460 lines
57 KiB
XML
460 lines
57 KiB
XML
<document id="5EAB3C2113B63BE4F288055CCA1FD0E1" ID-CLB-Dataset="244287" ID-DOI="10.1093/zoolinnean/zlac091" ID-GBIF-Dataset="87852b6b-9e57-46bc-97af-35a7a8aee100" ID-ISSN="0024-4082" ID-Zenodo-Dep="7814306" ID-ZooBank="6DBA2650-DB10-4BDC-AEDB-2EF08D82815E" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1680598078240" checkinUser="plazi" docAuthor="Fassio, Giulia, Stefani, Matteo, Russini, Valeria, Buge, Barbara, Bouchet, Philippe, Treneman, Nancy, Malaquias, Manuel António E., Schiaparelli, Stefano, Modica, Maria Vittoria & Oliverio, Marco" docDate="2023" docId="8C6587D7FFA8FFEE168A7420FD685D14" docLanguage="en" docName="zlac091.pdf" docOrigin="Zoological Journal of the Linnean Society 197 (4)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlac091" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Variolipallium Fassio, Bouchet & Oliverio 2023" docType="treatment" docVersion="5" lastPageNumber="953" masterDocId="705CFFAFFFB4FFF3164A7701FFBC587C" masterDocTitle="Neither slugs nor snails: a molecular reappraisal of the gastropod family Velutinidae" masterLastPageNumber="964" masterPageNumber="924" pageNumber="952" updateTime="1699513426530" updateUser="ExternalLinkService">
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<mods:title id="848642616EF1E258B045D269A7753D87">Neither slugs nor snails: a molecular reappraisal of the gastropod family Velutinidae</mods:title>
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<treatment id="8C6587D7FFA8FFEE168A7420FD685D14" ID-DOI="http://doi.org/10.5281/zenodo.7814353" ID-GBIF-Taxon="209495729" ID-Zenodo-Dep="7814353" LSID="urn:lsid:plazi:treatment:8C6587D7FFA8FFEE168A7420FD685D14" httpUri="http://treatment.plazi.org/id/8C6587D7FFA8FFEE168A7420FD685D14" lastPageId="29" lastPageNumber="953" pageId="28" pageNumber="952">
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<subSubSection id="4CD6654AFFA8FFEF168A7420FDAE5B2B" pageId="28" pageNumber="952" type="nomenclature">
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<taxonomicName id="C3CC4D42FFA8FFEF168A7420FD515B46" ID-CoL="e9ca0fb1-9c66-442a-a408-aee85655197e" authority="FASSIO, BOUCHET & OLIVERIO" authorityName="Fassio, Bouchet & Oliverio" authorityYear="2023" box="[192,749,801,826]" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="36B8EAD3FFA8FFEF168A7420FE355B44" bold="true" box="[192,393,801,824]" italics="true" pageId="28" pageNumber="952">VARIOLIPALLIUM</emphasis>
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FASSIO, BOUCHET & OLIVERIO
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<heading id="5F3B81ADFFA8FFEF17D17442FDAE5B2B" box="[411,530,831,855]" centered="true" fontSize="9" level="2" pageId="28" pageNumber="952" reason="3">
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<emphasis id="36B8EAD3FFA8FFEF17D17442FDAE5B2B" bold="true" box="[411,530,831,855]" pageId="28" pageNumber="952">GEN. NOV.</emphasis>
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<subSubSection id="4CD6654AFFA8FFEF16BB7467FD415BBF" pageId="28" pageNumber="952" type="description">
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<paragraph id="047336C1FFA8FFEF16BB7467FD005B03" blockId="28.[241,700,870,895]" box="[241,700,870,895]" pageId="28" pageNumber="952">
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(
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<figureCitation id="9CF72A44FFA8FFEF16B37467FEF85B03" box="[249,324,870,895]" captionStart="Figure 2" captionStartId="6.[163,244,1709,1731]" captionTargetBox="[241,1358,197,1670]" captionTargetId="figure-27@6.[240,1360,195,1672]" captionTargetPageId="6" captionText="Figure 2. Phylogenetic relationships of the family Velutinidae (Bayesian tree based on the combined dataset), with clades (A–L) collapsed by genera. Numbers at nodes indicate branch support values [posterior probability (PP) and ultrafast bootstrap (UFb) values, respectively]; support values are shown only when at least one of them is ≥ 80%; white circles at nodes indicate maximum support (PP = 1, Ufb = 100). On the right side of the tree, for each genus is reported a representative photograph of the shell, a schematic representation of the radula, the depth range (in metres) and the geographical distribution." figureDoi="http://doi.org/10.5281/zenodo.7814312" httpUri="https://zenodo.org/record/7814312/files/figure.png" pageId="28" pageNumber="952">FIGS 2</figureCitation>
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,
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<figureCitation id="9CF72A44FFA8FFEF171B7467FEE35B03" box="[337,351,870,895]" captionStart="Figure 3" captionStartId="7.[145,225,1585,1607]" captionTargetBox="[271,1296,200,1554]" captionTargetId="figure-25@7.[269,1299,195,1557]" captionTargetPageId="7" captionText="Figure 3. Phylogenetic relationships of the family Velutinidae (Bayesian tree based on the combined dataset) showing clades A–L expanded. Numbers at nodes indicate branch support values [posterior probability (PP) and ultrafast bootstrap (UFb) values, respectively]; support values are shown only when at least one of them is ≥ 80%; white circles at nodes indicate maximum support (PP = 1, Ufb = 100). Boxes indicate hypothetical species; dashed lines indicate possible alternative species partitions. Coloured boxes on the right indicate genera, and white boxes indicate subfamilies. Letters at the end of sample codes broadly indicate the area of origin:AK, Alaska; ANT, Antarctica; AR, Argentina; BC, British Columbia; BRE, Bretagne; CA, California; CH, China; COR, Corse; CR, Costa Rica; FL, Florida; GF, French Guiana; GP, Guadeloupe; HI, Hawaii; IT, Italy; MG, Madagascar; MQ, Martinique; MX, Mexico; MZ, Mozambique; NC, New Caledonia; NL, Newfoundland; NO, Norway; OR, Oregon; PA, Panama; PF, French Polynesia; PG, Papua New Guinea; PH, Philippines; QLS, Queensland; RE, Reunion; SA, Saudi Arabia; SP, Spain; TAS, Tasmania; TW, Taiwan; VU, Vanuatu; WA, Washington State; ZNZ, Zanzibar." figureDoi="http://doi.org/10.5281/zenodo.7814314" httpUri="https://zenodo.org/record/7814314/files/figure.png" pageId="28" pageNumber="952">3</figureCitation>
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,
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<figureCitation id="9CF72A44FFA8FFEF17267467FE165B03" box="[364,426,870,895]" captionStart="Figure 4" captionStartId="11.[145,228,1695,1717]" captionTargetBox="[197,1373,195,1664]" captionTargetId="figure-25@11.[197,1373,195,1665]" captionTargetPageId="11" captionText="Figure 4. A, Marseniopsis sp. (M9), MNA 5376. B, Marseniopsis conica, MNA 5361. C, Marsenina zadei, LACM 182861. D, Marsenina rhombica, LACM 182862. E, Limneria prolongata, LACM 182858. F, Velutina sp., LACM 182869. G, Hainotis sharonae. H, Variolipallium regium sp. nov., MNHN-IM-2013-69285, holotype. I, Variolipallium sp. (L3), MNHN-IM-2013-69605. J, Variolipallium sp. (L10), MNHN-IM-2013-69280. Photograph credits: A, B, MNA, Stefano Schiaparelli; C–F, Nancy Treneman; G, iNaturalist, Robin Gwen Agarwal; H–J, MNHN, Philippe Maestrati. For the sequenced specimens, the corresponding species code (see Fig. 3) is reported in parentheses." figureDoi="http://doi.org/10.5281/zenodo.7814318" httpUri="https://zenodo.org/record/7814318/files/figure.png" pageId="28" pageNumber="952">4H–J</figureCitation>
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,
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<figureCitation id="9CF72A44FFA8FFEF17FD7467FE6A5B03" box="[439,470,870,895]" captionStart="Figure 7" captionStartId="14.[163,243,1623,1645]" captionTargetBox="[247,1359,199,1589]" captionTargetId="figure-119@14.[243,1363,195,1593]" captionTargetPageId="14" captionText="Figure 7. Velutinidae shells. A, Calyptoconcha pellucida, BAU 3758. B, Variolipallium cerebroides, NIWA 74086.A. C, Pacifica lentiginosa sp. nov., MNHN-IM-2009-16140, holotype. D, Pacifica sp. (L16), MNHN-IM-2013-58543. Scale bars: A, B, 5 mm; C, D, 1 mm. For the sequenced specimens, the corresponding species code (see Fig. 3) is reported in parentheses." figureDoi="http://doi.org/10.5281/zenodo.7814328" httpUri="https://zenodo.org/record/7814328/files/figure.png" pageId="28" pageNumber="952">7B</figureCitation>
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,
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<figureCitation id="9CF72A44FFA8FFEF17A97467FDB85B03" box="[483,516,870,895]" captionStart="Figure 9" captionStartId="16.[163,246,1664,1686]" captionTargetBox="[230,1376,198,1631]" captionTargetId="figure-26@16.[227,1379,195,1634]" captionTargetPageId="16" captionText="Figure 9. Velutinidae protoconchs. A, Marseniopsis conica, BAS 03-764. B, Marsenina rhombica, BAU 4229. C, Calyptoconcha pellucida, BAU 3758. D, Variolipallium cerebroides, NIWA 74086.A. E, Pacifica lentiginosa sp. nov., MNHN-IM-2009-16140, holotype.F, Pacifica sp. (L16), MNHN-IM-2013-58543. G, Djiboutia sp. (L22), MNHN-IM-2019-9053. H, Coriocella sp. (L27), MNHN-IM-2013-84230. I, Lamellaria latens complex (L41), MNHN-IM-2019-14179. J, Lamellaria mopsicolor, MNHN-IM-2013-73086. K, Marsenia perspicua complex (L55), MNHN-IM-2019-14574. L, Marsenia sp. (L42), MNHN-IM-2019-7641. Scale bars: A–L, 500 µm. For the sequenced specimens, the corresponding species code (see Fig. 3) is reported in parentheses. Arrows indicate protoconch I – protoconch II boundary." figureDoi="http://doi.org/10.5281/zenodo.7814334" httpUri="https://zenodo.org/record/7814334/files/figure.png" pageId="28" pageNumber="952">9D</figureCitation>
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,
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<figureCitation id="9CF72A44FFA8FFEF145B7467FDE75B03" box="[529,603,870,895]" captionStart="Figure 10" captionStartId="17.[145,226,1664,1686]" captionTargetBox="[212,1358,198,1631]" captionTargetId="figure-24@17.[209,1361,195,1634]" captionTargetPageId="17" captionText="Figure 10. Velutinidae radulae. A, Marseniopsis mollis, CE 6137. B, Marsenina rhombica, BAU 4229. C, Velutina cf. coriacea, LACM 182870. D, Calyptoconcha pellucida, BAU 3751. E, Variolipallium regium sp. nov., MNHN-IM-2013-69285, holotype. F, Variolipallium sp. (L10), MNHN-IM-2013-69280. G, Pacifica lentiginosa sp. nov., MNHN-IM-2009-16140, holotype. H, Pacifica sp. (L15), MNHN-IM-2013-59969. I, Djiboutia sp. (L22), MNHN-IM-2019-9053. J, Coriocella sp. (L27), MNHN-IM-2013-84230. K, Lamellaria mopsicolor, MNHN-IM-2013-73086. L, Lamellaria latens complex (L41), MNHN-IM-2019-14113. M, Marsenia herberti, MNHN-IM-2019-8503. N, Marsenia sp. (L45), MNHN-IM-2013-59960. O, Marsenia perspicua complex (L55), MNHN-IM-2019-5980. Scale bars: A–O, 100 µm. For the sequenced specimens, the corresponding species code (see Fig. 3) is reported in parentheses." figureDoi="http://doi.org/10.5281/zenodo.7814338" httpUri="https://zenodo.org/record/7814338/files/figure.png" pageId="28" pageNumber="952">10E, F</figureCitation>
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,
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<figureCitation id="9CF72A44FFA8FFEF14227467FD085B03" box="[616,692,870,895]" captionStart="Figure 11" captionStartId="18.[163,243,1664,1686]" captionTargetBox="[230,1376,198,1631]" captionTargetId="figure-26@18.[227,1379,195,1634]" captionTargetPageId="18" captionText="Figure 11. Velutinidae jaws. A, M, Marseniopsis mollis, CE 6152. B, N, O, Marsenina rhombica, BAU 2339. C, Limneria prolongata, BAU 4228. D, Calyptoconcha pellucida, BAU 3758. E, Variolipallium regium sp. nov., MNHN-IM-2013-69285, holotype. F, Variolipallium sp. (L10), MNHN-IM-2013-69280. G, Pacifica lentiginosa sp. nov., MNHN-IM-2009-16140, holotype.H, Pacifica sp. (L19), MNHN-IM-2009-16136.I,Djiboutia sp.(L22), MNHN-IM-2019-9053.J,Coriocella cf.jayi (L28), MNHN-IM-2009-16119. K, Lamellaria latens complex (L40), BAU 2536.2. L, Marsenia sp. (L45), MNHN-IM-2013-59960. Scale bars: H, M–O, 100 µm; A–G, I–L, 500 µm. For the sequenced specimens, the corresponding species code (see Fig. 3) is reported in parentheses." figureDoi="http://doi.org/10.5281/zenodo.7814342" httpUri="https://zenodo.org/record/7814342/files/figure.png" pageId="28" pageNumber="952">11E, F</figureCitation>
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)
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<emphasis id="36B8EAD3FFA8FFEF16E9748FFE545BD8" box="[163,488,910,932]" italics="true" pageId="28" pageNumber="952">Zoobank registration:</emphasis>
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urn: lsid: zoobank. org:act:
|
||
<uuid id="706A0C14FFA8FFEF16A574ACFD415BBF" box="[239,765,941,963]" pageId="28" pageNumber="952">170D2A4B-E62D-4612-9737-FFC39B639EFA</uuid>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4CD6654AFFA8FFEF16E974ECFEE45C5C" pageId="28" pageNumber="952" type="type_taxon">
|
||
<paragraph id="047336C1FFA8FFEF16E974ECFEE45C5C" blockId="28.[163,779,1004,1056]" pageId="28" pageNumber="952">
|
||
<emphasis id="36B8EAD3FFA8FFEF16E974ECFE8E5C7D" box="[163,306,1004,1026]" italics="true" pageId="28" pageNumber="952">
|
||
<typeStatus id="DB778863FFA8FFEF16E974ECFF645C7E" box="[163,216,1005,1026]" pageId="28" pageNumber="952">Type</typeStatus>
|
||
species:
|
||
</emphasis>
|
||
<taxonomicName id="C3CC4D42FFA8FFEF177774EDFF425C5C" ID-CoL="1b27cd9a-2df2-4f4a-8122-6c83160d8a7a" authority="Fassio, Bouchet & Oliverio" authorityName="Fassio, Bouchet & Oliverio" authorityYear="2023" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="regium" status="sp. nov.">
|
||
<emphasis id="36B8EAD3FFA8FFEF177774EDFD875C7D" box="[317,571,1004,1025]" italics="true" pageId="28" pageNumber="952">Variolipallium regium</emphasis>
|
||
Fassio, Bouchet & Oliverio
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2D8B57A8FFA8FFEF174F730AFEE45C5C" box="[261,344,1035,1056]" pageId="28" pageNumber="952" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4CD6654AFFA8FFEF16E97348FDAA5D6B" pageId="28" pageNumber="952" type="synonymic_list">
|
||
<paragraph id="047336C1FFA8FFEF16E97348FDAA5D6B" blockId="28.[163,779,1097,1303]" pageId="28" pageNumber="952">
|
||
<emphasis id="36B8EAD3FFA8FFEF16E97348FEDF5C23" box="[163,355,1097,1119]" italics="true" pageId="28" pageNumber="952">Included species:</emphasis>
|
||
<taxonomicName id="C3CC4D42FFA8FFEF173F7348FF595C01" authority="(Hutton, 1882)" authorityName="Fassio & Stefani & Russini & Buge & Bouchet & Treneman & Malaquias & Schiaparelli & Modica & Oliverio" authorityYear="2023" baseAuthorityName="Hutton" baseAuthorityYear="1882" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="cerebroides" status="comb. nov.">
|
||
<emphasis id="36B8EAD3FFA8FFEF173F7348FD1F5C22" box="[373,675,1097,1118]" italics="true" pageId="28" pageNumber="952">Variolipallium cerebroides</emphasis>
|
||
(Hutton, 1882)
|
||
</taxonomicName>
|
||
<emphasis id="36B8EAD3FFA8FFEF16A47369FEC05C01" bold="true" box="[238,380,1128,1149]" pageId="28" pageNumber="952">
|
||
<taxonomicNameLabel id="2D8B57A8FFA8FFEF16A47369FEC05C01" box="[238,380,1128,1149]" pageId="28" pageNumber="952" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="C3CC4D42FFA8FFEF17C77369FF5E5CE0" authority="(Strebel, 1906)" authorityName="Fassio & Stefani & Russini & Buge & Bouchet & Treneman & Malaquias & Schiaparelli & Modica & Oliverio" authorityYear="2023" baseAuthorityName="Strebel" baseAuthorityYear="1906" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="elatum" status="comb. nov.">
|
||
<emphasis id="36B8EAD3FFA8FFEF17C77369FD275C01" box="[397,667,1128,1149]" italics="true" pageId="28" pageNumber="952">Variolipallium elatum</emphasis>
|
||
(Strebel, 1906)
|
||
</taxonomicName>
|
||
<emphasis id="36B8EAD3FFA8FFEF16A37386FED35CE0" bold="true" box="[233,367,1159,1180]" pageId="28" pageNumber="952">
|
||
<taxonomicNameLabel id="2D8B57A8FFA8FFEF16A37386FED35CE0" box="[233,367,1159,1180]" pageId="28" pageNumber="952" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="C3CC4D42FFA8FFEF17367387FF5E5CC6" authority="(Bergh, 1907)" authorityName="Fassio & Stefani & Russini & Buge & Bouchet & Treneman & Malaquias & Schiaparelli & Modica & Oliverio" authorityYear="2023" baseAuthorityName="Bergh" baseAuthorityYear="1907" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="leptoconcha" status="comb. nov.">
|
||
<emphasis id="36B8EAD3FFA8FFEF17367387FD0C5CE7" box="[380,688,1158,1179]" italics="true" pageId="28" pageNumber="952">Variolipallium leptoconcha</emphasis>
|
||
(
|
||
<bibRefCitation id="605D4B30FFA8FFEF14F57387FF665CC6" author="Bergh LSR" pageId="28" pageNumber="952" pagination="1 - 144" refId="ref25396" refString="Bergh LSR. 1907. The Opisthobranchiata of South Africa. In: Marine InVestigations of South Africa 5 (1) [Dated 1908]. Page (s): 106, pl. 9, figs 17 - 20; pl. 10, figs 1, 2. Transactions of the South African Philosophical Society 17: 1 - 144, pls 1 - 14." type="journal article" year="1907">Bergh, 1907</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="36B8EAD3FFA8FFEF16AF73A4FEDA5CC6" bold="true" box="[229,358,1189,1210]" pageId="28" pageNumber="952">
|
||
<taxonomicNameLabel id="2D8B57A8FFA8FFEF16AF73A4FEDA5CC6" box="[229,358,1189,1210]" pageId="28" pageNumber="952" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="36B8EAD3FFA8FFEF172573A4FD3C5CC6" box="[367,640,1189,1210]" italics="true" pageId="28" pageNumber="952">Variolipalliumnodosum</emphasis>
|
||
(Ev.Marcus, 1987)
|
||
<emphasis id="36B8EAD3FFA8FFEF16A073C5FECD5CA5" bold="true" box="[234,369,1220,1241]" pageId="28" pageNumber="952">
|
||
<taxonomicNameLabel id="2D8B57A8FFA8FFEF16A073C5FECD5CA5" box="[234,369,1220,1241]" pageId="28" pageNumber="952" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="C3CC4D42FFA8FFEF173473C5FE845C84" authority="(E. A. Smith, 1881)" authorityName="Fassio & Stefani & Russini & Buge & Bouchet & Treneman & Malaquias & Schiaparelli & Modica & Oliverio" authorityYear="2023" baseAuthorityName="E. A. Smith" baseAuthorityYear="1881" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="patagonicum" status="comb. nov.">
|
||
<emphasis id="36B8EAD3FFA8FFEF173473C5FD785CA5" box="[382,708,1220,1241]" italics="true" pageId="28" pageNumber="952">Variolipallium patagonicum</emphasis>
|
||
(E. A. Smith, 1881)
|
||
</taxonomicName>
|
||
<emphasis id="36B8EAD3FFA8FFEF170B73E2FE755C84" bold="true" box="[321,457,1251,1272]" pageId="28" pageNumber="952">
|
||
<taxonomicNameLabel id="2D8B57A8FFA8FFEF170B73E2FE755C84" box="[321,457,1251,1272]" pageId="28" pageNumber="952" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="C3CC4D42FFA8FFEF144F73E3FE075D6B" authority="Fassio, Bouchet, Oliverio" authorityName="Fassio, Bouchet, Oliverio" authorityYear="2023" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="regium" status="sp. nov.">
|
||
<emphasis id="36B8EAD3FFA8FFEF144F73E3FCB75C84" box="[517,779,1250,1272]" italics="true" pageId="28" pageNumber="952">Variolipallium regium</emphasis>
|
||
Fassio, Bouchet, Oliverio
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2D8B57A8FFA8FFEF17897203FDAA5D6B" box="[451,534,1282,1303]" pageId="28" pageNumber="952" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4CD6654AFFA8FFEF16E97241FAB25A16" pageId="28" pageNumber="952" type="description">
|
||
<paragraph id="047336C1FFA8FFEF16E97241FE035E72" blockId="28.[163,779,1344,1887]" pageId="28" pageNumber="952">
|
||
<emphasis id="36B8EAD3FFA8FFEF16E97241FE8F5D29" box="[163,307,1344,1365]" italics="true" pageId="28" pageNumber="952">Description:</emphasis>
|
||
Body of small to medium size for the subfamily, 0.5–3.0 cm total length. Shell very thin, weakly calcified to membranaceus; ear shaped, high spired, with expanded aperture; smooth or weakly sculptured by axial growth lines; completely enclosed by the mantle or presenting a small dorsal fissure. Periostracum not visible.
|
||
</paragraph>
|
||
<paragraph id="047336C1FFA8FFEF16F17117FEEA5EDB" blockId="28.[163,779,1344,1887]" pageId="28" pageNumber="952">Protoconch of 1.24–1.70 whorls; protoconch I of 0.48– 1.20 whorls, nucleus diameter 100–150 μm, smooth, with subsutural axial folds; protoconch II with axial growth lines; protoconch–teleoconch boundary not always distinct.</paragraph>
|
||
<paragraph id="047336C1FFA8FFEF16F171B1FBA25964" blockId="28.[163,779,1344,1887]" lastBlockId="28.[827,1443,197,618]" pageId="28" pageNumber="952">Mantle flat or dome shaped, outline rounded or polygonal; thick to thin, often with a few to several dorsal tubercles; with anterior siphon fold; texture smooth to wrinkled (resembling the convolutions of a brain); colour variable, almost transparent to white, grey, yellow, orange to red, pink to violet, light blue, brown, dark green, occasionally patterned with spots. Cephalic tentacle tips can be white, lime yellow or almost transparent.</paragraph>
|
||
<paragraph id="047336C1FFA8FFEF15197620FB0F5908" blockId="28.[827,1443,197,618]" pageId="28" pageNumber="952">Penis to the right of the right cephalic tentacle, with or without a lateral subterminal papilla. Vas deferens without a free loop in haemocoel.</paragraph>
|
||
<paragraph id="047336C1FFA8FFEF1519767CFC425A37" blockId="28.[827,1443,197,618]" pageId="28" pageNumber="952">Radula reduced taenioglossate, formula 0:1:1:1:0; rachidian tooth base bifurcated; rachidian cusp with few to several, very small to pronounced external denticles; lateral teeth elongated, with a pointed triangular external cusp, with few to several, very small to pronounced denticles on both sides or only on the internal side.</paragraph>
|
||
<paragraph id="047336C1FFA8FFEF15197555FAB25A16" blockId="28.[827,1443,197,618]" box="[851,1294,596,618]" pageId="28" pageNumber="952">Jaws homogeneous, short to elongated.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4CD6654AFFA8FFEF15717593FAD75B78" pageId="28" pageNumber="952" type="distribution">
|
||
<paragraph id="047336C1FFA8FFEF15717593FAD75B78" blockId="28.[827,1442,658,772]" pageId="28" pageNumber="952">
|
||
<emphasis id="36B8EAD3FFA8FFEF15717593FC705ADB" box="[827,972,658,679]" italics="true" pageId="28" pageNumber="952">Distribution:</emphasis>
|
||
Tropical West Pacific (South
|
||
<collectingCountry id="7CDB7651FFA8FFEF136C7593FAD75AD4" box="[1318,1387,658,680]" name="China" pageId="28" pageNumber="952">China</collectingCountry>
|
||
Sea,
|
||
<collectingCountry id="7CDB7651FFA8FFEF157175B0FB985ABB" box="[827,1060,689,711]" name="Papua New Guinea" pageId="28" pageNumber="952">Papua New Guinea</collectingCountry>
|
||
,
|
||
<collectingCountry id="7CDB7651FFA8FFEF127875B0FB575ABB" box="[1074,1259,689,711]" name="New Caledonia" pageId="28" pageNumber="952">New Caledonia</collectingCountry>
|
||
),
|
||
<collectingCountry id="7CDB7651FFA8FFEF12B575B0FA225ABB" box="[1279,1438,689,711]" name="New Zealand" pageId="28" pageNumber="952">New Zealand</collectingCountry>
|
||
; Caribbean; tip of South America (
|
||
<collectingCountry id="7CDB7651FFA8FFEF129E75D1FAAA5A9A" box="[1236,1302,720,742]" name="Chile" pageId="28" pageNumber="952">Chile</collectingCountry>
|
||
,
|
||
<collectingCountry id="7CDB7651FFA8FFEF136975D1FA225A9A" box="[1315,1438,720,742]" name="Argentina" pageId="28" pageNumber="952">Argentina</collectingCountry>
|
||
,
|
||
<collectingCountry id="7CDB7651FFA8FFEF157175EFFC405B78" box="[827,1020,750,772]" name="Falkland Islands" pageId="28" pageNumber="952">Falkland Islands</collectingCountry>
|
||
),
|
||
<collectingCountry id="7CDB7651FFA8FFEF125A75EFFB1D5B78" box="[1040,1185,750,772]" name="South Africa" pageId="28" pageNumber="952">South Africa</collectingCountry>
|
||
;
|
||
<quantity id="C3349B24FFA8FFEF12E675EFFA945B78" box="[1196,1320,750,772]" metricMagnitude="2" metricUnit="m" metricValue="8.075" metricValueMax="15.73" metricValueMin="0.42" pageId="28" pageNumber="952" unit="m" value="807.5" valueMax="1573.0" valueMin="42.0">42–1573 m</quantity>
|
||
deep.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4CD6654AFFA8FFEF1571742CFA185BE2" pageId="28" pageNumber="952" type="etymology">
|
||
<paragraph id="047336C1FFA8FFEF1571742CFA185BE2" blockId="28.[827,1444,813,927]" pageId="28" pageNumber="952">
|
||
<emphasis id="36B8EAD3FFA8FFEF1571742CFC755B3E" box="[827,969,813,834]" italics="true" pageId="28" pageNumber="952">Etymology:</emphasis>
|
||
From
|
||
<taxonomicName id="C3CC4D42FFA8FFEF1279742CFB2D5B3E" authorityName="Swainson" authorityYear="1839" box="[1075,1169,813,834]" class="Actinopterygii" family="Serranidae" genus="Variola" kingdom="Animalia" order="Perciformes" pageId="28" pageNumber="952" phylum="Chordata" rank="genus">
|
||
<emphasis id="36B8EAD3FFA8FFEF1279742CFB2D5B3E" box="[1075,1169,813,834]" italics="true" pageId="28" pageNumber="952">Variola</emphasis>
|
||
</taxonomicName>
|
||
, the Latin name for smallpox, and
|
||
<emphasis id="36B8EAD3FFA8FFEF15A8744DFB815B1D" box="[994,1085,844,865]" italics="true" pageId="28" pageNumber="952">pallium</emphasis>
|
||
, meaning ‘mantle’, referring to the coloured small tubercles on the mantle of the
|
||
<typeStatus id="DB778863FFA8FFEF133B746AFA1E5BFC" box="[1393,1442,875,896]" pageId="28" pageNumber="952">type</typeStatus>
|
||
species and other members of the group. Gender neuter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4CD6654AFFA8FFEE157174C9FD685D14" lastPageId="29" lastPageNumber="953" pageId="28" pageNumber="952" type="discussion">
|
||
<paragraph id="047336C1FFA8FFEF157174C9FBC85CEA" blockId="28.[827,1444,968,1879]" pageId="28" pageNumber="952">
|
||
<emphasis id="36B8EAD3FFA8FFEF157174C9FB365BA1" box="[827,1162,968,989]" italics="true" pageId="28" pageNumber="952">
|
||
Remarks:
|
||
<taxonomicName id="C3CC4D42FFA8FFEF158674C9FB365BA1" authorityName="Fassio, Bouchet & Oliverio" authorityYear="2023" box="[972,1162,968,989]" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="genus">Variolipallium</taxonomicName>
|
||
</emphasis>
|
||
can be diagnosed by a membranaceous to weakly calcified shell with high spire (similar to that of
|
||
<taxonomicName id="C3CC4D42FFA8FFEF12FC7304FAD65C66" authorityName="Bouchet & Waren" authorityYear="1993" box="[1206,1386,1029,1050]" class="Gastropoda" family="Velutinidae" genus="Calyptoconcha" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="genus">
|
||
<emphasis id="36B8EAD3FFA8FFEF12FC7304FAD65C66" box="[1206,1386,1029,1050]" italics="true" pageId="28" pageNumber="952">Calyptoconcha</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C3CC4D42FFA8FFEF15717325FC695C45" authorityName="Bergh" authorityYear="1866" box="[827,981,1060,1081]" class="Gastropoda" family="Velutinidae" genus="Marseniopsis" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="genus">
|
||
<emphasis id="36B8EAD3FFA8FFEF15717325FC695C45" box="[827,981,1060,1081]" italics="true" pageId="28" pageNumber="952">Marseniopsis</emphasis>
|
||
</taxonomicName>
|
||
) and a bifurcated rachidian tooth (like
|
||
<taxonomicName id="C3CC4D42FFA8FFEF15717343FC065C2B" authorityName="Montagu" authorityYear="1816" box="[827,954,1090,1111]" class="Gastropoda" family="Velutinidae" genus="Lamellaria" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="genus">
|
||
<emphasis id="36B8EAD3FFA8FFEF15717343FC065C2B" box="[827,954,1090,1111]" italics="true" pageId="28" pageNumber="952">Lamellaria</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C3CC4D42FFA8FFEF158F7342FB8C5C24" authorityName="Oken" authorityYear="1823" box="[965,1072,1091,1112]" class="Gastropoda" family="Velutinidae" genus="Marsenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="genus">
|
||
<emphasis id="36B8EAD3FFA8FFEF158F7342FB8C5C24" box="[965,1072,1091,1112]" italics="true" pageId="28" pageNumber="952">Marsenia</emphasis>
|
||
</taxonomicName>
|
||
, etc.), although in
|
||
<taxonomicName id="C3CC4D42FFA8FFEF12B17343FA1F5C2B" authorityName="Fassio, Bouchet & Oliverio" authorityYear="2023" box="[1275,1443,1090,1111]" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="genus">
|
||
<emphasis id="36B8EAD3FFA8FFEF12B17343FA1F5C2B" box="[1275,1443,1090,1111]" italics="true" pageId="28" pageNumber="952">Variolipallium</emphasis>
|
||
</taxonomicName>
|
||
the ‘V’-shaped base of the rachidian tooth can be comparatively less marked.
|
||
</paragraph>
|
||
<paragraph id="047336C1FFA8FFEF1519739FFBF55CAE" blockId="28.[827,1444,968,1879]" pageId="28" pageNumber="952">
|
||
All the species included in this genus have been found at depths>
|
||
<specimenCount id="12CAFD48FFA8FFEF124073BCFBF55CAE" box="[1034,1097,1213,1235]" pageId="28" pageNumber="952" type="generic">
|
||
<quantity id="C3349B24FFA8FFEF124073BCFBF95CAE" box="[1034,1093,1213,1235]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="28" pageNumber="952" unit="m" value="95.0">95 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
<paragraph id="047336C1FFA8FFEF151973DDFAA45E3F" blockId="28.[827,1444,968,1879]" pageId="28" pageNumber="952">
|
||
The dorsal appearance of the mantle varies from wrinkled, with several tubercles (resembling the convolutions of a brain), to rather smooth and studded with fewer small tubercles of different colour, to completely smooth. Several species (
|
||
<taxonomicName id="C3CC4D42FFA8FFEF12B97257FC6D5DF7" authorityName="Fassio & Stefani & Russini & Buge & Bouchet & Treneman & Malaquias & Schiaparelli & Modica & Oliverio" authorityYear="2023" baseAuthorityName="Hutton" baseAuthorityYear="1882" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="cerebroides">
|
||
<emphasis id="36B8EAD3FFA8FFEF12B97257FC6D5DF7" italics="true" pageId="28" pageNumber="952">Variolipallium cerebroides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C3CC4D42FFA8FFEF15A37274FA265DF7" authorityName="Fassio & Stefani & Russini & Buge & Bouchet & Treneman & Malaquias & Schiaparelli & Modica & Oliverio" authorityYear="2023" baseAuthorityName="E. A. Smith" baseAuthorityYear="1881" box="[1001,1434,1397,1419]" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" isUncertain="true" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="patagonicum">
|
||
<emphasis id="36B8EAD3FFA8FFEF15A37274FB0D5DF7" box="[1001,1201,1397,1419]" italics="true" pageId="28" pageNumber="952">Variolipallium</emphasis>
|
||
cf.
|
||
<emphasis id="36B8EAD3FFA8FFEF12A47277FA265DF7" box="[1262,1434,1398,1419]" italics="true" pageId="28" pageNumber="952">patagonicum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C3CC4D42FFA8FFEF15717295FB295DD5" authorityName="Fassio & Stefani & Russini & Buge & Bouchet & Treneman & Malaquias & Schiaparelli & Modica & Oliverio" authorityYear="2023" baseAuthorityName="Strebel" baseAuthorityYear="1906" box="[827,1173,1428,1450]" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" isUncertain="true" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="elatum">
|
||
<emphasis id="36B8EAD3FFA8FFEF15717295FC435DD5" box="[827,1023,1428,1449]" italics="true" pageId="28" pageNumber="952">Variolipallium</emphasis>
|
||
cf.
|
||
<emphasis id="36B8EAD3FFA8FFEF12707295FB295DD5" box="[1082,1173,1428,1449]" italics="true" pageId="28" pageNumber="952">elatum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C3CC4D42FFA8FFEF12E17295FA1F5DD5" box="[1195,1443,1428,1449]" class="Gastropoda" family="Velutinidae" genus="Variolipallium" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="36B8EAD3FFA8FFEF12E17295FACC5DD5" box="[1195,1392,1428,1449]" italics="true" pageId="28" pageNumber="952">Variolipallium</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
L10,
|
||
<taxonomicName id="C3CC4D42FFA8FFEF153D72B3FBF15DB4" box="[887,1101,1458,1480]" class="Gastropoda" family="Velutinidae" genus="Variolipallium" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="36B8EAD3FFA8FFEF153D72B3FB985DBB" box="[887,1060,1458,1479]" italics="true" pageId="28" pageNumber="952">Variolipallium</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
L11,
|
||
<taxonomicName id="C3CC4D42FFA8FFEF12DB72B3FADB5DB4" box="[1169,1383,1458,1480]" class="Gastropoda" family="Velutinidae" genus="Variolipallium" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="36B8EAD3FFA8FFEF12DB72B3FA825DBB" box="[1169,1342,1458,1479]" italics="true" pageId="28" pageNumber="952">Variolipallium</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
L12,
|
||
<taxonomicName id="C3CC4D42FFA8FFEF157172D0FBAA5D9A" box="[827,1046,1489,1510]" class="Gastropoda" family="Velutinidae" genus="Variolipallium" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="36B8EAD3FFA8FFEF157172D0FC505D9A" box="[827,1004,1489,1510]" italics="true" pageId="28" pageNumber="952">Variolipallium</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
L14 and
|
||
<taxonomicName id="C3CC4D42FFA8FFEF12C672D0FA265D9A" authorityName="Fassio, Bouchet, Oliverio" authorityYear="2023" box="[1164,1434,1489,1510]" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="regium">
|
||
<emphasis id="36B8EAD3FFA8FFEF12C672D0FA265D9A" box="[1164,1434,1489,1510]" italics="true" pageId="28" pageNumber="952">Variolipallium regium</emphasis>
|
||
</taxonomicName>
|
||
) include specimens with a small to very small dorsal mantle fissure, occasionally marked by a small black spot (visible also in preserved specimens).
|
||
</paragraph>
|
||
<paragraph id="047336C1FFA8FFEE1519714AFDC05885" blockId="28.[827,1444,968,1879]" lastBlockId="29.[145,761,197,1384]" lastPageId="29" lastPageNumber="953" pageId="28" pageNumber="952">
|
||
<taxonomicName id="C3CC4D42FFA8FFEF1519714AFAE55E1D" authority="E. A. Smith, 1881" authorityName="E. A. Smith" authorityYear="1881" box="[851,1369,1611,1633]" class="Gastropoda" family="Velutinidae" genus="Lamellaria" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="patagonica">
|
||
<emphasis id="36B8EAD3FFA8FFEF1519714AFBCC5E1D" box="[851,1136,1611,1633]" italics="true" pageId="28" pageNumber="952">Lamellaria patagonica</emphasis>
|
||
E. A. Smith, 1881
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="DB778863FFA8FFEF1327714DFA1E5E1D" box="[1389,1442,1612,1633]" pageId="28" pageNumber="952">type</typeStatus>
|
||
locality:
|
||
<collectingCountry id="7CDB7651FFA8FFEF15E1716BFBA65EFC" box="[939,1050,1642,1664]" name="Trinidad and Tobago" pageId="28" pageNumber="952">Trinidad</collectingCountry>
|
||
Channel,
|
||
<collectingCountry id="7CDB7651FFA8FFEF12EE716BFB555EFC" box="[1188,1257,1642,1664]" name="Chile" pageId="28" pageNumber="952">Chile</collectingCountry>
|
||
, in
|
||
<quantity id="C3349B24FFA8FFEF1355716BFADC5E03" box="[1311,1376,1642,1664]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="28" pageNumber="952" unit="m" value="54.0">54 m</quantity>
|
||
) and
|
||
<taxonomicName id="C3CC4D42FFA8FFEF15717188FB1F5EE3" authority="Strebel, 1906" authorityName="Strebel" authorityYear="1906" box="[827,1187,1673,1695]" class="Gastropoda" family="Velutinidae" genus="Lamellaria" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="elata">
|
||
<emphasis id="36B8EAD3FFA8FFEF15717188FBBC5EE2" box="[827,1024,1673,1694]" italics="true" pageId="28" pageNumber="952">Lamellaria elata</emphasis>
|
||
Strebel, 1906
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="DB778863FFA8FFEF12FE7188FB5B5EE2" box="[1204,1255,1673,1694]" pageId="28" pageNumber="952">type</typeStatus>
|
||
locality: Puerto Condor,
|
||
<collectingCountry id="7CDB7651FFA8FFEF15DE71A6FC6D5EC1" box="[916,977,1703,1725]" name="Chile" pageId="28" pageNumber="952">Chile</collectingCountry>
|
||
) belong to this genus based on their fragile and high-spired shells, smooth mantle and radular formula and shape.
|
||
<taxonomicName id="C3CC4D42FFA8FFEF126E71E4FAD35E86" authorityName="Fassio & Stefani & Russini & Buge & Bouchet & Treneman & Malaquias & Schiaparelli & Modica & Oliverio" authorityYear="2023" baseAuthorityName="E. A. Smith" baseAuthorityYear="1881" box="[1060,1391,1765,1786]" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="patagonicum">
|
||
<emphasis id="36B8EAD3FFA8FFEF126E71E4FAD35E86" box="[1060,1391,1765,1786]" italics="true" pageId="28" pageNumber="952">Variolipallium patagonicum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C3CC4D42FFA8FFEF15717002FBEE5F64" authorityName="Fassio & Stefani & Russini & Buge & Bouchet & Treneman & Malaquias & Schiaparelli & Modica & Oliverio" authorityYear="2023" baseAuthorityName="Strebel" baseAuthorityYear="1906" box="[827,1106,1795,1816]" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="952" phylum="Mollusca" rank="species" species="elatum">
|
||
<emphasis id="36B8EAD3FFA8FFEF15717002FBEE5F64" box="[827,1106,1795,1816]" italics="true" pageId="28" pageNumber="952">Variolipallium elatum</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by a more rapidly expanding first whorl in the former and, consequently, a higher shell in the latter, also resulting in a flatter body in
|
||
<taxonomicName id="C3CC4D42FFA9FFEE172477C4FD0D58A7" authorityName="Fassio & Stefani & Russini & Buge & Bouchet & Treneman & Malaquias & Schiaparelli & Modica & Oliverio" authorityYear="2023" baseAuthorityName="E. A. Smith" baseAuthorityYear="1881" box="[366,689,197,219]" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="953" phylum="Mollusca" rank="species" species="patagonicum">
|
||
<emphasis id="36B8EAD3FFA9FFEE172477C4FD0D58A7" box="[366,689,197,219]" italics="true" pageId="29" pageNumber="953">Variolipallium patagonicum</emphasis>
|
||
</taxonomicName>
|
||
and a more globose one in
|
||
<taxonomicName id="C3CC4D42FFA9FFEE173D77E5FDC95885" authorityName="Fassio & Stefani & Russini & Buge & Bouchet & Treneman & Malaquias & Schiaparelli & Modica & Oliverio" authorityYear="2023" baseAuthorityName="Strebel" baseAuthorityYear="1906" box="[375,629,228,249]" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="953" phylum="Mollusca" rank="species" species="elatum">
|
||
<emphasis id="36B8EAD3FFA9FFEE173D77E5FDC95885" box="[375,629,228,249]" italics="true" pageId="29" pageNumber="953">Variolipallium elatum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="047336C1FFA9FFEE16E37603FD225993" blockId="29.[145,761,197,1384]" pageId="29" pageNumber="953">
|
||
<emphasis id="36B8EAD3FFA9FFEE16E37603FEC8596B" box="[169,372,258,279]" italics="true" pageId="29" pageNumber="953">Lamellariaampla</emphasis>
|
||
Strebel, 1906 (Ushuaia,
|
||
<collectingCountry id="7CDB7651FFA9FFEE14377603FD4D5964" box="[637,753,258,280]" name="Argentina" pageId="29" pageNumber="953">Argentina</collectingCountry>
|
||
) might also belong here (based on shell shape and shell consistency), but further analysis of the
|
||
<typeStatus id="DB778863FFA9FFEE14147641FD335929" box="[606,655,320,341]" pageId="29" pageNumber="953">type</typeStatus>
|
||
material is necessary, because the radula, in particular, is not described in the original description. However, the general description, in particular the jelly-like mantle, wrinkled, grey with darker spots, would also be compatible with a position in
|
||
<taxonomicName id="C3CC4D42FFA9FFEE17A876D8FD265993" authorityName="Fassio, Bouchet, Schiaparelli & Oliverio" authorityYear="2023" box="[482,666,473,495]" class="Gastropoda" family="Velutinidae" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="943" phylum="Mollusca" rank="subFamily" subFamily="Marseniopsinae">Marseniopsinae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="047336C1FFA9FFEE16E376F9FEC55B3C" blockId="29.[145,761,197,1384]" pageId="29" pageNumber="953">
|
||
<taxonomicName id="C3CC4D42FFA9FFEE16E376F9FE0E5A71" baseAuthorityName="Bergh" baseAuthorityYear="1907" box="[169,434,504,525]" class="Gastropoda" family="Velutinidae" genus="Lamellaria" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="953" phylum="Mollusca" rank="species" species="leptoconcha">
|
||
<emphasis id="36B8EAD3FFA9FFEE16E376F9FE0E5A71" box="[169,434,504,525]" italics="true" pageId="29" pageNumber="953">Lamellaria leptoconcha</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="7CDB7651FFA9FFEE178A76F9FDED5A72" box="[448,593,504,526]" name="South Africa" pageId="29" pageNumber="953">South Africa</collectingCountry>
|
||
,
|
||
<collectingRegion id="C608F823FFA9FFEE141076F9FD285A71" box="[602,660,504,525]" country="South Africa" name="Western Cape" pageId="29" pageNumber="953">Cape</collectingRegion>
|
||
Point, in
|
||
<quantity id="C3349B24FFA9FFEE16DB7516FE915A50" box="[145,301,535,556]" metricMagnitude="3" metricUnit="m" metricValue="1.1885" metricValueMax="1.28" metricValueMin="1.097" pageId="29" pageNumber="953" unit="m" value="1188.5" valueMax="1280.0" valueMin="1097.0">1097–1280 m</quantity>
|
||
) is included in
|
||
<taxonomicName id="C3CC4D42FFA9FFEE17AB7517FD305A57" authorityName="Fassio, Bouchet & Oliverio" authorityYear="2023" box="[481,652,534,555]" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="953" phylum="Mollusca" rank="genus">
|
||
<emphasis id="36B8EAD3FFA9FFEE17AB7517FD305A57" box="[481,652,534,555]" italics="true" pageId="29" pageNumber="953">Variolipallium</emphasis>
|
||
</taxonomicName>
|
||
based on the membranaceus texture of the shell and the shape of the rachidian tooth.
|
||
<bibRefCitation id="605D4B30FFA9FFEE17E67555FDFC5A15" author="Bergh LSR" box="[428,576,596,618]" pageId="29" pageNumber="953" pagination="1 - 144" refId="ref25396" refString="Bergh LSR. 1907. The Opisthobranchiata of South Africa. In: Marine InVestigations of South Africa 5 (1) [Dated 1908]. Page (s): 106, pl. 9, figs 17 - 20; pl. 10, figs 1, 2. Transactions of the South African Philosophical Society 17: 1 - 144, pls 1 - 14." type="journal article" year="1907">Bergh (1907</bibRefCitation>
|
||
: pl. IX, fig. 18) sketched a rachidian tooth with a not marked but visible ‘V’-shaped base (similar to our SEM photographs;
|
||
<figureCitation id="9CF72A44FFA9FFEE16DB75B1FEAF5AB9" box="[145,275,688,710]" captionStart="Figure 10" captionStartId="17.[145,226,1664,1686]" captionTargetBox="[212,1358,198,1631]" captionTargetId="figure-24@17.[209,1361,195,1634]" captionTargetPageId="17" captionText="Figure 10. Velutinidae radulae. A, Marseniopsis mollis, CE 6137. B, Marsenina rhombica, BAU 4229. C, Velutina cf. coriacea, LACM 182870. D, Calyptoconcha pellucida, BAU 3751. E, Variolipallium regium sp. nov., MNHN-IM-2013-69285, holotype. F, Variolipallium sp. (L10), MNHN-IM-2013-69280. G, Pacifica lentiginosa sp. nov., MNHN-IM-2009-16140, holotype. H, Pacifica sp. (L15), MNHN-IM-2013-59969. I, Djiboutia sp. (L22), MNHN-IM-2019-9053. J, Coriocella sp. (L27), MNHN-IM-2013-84230. K, Lamellaria mopsicolor, MNHN-IM-2013-73086. L, Lamellaria latens complex (L41), MNHN-IM-2019-14113. M, Marsenia herberti, MNHN-IM-2019-8503. N, Marsenia sp. (L45), MNHN-IM-2013-59960. O, Marsenia perspicua complex (L55), MNHN-IM-2019-5980. Scale bars: A–O, 100 µm. For the sequenced specimens, the corresponding species code (see Fig. 3) is reported in parentheses." figureDoi="http://doi.org/10.5281/zenodo.7814338" httpUri="https://zenodo.org/record/7814338/files/figure.png" pageId="29" pageNumber="953">Fig. 10E, F</figureCitation>
|
||
), and in the description underlined that the difference between this species and
|
||
<taxonomicName id="C3CC4D42FFA9FFEE14CC75CEFEA95B7E" authorityName="Bergh" authorityYear="1899" class="Gastropoda" family="Velutinidae" genus="Marsenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="953" phylum="Mollusca" rank="species" species="leptolemma">
|
||
<emphasis id="36B8EAD3FFA9FFEE14CC75CEFEA95B7E" italics="true" pageId="29" pageNumber="953">Marsenia leptolemma</emphasis>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="C3CC4D42FFA9FFEE177D75ECFDF75B7E" baseAuthorityName="A. E. Verrill" baseAuthorityYear="1880" box="[311,587,749,770]" class="Gastropoda" family="Velutinidae" genus="Calyptoconcha" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="953" phylum="Mollusca" rank="species" species="pellucida">
|
||
<emphasis id="36B8EAD3FFA9FFEE177D75ECFDF75B7E" box="[311,587,749,770]" italics="true" pageId="29" pageNumber="953">Calyptoconcha pellucida</emphasis>
|
||
</taxonomicName>
|
||
) was the shape of the rachidian tooth (that in
|
||
<taxonomicName id="C3CC4D42FFA9FFEE17B3740DFD1F5B5D" authorityName="Bouchet & Waren" authorityYear="1993" box="[505,675,780,801]" class="Gastropoda" family="Velutinidae" genus="Calyptoconcha" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="953" phylum="Mollusca" rank="genus">
|
||
<emphasis id="36B8EAD3FFA9FFEE17B3740DFD1F5B5D" box="[505,675,780,801]" italics="true" pageId="29" pageNumber="953">Calyptoconcha</emphasis>
|
||
</taxonomicName>
|
||
indeed has a squared base).
|
||
</paragraph>
|
||
<paragraph id="047336C1FFA9FFEE16E37448FDD25CEE" blockId="29.[145,761,197,1384]" pageId="29" pageNumber="953">
|
||
<taxonomicName id="C3CC4D42FFA9FFEE16E37448FE015B22" authorityName="Hutton" authorityYear="1882" box="[169,445,841,862]" class="Gastropoda" family="Velutinidae" genus="Lamellaria" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="953" phylum="Mollusca" rank="species" species="cerebroides">
|
||
<emphasis id="36B8EAD3FFA9FFEE16E37448FE015B22" box="[169,445,841,862]" italics="true" pageId="29" pageNumber="953">Lamellaria cerebroides</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectingRegion id="C608F823FFA9FFEE17857448FDF85B23" box="[463,580,841,863]" country="New Zealand" name="Auckland" pageId="29" pageNumber="953">Auckland</collectingRegion>
|
||
,
|
||
<collectingCountry id="7CDB7651FFA9FFEE141B744BFD4C5B23" box="[593,752,841,863]" name="New Zealand" pageId="29" pageNumber="953">New Zealand</collectingCountry>
|
||
) is included in
|
||
<taxonomicName id="C3CC4D42FFA9FFEE17757469FE4C5B01" authorityName="Fassio, Bouchet & Oliverio" authorityYear="2023" box="[319,496,872,893]" class="Gastropoda" family="Velutinidae" genus="Variolipallium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="953" phylum="Mollusca" rank="genus">
|
||
<emphasis id="36B8EAD3FFA9FFEE17757469FE4C5B01" box="[319,496,872,893]" italics="true" pageId="29" pageNumber="953">Variolipallium</emphasis>
|
||
</taxonomicName>
|
||
based on the radular formula and shape, and the wrinkled dorsum appearance.
|
||
<taxonomicName id="C3CC4D42FFA9FFEE177B74A4FD835BC6" authorityName="Odhner" authorityYear="1924" box="[305,575,933,954]" class="Gastropoda" family="Velutinidae" genus="Lamellaria" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="953" phylum="Mollusca" rank="species" species="verrucosa">
|
||
<emphasis id="36B8EAD3FFA9FFEE177B74A4FD835BC6" box="[305,575,933,954]" italics="true" pageId="29" pageNumber="953">Lamellaria Ʋerrucosa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="36B8EAD3FFA9FFEE141E74A7FF585BA5" italics="true" pageId="29" pageNumber="953">
|
||
=
|
||
<taxonomicName id="C3CC4D42FFA9FFEE142474A4FF585BA5" authorityName="Ev. Marcus" authorityYear="1987" class="Gastropoda" family="Velutinidae" genus="Lamellaria" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="953" phylum="Mollusca" rank="species" species="nodosa">Lamellaria nodosa</taxonomicName>
|
||
</emphasis>
|
||
) (
|
||
<collectingRegion id="C608F823FFA9FFEE16B474C5FECD5BA6" box="[254,369,964,986]" country="New Zealand" name="Auckland" pageId="29" pageNumber="953">Auckland</collectingRegion>
|
||
Islands) is also included in this genus based on the membranaceus texture of the shell, the rugose appearance of the dorsum, and the radular formula and shape.
|
||
<bibRefCitation id="605D4B30FFA9FFEE17C37321FD865C49" author="Odhner NH" box="[393,570,1056,1078]" pageId="29" pageNumber="953" pagination="1 - 90" refId="ref27413" refString="Odhner NH. 1924. Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916. XIX. New Zealand Mollusca. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i KjobenhaVn 77: 1 - 90, pls 1, 2. Page: 31" type="book chapter" year="1924">Odhner (1924)</bibRefCitation>
|
||
described it as being similar to
|
||
<taxonomicName id="C3CC4D42FFA9FFEE171A733EFD8B5C2F" box="[336,567,1086,1108]" class="Gastropoda" family="Velutinidae" genus="Marseniopsis" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="953" phylum="Mollusca" rank="species" species="mollis">
|
||
<emphasis id="36B8EAD3FFA9FFEE171A733EFD8B5C2F" box="[336,567,1086,1108]" italics="true" pageId="29" pageNumber="953">Marseniopsis mollis</emphasis>
|
||
</taxonomicName>
|
||
, but the radular formula of
|
||
<taxonomicName id="C3CC4D42FFA9FFEE1747735CFDBC5C0E" authorityName="Odhner" authorityYear="1924" box="[269,512,1117,1138]" class="Gastropoda" family="Velutinidae" genus="Lamellaria" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="953" phylum="Mollusca" rank="species" species="verrucosa">
|
||
<emphasis id="36B8EAD3FFA9FFEE1747735CFDBC5C0E" box="[269,512,1117,1138]" italics="true" pageId="29" pageNumber="953">Lamellaria Ʋerrucosa</emphasis>
|
||
</taxonomicName>
|
||
clearly indicates that it belongs to the subfamily
|
||
<taxonomicName id="C3CC4D42FFA9FFEE178C737DFDD65CEE" authorityName="d'Orbigny" authorityYear="1841" box="[454,618,1148,1170]" class="Gastropoda" family="Velutinidae" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="953" phylum="Mollusca" rank="subFamily" subFamily="Lamellariinae">Lamellariinae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="047336C1FFA9FFEE16E3739BFD685D14" blockId="29.[145,761,197,1384]" pageId="29" pageNumber="953">
|
||
The original description of
|
||
<taxonomicName id="C3CC4D42FFA9FFEE1793739BFF745CB3" authority="Gray, 1850" authorityName="Gray" authorityYear="1850" class="Gastropoda" family="Velutinidae" genus="Lamellaria" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="953" phylum="Mollusca" rank="species" species="ophione">
|
||
<emphasis id="36B8EAD3FFA9FFEE1793739BFD0B5CD3" box="[473,695,1178,1199]" italics="true" pageId="29" pageNumber="953">Lamellaria ophione</emphasis>
|
||
Gray, 1850
|
||
</taxonomicName>
|
||
is general (fitting most lamellariine genera) and does not report any information regarding diagnostic characters. However, considering the
|
||
<typeStatus id="DB778863FFA9FFEE142B73F6FD295D70" box="[609,661,1271,1292]" pageId="29" pageNumber="953">type</typeStatus>
|
||
locality (
|
||
<collectingRegion id="C608F823FFA9FFEE16D27214FEB65D57" box="[152,266,1301,1323]" country="New Zealand" name="Auckland" pageId="29" pageNumber="953">Auckland</collectingRegion>
|
||
,
|
||
<collectingCountry id="7CDB7651FFA9FFEE175C7214FE0E5D57" box="[278,434,1301,1323]" name="New Zealand" pageId="29" pageNumber="953">New Zealand</collectingCountry>
|
||
), there is a chance that this species also belongs to this genus, because this is the only lamellarine genus recorded in this area so far.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |