BF3E1D657CC02</mods:identifier>
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<mods:identifier id="EA385134B7635A5391C3007E00623A9C" type="Pensoft-UUID">9FB8610C17435068844C1BEBF3EE409F</mods:identifier>
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</mods:mods>
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<treatment id="4FE7FB7679865ABAB75606F44AE07506" ID-GBIF-Taxon="181019329" LSID="urn:lsid:zoobank.org:act:3A17CF00-CFE6-482A-89B7-2CF243171883" httpUri="http://treatment.plazi.org/id/4FE7FB7679865ABAB75606F44AE07506" lastPageNumber="143" pageId="0" pageNumber="143" scope_family="Ochyroceratidae" scope_order="Araneae">
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<subSubSection id="9BF60B9915C00FE2B2081E17E9803AA9" pageId="0" pageNumber="143" type="nomenclature">
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<paragraph id="40033F51D71C43833592A5D3DE833650" pageId="0" pageNumber="143">
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<taxonomicName id="F8A753E89A3B83AB58FAD77D63193CC8" ID-CoL="98YXG" LSID="http://zoobank.org/3A17CF00-CFE6-482A-89B7-2CF243171883" authority="Brescovit & Zampaulo & Cizauskas, 2021" authorityName="Brescovit & Zampaulo & Cizauskas" authorityYear="2021" class="Arachnida" family="Ochyroceratidae" genus="Ochyrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ochyrocera ritxoco" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ritxoco" status="sp. nov.">Ochyrocera ritxoco</taxonomicName>
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<taxonomicNameLabel id="A8C97157E4CC0A99AFF312D3F9BB5165" pageId="0" pageNumber="143">sp. nov.</taxonomicNameLabel>
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<figureCitation id="3195A86DEE6DF24C16F2649C6D6BDE30" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ochyrocera ritxoco sp. nov., male IBSP 186160 (A, C-F), female IBSP 186146 (B) A, B habitus, dorsal view C left male palp, retrolateral view D same, prolateral view E right male palp, ventral view F same, dorsal view. Abbreviations: C = cymbium, CE = cymbial extension, E = embolus, T = tegulum." figureDoi="10.3897/zookeys.1031.62181.figure1" httpUri="https://binary.pensoft.net/fig/532001" pageId="0" pageNumber="143">Figs 1</figureCitation>
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<figureCitation id="0ECA3D2E7FD71DE5D788E1D9ECA3B1AA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ochyrocera ritxoco sp. nov. (A, B) A left male palp IBSP 186149, retrolateral view B same, prolateral view (C, D) C female genitalia IBSP 186149, enzyme cleared, dorsal view D same, dorsal view. Abbreviations: C = cymbium, CUE = columnar uterus externus, E = embolus, NUE = neck of uterus externus, PP = pore-plate, SP = spermathecae, T = tegulum, UE = uterus externus." figureDoi="10.3897/zookeys.1031.62181.figure2" httpUri="https://binary.pensoft.net/fig/532002" pageId="0" pageNumber="143">, 2</figureCitation>
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<figureCitation id="D945BAA46007FDEEF5A53346B236D09F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Ochyrocera ritxoco sp. nov., male IBSP 260307 (A-F) A carapace, dorsal view B chelicerae, frontal view C endites and labium, ventral view D crosier-like macrosetae (arrow, detail) E epiandrous area, abdomen, ventral view F male palp, tibia (arrows, trichobothria), dorsal view." figureDoi="10.3897/zookeys.1031.62181.figure3" httpUri="https://binary.pensoft.net/fig/532003" pageId="0" pageNumber="143">, 3</figureCitation>
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<figureCitation id="9D81B371A8986063FA158EDB21A11D21" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of Ochyrocera ritxoco sp. nov., male IBSP 260307 (A-G) A cymbium, retrolateral view B same, basal macrosetae, retrolateral view C same, tarsal organ, detail, retrolateral view D same, apex detail E-G male palp, arrows indicating the embolus E retrolateral view F same, prolateral view G same, frontal view." figureDoi="10.3897/zookeys.1031.62181.figure4" httpUri="https://binary.pensoft.net/fig/532004" pageId="0" pageNumber="143">, 4</figureCitation>
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<figureCitation id="234E602CC4F80EFA3525A8B81830D90A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM images of Ochyrocera ritxoco sp. nov., female IBSP 260307 (A-F) A carapace, dorsal view B chelicerae, frontal view C colulus, ventral view D pedipalp, distal, prolateral view E same, tarsal organ F leg IV, claw, prolateral view." figureDoi="10.3897/zookeys.1031.62181.figure5" httpUri="https://binary.pensoft.net/fig/532005" pageId="0" pageNumber="143">, 5</figureCitation>
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<figureCitation id="7CEF0994883737B3B341E25260FC298E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Ochyrocera ritxoco sp. nov., female in webs under rocks on the ground in the Cave N 4 WS _ 0067." figureDoi="10.3897/zookeys.1031.62181.figure6" httpUri="https://binary.pensoft.net/fig/532006" pageId="0" pageNumber="143">, 6</figureCitation>
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<figureCitation id="323FD3BA63F57E20A036627A4B529914" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution map of Ochyrocera ritxoco sp. nov., yellow triangles, and Ochyrocera ritxoo sp. nov., red plus signs, in FLONA de Carajas, Para, Brazil." figureDoi="10.3897/zookeys.1031.62181.figure10" httpUri="https://binary.pensoft.net/fig/532010" pageId="0" pageNumber="143">, 10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="8EB9643CE7171B45EC24BA14034E4FBF" pageId="0" pageNumber="143" type="type material">
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<paragraph id="D3852B210F2EA110A3679A9EB4C79E42" pageId="0" pageNumber="143">Type material.</paragraph>
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<paragraph id="50F3CAFEA3D7CDAC90E8ECDCB0D081FD" pageId="0" pageNumber="143">
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<emphasis id="B4FB3A73C76D5A4F70AA474AC673FC82" bold="true" italics="true" pageId="0" pageNumber="143">Holotype</emphasis>
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male from Cave N1_0103 (GEM-1301 or
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<normalizedToken id="75C9C2975E73DC8E0815F2B2F691B4D6" originalValue="Cipó">Cipo</normalizedToken>
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cave;
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<geoCoordinate id="BD5FD18899E2507F38DE75CE8A8ABD1E" degrees="6" direction="south" minutes="0" orientation="latitude" precision="15" seconds="13" value="-6.003611">6°0'13"S</geoCoordinate>
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,
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<geoCoordinate id="BAFE6664045EA86220731344D1653ED6" degrees="50" direction="west" minutes="17" orientation="longitude" precision="15" seconds="55" value="-50.29861">50°17'55"W</geoCoordinate>
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), FLONA de
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<normalizedToken id="944F374FA0B5266E0314051324423B3A" originalValue="Carajás">Carajas</normalizedToken>
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, Parauapebas,
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<normalizedToken id="76C960B7F0D55C0260C424F6CF0C4066" originalValue="Pará">Para</normalizedToken>
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, Brazil, 28/XI-03/X/2007, R. Andrade et al. col. (IBSP 115497).
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<emphasis id="7FB61B9D6D273D73C9D0B8526857D8CC" bold="true" italics="true" pageId="0" pageNumber="143">Paratype</emphasis>
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female from Cave N1_0075 (GEM_1273 or Piranha cave;
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<geoCoordinate id="BF35DEF9F2DBD223C4BC8FBCDDDE170D" degrees="6" direction="south" minutes="1" orientation="latitude" precision="15" seconds="14" value="-6.020556">6°1'14"S</geoCoordinate>
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,
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<geoCoordinate id="AF4F74FF4C57CF25BEF9A1C9A02D9199" degrees="50" direction="west" minutes="16" orientation="longitude" precision="15" seconds="49" value="-50.280277">50°16'49"W</geoCoordinate>
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), FLONA de
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<normalizedToken id="99EFB0DF4EF5D7F7B139BA520D2CEB53" originalValue="Carajás">Carajas</normalizedToken>
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, Parauapebas,
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<normalizedToken id="073E09DCACD3504F8EFDF9D1D973FBA5" originalValue="Pará">Para</normalizedToken>
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, Brazil, 28/XI-03/X/2007, R. Andrade et al. col. (IBSP 115499).
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</paragraph>
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</subSubSection>
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<subSubSection id="2B4240A2AF581D3CBDC30D7CB831B8D7" pageId="0" pageNumber="143" type="materials_examined">
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<paragraph id="8668900CEED91D1CC453350D1149621E" pageId="0" pageNumber="143">Other material examined.</paragraph>
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<paragraph id="B5436FC219EB86422F019ED4D2B28C90" pageId="0" pageNumber="143">
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<materialsCitation id="38309B380DB0ACD81086FEF879F40D09" ID-GBIF-Occurrence="3075953301" collectingDate="2014-06-11" collectingDateMax="2014-07-02" collectingDateMin="2014-06-11" collectorName="Parauapebas, de Carajas, Cave N" country="Brazil" latitude="-6.033889" location="Para" longLatPrecision="21" longitude="-50.27111" specimenCount="1" stateProvince="Para">
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<collectingCountry id="A6CB8F71064C976D9808C8017B126C2B" name="Brazil">Brazil</collectingCountry>
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.
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<normalizedToken id="7098CF62BC287C4B04051B050433B43C" originalValue="Pará">
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<collectingRegion id="B3DDCA6CA879A860A51940288FB0664B" country="Brazil" name="Para">Para</collectingRegion>
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</normalizedToken>
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:
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<collectorName id="3AFECD078B26C7EAA0DDCD968BF6E384">Parauapebas</collectorName>
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, FLONA
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<collectorName id="4CFF035865D3A1F9DD97F1E34A4E0770">
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de
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<normalizedToken id="F7E2C2B22FB616E032E1C612E6BEB1EA" originalValue="Carajás">Carajas</normalizedToken>
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</collectorName>
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,
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<collectorName id="51C8613498640CC11B884BC973C21B48">Cave N</collectorName>
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1_0015 (GEM-1211) (
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<geoCoordinate id="5002FB94D231819F1750DC23CB8517FB" degrees="6" direction="south" minutes="2" orientation="latitude" precision="15" seconds="2" value="-6.033889">6°2'2"S</geoCoordinate>
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,
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<geoCoordinate id="637888B4F4143EDBE946C27B52B49767" degrees="50" direction="west" minutes="16" orientation="longitude" precision="15" seconds="16" value="-50.27111">50°16'16"W</geoCoordinate>
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),
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<collectingDate id="D4E4BE205A839A195A32E19C6CD2FB67" value="2014-06-11" valueMax="2014-07-02" valueMin="2014-06-11">11/VI-02/VII/2014</collectingDate>
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</materialsCitation>
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,
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<specimenCount id="83632233DFF17A59AC40DDF9D76F02C0" type="female">1♀</specimenCount>
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(IBSP 186123);
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<specimenCount id="CEDE08906DCF96711208AD87FEEF3431" type="female">1♀</specimenCount>
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(IBSP 186124);
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<specimenCount id="604D7E17A2C7E6306C8E7730CD68E898" type="male">1♂</specimenCount>
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(IBSP 186125);
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<specimenCount id="3902D89E8F3690C0C29CA43D22A552ED" type="female">2♀</specimenCount>
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(IBSP 186126); Cave N1_0016 (GEM-1212) (
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<geoCoordinate id="7AF0819A38A905CFF3BF6575821576E9" degrees="6" direction="south" minutes="1" orientation="latitude" precision="15" seconds="10" value="-6.0194445">6°1'10"S</geoCoordinate>
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,
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<geoCoordinate id="E15840B9A44010752DFB2E104135AEAC" degrees="50" direction="west" minutes="16" orientation="longitude" precision="15" seconds="41" value="-50.278057">50°16'41"W</geoCoordinate>
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),
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<date id="40A74CD288FAF1B12E87507CB591A2C8" value="2014-09-04" valueMax="2014-10-06" valueMin="2014-09-04">04/IX-06/X/2014</date>
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,
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<specimenCount id="4E888CAD8737929C93F3847221B7C863" type="female">1♀</specimenCount>
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(IBSP 186130);
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<date id="EED1E9D347AD8FE849ABD3BE404473F2" value="2015-04-02" valueMax="2015-04-29" valueMin="2015-04-02">02-29/IV/2015</date>
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,
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<specimenCount id="4434FE340B77584C544EF6253E5946E6" type="female">2♀</specimenCount>
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(IBSP 186153);
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<specimenCount id="97CB70E6BD80D9954B51B67F5B425335" type="female">3♀</specimenCount>
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(IBSP 186154);
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<specimenCount id="D85FFD89B0721843808FDCFE276FF83A" type="female">1♀</specimenCount>
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(IBSP 186155); Cave N1_0055 (GEM-1253) (
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<geoCoordinate id="8E7DE964BAF03559695872243EAF0467" degrees="6" direction="south" minutes="1" orientation="latitude" precision="15" seconds="12" value="-6.0200005">6°1'12"S</geoCoordinate>
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,
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<geoCoordinate id="A4C4C23837D3F6307DDF44CEC9270C7F" degrees="50" direction="west" minutes="16" orientation="longitude" precision="15" seconds="43" value="-50.27861">50°16'43"W</geoCoordinate>
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),
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<date id="6DE9BCB7319D6278A9400BD21229AEE2" value="2015-01-07" valueMax="2015-01-28" valueMin="2015-01-07">07-28/I/2015</date>
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,
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<specimenCount id="80C4E711D3547CB841368E84D8B67C9D" type="female">1♀</specimenCount>
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(MPEG 37086, ex IBSP 186147); Cave N1_0056 (GEM-1254) (
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<geoCoordinate id="95799CE8574F27817DD4B6D9E0585268" degrees="6" direction="south" minutes="1" orientation="latitude" precision="15" seconds="11" value="-6.0197225">6°1'11"S</geoCoordinate>
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,
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<geoCoordinate id="AA2067079EAED4A5F2D4648E99A4F647" degrees="50" direction="west" minutes="16" orientation="longitude" precision="15" seconds="44" value="-50.27889">50°16'44"W</geoCoordinate>
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),
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<date id="D4932D2046D1C3D2C52E9CA668EA2A7C" value="2015-01-07" valueMax="2015-01-28" valueMin="2015-01-07">07-28/I/2015</date>
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,
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<specimenCount id="19438779E677C9E03B2065A11A4456A6" type="male">1♂</specimenCount>
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(MPEG 37087, ex IBSP 186148);
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<specimenCount id="56695190699149461F4C66E685BC01EF" type="male">1♂</specimenCount>
|
|
<specimenCount id="0515123334975F623BFA9956462A9D74" type="female">1♀</specimenCount>
|
|
(IBSP 186149); Cave N1_0060 (GEM-1258) (
|
|
<geoCoordinate id="5B73FCD95F6062B33540BF794B9F20F7" degrees="6" direction="south" minutes="1" orientation="latitude" precision="15" seconds="12" value="-6.0200005">6°1'12"S</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="90D6BC69D0B0FC1084F5FC061483EACF" degrees="50" direction="west" minutes="16" orientation="longitude" precision="15" seconds="41" value="-50.278057">50°16'41"W</geoCoordinate>
|
|
),
|
|
<date id="074C89EC5D465E44E03098EAF7982A01" value="2014-06-11" valueMax="2014-07-02" valueMin="2014-06-11">11/VI-02/VII/2014</date>
|
|
,
|
|
<specimenCount id="B62EB0B4F96DEF5484F7846393FBA40C" type="female">1♀</specimenCount>
|
|
(IBSP 186127);
|
|
<specimenCount id="468E2B6CC0FBD4B40ED5E5F991A8A454" type="female">1♀</specimenCount>
|
|
(IBSP 186128);
|
|
<date id="9CABE8A2544B19F783C8332B8577CC45" value="2015-01-07" valueMax="2015-01-28" valueMin="2015-01-07">07-28/I/2015</date>
|
|
,
|
|
<specimenCount id="9223729C92384125180DEC83B0D79895" type="female">1♀</specimenCount>
|
|
(IBSP 186150);
|
|
<specimenCount id="DB501137D641FAC8595821FB30016231" type="female">1♀</specimenCount>
|
|
(IBSP 186151);
|
|
<specimenCount id="B1F34830B8924FCB7CB6FE6070FEF79B" type="male">1♂</specimenCount>
|
|
<specimenCount id="6FBF5E62B6DD12AA48EBC931E0F906F3" type="female">1♀</specimenCount>
|
|
(MPEG 37088, ex IBSP 186152); Cave N1_0062 (GEM-1260) (
|
|
<geoCoordinate id="9D8524507ECBFDC0A892EC05377CB4ED" degrees="6" direction="south" minutes="1" orientation="latitude" precision="15" seconds="10" value="-6.0194445">6°1'10"S</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="92B9FEEEFA5776BC7ED7587A7D447FD2" degrees="50" direction="west" minutes="16" orientation="longitude" precision="15" seconds="44" value="-50.27889">50°16'44"W</geoCoordinate>
|
|
),
|
|
<date id="BAFC00EB0721D8BF760531BD4804692D" value="2014-09-04" valueMax="2014-10-06" valueMin="2014-09-04">04/IX-06/X/2014</date>
|
|
,
|
|
<specimenCount id="B1C26C31D03519E845720AC0EAAA2AB9" type="female">1♀</specimenCount>
|
|
(IBSP 186131);
|
|
<specimenCount id="9133D77447798D42592C4F7AF3E2513A" type="female">1♀</specimenCount>
|
|
(IBSP 186132);
|
|
<specimenCount id="BFFC0959B932C1B22C426883B7B8A2AB" type="female">2♀</specimenCount>
|
|
(IBSP 186133);
|
|
<specimenCount id="1790A535595D84CBE40BA63566F1050D" type="female">1♀</specimenCount>
|
|
(IBSP 186134);
|
|
<specimenCount id="9AFAAC9089C27F2525EFC267D8735855" type="male">1♂</specimenCount>
|
|
<specimenCount id="674489CD9DDFC5042C865191BBCDA877" type="female">2♀</specimenCount>
|
|
(IBSP 186135);
|
|
<date id="FBACD123A8B9EE81D0E70EFFA6ADDB4C" value="2015-04-02" valueMax="2015-04-29" valueMin="2015-04-02">02-29/IV/2015</date>
|
|
,
|
|
<specimenCount id="8F93F03A8D8CC365207F9B213031DE3E" type="female">2♀</specimenCount>
|
|
(IBSP 186157);
|
|
<specimenCount id="6CA7DD7C8BCD770CD50BAAAD5A6BEB21" type="male">3♂</specimenCount>
|
|
<specimenCount id="798ECD8FF37F425D7DC944C394AA077D" type="female">1♀</specimenCount>
|
|
(IBSP 186156);
|
|
<specimenCount id="71C25FDCBFE4C5FAF2822B8B9B2A5DD7" type="female">1♀</specimenCount>
|
|
(IBSP 186158); Cave N1_0073 (GEM-1271) (
|
|
<geoCoordinate id="488738162807C8CB1BD39E4B3C7E7D95" degrees="6" direction="south" minutes="1" orientation="latitude" precision="15" seconds="13" value="-6.020278">6°1'13"S</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="5144805A4BDE21BAC1EA6AADEEBBF8FD" degrees="50" direction="west" minutes="17" orientation="longitude" precision="15" seconds="17" value="-50.288055">50°17'17"W</geoCoordinate>
|
|
),
|
|
<date id="B581B6344EE14053A40ECC0502AC0A0B" value="2015-04-02" valueMax="2015-04-29" valueMin="2015-04-02">02-29/IV/2015</date>
|
|
,
|
|
<specimenCount id="06FD4111D321D78B9B29C87DA916691F" type="female">1♀</specimenCount>
|
|
(IBSP 186159);
|
|
<specimenCount id="2F792AC3A146111F2CF0E1DBD1305F09" type="male">1♂</specimenCount>
|
|
(IBSP 186160);
|
|
<specimenCount id="BF722F10726C7B03F7607BB2FE123AFE" type="female">1♀</specimenCount>
|
|
(IBSP 186161);
|
|
<specimenCount id="BB1353B052F7810C9ECC6597EE17613A" type="female">1♀</specimenCount>
|
|
(IBSP 186162); Cave N1_0084 (GEM-1282) (
|
|
<geoCoordinate id="E59F4E79F01AD2F044F775663F31AE45" degrees="6" direction="south" minutes="1" orientation="latitude" precision="15" seconds="7" value="-6.0186114">6°1'7"S</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="B9C6A4AA13E784B32C0300395078DD60" degrees="50" direction="west" minutes="17" orientation="longitude" precision="15" seconds="1" value="-50.28361">50°17'1"W</geoCoordinate>
|
|
),
|
|
<date id="89B31B67A32BAAE1AD3B8382FE958091" value="2014-06-11" valueMax="2014-07-02" valueMin="2014-06-11">11/VI-02/VII/2014</date>
|
|
,
|
|
<specimenCount id="9024B544F527BBA7C7913F5C2F455D0D" type="female">1♀</specimenCount>
|
|
(IBSP 186129); Cave N1_0101 (GEM-1299) (
|
|
<geoCoordinate id="96423CB8A266E19A345BB41F8ED6B0ED" degrees="6" direction="south" minutes="1" orientation="latitude" precision="15" seconds="9" value="-6.019167">6°1'9"S</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="8169632D0EA09E04AF082E29C5021054" degrees="50" direction="west" minutes="16" orientation="longitude" precision="15" seconds="46" value="-50.279446">50°16'46"W</geoCoordinate>
|
|
),
|
|
<date id="068AB8030BB7BF337110DFEFC157667F" value="2014-09-04" valueMax="2014-10-06" valueMin="2014-09-04">04/IX-06/X/2014</date>
|
|
,
|
|
<specimenCount id="312F08CC3B5B143863EDF07397CAF067" type="male">1♂</specimenCount>
|
|
(IBSP 186136);
|
|
<specimenCount id="DDEC567DDED0738C0C69A6C6F7CC8615" type="female">1♀</specimenCount>
|
|
(IBSP 186137);
|
|
<specimenCount id="3F1B3687AABB786C09227324C9BD1B5E" type="female">1♀</specimenCount>
|
|
(IBSP 186138);
|
|
<specimenCount id="5E33AD610FFAAB9E786382A07666E0E8" type="male">1♂</specimenCount>
|
|
(IBSP 186139);
|
|
<specimenCount id="EB8702BF69CA45D74D69C5775A55260D" type="female">2♀</specimenCount>
|
|
(IBSP 186140);
|
|
<specimenCount id="37B893442837E370B8378D5B1A699D0C" type="male">1♂</specimenCount>
|
|
(IBSP 186141);
|
|
<specimenCount id="3FA7AA86393470DEE5D34E3CFDA1A9C3" type="male">1♂</specimenCount>
|
|
(IBSP 186142); Cave N1_0240 (
|
|
<geoCoordinate id="4087D704156EAE938FDF2F1251539696" degrees="6" direction="south" minutes="1" orientation="latitude" precision="15" seconds="19" value="-6.0219445">6°1'19"S</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="FA2178A6127417FFEBC0E9F749C832DC" degrees="50" direction="west" minutes="16" orientation="longitude" precision="15" seconds="26" value="-50.273888">50°16'26"W</geoCoordinate>
|
|
),
|
|
<date id="73F419F38712710E0479CF2BA6532501" value="2014-09-04" valueMax="2014-10-06" valueMin="2014-09-04">04/IX-06/X/2014</date>
|
|
,
|
|
<specimenCount id="2C3EE813758D3242EE6735E5F27737D4" type="male">1♂</specimenCount>
|
|
(IBSP 186143);
|
|
<specimenCount id="B1B28F6E8B2C07963500C00D9E538866" type="female">1♀</specimenCount>
|
|
(IBSP 186144);
|
|
<specimenCount id="CC25684DDA5CF63C72F8C5B255FA82EE" type="male">1♂</specimenCount>
|
|
(IBSP 186145);
|
|
<specimenCount id="71C98177FFFE90022748C7DB25F8579C" type="female">1♀</specimenCount>
|
|
(IBSP 186146);
|
|
<date id="E0011953ED592EC8A4A5A054BD8F9DCB" value="2015-04-02" valueMax="2015-04-29" valueMin="2015-04-02">02-29/IV/2015</date>
|
|
,
|
|
<specimenCount id="F4DD3E1C52DE06DD28445F192261AD6A" type="female">1♀</specimenCount>
|
|
(IBSP 186163);
|
|
<specimenCount id="FA3C92886EA99BF0A483CA5E0D7D28E6" type="male">1♂</specimenCount>
|
|
<specimenCount id="1B403A76031C9F99F5F9910B0525D42D" type="female">1♀</specimenCount>
|
|
(IBSP 186164); all collected by Equipe Carste; Cave N4WS_0067 (GEM-1846) (
|
|
<geoCoordinate id="CCF2701309AC01C111AB5FB3156AEAD8" degrees="6" direction="south" minutes="04" orientation="latitude" precision="15" seconds="22" value="-6.0727777">6°04'22"S</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="4185B79283399C6760C70DD381DD526D" degrees="50" direction="west" minutes="11" orientation="longitude" precision="15" seconds="30" value="-50.19167">50°11'30"W</geoCoordinate>
|
|
),
|
|
<date id="1AE55810E952C206EBE0C8D16865DE3D" value="2010-11-18" valueMax="2010-12-01" valueMin="2010-11-18">18/XI-01/XII/2010</date>
|
|
,
|
|
<materialsCitation id="5C33F8C41721208A10F00CF230CA9038" ID-GBIF-Occurrence="3075953302" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="F. P. Franco" country="Brazil" location="Souza" specimenCount="1" stateProvince="Para">
|
|
<collectorName id="099705D9337DACB31F9900F19CC85F36">F.P. Franco</collectorName>
|
|
& C.A.R.
|
|
<location id="7CEBA761021C3020BB022110C48B0906" LSID="urn:lsid:plazi:treatment:4FE7FB7679865ABAB75606F44AE07506:7CEBA761021C3020BB022110C48B0906" country="Brazil" name="Souza" stateProvince="Para">Souza</location>
|
|
et al. col
|
|
</materialsCitation>
|
|
.,
|
|
<specimenCount id="843C007E827B679E487C0D4B4EF2D3B5" type="male">1♂</specimenCount>
|
|
(IBSP 174069);
|
|
<specimenCount id="22D4E355F66A77B73B3D7D158B8BDEDF" type="female">2♀</specimenCount>
|
|
(IBSP 174070); Piranha, Cave N1-75 (
|
|
<geoCoordinate id="9692A5469B60FB9B7B907922A43A8F44" degrees="6" direction="south" minutes="1" orientation="latitude" precision="15" seconds="14" value="-6.020556">6°1'14"S</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="0A7C237434EB3B1E7EA64542E4782A36" degrees="50" direction="west" minutes="16" orientation="longitude" precision="15" seconds="49" value="-50.280277">50°16'49"W</geoCoordinate>
|
|
),
|
|
<date id="692B8D33CF90D8CA6B2260726AFD0527" value="2007-09-28" valueMax="2007-10-03" valueMin="2007-09-28">28/IX-03/X/2007</date>
|
|
,
|
|
<specimenCount id="41E902D436AC980852FCE96C9194266C" type="male">1♂</specimenCount>
|
|
<specimenCount id="C2EDC40CF96BBC5448EACCBFF134AE4A" type="female">2♀</specimenCount>
|
|
(IBSP 260307;
|
|
<specimenCount id="7EC888D6B9086DEB0087E27EA917A35C" type="male">1♂</specimenCount>
|
|
SEM;
|
|
<specimenCount id="BAB2ACA13798E1A734FCF339E668E12D" type="female">1♀</specimenCount>
|
|
SEM, ex IBSP 115499); all collected by R. Andrade et al.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="1DAF9029874D7092B0CA964B16C3CBF7" pageId="0" pageNumber="143" type="diagnosis">
|
|
<paragraph id="2CF0A30DD553561BA5113C6EA43F4A29" pageId="0" pageNumber="143">Diagnosis.</paragraph>
|
|
<paragraph id="42282A9684FC2E95D35E3DF69A335576" pageId="0" pageNumber="143">
|
|
<taxonomicName id="69AB233124A8350FB6D34F1A404DA3E3" authorityName="Brescovit & Zampaulo & Cizauskas" authorityYear="2021" class="Arachnida" family="Ochyroceratidae" genus="Ochyrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ochyrocera ritxoco" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ritxoco">
|
|
<emphasis id="15E2173DAFCF54EEA672B90B337F4A3D" italics="true" pageId="0" pageNumber="143">Ochyrocera ritxoco</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. is distinguished from
|
|
<taxonomicName id="85ED4113F94ABDEA6AEA800193150234" lsidName="O. ritxoo" pageId="0" pageNumber="143" rank="species" species="ritxoo">
|
|
<emphasis id="1803917540CA3E24187B4F008FB8FE96" italics="true" pageId="0" pageNumber="143">O. ritxoo</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by having an elongated embolus, which is two times longer than the bulb in the male palp (Figs
|
|
<figureCitation id="80FA020A19A438F7294354A98E184430" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ochyrocera ritxoco sp. nov., male IBSP 186160 (A, C-F), female IBSP 186146 (B) A, B habitus, dorsal view C left male palp, retrolateral view D same, prolateral view E right male palp, ventral view F same, dorsal view. Abbreviations: C = cymbium, CE = cymbial extension, E = embolus, T = tegulum." figureDoi="10.3897/zookeys.1031.62181.figure1" httpUri="https://binary.pensoft.net/fig/532001" pageId="0" pageNumber="143">1F</figureCitation>
|
|
,
|
|
<figureCitation id="C35957E2BB7259A8CCCCB515166F0104" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ochyrocera ritxoco sp. nov. (A, B) A left male palp IBSP 186149, retrolateral view B same, prolateral view (C, D) C female genitalia IBSP 186149, enzyme cleared, dorsal view D same, dorsal view. Abbreviations: C = cymbium, CUE = columnar uterus externus, E = embolus, NUE = neck of uterus externus, PP = pore-plate, SP = spermathecae, T = tegulum, UE = uterus externus." figureDoi="10.3897/zookeys.1031.62181.figure2" httpUri="https://binary.pensoft.net/fig/532002" pageId="0" pageNumber="143">2A, B</figureCitation>
|
|
,
|
|
<figureCitation id="89AF95736F1DE40610D1D45AA77D032B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of Ochyrocera ritxoco sp. nov., male IBSP 260307 (A-G) A cymbium, retrolateral view B same, basal macrosetae, retrolateral view C same, tarsal organ, detail, retrolateral view D same, apex detail E-G male palp, arrows indicating the embolus E retrolateral view F same, prolateral view G same, frontal view." figureDoi="10.3897/zookeys.1031.62181.figure4" httpUri="https://binary.pensoft.net/fig/532004" pageId="0" pageNumber="143">4E-G</figureCitation>
|
|
), while the embolus is one times longer than the bulb in
|
|
<taxonomicName id="041AC841D39614BAF1D0F61B991688A2" lsidName="O. ritxoo" pageId="0" pageNumber="143" rank="species" species="ritxoo">
|
|
<emphasis id="9290CF8AB5E0C40353700C73152914FF" italics="true" pageId="0" pageNumber="143">O. ritxoo</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (Fig.
|
|
<figureCitation id="3470A6767492A6ABAF3EF2A011E1A92A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ochyrocera ritxoo sp. nov., male IBSP 193194 (A, C-F), female IBSP 193196 (B) A, B habitus, dorsal view C left male palp, retrolateral view D same, prolateral view E right male palp, dorsal view F same, retro-ventral view. Abbreviations: C = cymbium, CE = cymbial extension, E = embolus, T = tegulum." figureDoi="10.3897/zookeys.1031.62181.figure7" httpUri="https://binary.pensoft.net/fig/532007" pageId="0" pageNumber="143">7C, D</figureCitation>
|
|
); a long and globose distal area in the spermathecae and an elongated columnar uterus externus with approximately eight internal chambers in the female genitalia (Fig.
|
|
<figureCitation id="7EA6421D07EF01B2E352FC2C81497576" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ochyrocera ritxoco sp. nov. (A, B) A left male palp IBSP 186149, retrolateral view B same, prolateral view (C, D) C female genitalia IBSP 186149, enzyme cleared, dorsal view D same, dorsal view. Abbreviations: C = cymbium, CUE = columnar uterus externus, E = embolus, NUE = neck of uterus externus, PP = pore-plate, SP = spermathecae, T = tegulum, UE = uterus externus." figureDoi="10.3897/zookeys.1031.62181.figure2" httpUri="https://binary.pensoft.net/fig/532002" pageId="0" pageNumber="143">2C, D</figureCitation>
|
|
); while
|
|
<taxonomicName id="A4507EA0885A04BD05C3EFE84625C60B" lsidName="O. ritxoo" pageId="0" pageNumber="143" rank="species" species="ritxoo">
|
|
<emphasis id="5CFFAA900B8BA33C2EFA602123FACE5D" italics="true" pageId="0" pageNumber="143">O. ritxoo</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. have a triangular distal area in the spermathecae and a shorter columnar uterus externus with 3-4 internal chambers in the female genitalia (Fig.
|
|
<figureCitation id="612889FF9414539F13A360981FDFA0DE" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Ochyrocera ritxoo sp. nov. (A, B) A left male palp IBSP 193194, retrolateral view B same, prolateral view (C, D) C female genitalia IBSP 1741071, enzyme cleared, dorsal view D same, dorsal view. Abbreviations: C = cymbium, CE = cymbial extension, CUE = columnar uterus externus, E = embolus, NUE = neck of uterus externus, PP = pore-plate, SP = spermathecae, T = tegulum, UE = uterus externus." figureDoi="10.3897/zookeys.1031.62181.figure8" httpUri="https://binary.pensoft.net/fig/532008" pageId="0" pageNumber="143">8C, D</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="CE13D6D8E8C615D92A6C3616063C119C" doi="10.3897/zookeys.1031.62181.figure1" httpUri="https://binary.pensoft.net/fig/532001" pageId="0" pageNumber="143" start="Figure 1" startId="F1">
|
|
<paragraph id="D755153D88E22250BFFC2960D637A361" pageId="0" pageNumber="143">
|
|
<emphasis id="9CFE76DAB8A646B2B3EA875524208BA5" bold="true" pageId="0" pageNumber="143">Figure 1.</emphasis>
|
|
<taxonomicName id="DAFD845C08C838418D7D01B23C4B14BE" authorityName="Brescovit & Zampaulo & Cizauskas" authorityYear="2021" class="Arachnida" family="Ochyroceratidae" genus="Ochyrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ochyrocera ritxoco" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ritxoco">
|
|
<emphasis id="8F9DA36FA4342911F8559F72BEDF6627" italics="true" pageId="0" pageNumber="143">Ochyrocera ritxoco</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., male IBSP 186160 (
|
|
<emphasis id="6B351A396EF0FEDA5E3B8AB718C7E711" bold="true" pageId="0" pageNumber="143">A, C-F</emphasis>
|
|
), female IBSP 186146 (
|
|
<emphasis id="78B2538F310C5CA107F3C313F2A2D1AF" bold="true" pageId="0" pageNumber="143">B</emphasis>
|
|
)
|
|
<emphasis id="E94C60265C340AAF126CE52A6F5446E9" bold="true" pageId="0" pageNumber="143">A, B</emphasis>
|
|
habitus, dorsal view
|
|
<emphasis id="1D3A4ABF394BFFC0155B6D0B61BD18BF" bold="true" pageId="0" pageNumber="143">C</emphasis>
|
|
left male palp, retrolateral view
|
|
<emphasis id="515CAB303180FA5061622553EAE67130" bold="true" pageId="0" pageNumber="143">D</emphasis>
|
|
same, prolateral view
|
|
<emphasis id="EB21600BE8469C5CE51F14AB4E2CB572" bold="true" pageId="0" pageNumber="143">E</emphasis>
|
|
right male palp, ventral view
|
|
<emphasis id="EC46AAAD806B0BE32CA6280FC4C34CB7" bold="true" pageId="0" pageNumber="143">F</emphasis>
|
|
same, dorsal view. Abbreviations: C = cymbium, CE = cymbial extension, E = embolus, T = tegulum.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="5E3A5B1F193EDEE87CF3E333AB1C2184" doi="10.3897/zookeys.1031.62181.figure2" httpUri="https://binary.pensoft.net/fig/532002" pageId="0" pageNumber="143" start="Figure 2" startId="F2">
|
|
<paragraph id="731E92B726CAC6A1C832945F602117BC" pageId="0" pageNumber="143">
|
|
<emphasis id="64A3205FC4D40685415A7FF121D6F4DF" bold="true" pageId="0" pageNumber="143">Figure 2.</emphasis>
|
|
<taxonomicName id="7FF4AA9B6D2EAE1D2D3D2EA1238E1070" authorityName="Brescovit & Zampaulo & Cizauskas" authorityYear="2021" class="Arachnida" family="Ochyroceratidae" genus="Ochyrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ochyrocera ritxoco" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ritxoco">
|
|
<emphasis id="35F06C71426CA40E2C566393C8CD0DC1" italics="true" pageId="0" pageNumber="143">Ochyrocera ritxoco</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (
|
|
<emphasis id="63F4D3425E1BE0D3ADFD76C26423108D" bold="true" pageId="0" pageNumber="143">A, B</emphasis>
|
|
)
|
|
<emphasis id="E339F1F31DCE3B1BCF1CA5B7BE5CC79F" bold="true" pageId="0" pageNumber="143">A</emphasis>
|
|
left male palp IBSP 186149, retrolateral view
|
|
<emphasis id="FE1A8CB7C9089B18DE1D0905F1A100BA" bold="true" pageId="0" pageNumber="143">B</emphasis>
|
|
same, prolateral view (
|
|
<emphasis id="A1D56F00EA7E5FE70768C7FEBE388891" bold="true" pageId="0" pageNumber="143">C, D) C</emphasis>
|
|
female genitalia IBSP 186149, enzyme cleared, dorsal view
|
|
<emphasis id="0E6B5EF70EBE38E6CF44ABC85E158C93" bold="true" pageId="0" pageNumber="143">D</emphasis>
|
|
same, dorsal view. Abbreviations: C = cymbium, CUE = columnar uterus externus, E = embolus, NUE = neck of uterus externus, PP = pore-plate, SP = spermathecae, T = tegulum, UE = uterus externus.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="51693CD453A8C5C11CAD0312BB918551" pageId="0" pageNumber="143" type="description">
|
|
<paragraph id="A635680BFFFB3A7ED9F570B6D836AC95" pageId="0" pageNumber="143">Description.</paragraph>
|
|
<paragraph id="37F5D0C560FFD3A00774B95F4EAF41C0" pageId="0" pageNumber="143">
|
|
<emphasis id="637DB166386C9D9CA2B84D8A6B27CA64" bold="true" pageId="0" pageNumber="143">Male</emphasis>
|
|
(Holotype). Total length 1.1. Carapace length 0.45; ovoid, narrowing gradually anteriorly, yellowish and bright, pars cephalic flat, fovea absent (Figs
|
|
<figureCitation id="41E0E91B5910723FD30ADE8CADB29039" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ochyrocera ritxoco sp. nov., male IBSP 186160 (A, C-F), female IBSP 186146 (B) A, B habitus, dorsal view C left male palp, retrolateral view D same, prolateral view E right male palp, ventral view F same, dorsal view. Abbreviations: C = cymbium, CE = cymbial extension, E = embolus, T = tegulum." figureDoi="10.3897/zookeys.1031.62181.figure1" httpUri="https://binary.pensoft.net/fig/532001" pageId="0" pageNumber="143">1A</figureCitation>
|
|
,
|
|
<figureCitation id="F67E55A2ACCD15B7623BC65BAA94D8E8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Ochyrocera ritxoco sp. nov., male IBSP 260307 (A-F) A carapace, dorsal view B chelicerae, frontal view C endites and labium, ventral view D crosier-like macrosetae (arrow, detail) E epiandrous area, abdomen, ventral view F male palp, tibia (arrows, trichobothria), dorsal view." figureDoi="10.3897/zookeys.1031.62181.figure3" httpUri="https://binary.pensoft.net/fig/532003" pageId="0" pageNumber="143">3A</figureCitation>
|
|
). Clypeus with two pairs of long bristles (Fig.
|
|
<figureCitation id="F5C957C9C3050C5AA7086B55246DB740" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Ochyrocera ritxoco sp. nov., male IBSP 260307 (A-F) A carapace, dorsal view B chelicerae, frontal view C endites and labium, ventral view D crosier-like macrosetae (arrow, detail) E epiandrous area, abdomen, ventral view F male palp, tibia (arrows, trichobothria), dorsal view." figureDoi="10.3897/zookeys.1031.62181.figure3" httpUri="https://binary.pensoft.net/fig/532003" pageId="0" pageNumber="143">3A</figureCitation>
|
|
). Eyes absent. Chelicerae light yellow with orange fang, promargin with three teeth attached to a very long lamina (Fig.
|
|
<figureCitation id="3C05E36232811974AC8E1243ECA0E0D5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Ochyrocera ritxoco sp. nov., male IBSP 260307 (A-F) A carapace, dorsal view B chelicerae, frontal view C endites and labium, ventral view D crosier-like macrosetae (arrow, detail) E epiandrous area, abdomen, ventral view F male palp, tibia (arrows, trichobothria), dorsal view." figureDoi="10.3897/zookeys.1031.62181.figure3" httpUri="https://binary.pensoft.net/fig/532003" pageId="0" pageNumber="143">3B</figureCitation>
|
|
), retromargin without teeth. Endites light yellow with large serrula with more than 30 denticles, distal macrosetae paired and crosier-like, many multifid macrosetae present (Fig.
|
|
<figureCitation id="35BA18A78E62569A908EC2418E95A624" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Ochyrocera ritxoco sp. nov., male IBSP 260307 (A-F) A carapace, dorsal view B chelicerae, frontal view C endites and labium, ventral view D crosier-like macrosetae (arrow, detail) E epiandrous area, abdomen, ventral view F male palp, tibia (arrows, trichobothria), dorsal view." figureDoi="10.3897/zookeys.1031.62181.figure3" httpUri="https://binary.pensoft.net/fig/532003" pageId="0" pageNumber="143">3C, D</figureCitation>
|
|
). Labium cream-colored, rounded with 8-10 setae with an enlarged basally (Fig.
|
|
<figureCitation id="EDB56A3A69B87A58C88EF7600A8B06AB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Ochyrocera ritxoco sp. nov., male IBSP 260307 (A-F) A carapace, dorsal view B chelicerae, frontal view C endites and labium, ventral view D crosier-like macrosetae (arrow, detail) E epiandrous area, abdomen, ventral view F male palp, tibia (arrows, trichobothria), dorsal view." figureDoi="10.3897/zookeys.1031.62181.figure3" httpUri="https://binary.pensoft.net/fig/532003" pageId="0" pageNumber="143">3C</figureCitation>
|
|
). Sternum light yellow. Legs cream-colored, formula 1423, total length I 4.5, II 3.8, III 3.1, IV 4.0. Male palp with palpal femur length 0.04, palpal tibia almost as long as wide with two long dorsal trichobothria (Fig.
|
|
<figureCitation id="EEB6C050E9C5343F3C11E221ED2DDD9E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Ochyrocera ritxoco sp. nov., male IBSP 260307 (A-F) A carapace, dorsal view B chelicerae, frontal view C endites and labium, ventral view D crosier-like macrosetae (arrow, detail) E epiandrous area, abdomen, ventral view F male palp, tibia (arrows, trichobothria), dorsal view." figureDoi="10.3897/zookeys.1031.62181.figure3" httpUri="https://binary.pensoft.net/fig/532003" pageId="0" pageNumber="143">3F</figureCitation>
|
|
), cymbium enlarged basally, narrowed distally, bearing elongated cuspule (Fig.
|
|
<figureCitation id="292FDEB2B30F7B9EE866DB1BE1204F29" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of Ochyrocera ritxoco sp. nov., male IBSP 260307 (A-G) A cymbium, retrolateral view B same, basal macrosetae, retrolateral view C same, tarsal organ, detail, retrolateral view D same, apex detail E-G male palp, arrows indicating the embolus E retrolateral view F same, prolateral view G same, frontal view." figureDoi="10.3897/zookeys.1031.62181.figure4" httpUri="https://binary.pensoft.net/fig/532004" pageId="0" pageNumber="143">4D</figureCitation>
|
|
), three setae on projected bases retrolaterally (Fig.
|
|
<figureCitation id="17B9908F91B54D7210D9836C7033F15C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of Ochyrocera ritxoco sp. nov., male IBSP 260307 (A-G) A cymbium, retrolateral view B same, basal macrosetae, retrolateral view C same, tarsal organ, detail, retrolateral view D same, apex detail E-G male palp, arrows indicating the embolus E retrolateral view F same, prolateral view G same, frontal view." figureDoi="10.3897/zookeys.1031.62181.figure4" httpUri="https://binary.pensoft.net/fig/532004" pageId="0" pageNumber="143">4A, B</figureCitation>
|
|
), elongated tarsal organ subdistally (Fig.
|
|
<figureCitation id="4F1DC26019C19D50D43DC3CF09EBA737" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of Ochyrocera ritxoco sp. nov., male IBSP 260307 (A-G) A cymbium, retrolateral view B same, basal macrosetae, retrolateral view C same, tarsal organ, detail, retrolateral view D same, apex detail E-G male palp, arrows indicating the embolus E retrolateral view F same, prolateral view G same, frontal view." figureDoi="10.3897/zookeys.1031.62181.figure4" httpUri="https://binary.pensoft.net/fig/532004" pageId="0" pageNumber="143">4C</figureCitation>
|
|
), no basal setae on the rounded cymbial prolateral extension, bulb oval, embolus filiform, sinuous and at least twice as long as the cymbium (Figs
|
|
<figureCitation id="A113BF58A0662C595F1E7D6C8D99C780" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ochyrocera ritxoco sp. nov., male IBSP 186160 (A, C-F), female IBSP 186146 (B) A, B habitus, dorsal view C left male palp, retrolateral view D same, prolateral view E right male palp, ventral view F same, dorsal view. Abbreviations: C = cymbium, CE = cymbial extension, E = embolus, T = tegulum." figureDoi="10.3897/zookeys.1031.62181.figure1" httpUri="https://binary.pensoft.net/fig/532001" pageId="0" pageNumber="143">1F</figureCitation>
|
|
,
|
|
<figureCitation id="FB1F3D0C5B509087D80948C0862EF43D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ochyrocera ritxoco sp. nov. (A, B) A left male palp IBSP 186149, retrolateral view B same, prolateral view (C, D) C female genitalia IBSP 186149, enzyme cleared, dorsal view D same, dorsal view. Abbreviations: C = cymbium, CUE = columnar uterus externus, E = embolus, NUE = neck of uterus externus, PP = pore-plate, SP = spermathecae, T = tegulum, UE = uterus externus." figureDoi="10.3897/zookeys.1031.62181.figure2" httpUri="https://binary.pensoft.net/fig/532002" pageId="0" pageNumber="143">2A, B</figureCitation>
|
|
,
|
|
<figureCitation id="3527BD62278EF5D091CB103CFE7AFB18" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM images of Ochyrocera ritxoco sp. nov., male IBSP 260307 (A-G) A cymbium, retrolateral view B same, basal macrosetae, retrolateral view C same, tarsal organ, detail, retrolateral view D same, apex detail E-G male palp, arrows indicating the embolus E retrolateral view F same, prolateral view G same, frontal view." figureDoi="10.3897/zookeys.1031.62181.figure4" httpUri="https://binary.pensoft.net/fig/532004" pageId="0" pageNumber="143">4E-G</figureCitation>
|
|
). Abdomen length 0.50, oval, uniformly gray. Six epiandrous spigots with a short base (Fig.
|
|
<figureCitation id="15A213F175B409EE0581AD65549831F5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM images of Ochyrocera ritxoco sp. nov., male IBSP 260307 (A-F) A carapace, dorsal view B chelicerae, frontal view C endites and labium, ventral view D crosier-like macrosetae (arrow, detail) E epiandrous area, abdomen, ventral view F male palp, tibia (arrows, trichobothria), dorsal view." figureDoi="10.3897/zookeys.1031.62181.figure3" httpUri="https://binary.pensoft.net/fig/532003" pageId="0" pageNumber="143">3E</figureCitation>
|
|
)
|
|
</paragraph>
|
|
<caption id="40902C0E651867651B32A5062ACAE2AE" doi="10.3897/zookeys.1031.62181.figure3" httpUri="https://binary.pensoft.net/fig/532003" pageId="0" pageNumber="143" start="Figure 3" startId="F3">
|
|
<paragraph id="EB91894E1D3412E194E4F0A2BB980687" pageId="0" pageNumber="143">
|
|
<emphasis id="4FB15C5F648F5FEDE18E32B3521F1AE3" bold="true" pageId="0" pageNumber="143">Figure 3.</emphasis>
|
|
SEM images of
|
|
<taxonomicName id="5B973DCFFBD6B685E0C0705029A2C942" authorityName="Brescovit & Zampaulo & Cizauskas" authorityYear="2021" class="Arachnida" family="Ochyroceratidae" genus="Ochyrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ochyrocera ritxoco" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ritxoco">
|
|
<emphasis id="1FF6848CFA0C5DEEA1C27BB0CB955704" italics="true" pageId="0" pageNumber="143">Ochyrocera ritxoco</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., male IBSP 260307 (
|
|
<emphasis id="95B74513B5EAD28308DAD525111ECEE2" bold="true" pageId="0" pageNumber="143">A-F</emphasis>
|
|
)
|
|
<emphasis id="54ABB85B387976B5C0389476C3103611" bold="true" pageId="0" pageNumber="143">A</emphasis>
|
|
carapace, dorsal view
|
|
<emphasis id="E6A2D0ED773FE1EFFE296B024C85ECE8" bold="true" pageId="0" pageNumber="143">B</emphasis>
|
|
chelicerae, frontal view
|
|
<emphasis id="E334F396785BB801EA95EFC3F42409F5" bold="true" pageId="0" pageNumber="143">C</emphasis>
|
|
endites and labium, ventral view
|
|
<emphasis id="859172217B08125DD3566D3D85B6471C" bold="true" pageId="0" pageNumber="143">D</emphasis>
|
|
crosier-like macrosetae (arrow, detail)
|
|
<emphasis id="90CE4615AC0810E90F7E6FEA796F8313" bold="true" pageId="0" pageNumber="143">E</emphasis>
|
|
epiandrous area, abdomen, ventral view
|
|
<emphasis id="38600F5DD54C19C046E09270D42DBC31" bold="true" pageId="0" pageNumber="143">F</emphasis>
|
|
male palp, tibia (arrows, trichobothria), dorsal view.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="3C5DB982EEA8DCB939B6478E3639EBF2" doi="10.3897/zookeys.1031.62181.figure4" httpUri="https://binary.pensoft.net/fig/532004" pageId="0" pageNumber="143" start="Figure 4" startId="F4">
|
|
<paragraph id="2487F2C20E47857DCD9CA91C6B3450B4" pageId="0" pageNumber="143">
|
|
<emphasis id="79DEC07A895CDAEE56F05CA922D24B0E" bold="true" pageId="0" pageNumber="143">Figure 4.</emphasis>
|
|
SEM images of
|
|
<taxonomicName id="C6704770E403E8568DFF6E4AD5A4E645" authorityName="Brescovit & Zampaulo & Cizauskas" authorityYear="2021" class="Arachnida" family="Ochyroceratidae" genus="Ochyrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ochyrocera ritxoco" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ritxoco">
|
|
<emphasis id="AE1FDA91822221E623734A236C9CEF2D" italics="true" pageId="0" pageNumber="143">Ochyrocera ritxoco</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., male IBSP 260307 (
|
|
<emphasis id="150E4E585F45318E689E789EC5C440BB" bold="true" pageId="0" pageNumber="143">A-G</emphasis>
|
|
)
|
|
<emphasis id="47E8BBB1F339ED95B8643E29D959C811" bold="true" pageId="0" pageNumber="143">A</emphasis>
|
|
cymbium, retrolateral view
|
|
<emphasis id="82E4685A24CFE1D3467F8761079A6AF9" bold="true" pageId="0" pageNumber="143">B</emphasis>
|
|
same, basal macrosetae, retrolateral view
|
|
<emphasis id="70F502520483A80AFCC4F300CA7C56CB" bold="true" pageId="0" pageNumber="143">C</emphasis>
|
|
same, tarsal organ, detail, retrolateral view
|
|
<emphasis id="63BEDB49AC8174E8D49720219582B978" bold="true" pageId="0" pageNumber="143">D</emphasis>
|
|
same, apex detail
|
|
<emphasis id="1498EAA0D4DF512D9AC49D7D1EED9F51" bold="true" pageId="0" pageNumber="143">E-G</emphasis>
|
|
male palp, arrows indicating the embolus
|
|
<emphasis id="116D98BA3A0BF3131D04068B2C93842F" bold="true" pageId="0" pageNumber="143">E</emphasis>
|
|
retrolateral view
|
|
<emphasis id="C489B10C3E6D52FB4059C99BAA69199A" bold="true" pageId="0" pageNumber="143">F</emphasis>
|
|
same, prolateral view
|
|
<emphasis id="08C94DB79C5CADB165D19B45C1278EEA" bold="true" pageId="0" pageNumber="143">G</emphasis>
|
|
same, frontal view.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="45C26CA0B1E6E5CBF16C0CD45B7E9B7A" pageId="0" pageNumber="143">
|
|
<emphasis id="21C72336EC161CF0BD2B147208CEA65E" bold="true" pageId="0" pageNumber="143">Female</emphasis>
|
|
(Paratype IBSP 115499). Total length 1.15. Carapace length 0.55 as in male with light yellowish pattern (Figs
|
|
<figureCitation id="F1EC13701BAEED0AC082973A744CAA50" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ochyrocera ritxoco sp. nov., male IBSP 186160 (A, C-F), female IBSP 186146 (B) A, B habitus, dorsal view C left male palp, retrolateral view D same, prolateral view E right male palp, ventral view F same, dorsal view. Abbreviations: C = cymbium, CE = cymbial extension, E = embolus, T = tegulum." figureDoi="10.3897/zookeys.1031.62181.figure1" httpUri="https://binary.pensoft.net/fig/532001" pageId="0" pageNumber="143">1B</figureCitation>
|
|
,
|
|
<figureCitation id="326ED9B00983E0BE3ABB2E94B811DFA8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM images of Ochyrocera ritxoco sp. nov., female IBSP 260307 (A-F) A carapace, dorsal view B chelicerae, frontal view C colulus, ventral view D pedipalp, distal, prolateral view E same, tarsal organ F leg IV, claw, prolateral view." figureDoi="10.3897/zookeys.1031.62181.figure5" httpUri="https://binary.pensoft.net/fig/532005" pageId="0" pageNumber="143">5A</figureCitation>
|
|
). Pedipalp without claw, with conical tip and subdistal trichobothrium (Fig.
|
|
<figureCitation id="7C1FA89074EFA99C3D01735BAB82C9A2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM images of Ochyrocera ritxoco sp. nov., female IBSP 260307 (A-F) A carapace, dorsal view B chelicerae, frontal view C colulus, ventral view D pedipalp, distal, prolateral view E same, tarsal organ F leg IV, claw, prolateral view." figureDoi="10.3897/zookeys.1031.62181.figure5" httpUri="https://binary.pensoft.net/fig/532005" pageId="0" pageNumber="143">5D-E</figureCitation>
|
|
). Clypeus, eyes, chelicerae (Fig.
|
|
<figureCitation id="9DB32CC5C30F19EB871E12C2ECF4FF47" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM images of Ochyrocera ritxoco sp. nov., female IBSP 260307 (A-F) A carapace, dorsal view B chelicerae, frontal view C colulus, ventral view D pedipalp, distal, prolateral view E same, tarsal organ F leg IV, claw, prolateral view." figureDoi="10.3897/zookeys.1031.62181.figure5" httpUri="https://binary.pensoft.net/fig/532005" pageId="0" pageNumber="143">5B</figureCitation>
|
|
), sternum, endites, and labium as in male. Legs as in male, formula 4123, total length I 4.1, II 3.6, III 2.4, IV 4.3. Claw of leg with five teeth (Fig.
|
|
<figureCitation id="BA1820A509B217004822D1FFCEA4EBB0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM images of Ochyrocera ritxoco sp. nov., female IBSP 260307 (A-F) A carapace, dorsal view B chelicerae, frontal view C colulus, ventral view D pedipalp, distal, prolateral view E same, tarsal organ F leg IV, claw, prolateral view." figureDoi="10.3897/zookeys.1031.62181.figure5" httpUri="https://binary.pensoft.net/fig/532005" pageId="0" pageNumber="143">5F</figureCitation>
|
|
). Abdomen length 0.65. Colulus rectangular with five long bristles (Fig.
|
|
<figureCitation id="8F38ABF4CD4290837A866F267F3BCC86" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM images of Ochyrocera ritxoco sp. nov., female IBSP 260307 (A-F) A carapace, dorsal view B chelicerae, frontal view C colulus, ventral view D pedipalp, distal, prolateral view E same, tarsal organ F leg IV, claw, prolateral view." figureDoi="10.3897/zookeys.1031.62181.figure5" httpUri="https://binary.pensoft.net/fig/532005" pageId="0" pageNumber="143">5C</figureCitation>
|
|
). Internal genitalia with long spermathecae narrowed at tip, conspicuous pore-plate at base; medial columnar uterus externus long, with visible internal chambers. Uterus externus shorter than spermathecae. Oval pore-plates on spermathecae with approximately 20-30 glandular ducts (Fig.
|
|
<figureCitation id="52C3CA0ABCFD081A417FFEBABE9568F4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ochyrocera ritxoco sp. nov. (A, B) A left male palp IBSP 186149, retrolateral view B same, prolateral view (C, D) C female genitalia IBSP 186149, enzyme cleared, dorsal view D same, dorsal view. Abbreviations: C = cymbium, CUE = columnar uterus externus, E = embolus, NUE = neck of uterus externus, PP = pore-plate, SP = spermathecae, T = tegulum, UE = uterus externus." figureDoi="10.3897/zookeys.1031.62181.figure2" httpUri="https://binary.pensoft.net/fig/532002" pageId="0" pageNumber="143">2C, D</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="73334586D3FD58B9BC4FDA50DF6123A0" doi="10.3897/zookeys.1031.62181.figure5" httpUri="https://binary.pensoft.net/fig/532005" pageId="0" pageNumber="143" start="Figure 5" startId="F5">
|
|
<paragraph id="E4EC756EF5485C50E383FEE22F76565C" pageId="0" pageNumber="143">
|
|
<emphasis id="079A7FF136F104A1C6E4FE37EDCD6EBF" bold="true" pageId="0" pageNumber="143">Figure 5.</emphasis>
|
|
SEM images of
|
|
<taxonomicName id="334A60C29F3E4C37A9C7480CB199E16A" authorityName="Brescovit & Zampaulo & Cizauskas" authorityYear="2021" class="Arachnida" family="Ochyroceratidae" genus="Ochyrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ochyrocera ritxoco" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ritxoco">
|
|
<emphasis id="EAFC5C9B1124C8D22E3CF7ED5451BC0D" italics="true" pageId="0" pageNumber="143">Ochyrocera ritxoco</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., female IBSP 260307 (
|
|
<emphasis id="11380BFED53D2B33EEEB0D1F8A1CD771" bold="true" pageId="0" pageNumber="143">A-F</emphasis>
|
|
)
|
|
<emphasis id="BFE3C357DC845294CF5ABA67558EB641" bold="true" pageId="0" pageNumber="143">A</emphasis>
|
|
carapace, dorsal view
|
|
<emphasis id="85EB6DAC1C2772EAF5E699B149FABAFB" bold="true" pageId="0" pageNumber="143">B</emphasis>
|
|
chelicerae, frontal view
|
|
<emphasis id="37658B013B793A7CDA328223FB22E220" bold="true" pageId="0" pageNumber="143">C</emphasis>
|
|
colulus, ventral view
|
|
<emphasis id="7AE509557F98AF54259E908479FCECF0" bold="true" pageId="0" pageNumber="143">D</emphasis>
|
|
pedipalp, distal, prolateral view
|
|
<emphasis id="152FF53E41BA313DDFF3FF20E5805965" bold="true" pageId="0" pageNumber="143">E</emphasis>
|
|
same, tarsal organ
|
|
<emphasis id="BE1CDBC232545F022632682041D4461F" bold="true" pageId="0" pageNumber="143">F</emphasis>
|
|
leg IV, claw, prolateral view.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="06058E18C44E9F2AC6E334A132407D36" pageId="0" pageNumber="143" type="variation">
|
|
<paragraph id="9D5395628FAE0BBF4DC62C31AA67940F" pageId="0" pageNumber="143">Variation.</paragraph>
|
|
<paragraph id="2226A983377C27F9BED33186EBA9E9AE" pageId="0" pageNumber="143">Ten males: total length 1-1.25; carapace 0.4-0.5; femur I 1.05-1.4; ten females: total length 1-1.3; carapace 0.4-0.5; femur I 1-1.3.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="7228EB9D62C515B47ED73A8A2BB6501A" pageId="0" pageNumber="143" type="etymology">
|
|
<paragraph id="60EF79E4A4EEF826AE287B1611A258D3" pageId="0" pageNumber="143">Etymology.</paragraph>
|
|
<paragraph id="9D962FC5C5D2D03E88378AFACF158F76" pageId="0" pageNumber="143">
|
|
The specific name
|
|
<normalizedToken id="C54B33A67A0C6019B80A292E8B552CDB" originalValue="Ritxòkò">Ritxoko</normalizedToken>
|
|
means "ceramic dolls" in the female language of the
|
|
<normalizedToken id="2AB1B1A24E1758C062648A0D9F276444" originalValue="Karajá">Karaja</normalizedToken>
|
|
people, an indigenous population of the region. The dolls are produced by
|
|
<normalizedToken id="5614C39D0B2B9AB993A0C553D3A0BD65" originalValue="Karajá">Karaja</normalizedToken>
|
|
women, who model, burn, paint, and sell them (
|
|
<bibRefCitation id="84680AA2B657668331040924FB6268FB" author="Silva, TC" journalOrPublisher="Goiania, Editora Canone" pageId="0" pageNumber="143" refId="B13" refString="Silva, TC, 2015. Ritxoko. Goiania, Editora Canone" title="Ritxoko." year="2015">Silva 2015</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="93C9B890A26EC581B7ABDF017E99513C" pageId="0" pageNumber="143" type="natural history">
|
|
<paragraph id="3DB14BD6FDA85EFFFB9AD083CAE70BA8" pageId="0" pageNumber="143">Natural history.</paragraph>
|
|
<paragraph id="D968EB311BE535A0902B76495AD58F57" pageId="0" pageNumber="143">
|
|
<taxonomicName id="1331F0FD6C8826B8D4CA24FD2FD4B772" authorityName="Brescovit & Zampaulo & Cizauskas" authorityYear="2021" class="Arachnida" family="Ochyroceratidae" genus="Ochyrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ochyrocera ritxoco" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ritxoco">
|
|
<emphasis id="8B599C03E3B94B9DB09F18D7EB34A1D7" italics="true" pageId="0" pageNumber="143">Ochyrocera ritxoco</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. is a small troglobitic spider that is exclusive to caves in the
|
|
<normalizedToken id="9E641D554114FDB698E6C898E7628C99" originalValue="Carajás">Carajas</normalizedToken>
|
|
karst region. Specimens were found only in aphotic zones of caves. They build small, tangled webs under blocks of stone on the ground or in slits (Fig.
|
|
<figureCitation id="744760740FE1F10F090AA2EF7FEE2A23" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Ochyrocera ritxoco sp. nov., female in webs under rocks on the ground in the Cave N 4 WS _ 0067." figureDoi="10.3897/zookeys.1031.62181.figure6" httpUri="https://binary.pensoft.net/fig/532006" pageId="0" pageNumber="143">6</figureCitation>
|
|
). The observed sex ratio for the species was 2.4F:1M (
|
|
<emphasis id="BD6B9216CCC5BEBF49175AB1EE6FD437" italics="true" pageId="0" pageNumber="143">N</emphasis>
|
|
= 66).
|
|
<taxonomicName id="FA2CEBBA4A442D2BD67D972550187A7B" authorityName="Brescovit & Zampaulo & Cizauskas" authorityYear="2021" class="Arachnida" family="Ochyroceratidae" genus="Ochyrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ochyrocera ritxoco" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ritxoco">
|
|
<emphasis id="DF2BBAA1CDBAF0049276B8B0722FAAC7" italics="true" pageId="0" pageNumber="143">Ochyrocera ritxoco</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. was generally found in large cavities with horizontal projections varying from 9.5 to 216 meters (
|
|
<emphasis id="AE1CA4F5F7197576E95FE6A58FBDFE02" italics="true" pageId="0" pageNumber="143">N</emphasis>
|
|
= 13, mean = 107 m). All caves where the species was found have only one entrance and are located only in the middle and high slopes of Serra Norte. Most caves have aphotic zones or twilight zones (except for cavities N1_0103 and N1_0084) and high humidity, thus explaining the observed presence of small bodies of water in almost all cavities, especially during the wet season. The number of troglobitic species in these caves varied from one to ten (average 5.3 per cave), with species of the following taxa: spiders -
|
|
<taxonomicName id="E3C331B3825342346A546B22E9031591" class="Arachnida" family="Oonopidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="family">Oonopidae</taxonomicName>
|
|
(many species),
|
|
<taxonomicName id="BC0C366714EDB9025AB5AAE04B3FA776" authorityName="Simon" authorityYear="1890" class="Arachnida" family="Caponiidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="family">Caponiidae</taxonomicName>
|
|
(
|
|
<taxonomicName id="FB74272DA2C243F5AA3DB0412600C88F" authorityName="Brescovit & Sanchez-Ruiz" authorityYear="2016" class="Arachnida" family="Caponiidae" genus="Carajas" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Carajas paraua" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="paraua">
|
|
<emphasis id="52F745E2D239D8717FA477322273849B" italics="true" pageId="0" pageNumber="143">Carajas paraua</emphasis>
|
|
</taxonomicName>
|
|
Brescovit &
|
|
<normalizedToken id="92AC10ACB843B463FBEEFB66D7450DFC" originalValue="Sánchez-Ruiz">Sanchez-Ruiz</normalizedToken>
|
|
, 2016) and
|
|
<taxonomicName id="B0FF18CA4906C9ADCBB9DC7648ED16B4" authorityName="Fage" authorityYear="1912" class="Arachnida" family="Ochyroceratidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="family">Ochyroceratidae</taxonomicName>
|
|
(
|
|
<taxonomicName id="160306423B1B728AEDE793095AEA656E" authorityName="Berland" authorityYear="1914" class="Arachnida" family="Ochyroceratidae" genus="Speocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Speocera" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2287EF9964393533A17DCE590C31F13C" italics="true" pageId="0" pageNumber="143">Speocera</emphasis>
|
|
</taxonomicName>
|
|
spp.); pseudoscorpions -
|
|
<taxonomicName id="08B8F6A8180234824C78291E0492ACA7" authorityName="J.C.Chamberlin" authorityYear="1930" class="Arachnida" family="Bochicidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pseudoscorpiones" pageId="0" pageNumber="143" phylum="Arthropoda" rank="family">Bochicidae</taxonomicName>
|
|
,
|
|
<taxonomicName id="6522E5CA685B89846569D2DE1161C6CB" class="Arachnida" family="Chthoniidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pseudoscorpiones" pageId="0" pageNumber="143" phylum="Arthropoda" rank="family">Chthoniidae</taxonomicName>
|
|
and
|
|
<taxonomicName id="0492E2B3A9711D40F6F47310293BB3E1" authorityName="J.C.Chamberlin" authorityYear="1930" class="Arachnida" family="Ideoroncidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pseudoscorpiones" pageId="0" pageNumber="143" phylum="Arthropoda" rank="family">Ideoroncidae</taxonomicName>
|
|
; springtails -
|
|
<taxonomicName id="5F801E5821C2B084D7809D881657226F" authorityName="Borner" authorityYear="1913" class="Entognatha" family="Paronellidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Entomobryomorpha" pageId="0" pageNumber="143" rank="family">Paronellidae</taxonomicName>
|
|
(
|
|
<taxonomicName id="169E79C234D6C30DD4CA2FB4C5FE7FD5" authorityName="Mills" authorityYear="1938" class="Entognatha" family="Paronellidae" genus="Trogolaphysa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Trogolaphysa" order="Collembola" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="12233E9AFFEF0D7C16590A3637B50B0D" italics="true" pageId="0" pageNumber="143">Trogolaphysa</emphasis>
|
|
</taxonomicName>
|
|
sp.,
|
|
<taxonomicName id="0AC6C984F05DC0C931CE58006D749AAC" class="Collembola" family="Entomobryidae" genus="Cyphoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyphoderus" order="Entomobryomorpha" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="C955BB4BD62980BC5B3C6F5F1CFFE899" italics="true" pageId="0" pageNumber="143">Cyphoderus</emphasis>
|
|
</taxonomicName>
|
|
sp.),
|
|
<taxonomicName id="C38A3C7FEE4D9B24BDC16B76D8D61D91" class="Entognatha" family="Entomobryidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Entomobryomorpha" pageId="0" pageNumber="143" rank="family">Entomobryidae</taxonomicName>
|
|
(
|
|
<taxonomicName id="F21ABA9A70252A9321203FD97B260A45" authorityName="Schaeffer" authorityYear="1897" class="Collembola" family="Entomobryidae" genus="Pseudosinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudosinella" order="Entomobryomorpha" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="57EEBD45F4F4A32075189B3656CFC795" italics="true" pageId="0" pageNumber="143">Pseudosinella</emphasis>
|
|
</taxonomicName>
|
|
sp.) and
|
|
<taxonomicName id="B8FCECCED250900B55627F8FAC9D196D" authorityName="Lubbock" authorityYear="1862" class="Collembola" family="Sminthuridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Symphypleona" pageId="0" pageNumber="143" phylum="Arthropoda" rank="family">Sminthuridae</taxonomicName>
|
|
(
|
|
<taxonomicName id="883070C82252B44E79AA1D5F1C5AE378" authorityName="Bonet & Tellez" authorityYear="1947" class="Collembola" family="Sminthuridae" genus="Pararrhopalites" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pararrhopalites" order="Symphypleona" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="BE3D39EC2360A88DE2649F2D0D645A55" italics="true" pageId="0" pageNumber="143">Pararrhopalites</emphasis>
|
|
</taxonomicName>
|
|
sp.); beetles -
|
|
<taxonomicName id="E3FB03E6EED4380567FD215C6F9C7327" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
|
|
(
|
|
<taxonomicName id="FCC2151209C61117D863B29F20E7E14B" class="Insecta" family="Carabidae" genus="Coarazuphium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Coarazuphium" order="Coleoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="775E0ECA8D2B19380B071FEC0EDF78C6" italics="true" pageId="0" pageNumber="143">Coarazuphium</emphasis>
|
|
</taxonomicName>
|
|
spp.),
|
|
<taxonomicName id="E1E4F9B349D75E6E3676EB77E63141A0" class="Insecta" family="Dytiscidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
|
|
(
|
|
<taxonomicName id="DE14D5FC50E30E65D89BF2B9B872B0FF" class="Hexapoda" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Copelatus cessaima" order="Coleoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="cessaima">
|
|
<emphasis id="24476C9D0EAE3258A9DA317D08830D07" italics="true" pageId="0" pageNumber="143">Copelatus cessaima</emphasis>
|
|
</taxonomicName>
|
|
Caetano, Bena & Vanin, 2013); isopods -
|
|
<taxonomicName id="FF74EDC85E7558B781832FB12CF076B8" authorityName="Verhoeff" authorityYear="1938" class="Arachnida" family="Scleropactidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="family">Scleropactidae</taxonomicName>
|
|
(
|
|
<taxonomicName id="CE1ED96C6F79BD9535D9789EDE9D4E78" class="Malacostraca" family="Scleropactidae" genus="Circoniscus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Circoniscus" order="Isopoda" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="D3C222DC86CF665CE7F3813D246C9E50" italics="true" pageId="0" pageNumber="143">Circoniscus</emphasis>
|
|
</taxonomicName>
|
|
spp.),
|
|
<taxonomicName id="2032C87F384A39563316C96E827EDBEB" class="Malacostraca" family="Calabozoidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="0" pageNumber="143" phylum="Arthropoda" rank="family">Calabozoidae</taxonomicName>
|
|
; amphipods -
|
|
<taxonomicName id="83A1CBC5A2E4235CC6CF588ACD8BF827" authorityName="Hertzog" authorityYear="1936" class="Malacostraca" family="Bogidiellidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Amphipoda" pageId="0" pageNumber="143" phylum="Arthropoda" rank="family">Bogidiellidae</taxonomicName>
|
|
(
|
|
<taxonomicName id="C2E05FDF9941836FED56362155FD200A" authorityName="Hertzog" authorityYear="1933" class="Malacostraca" family="Bogidiellidae" genus="Bogidiella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Bogidiella" order="Amphipoda" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="9282AF21E89E65C939EC8EFD16BCC736" italics="true" pageId="0" pageNumber="143">Bogidiella</emphasis>
|
|
</taxonomicName>
|
|
sp.); planarian -
|
|
<taxonomicName id="F6D33D77D5F2C66630DB93D772E079A0" authorityName="Hallez" authorityYear="1894" class="Cestoda" family="Prorhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Cyclophyllidea" pageId="0" pageNumber="143" phylum="Platyhelminthes" rank="family">Prorhynchidae</taxonomicName>
|
|
(
|
|
<taxonomicName id="0BFFA5D895A59FF3979AB67BBDCE14D1" class="Polycladidea" family="Prorhynchidae" genus="Geocentrophora" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Geocentrophora" order="Lecithoepiteliata" pageId="0" pageNumber="143" phylum="Platyhelminthes" rank="genus">
|
|
<emphasis id="7C56B9FD35527B8A815890FC6A95D008" italics="true" pageId="0" pageNumber="143">Geocentrophora</emphasis>
|
|
</taxonomicName>
|
|
sp.); and Harvestmen -
|
|
<taxonomicName id="FF258E06050A417EB7796AB800F4967F" authorityName="Kury & Perez" authorityYear="2003" class="Arachnida" family="Escadabiidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Opiliones" pageId="0" pageNumber="143" phylum="Arthropoda" rank="family">Escadabiidae</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<caption id="6EC97B43D249E3F90AC24C1D395216BC" doi="10.3897/zookeys.1031.62181.figure6" httpUri="https://binary.pensoft.net/fig/532006" pageId="0" pageNumber="143" start="Figure 6" startId="F6">
|
|
<paragraph id="6760E3DADEB9BC278553C63F8DCE635B" pageId="0" pageNumber="143">
|
|
<emphasis id="98A4F5A7426E952A34771738D2AC9997" bold="true" pageId="0" pageNumber="143">Figure 6.</emphasis>
|
|
<taxonomicName id="F871E9D2E832CDD4215EF69C407677BB" authorityName="Brescovit & Zampaulo & Cizauskas" authorityYear="2021" class="Arachnida" family="Ochyroceratidae" genus="Ochyrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ochyrocera ritxoco" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ritxoco">
|
|
<emphasis id="6F4E942F05E7FE350E34051D5F9CE57A" italics="true" pageId="0" pageNumber="143">Ochyrocera ritxoco</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., female in webs under rocks on the ground in the Cave N4WS_0067.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="72B3D08B593F97E7E792F0382D6B05F7" doi="10.3897/zookeys.1031.62181.figure7" httpUri="https://binary.pensoft.net/fig/532007" pageId="0" pageNumber="143" start="Figure 7" startId="F7">
|
|
<paragraph id="271BB064782CF39BDE54144C2E0A3379" pageId="0" pageNumber="143">
|
|
<emphasis id="D4A47F373F7B8F421483B94C72C65784" bold="true" pageId="0" pageNumber="143">Figure 7.</emphasis>
|
|
<taxonomicName id="C5890E5DDC15FB5A737FF225F58E13F5" authorityName="Brescovit & Zampaulo & Cizauskas" authorityYear="2021" class="Arachnida" family="Ochyroceratidae" genus="Ochyrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ochyrocera ritxoo" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ritxoo">
|
|
<emphasis id="B3F98AF180E2DBD91AA9642785A3C714" italics="true" pageId="0" pageNumber="143">Ochyrocera ritxoo</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., male IBSP 193194 (
|
|
<emphasis id="51C5C1F86477A75BDAB4D10286DB354B" bold="true" pageId="0" pageNumber="143">A, C-F</emphasis>
|
|
), female IBSP 193196 (
|
|
<emphasis id="81277FAEA30882E50836BB16625ACE7F" bold="true" pageId="0" pageNumber="143">B</emphasis>
|
|
)
|
|
<emphasis id="83B2BE8011DBB0C4514199512721CE3B" bold="true" pageId="0" pageNumber="143">A, B</emphasis>
|
|
habitus, dorsal view
|
|
<emphasis id="8FD2BCCEF32F1C385C03ED2B6EE1FF08" bold="true" pageId="0" pageNumber="143">C</emphasis>
|
|
left male palp, retrolateral view
|
|
<emphasis id="C9D1F474D89F053159D4C89EB7FA219E" bold="true" pageId="0" pageNumber="143">D</emphasis>
|
|
same, prolateral view
|
|
<emphasis id="5C28E669CD615470936C716682AEB236" bold="true" pageId="0" pageNumber="143">E</emphasis>
|
|
right male palp, dorsal view
|
|
<emphasis id="59300C52E92FA3DD49163C77033338ED" bold="true" pageId="0" pageNumber="143">F</emphasis>
|
|
same, retro-ventral view. Abbreviations: C = cymbium, CE = cymbial extension, E = embolus, T = tegulum.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="0711F5B4F0FEBBFD882BD62D5539D954" doi="10.3897/zookeys.1031.62181.figure8" httpUri="https://binary.pensoft.net/fig/532008" pageId="0" pageNumber="143" start="Figure 8" startId="F8">
|
|
<paragraph id="FACEC4F1963D19ABE37B8C59A8266272" pageId="0" pageNumber="143">
|
|
<emphasis id="5ACD68DF0CA31232C30F72849A29BBB3" bold="true" pageId="0" pageNumber="143">Figure 8.</emphasis>
|
|
<taxonomicName id="6D2517D62EDA464A207011EA7197F002" authorityName="Brescovit & Zampaulo & Cizauskas" authorityYear="2021" class="Arachnida" family="Ochyroceratidae" genus="Ochyrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ochyrocera ritxoo" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ritxoo">
|
|
<emphasis id="DDA7A5079382C00D1C50C85F5E5B9422" italics="true" pageId="0" pageNumber="143">Ochyrocera ritxoo</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (
|
|
<emphasis id="75204D45DD155EF3BCFD30ABA2AF9C7C" bold="true" pageId="0" pageNumber="143">A, B</emphasis>
|
|
)
|
|
<emphasis id="B1CA471445954CF191A7130D5B4C2412" bold="true" pageId="0" pageNumber="143">A</emphasis>
|
|
left male palp IBSP 193194, retrolateral view
|
|
<emphasis id="564B1DD5C33F7A267D1A20D5622F2F64" bold="true" pageId="0" pageNumber="143">B</emphasis>
|
|
same, prolateral view (
|
|
<emphasis id="937428C959E6721D982CC5F8B2E2A442" bold="true" pageId="0" pageNumber="143">C, D) C</emphasis>
|
|
female genitalia IBSP 1741071, enzyme cleared, dorsal view
|
|
<emphasis id="D473229C3EEAD520FAAF68F9BC4C8F6C" bold="true" pageId="0" pageNumber="143">D</emphasis>
|
|
same, dorsal view. Abbreviations: C = cymbium, CE = cymbial extension, CUE = columnar uterus externus, E = embolus, NUE = neck of uterus externus, PP = pore-plate, SP = spermathecae, T = tegulum, UE = uterus externus.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="6B0F5BB2E5FE999D227E15611D6DEA19" pageId="0" pageNumber="143" type="distribution">
|
|
<paragraph id="62DC15D0F381A10CC9C7AC95C7BEB2A9" pageId="0" pageNumber="143">Distribution.</paragraph>
|
|
<paragraph id="A7925F958276DC70B8E5A13EA2220B21" pageId="0" pageNumber="143">
|
|
Known exclusively from caves in a range of approximately 15 km of the Serra Norte (North Mountain), FLONA de
|
|
<normalizedToken id="D8A18494D0981BEC04C96F6E6875A141" originalValue="Carajás">Carajas</normalizedToken>
|
|
, Parauapebas, state of
|
|
<normalizedToken id="DFE2692406AEC5CC1225097621D098C7" originalValue="Pará">Para</normalizedToken>
|
|
, northern Brazil (Fig.
|
|
<figureCitation id="4CE85B7A3836F3AAA63EEA8357DCFB5B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution map of Ochyrocera ritxoco sp. nov., yellow triangles, and Ochyrocera ritxoo sp. nov., red plus signs, in FLONA de Carajas, Para, Brazil." figureDoi="10.3897/zookeys.1031.62181.figure10" httpUri="https://binary.pensoft.net/fig/532010" pageId="0" pageNumber="143">10</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |