treatments-xml/data/89/40/87/894087A5FFA3FFE7F5F1FB3DFDDB01D7.xml

<document ID-DOI="10.1655/d-18-00019" ID-GBIF-Dataset="4050ebbb-b790-47a8-a47d-21cb54e9770d" ID-ISSN="1938-5099" ID-Zenodo-Dep="7717262" checkinTime="1678363461378" checkinUser="felipe" docAuthor="Ospina-Sarria, Jhon Jairo" docDate="2019" docId="894087A5FFA3FFE7F5F1FB3DFDDB01D7" docLanguage="en" docName="Herpetologica.75.1.85-95.pdf" docOrigin="Herpetologica 75 (1)" docStyle="DocumentStyle:F3392F244A96CCB72E455B80E109DFAC.1:Herpetologica.2015-.journal_article.0cover" docStyleId="F3392F244A96CCB72E455B80E109DFAC" docStyleName="Herpetologica.2015-.journal_article.0cover" docStyleVersion="1" docTitle="Pristimantis sneiderni Ospina-Sarria, 2019, sp. nov." docType="treatment" docVersion="2" lastPageNumber="93" masterDocId="7579FFDDFFA7FFEFF61CFFDFFFBD0756" masterDocTitle="Two New Species of Pristimantis (Amphibia: Anura: Strabomantidae) from Southwestern Colombia" masterLastPageNumber="95" masterPageNumber="85" pageNumber="89" updateTime="1678468765219" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two New Species of Pristimantis (Amphibia: Anura: Strabomantidae) from Southwestern Colombia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ospina-Sarria, Jhon Jairo</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Herpetologica</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-03-05</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>75</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>85</mods:start>
<mods:end>95</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1655/d-18-00019</mods:identifier>
<mods:identifier type="GBIF-Dataset">4050ebbb-b790-47a8-a47d-21cb54e9770d</mods:identifier>
<mods:identifier type="ISSN">1938-5099</mods:identifier>
<mods:identifier type="Zenodo-Dep">7717262</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7717270" ID-Zenodo-Dep="7717270" LSID="urn:lsid:plazi:treatment:894087A5FFA3FFE7F5F1FB3DFDDB01D7" httpUri="http://treatment.plazi.org/id/894087A5FFA3FFE7F5F1FB3DFDDB01D7" lastPageId="8" lastPageNumber="93" pageId="4" pageNumber="89">
<subSubSection box="[1005,1353,1248,1273]" pageId="4" pageNumber="89" type="nomenclature">
<paragraph blockId="4.[1005,1353,1248,1273]" box="[1005,1353,1248,1273]" pageId="4" pageNumber="89">
<heading box="[1005,1353,1248,1273]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="89" reason="2">
<taxonomicName ID-CoL="4MJPC" authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" box="[1005,1258,1248,1273]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="93" phylum="Chordata" rank="species" species="sneiderni" status="sp. nov.">
<emphasis box="[1005,1258,1248,1273]" italics="true" pageId="4" pageNumber="89">
Pristimantis 
<emphasis bold="true" box="[1147,1258,1248,1272]" italics="true" pageId="4" pageNumber="89">sneiderni</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1266,1353,1249,1272]" pageId="4" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1026,1332,1278,1303]" pageId="4" pageNumber="89" type="description">
<paragraph blockId="4.[1026,1332,1278,1303]" box="[1026,1332,1278,1303]" pageId="4" pageNumber="89">
(
<figureCitation box="[1034,1154,1278,1302]" captionStart="FIG" captionStartId="5.[142,156,1840,1859]" captionTargetBox="[118,1490,191,1814]" captionTargetId="figure-0@5.[118,1490,191,1814]" captionTargetPageId="5" captionText="FIG. 5.—Dorsolateral, ventral, and lateral views of Pristimantis sneiderni (A, B; holotype, KU 145021, adult female, snout–vent length [SVL] 28 mm, head length [HL] 9.0 mm), P. boulengeri (C, D; paratype, KU 169062, adult female, SVL 34 mm, HL 10.6 mm), and P. schultei (E, F; paratype, KU 209504, adult female, SVL 33 mm, HL 10.5 mm). A color version of this figure is available online." figureDoi="http://doi.org/10.5281/zenodo.7717276" httpUri="https://zenodo.org/record/7717276/files/figure.png" pageId="4" pageNumber="89">Figs. 5A,B</figureCitation>
, 
<figureCitation box="[1168,1225,1278,1302]" captionStart="FIG" captionStartId="7.[142,156,1522,1541]" captionTargetBox="[118,1491,191,1496]" captionTargetId="figure-217@7.[118,1492,191,1496]" captionTargetPageId="7" captionText="FIG. 6.—Dorsolateral and ventral views of the male of Pristimantis sneiderni (A, B; paratype, KU 145019, adult male, snout–vent length [SVL] 23 mm) and P. deŋi (C, D; holotype, MUSM 8968, adult male, SVL 22.7 mm; photo Magnolia Press, reproduced with permission from copyright holder). A color version of this figure is available online." figureDoi="http://doi.org/10.5281/zenodo.7717278" httpUri="https://zenodo.org/record/7717278/files/figure.png" pageId="4" pageNumber="89">6A,B</figureCitation>
; 
<tableCitation box="[1238,1324,1278,1303]" captionStart="TABLE" captionStartId="6.[169,182,188,207]" captionTargetPageId="6" captionText="TABLE 2.—Selected meristic, mensural (in mm), and proportional (in percentages) data from the type series of Pristimantis sneiderni. Where n&gt; 1, values are given as range (mean ± 1 SD)." httpUri="http://table.plazi.org/id/5596663BFFA1FFE9F6B5FF63FDA207A9" pageId="4" pageNumber="89" tableUuid="5596663BFFA1FFE9F6B5FF63FDA207A9">Table 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="89" type="materials_examined">
<paragraph blockId="4.[844,1514,1324,2001]" pageId="4" pageNumber="89">
<materialsCitation collectingDate="1938-12" collectionCode="KU" collectorName="Kjell von Sneidern" country="Colombia" latitude="2.3833332" location="Cauca" longLatPrecision="1307" longitude="-76.316666" pageId="4" pageNumber="89" specimenCode="KU 145021" specimenCount="1" stateProvince="Cauca" typeStatus="holotype">
<emphasis bold="true" box="[871,990,1324,1348]" pageId="4" pageNumber="89">
<typeStatus box="[871,986,1324,1348]" pageId="4" pageNumber="89">Holotype</typeStatus>
.
</emphasis>
— 
<specimenCode box="[1018,1150,1324,1348]" collectionCode="KU" country="USA" name="Biodiversity Institute, University of Kansas" pageId="4" pageNumber="89" type="University or college">KU 145021</specimenCode>
, adult female from the Laguna San Rafael, 3450-m elevation (
<geoCoordinate box="[1200,1281,1350,1378]" degrees="2" direction="north" minutes="23" orientation="latitude" pageId="4" pageNumber="89" precision="925" value="2.3833332">
2°23 
<emphasis box="[1251,1256,1350,1367]" italics="true" pageId="4" pageNumber="89">
<superScript attach="right" box="[1251,1256,1350,1367]" fontSize="7" pageId="4" pageNumber="89">′</superScript>
</emphasis>
N
</geoCoordinate>
, 
<geoCoordinate box="[1297,1395,1350,1378]" degrees="76" direction="west" minutes="19" orientation="longitude" pageId="4" pageNumber="89" precision="925" value="-76.316666">
76°19 
<emphasis box="[1360,1365,1350,1367]" italics="true" pageId="4" pageNumber="89">
<superScript attach="right" box="[1360,1365,1350,1367]" fontSize="7" pageId="4" pageNumber="89">′</superScript>
</emphasis>
W
</geoCoordinate>
; datum ¼ WGS84), Departamento del 
<collectingRegion box="[1188,1260,1384,1408]" country="Colombia" name="Cauca" pageId="4" pageNumber="89">Cauca</collectingRegion>
, 
<collectingCountry box="[1279,1389,1384,1408]" name="Colombia" pageId="4" pageNumber="89">Colombia</collectingCountry>
, one of a series collected by 
<collectorName box="[1059,1268,1413,1437]" pageId="4" pageNumber="89">Kjell von Sneidern</collectorName>
in 
<date box="[1308,1490,1413,1437]" pageId="4" pageNumber="89" value="1938-12">
<collectingDate box="[1308,1490,1413,1437]" pageId="4" pageNumber="89" value="1938-12">December 1938</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph blockId="4.[844,1514,1324,2001]" pageId="4" pageNumber="89">
<materialsCitation collectionCode="KU" pageId="4" pageNumber="89" specimenCode="KU 145019-20, 145022" specimenCount="2" typeStatus="paratype">
<emphasis bold="true" box="[871,998,1443,1467]" pageId="4" pageNumber="89">
<typeStatus box="[871,994,1443,1467]" pageId="4" pageNumber="89">Paratypes</typeStatus>
.
</emphasis>
— 
<specimenCode box="[1026,1194,1443,1467]" collectionCode="KU" country="USA" name="Biodiversity Institute, University of Kansas" pageId="4" pageNumber="89" type="University or college">KU 145019–20</specimenCode>
, 
<specimenCode box="[1205,1287,1443,1467]" collectionCode="KU" country="USA" name="Biodiversity Institute, University of Kansas" pageId="4" pageNumber="89" type="University or college">145022</specimenCode>
, adult males collect-ed with the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="91" pageId="4" pageNumber="89" type="diagnosis">
<paragraph blockId="4.[844,1514,1324,2001]" lastBlockId="6.[142,813,996,2001]" lastPageId="6" lastPageNumber="91" pageId="4" pageNumber="89">
<emphasis bold="true" box="[871,997,1502,1526]" pageId="4" pageNumber="89">Diagnosis.</emphasis>
— 
<taxonomicName authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" box="[1024,1271,1503,1527]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="89" phylum="Chordata" rank="species" species="sneiderni">
<emphasis box="[1024,1271,1503,1527]" italics="true" pageId="4" pageNumber="89">Pristimantis sneiderni</emphasis>
</taxonomicName>
<materialsCitation collectionCode="III, IV, V" lastPageId="6" lastPageNumber="91" pageId="4" pageNumber="89" specimenCount="2" specimenCount-adult="2">
is diagnosed by the following combination of characters: (1) skin on dorsum bearing low warts (most obvious on flanks); ventral skin areolate; discoidal fold present, well anterior to groin; dorsolateral folds and ridges absent in scapula region; (2) tympanic membrane not evident; tympanic annulus present, its upper edge covered by supratympanic fold, extending only just posterior to tympanic annulus; (3) snout round in dorsal view and in profile, with papilla at the tip (most obvious in males); canthus rostralis concave; (4) upper eyelid warty, narrower than IOD (50.0–80.0% IOD); interocular tubercle absent; cranial crests absent; (5) choanae small, ovoid; not concealed by palatal shelf of maxillary arch; dentigerous processes of vomers absent; (6) males lacking vocal slits and nuptial pads; (7) finger I shorter than finger II; discs and circumferential grooves present on all fingers; discs truncate, except on finger I, which is round; disc of finger I smaller than that of finger II and this in turn smaller than discs on fingers 
<collectionCode box="[330,361,1026,1049]" pageId="6" pageNumber="91">III</collectionCode>
and 
<collectionCode box="[424,453,1026,1049]" pageId="6" pageNumber="91">IV</collectionCode>
; (8) fingers with narrow lateral fringes; palmar tubercle divided; thenar tubercle oval, same size as palmar tubercle; supernumerary tubercles low, distributed on all fleshy parts of palm; subarticular tubercles round, prominent, two on thumb and second finger and three on third and fourth fingers, and larger than supernumerary tubercles; (9) antebrachial tubercle present; other ulnar tubercles absent; (10) heel bearing one small tubercle; outer edge of tarsus bearing low tubercles; inner tarsal fold absent; (11) inner metatarsal tubercle elongate, its length twice its width; conical outer metatarsal tubercle onefourth the size of inner metatarsal tubercle; subarticular tubercles round, prominent, two on toes I and II, three on toes 
<collectionCode box="[197,228,1412,1435]" pageId="6" pageNumber="91">III</collectionCode>
and 
<collectionCode box="[289,308,1412,1435]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="91" type="Museum">V</collectionCode>
, and four on toe 
<collectionCode box="[512,541,1412,1435]" pageId="6" pageNumber="91">IV</collectionCode>
; supernumerary plantar tubercles low, rounded, on proximal segments of toes; (12) toes bearing narrow lateral fringes; webbing absent; toe 
<collectionCode box="[781,812,1472,1495]" pageId="6" pageNumber="91">III</collectionCode>
shorter than toe 
<collectionCode box="[338,357,1502,1525]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="91" type="Museum">V</collectionCode>
; toe 
<collectionCode box="[418,449,1502,1525]" pageId="6" pageNumber="91">III</collectionCode>
extending to distal edge of the antepenultimate subarticular tubercle of toe 
<collectionCode box="[692,722,1531,1554]" pageId="6" pageNumber="91">IV</collectionCode>
; toe 
<collectionCode box="[796,813,1531,1554]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="91" type="Museum">V</collectionCode>
reaching the edge distal to the penultimate subarticular tubercle of toe 
<collectionCode box="[312,341,1591,1614]" pageId="6" pageNumber="91">IV</collectionCode>
; discs and circumferential grooves present on all toes; discs of toes 
<collectionCode box="[410,437,1621,1644]" pageId="6" pageNumber="91">III</collectionCode>
– 
<collectionCode box="[451,470,1621,1644]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="91" type="Museum">V</collectionCode>
equal to discs on fingers II– 
<collectionCode box="[784,813,1621,1644]" pageId="6" pageNumber="91">IV</collectionCode>
and larger than discs of toe I; (13) color in preservative: dorsum and flanks tan with small dark brown markings; venter tan with brown mottling, mainly on chest, canthal stripe dark brown, throat tan with fine brown spots; posterior surfaces of thighs uniformly tan; (14) males smaller than females; SVL in 
<specimenCount box="[329,452,1799,1823]" pageId="6" pageNumber="91" type="adult">three adult</specimenCount>
males 21.0, 22.0, and 23.0 mm, in 
<specimenCount box="[173,278,1828,1852]" pageId="6" pageNumber="91" type="adult">one adult</specimenCount>
female 28.0 mm
</materialsCitation>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7717276" ID-Zenodo-Dep="7717276" httpUri="https://zenodo.org/record/7717276/files/figure.png" pageId="5" pageNumber="90" startId="5.[142,156,1840,1859]" targetBox="[118,1490,191,1814]" targetPageId="5">
<paragraph blockId="5.[118,1490,1839,1907]" pageId="5" pageNumber="90">
FIG. 5.—Dorsolateral, ventral, and lateral views of 
<taxonomicName authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" box="[585,777,1839,1858]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="90" phylum="Chordata" rank="species" species="sneiderni">
<emphasis box="[585,777,1839,1858]" italics="true" pageId="5" pageNumber="90">Pristimantis sneiderni</emphasis>
</taxonomicName>
(A, B; holotype, KU 145021, adult female, snout–vent length [SVL] 28 mm, head length [HL] 9.0 mm), 
<taxonomicName baseAuthorityName="Lynch" baseAuthorityYear="1981" box="[313,429,1863,1882]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="90" phylum="Chordata" rank="species" species="boulengeri">
<emphasis box="[313,429,1863,1882]" italics="true" pageId="5" pageNumber="90">P. boulengeri</emphasis>
</taxonomicName>
(C, D; paratype, KU 169062, adult female, SVL 34 mm, HL 10.6 mm), and 
<taxonomicName baseAuthorityName="Duellman" baseAuthorityYear="1990" box="[1096,1185,1863,1882]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="90" phylum="Chordata" rank="species" species="schultei">
<emphasis box="[1096,1185,1863,1882]" italics="true" pageId="5" pageNumber="90">P. schultei</emphasis>
</taxonomicName>
(E, F; paratype, KU 209504, adult female, SVL 33 mm, HL 10.5 mm). A color version of this figure is available online.
</paragraph>
</caption>
<caption ID-Table-UUID="5596663BFFA1FFE9F6B5FF63FDA207A9" httpUri="http://table.plazi.org/id/5596663BFFA1FFE9F6B5FF63FDA207A9" pageId="6" pageNumber="91" startId="6.[169,182,188,207]" targetBox="[142,812,290,937]" targetIsTable="true" targetPageId="6">
<paragraph blockId="6.[142,813,188,255]" pageId="6" pageNumber="91">
TABLE 2.—Selected meristic, mensural (in mm), and proportional (in percentages) data from the type series of 
<taxonomicName authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" box="[508,700,212,231]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="sneiderni">
<emphasis box="[508,700,212,231]" italics="true" pageId="6" pageNumber="91">Pristimantis sneiderni</emphasis>
</taxonomicName>
. Where 
<emphasis box="[777,788,213,231]" italics="true" pageId="6" pageNumber="91">n</emphasis>
&gt; 1, values are given as range (mean ± 1 SD).
</paragraph>
</caption>
<paragraph pageId="6" pageNumber="91">
<table box="[142,812,290,937]" gridcols="3" gridrows="27" pageId="6" pageNumber="91">
<tr box="[142,812,290,304]" gridrow="0" pageId="6" pageNumber="91" rowspan-0="1" rowspan-2="1">
<th box="[459,582,290,304]" gridcol="1" gridrow="0" pageId="6" pageNumber="91">Female (holotype:</th>
</tr>
<tr box="[142,812,308,323]" gridrow="1" pageId="6" pageNumber="91">
<th box="[142,429,308,323]" gridcol="0" gridrow="1" pageId="6" pageNumber="91">Trait</th>
<td box="[459,582,308,323]" gridcol="1" gridrow="1" pageId="6" pageNumber="91">KU 145021)</td>
<td box="[611,812,308,323]" gridcol="2" gridrow="1" pageId="6" pageNumber="91">
Males (
<emphasis box="[714,723,309,323]" italics="true" pageId="6" pageNumber="91">n</emphasis>
¼ 3)
</td>
</tr>
<tr box="[142,812,343,363]" gridrow="2" pageId="6" pageNumber="91">
<th box="[142,429,343,363]" gridcol="0" gridrow="2" pageId="6" pageNumber="91">Snout–vent length</th>
<td box="[459,582,343,363]" gridcol="1" gridrow="2" pageId="6" pageNumber="91">28.0</td>
<td box="[611,812,343,363]" gridcol="2" gridrow="2" pageId="6" pageNumber="91">21.0–23.0 (22.0 ± 1.0)</td>
</tr>
<tr box="[142,812,367,386]" gridrow="3" pageId="6" pageNumber="91">
<th box="[142,429,367,386]" gridcol="0" gridrow="3" pageId="6" pageNumber="91">Head width</th>
<td box="[459,582,367,386]" gridcol="1" gridrow="3" pageId="6" pageNumber="91">11.0</td>
<td box="[611,812,367,386]" gridcol="2" gridrow="3" pageId="6" pageNumber="91">8.0–8.5 (8.2 ± 0.2)</td>
</tr>
<tr box="[142,812,391,410]" gridrow="4" pageId="6" pageNumber="91">
<th box="[142,429,391,410]" gridcol="0" gridrow="4" pageId="6" pageNumber="91">Head length</th>
<td box="[459,582,391,410]" gridcol="1" gridrow="4" pageId="6" pageNumber="91">9.0</td>
<td box="[611,812,391,410]" gridcol="2" gridrow="4" pageId="6" pageNumber="91">6.0–7.0 (6.5 ± 0.5)</td>
</tr>
<tr box="[142,812,415,434]" gridrow="5" pageId="6" pageNumber="91">
<th box="[142,429,415,434]" gridcol="0" gridrow="5" pageId="6" pageNumber="91">Interorbital distance</th>
<td box="[459,582,415,434]" gridcol="1" gridrow="5" pageId="6" pageNumber="91">4.0</td>
<td box="[611,812,415,434]" gridcol="2" gridrow="5" pageId="6" pageNumber="91">2.5–3.0 (2.8 ± 0.2)</td>
</tr>
<tr box="[142,812,439,458]" gridrow="6" pageId="6" pageNumber="91">
<th box="[142,429,439,458]" gridcol="0" gridrow="6" pageId="6" pageNumber="91">Upper eyelid width</th>
<td box="[459,582,439,458]" gridcol="1" gridrow="6" pageId="6" pageNumber="91">2.0</td>
<td box="[611,812,439,458]" gridcol="2" gridrow="6" pageId="6" pageNumber="91">2.0</td>
</tr>
<tr box="[142,812,463,482]" gridrow="7" pageId="6" pageNumber="91">
<th box="[142,429,463,482]" gridcol="0" gridrow="7" pageId="6" pageNumber="91">Eye diameter</th>
<td box="[459,582,463,482]" gridcol="1" gridrow="7" pageId="6" pageNumber="91">3.0</td>
<td box="[611,812,463,482]" gridcol="2" gridrow="7" pageId="6" pageNumber="91">3.0</td>
</tr>
<tr box="[142,812,487,506]" gridrow="8" pageId="6" pageNumber="91">
<th box="[142,429,487,506]" gridcol="0" gridrow="8" pageId="6" pageNumber="91">Eye–nostril distance</th>
<td box="[459,582,487,506]" gridcol="1" gridrow="8" pageId="6" pageNumber="91">2.0</td>
<td box="[611,812,487,506]" gridcol="2" gridrow="8" pageId="6" pageNumber="91">1.4–1.5 (1.4 ± 0.1)</td>
</tr>
<tr box="[142,812,511,530]" gridrow="9" pageId="6" pageNumber="91">
<th box="[142,429,511,530]" gridcol="0" gridrow="9" pageId="6" pageNumber="91">Internarial distance</th>
<td box="[459,582,511,530]" gridcol="1" gridrow="9" pageId="6" pageNumber="91">2.0</td>
<td box="[611,812,511,530]" gridcol="2" gridrow="9" pageId="6" pageNumber="91">1.2–1.4 (1.3 ± 0.1)</td>
</tr>
<tr box="[142,812,534,554]" gridrow="10" pageId="6" pageNumber="91">
<th box="[142,429,534,554]" gridcol="0" gridrow="10" pageId="6" pageNumber="91">Tympanum diameter</th>
<td box="[459,582,534,554]" gridcol="1" gridrow="10" pageId="6" pageNumber="91">1.0</td>
<td box="[611,812,534,554]" gridcol="2" gridrow="10" pageId="6" pageNumber="91">0.6–0.7 (0.6 ± 0.1)</td>
</tr>
<tr box="[142,812,558,578]" gridrow="11" pageId="6" pageNumber="91">
<th box="[142,429,558,578]" gridcol="0" gridrow="11" pageId="6" pageNumber="91">Tibia length</th>
<td box="[459,582,558,578]" gridcol="1" gridrow="11" pageId="6" pageNumber="91">14.0</td>
<td box="[611,812,558,578]" gridcol="2" gridrow="11" pageId="6" pageNumber="91">10.5–11.0 (10.7 ± 0.2)</td>
</tr>
<tr box="[142,812,582,602]" gridrow="12" pageId="6" pageNumber="91">
<th box="[142,429,582,602]" gridcol="0" gridrow="12" pageId="6" pageNumber="91">Foot length</th>
<td box="[459,582,582,602]" gridcol="1" gridrow="12" pageId="6" pageNumber="91">14.0</td>
<td box="[611,812,582,602]" gridcol="2" gridrow="12" pageId="6" pageNumber="91">10.2–10.5 (10.3 ± 0.1)</td>
</tr>
<tr box="[142,812,606,625]" gridrow="13" pageId="6" pageNumber="91">
<th box="[142,429,606,625]" gridcol="0" gridrow="13" pageId="6" pageNumber="91">Head length/snout–vent length</th>
<td box="[459,582,606,625]" gridcol="1" gridrow="13" pageId="6" pageNumber="91">32.1</td>
<td box="[611,812,606,625]" gridcol="2" gridrow="13" pageId="6" pageNumber="91">27.2–30.9</td>
</tr>
<tr box="[142,812,630,649]" gridrow="14" pageId="6" pageNumber="91">
<th box="[142,429,630,649]" gridcol="0" gridrow="14" pageId="6" pageNumber="91">Head width/snout–vent length</th>
<td box="[459,582,630,649]" gridcol="1" gridrow="14" pageId="6" pageNumber="91">39.2</td>
<td box="[611,812,630,649]" gridcol="2" gridrow="14" pageId="6" pageNumber="91">27.2–30.9</td>
</tr>
<tr box="[142,812,654,674]" gridrow="15" pageId="6" pageNumber="91">
<th box="[142,429,654,674]" gridcol="0" gridrow="15" pageId="6" pageNumber="91">Eye diameter/head length</th>
<td box="[459,582,654,674]" gridcol="1" gridrow="15" pageId="6" pageNumber="91">33.3</td>
<td box="[611,812,654,674]" gridcol="2" gridrow="15" pageId="6" pageNumber="91">42.8–50.0</td>
</tr>
<tr box="[142,812,678,697]" gridrow="16" pageId="6" pageNumber="91">
<th box="[142,429,678,697]" gridcol="0" gridrow="16" pageId="6" pageNumber="91">Eye–nostril distance/</th>
<td box="[459,582,678,697]" gridcol="1" gridrow="16" pageId="6" pageNumber="91">66.6</td>
<td box="[611,812,678,697]" gridcol="2" gridrow="16" pageId="6" pageNumber="91">46.6–50.0</td>
</tr>
<tr box="[142,812,702,721]" gridrow="17" pageId="6" pageNumber="91" rowspan-1="1" rowspan-2="1">
<th box="[142,429,702,721]" gridcol="0" gridrow="17" pageId="6" pageNumber="91">eye diameter %</th>
</tr>
<tr box="[142,812,726,745]" gridrow="18" pageId="6" pageNumber="91">
<th box="[142,429,726,745]" gridcol="0" gridrow="18" pageId="6" pageNumber="91">Upper eyelid width/</th>
<td box="[459,582,726,745]" gridcol="1" gridrow="18" pageId="6" pageNumber="91">50.0</td>
<td box="[611,812,726,745]" gridcol="2" gridrow="18" pageId="6" pageNumber="91">66.6–80.0</td>
</tr>
<tr box="[142,812,750,769]" gridrow="19" pageId="6" pageNumber="91" rowspan-1="1" rowspan-2="1">
<th box="[142,429,750,769]" gridcol="0" gridrow="19" pageId="6" pageNumber="91">interorbital distance</th>
</tr>
<tr box="[142,812,774,793]" gridrow="20" pageId="6" pageNumber="91">
<th box="[142,429,774,793]" gridcol="0" gridrow="20" pageId="6" pageNumber="91">Interorbital distance/head width</th>
<td box="[459,582,774,793]" gridcol="1" gridrow="20" pageId="6" pageNumber="91">36.3</td>
<td box="[611,812,774,793]" gridcol="2" gridrow="20" pageId="6" pageNumber="91">29.4–37.5</td>
</tr>
<tr box="[142,812,798,817]" gridrow="21" pageId="6" pageNumber="91">
<th box="[142,429,798,817]" gridcol="0" gridrow="21" pageId="6" pageNumber="91">Internarial distance/</th>
<td box="[459,582,798,817]" gridcol="1" gridrow="21" pageId="6" pageNumber="91">1.0</td>
<td box="[611,812,798,817]" gridcol="2" gridrow="21" pageId="6" pageNumber="91">6.0–7.0</td>
</tr>
<tr box="[142,812,822,841]" gridrow="22" pageId="6" pageNumber="91" rowspan-1="1" rowspan-2="1">
<th box="[142,429,822,841]" gridcol="0" gridrow="22" pageId="6" pageNumber="91">interorbital distance</th>
</tr>
<tr box="[142,812,845,865]" gridrow="23" pageId="6" pageNumber="91">
<th box="[142,429,845,865]" gridcol="0" gridrow="23" pageId="6" pageNumber="91">Tympanum diameter/</th>
<td box="[459,582,845,865]" gridcol="1" gridrow="23" pageId="6" pageNumber="91">33.3</td>
<td box="[611,812,845,865]" gridcol="2" gridrow="23" pageId="6" pageNumber="91">20.0–23.3</td>
</tr>
<tr box="[142,812,869,889]" gridrow="24" pageId="6" pageNumber="91" rowspan-1="1" rowspan-2="1">
<th box="[142,429,869,889]" gridcol="0" gridrow="24" pageId="6" pageNumber="91">eye diameter</th>
</tr>
<tr box="[142,812,893,912]" gridrow="25" pageId="6" pageNumber="91">
<th box="[142,429,893,912]" gridcol="0" gridrow="25" pageId="6" pageNumber="91">Tibia length/snout–vent length</th>
<td box="[459,582,893,912]" gridcol="1" gridrow="25" pageId="6" pageNumber="91">50.0</td>
<td box="[611,812,893,912]" gridcol="2" gridrow="25" pageId="6" pageNumber="91">46.0–50.0</td>
</tr>
<tr box="[142,812,917,937]" gridrow="26" pageId="6" pageNumber="91">
<th box="[142,429,917,937]" gridcol="0" gridrow="26" pageId="6" pageNumber="91">Foot length/snout–vent length</th>
<td box="[459,582,917,937]" gridcol="1" gridrow="26" pageId="6" pageNumber="91">50.0</td>
<td box="[611,812,917,937]" gridcol="2" gridrow="26" pageId="6" pageNumber="91">44.4–50.0</td>
</tr>
</table>
</paragraph>
<paragraph blockId="6.[142,813,996,2001]" lastBlockId="6.[844,1514,188,2001]" pageId="6" pageNumber="91">
<emphasis bold="true" box="[169,336,1858,1882]" pageId="6" pageNumber="91">Comparisons.</emphasis>
— 
<taxonomicName authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" box="[362,610,1858,1882]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="sneiderni">
<emphasis box="[362,610,1858,1882]" italics="true" pageId="6" pageNumber="91">Pristimantis sneiderni</emphasis>
</taxonomicName>
differs from the other species in the genus by having subarticular tubercles beneath the joint between distal phalanges, dorsal skin with low warts, papilla at the tip of snout in adult males, and by lacking a tympanic membrane, dentigerous processes of vomers, and vocal slits. It is most similar to 
<taxonomicName baseAuthorityName="Lynch" baseAuthorityYear="1981" box="[1322,1466,188,212]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="boulengeri">
<emphasis box="[1322,1466,188,212]" italics="true" pageId="6" pageNumber="91">P. boulengeri</emphasis>
</taxonomicName>
and two Peruvian species, 
<taxonomicName box="[1093,1169,217,241]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="deni">
<emphasis box="[1093,1169,217,241]" italics="true" pageId="6" pageNumber="91">P. deŋi</emphasis>
</taxonomicName>
and 
<taxonomicName baseAuthorityName="Duellman" baseAuthorityYear="1990" box="[1228,1342,217,241]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="schultei">
<emphasis box="[1228,1342,217,241]" italics="true" pageId="6" pageNumber="91">P. schultei</emphasis>
</taxonomicName>
(
<figureCitation box="[1359,1437,217,241]" captionStart="FIG" captionStartId="5.[142,156,1840,1859]" captionTargetBox="[118,1490,191,1814]" captionTargetId="figure-0@5.[118,1490,191,1814]" captionTargetPageId="5" captionText="FIG. 5.—Dorsolateral, ventral, and lateral views of Pristimantis sneiderni (A, B; holotype, KU 145021, adult female, snout–vent length [SVL] 28 mm, head length [HL] 9.0 mm), P. boulengeri (C, D; paratype, KU 169062, adult female, SVL 34 mm, HL 10.6 mm), and P. schultei (E, F; paratype, KU 209504, adult female, SVL 33 mm, HL 10.5 mm). A color version of this figure is available online." figureDoi="http://doi.org/10.5281/zenodo.7717276" httpUri="https://zenodo.org/record/7717276/files/figure.png" pageId="6" pageNumber="91">Figs. 5</figureCitation>
, 
<figureCitation box="[1450,1465,217,241]" captionStart="FIG" captionStartId="7.[142,156,1522,1541]" captionTargetBox="[118,1491,191,1496]" captionTargetId="figure-217@7.[118,1492,191,1496]" captionTargetPageId="7" captionText="FIG. 6.—Dorsolateral and ventral views of the male of Pristimantis sneiderni (A, B; paratype, KU 145019, adult male, snout–vent length [SVL] 23 mm) and P. deŋi (C, D; holotype, MUSM 8968, adult male, SVL 22.7 mm; photo Magnolia Press, reproduced with permission from copyright holder). A color version of this figure is available online." figureDoi="http://doi.org/10.5281/zenodo.7717278" httpUri="https://zenodo.org/record/7717278/files/figure.png" pageId="6" pageNumber="91">6</figureCitation>
), by having tubercles beneath the joint between distal phalanges, dorsal skin with low scattered tubercles or warts, canthal stripe darker, papilla at tip of snout, tubercles or warts on upper eyelids, heel with one or more conical tubercles, low tubercles on medial surface of tarsus, dorsal coloration in preservative tan with darker brown markings (except 
<taxonomicName baseAuthorityName="Duellman" baseAuthorityYear="1990" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="schultei">
<emphasis italics="true" pageId="6" pageNumber="91">P. schultei</emphasis>
</taxonomicName>
), and by being relatively large (except 
<taxonomicName box="[1355,1429,423,447]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="deni">
<emphasis box="[1355,1429,423,447]" italics="true" pageId="6" pageNumber="91">P. deŋi</emphasis>
</taxonomicName>
). From 
<taxonomicName baseAuthorityName="Lynch" baseAuthorityYear="1981" box="[844,989,452,476]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="boulengeri">
<emphasis box="[844,989,452,476]" italics="true" pageId="6" pageNumber="91">P. boulengeri</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" box="[1003,1134,452,476]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="sneiderni">
<emphasis box="[1003,1134,452,476]" italics="true" pageId="6" pageNumber="91">P. sneiderni</emphasis>
</taxonomicName>
is distinguished in having a round snout in profile (rather acutely round; 
<figureCitation box="[1265,1377,481,505]" captionStart="FIG" captionStartId="5.[142,156,1840,1859]" captionTargetBox="[118,1490,191,1814]" captionTargetId="figure-0@5.[118,1490,191,1814]" captionTargetPageId="5" captionText="FIG. 5.—Dorsolateral, ventral, and lateral views of Pristimantis sneiderni (A, B; holotype, KU 145021, adult female, snout–vent length [SVL] 28 mm, head length [HL] 9.0 mm), P. boulengeri (C, D; paratype, KU 169062, adult female, SVL 34 mm, HL 10.6 mm), and P. schultei (E, F; paratype, KU 209504, adult female, SVL 33 mm, HL 10.5 mm). A color version of this figure is available online." figureDoi="http://doi.org/10.5281/zenodo.7717276" httpUri="https://zenodo.org/record/7717276/files/figure.png" pageId="6" pageNumber="91">Fig. 5B,D</figureCitation>
), dorsal skin with low warts (rather than finely granular), and in lacking dentigerous processes of vomers and vocal slits. 
<taxonomicName authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="sneiderni">
<emphasis italics="true" pageId="6" pageNumber="91">Pristimantis sneiderni</emphasis>
</taxonomicName>
differs from 
<taxonomicName baseAuthorityName="Duellman" baseAuthorityYear="1990" box="[1094,1207,569,593]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="schultei">
<emphasis box="[1094,1207,569,593]" italics="true" pageId="6" pageNumber="91">P. schultei</emphasis>
</taxonomicName>
by having a round snout in profile (rather than inclined posteroventrally; 
<figureCitation box="[1346,1453,599,623]" captionStart="FIG" captionStartId="5.[142,156,1840,1859]" captionTargetBox="[118,1490,191,1814]" captionTargetId="figure-0@5.[118,1490,191,1814]" captionTargetPageId="5" captionText="FIG. 5.—Dorsolateral, ventral, and lateral views of Pristimantis sneiderni (A, B; holotype, KU 145021, adult female, snout–vent length [SVL] 28 mm, head length [HL] 9.0 mm), P. boulengeri (C, D; paratype, KU 169062, adult female, SVL 34 mm, HL 10.6 mm), and P. schultei (E, F; paratype, KU 209504, adult female, SVL 33 mm, HL 10.5 mm). A color version of this figure is available online." figureDoi="http://doi.org/10.5281/zenodo.7717276" httpUri="https://zenodo.org/record/7717276/files/figure.png" pageId="6" pageNumber="91">Fig. 5B,F</figureCitation>
), and in lacking ulnar tubercles, dentigerous processes of vomers, and vocal slits. 
<taxonomicName authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" box="[1009,1250,657,681]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="sneiderni">
<emphasis box="[1009,1250,657,681]" italics="true" pageId="6" pageNumber="91">Pristimantis sneiderni</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="deni">
<emphasis italics="true" pageId="6" pageNumber="91">P. deŋi</emphasis>
</taxonomicName>
by having a snout round in profile (rather than protruding and inclined posteroventrally) and in lacking weakly defined dorsolateral folds, dentigerous processes of vomers, and subgular vocal sac (
<figureCitation box="[1228,1301,775,799]" captionStart="FIG" captionStartId="7.[142,156,1522,1541]" captionTargetBox="[118,1491,191,1496]" captionTargetId="figure-217@7.[118,1492,191,1496]" captionTargetPageId="7" captionText="FIG. 6.—Dorsolateral and ventral views of the male of Pristimantis sneiderni (A, B; paratype, KU 145019, adult male, snout–vent length [SVL] 23 mm) and P. deŋi (C, D; holotype, MUSM 8968, adult male, SVL 22.7 mm; photo Magnolia Press, reproduced with permission from copyright holder). A color version of this figure is available online." figureDoi="http://doi.org/10.5281/zenodo.7717278" httpUri="https://zenodo.org/record/7717278/files/figure.png" pageId="6" pageNumber="91">Fig. 6</figureCitation>
). From the genus 
<taxonomicName authorityName="Jimenez de la Espada" authorityYear="1870" box="[844,976,805,828]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="genus">
<emphasis box="[844,976,805,828]" italics="true" pageId="6" pageNumber="91">Pristimantis</emphasis>
</taxonomicName>
, other species with similar appearance to 
<taxonomicName authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="sneiderni">
<emphasis italics="true" pageId="6" pageNumber="91">P. sneiderni</emphasis>
</taxonomicName>
are 
<taxonomicName authority="(Shreve 1935)" baseAuthorityName="Shreve" baseAuthorityYear="1935" box="[1009,1348,833,857]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="acuminatus">
<emphasis box="[1009,1171,834,857]" italics="true" pageId="6" pageNumber="91">P. acuminatus</emphasis>
(
<bibRefCitation author="Shreve, B." box="[1192,1340,833,857]" pageId="6" pageNumber="91" pagination="209 - 218" refId="ref9511" refString="Shreve, B. 1935. On a new teiid and Amphibia from Panama, Ecuador, and Paraguay. Occasional Papers of the Boston Society of Natural History 8: 209 - 218." type="journal article" year="1935">Shreve 1935</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName authority="(Lynch 1981)" baseAuthorityName="Lynch" baseAuthorityYear="1981" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="brevifrons">
<emphasis box="[1367,1514,833,857]" italics="true" pageId="6" pageNumber="91">P. breƲifrons</emphasis>
(
<bibRefCitation author="Lynch, J. D." box="[851,988,863,887]" pageId="6" pageNumber="91" pagination="1 - 12" refId="ref8852" refString="Lynch, J. D. 1981. Two new species of Eleutherodactnlus from western Colombia (Amphibia: Anura: Leptodactylidae). Occasional Papers of the Museum of Zoology, University of Michigan 697: 1 - 12" type="journal article" year="1981">Lynch 1981</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName authority="Ortega-Andrade et al. (2015)" authorityName="Ortega-Andrade" authorityYear="2015" box="[1010,1510,862,887]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="enigmaticus">
<emphasis box="[1010,1172,863,886]" italics="true" pageId="6" pageNumber="91">P. enigmaticus</emphasis>
<bibRefCitation author="Ortega-Andrade, H. M. &amp; O. R. Rojas-Soto &amp; J. H. Valencia &amp; A. Espinosa &amp; J. J. Morrone &amp; S. R. Ron &amp; D. C. Cannatella" box="[1182,1510,862,887]" pageId="6" pageNumber="91" pagination="1 - 43" refId="ref9313" refString="Ortega-Andrade, H. M., O. R. Rojas-Soto, J. H. Valencia, A. Espinosa de los Monteros, J. J. Morrone, S. R. Ron, and D. C. Cannatella. 2015. Insights from integrative systematics reveal cryptic diversity in Pristimantis frogs (Anura: Craugastoridae) from the Upper Amazon Basin. PLoS One 10: 1 - 43." type="journal article" year="2015">Ortega-Andrade et al. (2015)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Ortega-Andrade et al. (2015)" authorityName="Ortega-Andrade" authorityYear="2015" box="[844,1383,892,916]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="limoncochensis">
<emphasis box="[844,1041,892,916]" italics="true" pageId="6" pageNumber="91">P. limoncochensis</emphasis>
<bibRefCitation author="Ortega-Andrade, H. M. &amp; O. R. Rojas-Soto &amp; J. H. Valencia &amp; A. Espinosa &amp; J. J. Morrone &amp; S. R. Ron &amp; D. C. Cannatella" box="[1052,1383,892,916]" pageId="6" pageNumber="91" pagination="1 - 43" refId="ref9313" refString="Ortega-Andrade, H. M., O. R. Rojas-Soto, J. H. Valencia, A. Espinosa de los Monteros, J. J. Morrone, S. R. Ron, and D. C. Cannatella. 2015. Insights from integrative systematics reveal cryptic diversity in Pristimantis frogs (Anura: Craugastoridae) from the Upper Amazon Basin. PLoS One 10: 1 - 43." type="journal article" year="2015">Ortega-Andrade et al. (2015)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Ortega-Andrade et al. (2015)" authorityName="Ortega-Andrade" authorityYear="2015" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="omeviridis">
<emphasis italics="true" pageId="6" pageNumber="91">P. omeƲiridis</emphasis>
<bibRefCitation author="Ortega-Andrade, H. M. &amp; O. R. Rojas-Soto &amp; J. H. Valencia &amp; A. Espinosa &amp; J. J. Morrone &amp; S. R. Ron &amp; D. C. Cannatella" box="[892,1218,921,945]" pageId="6" pageNumber="91" pagination="1 - 43" refId="ref9313" refString="Ortega-Andrade, H. M., O. R. Rojas-Soto, J. H. Valencia, A. Espinosa de los Monteros, J. J. Morrone, S. R. Ron, and D. C. Cannatella. 2015. Insights from integrative systematics reveal cryptic diversity in Pristimantis frogs (Anura: Craugastoridae) from the Upper Amazon Basin. PLoS One 10: 1 - 43." type="journal article" year="2015">Ortega-Andrade et al. (2015)</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName authority="(Shreve 1935)" baseAuthorityName="Shreve" baseAuthorityYear="1935" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="pseudoacuminatus">
<emphasis box="[1280,1514,921,945]" italics="true" pageId="6" pageNumber="91">P. pseudoacuminatus</emphasis>
(
<bibRefCitation author="Shreve, B." box="[851,996,950,975]" pageId="6" pageNumber="91" pagination="209 - 218" refId="ref9511" refString="Shreve, B. 1935. On a new teiid and Amphibia from Panama, Ecuador, and Paraguay. Occasional Papers of the Boston Society of Natural History 8: 209 - 218." type="journal article" year="1935">Shreve 1935</bibRefCitation>
)
</taxonomicName>
. However, 
<taxonomicName authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" box="[1135,1267,951,975]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="sneiderni">
<emphasis box="[1135,1267,951,975]" italics="true" pageId="6" pageNumber="91">P. sneiderni</emphasis>
</taxonomicName>
is distinguished from those species by lacking the dentigerous processes of vomers (dentigerous processes of vomers are present in those species). In addition, 
<taxonomicName authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" box="[1091,1224,1039,1063]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="sneiderni">
<emphasis box="[1091,1224,1039,1063]" italics="true" pageId="6" pageNumber="91">P. sneiderni</emphasis>
</taxonomicName>
differs from 
<taxonomicName class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="enigmaticus">
<emphasis italics="true" pageId="6" pageNumber="91">P. enigmaticus</emphasis>
</taxonomicName>
, 
<taxonomicName box="[898,1098,1068,1092]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="limoncochensis">
<emphasis box="[898,1098,1068,1092]" italics="true" pageId="6" pageNumber="91">P. limoncochensis</emphasis>
</taxonomicName>
, and 
<taxonomicName box="[1169,1317,1068,1092]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="omeviridis">
<emphasis box="[1169,1317,1068,1092]" italics="true" pageId="6" pageNumber="91">P. omeƲiridis</emphasis>
</taxonomicName>
by lacking vocal slits (character state present in those species) and by having a dorsal skin with low warts (rather than the dorsal skin shagreened as in 
<taxonomicName box="[1071,1245,1157,1180]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="enigmaticus">
<emphasis box="[1071,1245,1157,1180]" italics="true" pageId="6" pageNumber="91">P. enigmaticus</emphasis>
</taxonomicName>
, or smooth as in 
<taxonomicName class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="limoncochensis">
<emphasis italics="true" pageId="6" pageNumber="91">P. limoncochensis</emphasis>
</taxonomicName>
and 
<taxonomicName box="[1084,1237,1185,1209]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="omeviridis">
<emphasis box="[1084,1237,1185,1209]" italics="true" pageId="6" pageNumber="91">P. omeƲiridis</emphasis>
</taxonomicName>
). Additional characters that distinguish 
<taxonomicName authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" box="[1037,1174,1215,1239]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="sneiderni">
<emphasis box="[1037,1174,1215,1239]" italics="true" pageId="6" pageNumber="91">P. sneiderni</emphasis>
</taxonomicName>
from 
<taxonomicName baseAuthorityName="Shreve" baseAuthorityYear="1935" box="[1258,1422,1215,1238]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="acuminatus">
<emphasis box="[1258,1422,1215,1238]" italics="true" pageId="6" pageNumber="91">P. acuminatus</emphasis>
</taxonomicName>
and 
<taxonomicName baseAuthorityName="Lynch" baseAuthorityYear="1981" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="brevifrons">
<emphasis italics="true" pageId="6" pageNumber="91">P. breƲifrons</emphasis>
</taxonomicName>
are the round snout in dorsal view (rather acuminate or subacuminate in those species) and the absence of vocal slits (present in 
<taxonomicName baseAuthorityName="Lynch" baseAuthorityYear="1981" box="[1229,1373,1303,1327]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="brevifrons">
<emphasis box="[1229,1373,1303,1327]" italics="true" pageId="6" pageNumber="91">P. breƲifrons</emphasis>
</taxonomicName>
). Finally, 
<taxonomicName baseAuthorityName="Shreve" baseAuthorityYear="1935" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="pseudoacuminatus">
<emphasis italics="true" pageId="6" pageNumber="91">P. pseudoacuminatus</emphasis>
</taxonomicName>
is distinctively smaller (
<quantity box="[1322,1483,1332,1356]" metricMagnitude="-2" metricUnit="m" metricValue="1.515" metricValueMax="1.76" metricValueMin="1.27" pageId="6" pageNumber="91" unit="mm" value="15.15" valueMax="17.6" valueMin="12.7">12.7–17.6 mm</quantity>
in males, 
<quantity box="[933,1102,1361,1385]" metricMagnitude="-2" metricUnit="m" metricValue="2.025" metricValueMax="2.2399999999999998" metricValueMin="1.81" pageId="6" pageNumber="91" unit="mm" value="20.25" valueMax="22.4" valueMin="18.1">18.1–22.4 mm</quantity>
in females; 
<bibRefCitation author="Lynch, J. D." box="[1271,1427,1361,1385]" pageId="6" pageNumber="91" pagination="1 - 24" refId="ref8779" refString="Lynch, J. D. 1980 a. A taxonomic and distributional synopsis of the Amazonian frogs of the genus Eleutherodactnlus. American Museum Novitates 2696: 1 - 24." type="journal article" year="1980">Lynch 1980a</bibRefCitation>
) when compared with 
<taxonomicName authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" box="[1025,1159,1391,1415]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="sneiderni">
<emphasis box="[1025,1159,1391,1415]" italics="true" pageId="6" pageNumber="91">P. sneiderni</emphasis>
</taxonomicName>
(21.0–23.0 mm in males, 28.0 mm in the only known female).
</paragraph>
<paragraph blockId="6.[844,1514,188,2001]" pageId="6" pageNumber="91">
Among species distributed on the southern portions of the Cordillera Central of 
<collectingCountry box="[1083,1193,1479,1503]" name="Colombia" pageId="6" pageNumber="91">Colombia</collectingCountry>
, 
<taxonomicName authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" box="[1204,1445,1479,1503]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="sneiderni">
<emphasis box="[1204,1445,1479,1503]" italics="true" pageId="6" pageNumber="91">Pristimantis sneiderni</emphasis>
</taxonomicName>
could be confused with 
<taxonomicName authority="(Lynch and Duellman 1980)" baseAuthorityName="Lynch and Duellman" baseAuthorityYear="1980" box="[1057,1510,1508,1532]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="petersi">
<emphasis box="[1057,1165,1509,1532]" italics="true" pageId="6" pageNumber="91">P. petersi</emphasis>
(
<bibRefCitation author="Lynch, J. D. &amp; W. E. Duellman" box="[1184,1502,1508,1532]" pageId="6" pageNumber="91" pagination="1 - 86" refId="ref9062" refString="Lynch, J. D., and W. E. Duellman. 1980. The Eleutherodactnlus of the Amazonian slopes of the Ecuadorian Andes (Anura: Leptodactylidae). Miscellaneous Publication. Museum of Natural History, University of Kansas 69: 1 - 86." type="journal article" year="1980">Lynch and Duellman 1980</bibRefCitation>
)
</taxonomicName>
. However, 
<taxonomicName authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" box="[960,1091,1537,1561]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="sneiderni">
<emphasis box="[960,1091,1537,1561]" italics="true" pageId="6" pageNumber="91">P. sneiderni</emphasis>
</taxonomicName>
is larger (SVL 21.0–23.0 mm in adult males and 28.0 in 
<specimenCount box="[1063,1171,1567,1591]" pageId="6" pageNumber="91" type="adult">one adult</specimenCount>
female) than 
<taxonomicName baseAuthorityName="Lynch and Duellman" baseAuthorityYear="1980" box="[1338,1445,1567,1590]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="petersi">
<emphasis box="[1338,1445,1567,1590]" italics="true" pageId="6" pageNumber="91">P. petersi</emphasis>
</taxonomicName>
(SVL 
<quantity box="[844,979,1596,1620]" metricMagnitude="-1" metricUnit="m" metricValue="4.368799999999999" metricValueMax="5.0546" metricValueMin="3.683" pageId="6" pageNumber="91" unit="in" value="17.2" valueMax="19.9" valueMin="14.5">14.5–19.9 in</quantity>
adult males and 
<quantity box="[1167,1324,1596,1620]" metricMagnitude="-2" metricUnit="m" metricValue="2.17" metricValueMax="2.31" metricValueMin="2.03" pageId="6" pageNumber="91" unit="mm" value="21.7" valueMax="23.1" valueMin="20.3">20.3–23.1 mm</quantity>
in adult females; 
<bibRefCitation author="Lynch, J. D. &amp; W. E. Duellman" box="[844,1183,1625,1649]" pageId="6" pageNumber="91" pagination="1 - 86" refId="ref9062" refString="Lynch, J. D., and W. E. Duellman. 1980. The Eleutherodactnlus of the Amazonian slopes of the Ecuadorian Andes (Anura: Leptodactylidae). Miscellaneous Publication. Museum of Natural History, University of Kansas 69: 1 - 86." type="journal article" year="1980">Lynch and Duellman 1980</bibRefCitation>
), and it lacks dentigerous processes of vomers and vocal slits (character states present in 
<taxonomicName baseAuthorityName="Lynch and Duellman" baseAuthorityYear="1980" box="[874,979,1685,1708]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="petersi">
<emphasis box="[874,979,1685,1708]" italics="true" pageId="6" pageNumber="91">P. petersi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="93" pageId="6" pageNumber="91" type="description">
<paragraph blockId="6.[844,1514,188,2001]" lastBlockId="7.[118,789,1634,2001]" lastPageId="7" lastPageNumber="92" pageId="6" pageNumber="91">
<emphasis bold="true" box="[871,1234,1713,1737]" pageId="6" pageNumber="91">
Description of the 
<typeStatus box="[1121,1229,1713,1737]" pageId="6" pageNumber="91">holotype</typeStatus>
.
</emphasis>
—An adult female with head slightly wider than body; head width 39.2% of SVL; HL 32.1% of SVL; snout rounded in dorsal view and in profile; eye–nostril distance 66.6% of diameter of eye; nostrils not protuberant, directed slightly anterolaterally at level of lower jaw. Canthus rostralis curved, not elevated; canthal stripe dark brown; loreal region concave lacking tubercles; lips rounded; internarial region not depressed; top of head flat; upper eyelid warty, its width 50.0% of IOD; supratympanic fold slightly curved downward only just posterior to tympanic annulus, tympanic membrane not evident, tympanic annulus present, upper fifth covered by the supratympanic fold; two nonenlarged postrictal tubercles. Choanae small, nearly round, not obscured by palatal shelf; dentigerous processes of vomers absent; tongue as long as wide, its posterior border not notched, posterior third not adherent to floor of mouth.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7717278" ID-Zenodo-Dep="7717278" httpUri="https://zenodo.org/record/7717278/files/figure.png" pageId="7" pageNumber="92" startId="7.[142,156,1522,1541]" targetBox="[118,1491,191,1496]" targetPageId="7">
<paragraph blockId="7.[118,1489,1522,1589]" pageId="7" pageNumber="92">
FIG. 6.—Dorsolateral and ventral views of the male of 
<taxonomicName authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" box="[615,806,1522,1541]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="92" phylum="Chordata" rank="species" species="sneiderni">
<emphasis box="[615,806,1522,1541]" italics="true" pageId="7" pageNumber="92">Pristimantis sneiderni</emphasis>
</taxonomicName>
(A, B; paratype, KU 145019, adult male, snout–vent length [SVL] 23 mm) and 
<taxonomicName box="[118,180,1546,1565]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="92" phylum="Chordata" rank="species" species="deni">
<emphasis box="[118,180,1546,1565]" italics="true" pageId="7" pageNumber="92">P. deŋi</emphasis>
</taxonomicName>
(C, D; holotype, MUSM 8968, adult male, SVL 22.7 mm; photo Magnolia Press, reproduced with permission from copyright holder). A color version of this figure is available online.
</paragraph>
</caption>
<paragraph blockId="7.[118,789,1634,2001]" lastBlockId="8.[142,813,188,1665]" lastPageId="8" lastPageNumber="93" pageId="7" pageNumber="92">Skin on dorsum bearing low warts (most obvious on flanks); skin on belly and throat areolate; skin on the ventral surface of the thighs, just below the cloaca bearing low warts; skin on posteroventral surfaces of thighs smooth; discoidal fold present, well anterior to groin; dorsolateral folds absent; cloacal sheath short. Antebrachial tubercle present; other ulnar tubercles absent; palmar tubercle deeply bifid; thenar tubercle oval, as wide as palmar tubercle; subarticular tubercles round, prominent, larger than supernumerary tubercles, two on thumb and second finger and three on fingers III and IV; supernumerary tubercles low, on all the fleshy parts of the palm; fingers bearing lateral fringes; relative lengths of fingers I &lt;II &lt;IV &lt;III, all fingers having terminal ventral pads well defined by circumferential grooves; disc on thumb narrow, that on finger II slightly smaller than tympanic annulus; discs on fingers III and IV slightly larger than tympanic annulus. Hind limbs moderately robust; when hind limbs flexed perpendicular to axis of body, heels overlap; tibia length 50.0% of SVL; foot length 50.0% of SVL; heel bearing one low tubercle; outer edge and medial surface of tarsus bearing low tubercles; tarsal fold absent; inner metatarsal tubercle elongate, its length twice its width; conical outer metatarsal tubercle one-fourth size of inner; toes slender, bearing lateral fringes and discs (about as wide as long) on expanded pads; webbing absent; relative lengths of toes I &lt;II &lt;III &lt;V &lt;IV; toe III much shorter than toe V; toe III extending to distal edge of the antepenultimate subarticular tubercle on toe IV; toe V extending to edge distal of the penultimate subarticular tubercle on toe IV; all toes having terminal ventral pads well defined by circumferential grooves; discs of toes III–V equal to discs on fingers II–IV and larger than discs of toe I; subarticular tubercles round, prominent, two on toes I and II, three on toes III and V, and four on toe IV; supernumerary plantar tubercles low, rounded, numerous on proximal segments of digits.</paragraph>
<paragraph blockId="8.[142,813,188,1665]" pageId="8" pageNumber="93">
<emphasis bold="true" box="[169,601,721,746]" pageId="8" pageNumber="93">
Measurements of 
<typeStatus box="[407,513,722,746]" pageId="8" pageNumber="93">holotype</typeStatus>
(mm).
</emphasis>
—SVL 28.0, tibia length 14.0, foot length 14.0, HL 9.0, head width 11.0, IOD 4.0, internarial distance 2.0, width of upper eyelid 2.0, diameter of eye 3.0, eye–nostril distance 2.0.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7717280" ID-Zenodo-Dep="7717280" httpUri="https://zenodo.org/record/7717280/files/figure.png" pageId="8" pageNumber="93" startId="8.[868,882,779,798]" targetBox="[844,1515,191,753]" targetPageId="8">
<paragraph blockId="8.[844,1515,779,869]" pageId="8" pageNumber="93">
FIG. 7.—Palmar view of hand of 
<taxonomicName baseAuthorityName="Lynch" baseAuthorityYear="1998" box="[1171,1417,779,798]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="angustilineatus">
<emphasis box="[1171,1417,779,798]" italics="true" pageId="8" pageNumber="93">Pristimantis angustilineatus</emphasis>
</taxonomicName>
(paratype, ICN 19164, adult male, snout–vent length 18.4 mm) showing the basal, distal, and hyperdistal subarticular tubercles. Scale bar ¼ 2 mm. A color version of this figure is available online.
</paragraph>
</caption>
<subSubSection pageId="8" pageNumber="93" type="distribution">
<paragraph blockId="8.[142,813,188,1665]" pageId="8" pageNumber="93">
<emphasis bold="true" box="[169,326,840,864]" pageId="8" pageNumber="93">Distribution.</emphasis>
— 
<taxonomicName authorityName="Ospina-Sarria &amp; Duellman" authorityYear="2019" box="[353,594,840,864]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="sneiderni">
<emphasis box="[353,594,840,864]" italics="true" pageId="8" pageNumber="93">Pristimantis sneiderni</emphasis>
</taxonomicName>
is known only from the 
<typeStatus box="[192,239,871,895]" pageId="8" pageNumber="93">type</typeStatus>
locality, the vicinity the Laguna de San Rafael, Páramo de Puracé, 
<collectingRegion box="[356,428,900,924]" country="Colombia" name="Cauca" pageId="8" pageNumber="93">Cauca</collectingRegion>
, 
<collectingCountry box="[439,546,900,924]" name="Colombia" pageId="8" pageNumber="93">Colombia</collectingCountry>
(
<figureCitation box="[560,627,900,924]" captionStart="FIG" captionStartId="4.[166,180,1088,1107]" captionTargetBox="[142,813,191,1062]" captionTargetId="figure-774@4.[142,813,191,1062]" captionTargetPageId="4" captionText="FIG. 4.—Map of southwestern Colombia (inset) showing the type localities of Pristimantis calima (star) and P. sneiderni (triangle). Lines indicate boundaries of the two departments within which the new species are known to occur." figureDoi="http://doi.org/10.5281/zenodo.7717274" httpUri="https://zenodo.org/record/7717274/files/figure.png" pageId="8" pageNumber="93">Fig. 4</figureCitation>
). The Laguna de San Rafael is located within the Puracé National Park, Cordillera Central, 
<collectingCountry box="[362,473,959,983]" name="Colombia" pageId="8" pageNumber="93">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="93" type="etymology">
<paragraph blockId="8.[142,813,188,1665]" pageId="8" pageNumber="93">
<emphasis bold="true" box="[169,307,989,1013]" pageId="8" pageNumber="93">Etymology.</emphasis>
—The specific name is a noun in the genitive case and is a patronym for the naturalist, taxidermist, and collector Kjell von Sneidern (1910–1999), who first found the species during his explorations of Puracé, Cauca. Originally from 
<collectingCountry box="[327,417,1107,1131]" name="Sweden" pageId="8" pageNumber="93">Sweden</collectingCountry>
, von Sneidern was director of the Museo de Historia Natural de la Universidad del Cauca from 1940–1960, and founder in 1954 of 
<emphasis box="[536,805,1166,1190]" italics="true" pageId="8" pageNumber="93">NoƲedades Colombianas</emphasis>
, the journal published by the Museo de Historia Natural de la Universidad del Cauca.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="93" type="discussion">
<paragraph blockId="8.[142,813,188,1665]" pageId="8" pageNumber="93">
<emphasis bold="true" box="[169,285,1256,1280]" pageId="8" pageNumber="93">Remarks.</emphasis>
—Kjell von Sneidern visited the Laguna de San Rafael (southeast of Popayán) at the end of 1938 and the beginning of 1939. Together with the new species, von Sneidern collected 
<taxonomicName authority="Rivero 1963" authorityName="Rivero" authorityYear="1963" box="[357,711,1344,1368]" class="Amphibia" family="Bufonidae" genus="Atelopus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="ebenoides">
<emphasis box="[357,567,1344,1368]" italics="true" pageId="8" pageNumber="93">Atelopus ebenoides</emphasis>
<bibRefCitation author="Rivero, J. A." box="[574,711,1344,1368]" pageId="8" pageNumber="93" pagination="103 - 124" refId="ref9434" refString="Rivero, J. A. 1963. Five new species of Atelopus from Colombia, with notes on other forms from Colombia and Ecuador. Caribbean Journal of Science 3: 103 - 124." type="journal article" year="1963">Rivero 1963</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="(Peracca 1904)" baseAuthorityName="Peracca" baseAuthorityYear="1904" family="Strabomantidae" genus="Osornophrnne" pageId="8" pageNumber="93" rank="species" species="bufoniformis">
<emphasis italics="true" pageId="8" pageNumber="93">Osornophrŋne bufoniformis</emphasis>
(
<bibRefCitation author="Peracca, M. G." box="[408,571,1374,1398]" pageId="8" pageNumber="93" pagination="1 - 41" refId="ref9389" refString="Peracca, M. G. 1904. Viaggio del Dr. Enrico Festa nell ̕ Ecuador e regioni vicine. Rettili ed anfibi. Bollettino dei Musei di Zoologia e Anatomia Comparata della R. Universita di Torino 19: 1 - 41." type="journal article" year="1904">Peracca 1904</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName authority="(Rivero and Granado-Diaz 1990)" baseAuthorityName="Rivero and Granado-Diaz" baseAuthorityYear="1990" family="Strabomantidae" genus="Hnloxalus" pageId="8" pageNumber="93" rank="species" species="pinguis">
<emphasis box="[602,812,1374,1398]" italics="true" pageId="8" pageNumber="93">Hŋloxalus pinguis</emphasis>
(Rivero and Granado-Díaz 1990)
</taxonomicName>
, 
<taxonomicName authority="(Boulenger 1882)" baseAuthorityName="Boulenger" baseAuthorityYear="1882" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="buckleni">
<emphasis box="[567,812,1404,1428]" italics="true" pageId="8" pageNumber="93">Pristimantis buckleŋi</emphasis>
(
<bibRefCitation author="Boulenger, G. A." box="[150,343,1433,1457]" pageId="8" pageNumber="93" refId="ref8058" refString="Boulenger, G. A. 1882. Catalogue of the Batrachia Salientia s. Ecaudata in the Collection of the British Museum, 2 nd ed. Taylor and Francis, USA." type="book" year="1882">Boulenger 1882</bibRefCitation>
)
</taxonomicName>
, and 
<taxonomicName authority="(Lynch 1980 b)" baseAuthorityName="Lynch" baseAuthorityYear="1980" box="[433,809,1433,1457]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="obmutescens">
<emphasis box="[433,612,1433,1457]" italics="true" pageId="8" pageNumber="93">P. obmutescens</emphasis>
(
<bibRefCitation author="Lynch, J. D." box="[638,801,1433,1457]" pageId="8" pageNumber="93" pagination="165 - 188" refId="ref8811" refString="Lynch, J. D. 1980 b. New species of Eleutherodactnlus of Colombia (Amphibia: Leptodactylidae). I: Five new species from the paramos of the Cordillera Central. Caldasia 13: 165 - 188." type="journal article" year="1980">Lynch 1980b</bibRefCitation>
)
</taxonomicName>
. WED (with L. Trueb and J.E. Simmons) in 
<date pageId="8" pageNumber="93" value="1974-09">September 1974</date>
visited several sites in Puracé, vicinity of the 
<typeStatus box="[765,812,1493,1517]" pageId="8" pageNumber="93">type</typeStatus>
locality. They found two species of bufonids (
<taxonomicName authorityName="Rivero" authorityYear="1963" box="[670,812,1522,1546]" class="Amphibia" family="Bufonidae" genus="Atelopus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="ebenoides">
<emphasis box="[670,812,1522,1546]" italics="true" pageId="8" pageNumber="93">A. ebenoides</emphasis>
</taxonomicName>
and 
<emphasis box="[198,380,1552,1576]" italics="true" pageId="8" pageNumber="93">
O. 
<taxonomicName baseAuthorityName="Peracca" baseAuthorityYear="1904" box="[240,380,1552,1576]" family="Strabomantidae" genus="Osornophrnne" pageId="8" pageNumber="93" rank="species" species="bufoniformis">bufoniformis</taxonomicName>
</emphasis>
) and six species of 
<emphasis italics="true" pageId="8" pageNumber="93">
<taxonomicName authorityName="Jimenez de la Espada" authorityYear="1870" box="[632,763,1553,1576]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="93" phylum="Chordata" rank="genus">Pristimantis</taxonomicName>
— 
<taxonomicName baseAuthorityName="Lynch" baseAuthorityYear="1981" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="boulengeri">P. boulengeri</taxonomicName>
</emphasis>
, 
<taxonomicName baseAuthorityName="Boulenger" baseAuthorityYear="1882" box="[279,405,1581,1605]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="buckleni">
<emphasis box="[279,405,1581,1605]" italics="true" pageId="8" pageNumber="93">P. buckleŋi</emphasis>
</taxonomicName>
, 
<taxonomicName authority="(Lynch 1980 b)" baseAuthorityName="Lynch" baseAuthorityYear="1980" box="[426,773,1581,1605]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="leptolophus">
<emphasis box="[426,588,1581,1605]" italics="true" pageId="8" pageNumber="93">P. leptolophus</emphasis>
(
<bibRefCitation author="Lynch, J. D." box="[609,765,1581,1605]" pageId="8" pageNumber="93" pagination="165 - 188" refId="ref8811" refString="Lynch, J. D. 1980 b. New species of Eleutherodactnlus of Colombia (Amphibia: Leptodactylidae). I: Five new species from the paramos of the Cordillera Central. Caldasia 13: 165 - 188." type="journal article" year="1980">Lynch 1980b</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName authority="(Goin and Cochran 1963)" baseAuthorityName="Goin and Cochran" baseAuthorityYear="1963" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="mnersi">
<emphasis italics="true" pageId="8" pageNumber="93">P. mŋersi</emphasis>
(
<bibRefCitation author="Goin, C. J. &amp; D. M. Cochran" box="[236,522,1611,1635]" pageId="8" pageNumber="93" pagination="499 - 505" refId="ref8501" refString="Goin, C. J., and D. M. Cochran. 1963. Two new genera of leptodactylid frogs from Colombia. Proceedings of the California Academy of Sciences 31: 499 - 505." type="journal article" year="1963">Goin and Cochran 1963</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName baseAuthorityName="Lynch" baseAuthorityYear="1980" box="[546,718,1611,1635]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="obmutescens">
<emphasis box="[546,718,1611,1635]" italics="true" pageId="8" pageNumber="93">P. obmutescens</emphasis>
</taxonomicName>
, and 
<taxonomicName authority="(Lynch and Ruiz-Carranza 1983)" baseAuthorityName="Lynch and Ruiz-Carranza" baseAuthorityYear="1983" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="vicarius">
<emphasis italics="true" pageId="8" pageNumber="93">P. Ʋicarius</emphasis>
(
<bibRefCitation author="Lynch, J. D. &amp; P. M. Ruiz-Carranza" box="[246,602,1641,1665]" pageId="8" pageNumber="93" pagination="99 - 112" refId="ref9157" refString="Lynch, J. D., and P. M. Ruiz-Carranza. 1983. New frogs of the genus Eleutherodactnlus from the Andes of southern Colombia. Transactions of the Kansas Academy of Science 86: 99 - 112." type="journal article" year="1983">Lynch and Ruiz-Carranza 1983</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>