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<document id="CE7516880C10A1D57B890CC4F4E69E07" ID-DOI="10.1111/zoj.12362" ID-GBIF-Dataset="701ee46e-4e18-4f6e-a04f-28e1598f1dfd" ID-Zenodo-Dep="269748" IM.bibliography_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461051174125" checkinUser="plazi" docAuthor="Pablo M. Dellapé, María C. Melo &amp; Thomas J. Henry" docDate="2016" docId="6F64DB1B5E63FF8BFC0CFA25FB56FE24" docLanguage="en" docName="Dellap-_et_al-2016-Zoological_Journal_of_the_Linnean_Society.pdf" docOrigin="Zoological Journal of the Linnean Society 177 (177)" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Heraeus antennalis" docType="treatment" docVersion="7" lastPageNumber="65" masterDocId="935DA3635E40FFAFFFBBFFD2FFDAFFB7" masterDocTitle="A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species" masterLastPageNumber="134" masterPageNumber="29" pageNumber="64" updateTime="1698578884425" updateUser="plazi">
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<mods:title id="F742DF295DB709BDB470289935589665">A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species</mods:title>
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<mods:namePart id="7DA53EF47CBB5B56340CD47E3482531B">Pablo M. Dellapé</mods:namePart>
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<mods:namePart id="EC3954EFA70A88A73B7FB3079649DB2D">María C. Melo</mods:namePart>
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<mods:namePart id="543E906D39BD10C2B40F4B146A7B15DE">Thomas J. Henry</mods:namePart>
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<mods:title id="54DDA81438F64997F1886E281DC52AE6">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="D6337B9446D21D834D92714F28031338">2016</mods:date>
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<mods:number id="19CEC438679D9624B18366E31308A9C5">177</mods:number>
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<treatment id="6F64DB1B5E63FF8BFC0CFA25FB56FE24" ID-DOI="http://doi.org/10.5281/zenodo.5611135" ID-GBIF-Taxon="121066314" ID-Zenodo-Dep="5611135" LSID="urn:lsid:plazi:treatment:6F64DB1B5E63FF8BFC0CFA25FB56FE24" httpUri="http://treatment.plazi.org/id/6F64DB1B5E63FF8BFC0CFA25FB56FE24" lastPageId="36" lastPageNumber="65" pageId="35" pageNumber="64">
<subSubSection id="AFD739865E63FF8CFC0CFA25FAFEF9BB" box="[951,1316,1527,1548]" pageId="35" pageNumber="64" type="nomenclature">
<paragraph id="E7726A0D5E63FF8CFC0CFA25FAFEF9BB" blockId="35.[826,1442,1527,1902]" box="[951,1316,1527,1548]" pageId="35" pageNumber="64">
<heading id="BC3ADD615E63FF8CFC0CFA25FAFEF9BB" allCaps="true" bold="true" box="[951,1316,1527,1548]" centered="true" fontSize="8" level="3" pageId="35" pageNumber="64" reason="4">
<taxonomicName id="20CD118E5E63FF8CFC0CFA25FB18F9BB" ID-CoL="3KXLP" box="[951,1218,1527,1548]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="65" phylum="Arthropoda" rank="species" species="antennalis" status="sp. nov.">
<emphasis id="D5B9B61F5E63FF8CFC0CFA25FB18F9BB" bold="true" box="[951,1218,1527,1548]" italics="true" pageId="35" pageNumber="64">HERAEUS ANTENNALIS</emphasis>
</taxonomicName>
SP
<emphasis id="D5B9B61F5E63FF8CFB59FA25FAFEF9BB" bold="true" box="[1250,1316,1527,1548]" pageId="35" pageNumber="64">. NOV.</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AFD739865E63FF8CFC70F9CCFACBF983" box="[971,1297,1566,1588]" pageId="35" pageNumber="64" type="description">
<paragraph id="E7726A0D5E63FF8CFC70F9CCFACBF983" blockId="35.[826,1442,1527,1902]" box="[971,1297,1566,1588]" pageId="35" pageNumber="64">
<heading id="BC3ADD615E63FF8CFC70F9CCFACBF983" allCaps="true" box="[971,1297,1566,1588]" centered="true" fontSize="9" level="6" pageId="35" pageNumber="64" reason="4">
(
<figureCitation id="7FF676885E63FF8CFC69F9CCFBF6F983" box="[978,1068,1566,1588]" captionStart="Figure 15" captionStartId="32.[144,229,1610,1629]" captionTargetBox="[144,1422,192,1580]" captionTargetId="figure@32.[144,1424,192,1580]" captionTargetPageId="32" captionText="Figure 15. Habitus dorsal: A, Heraeus annulatus sp. nov.; B, Heraeus antennalis sp. nov.; C, Heraeus chamamecinus sp. nov.; D, Heraeus illitus Distant, 1882; E, Heraeus inca sp. nov.; F, Heraeus panamaensis sp. nov.; G, Heraeus similis sp. nov.; H, Heraeus spinosus sp. nov. I, Heraeus splendens sp. nov.; J, Heraeus tiputini sp. nov." httpUri="https://zenodo.org/record/269763/files/figure.png" pageId="35" pageNumber="64">FIGS 15</figureCitation>
B, 16B, 17E–H, 18)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AFD739865E63FF8BFC81F996FE6AFE24" lastPageId="36" lastPageNumber="65" pageId="35" pageNumber="64" type="diagnosis">
<paragraph id="E7726A0D5E63FF8CFC81F996FC70F9EE" blockId="35.[826,1442,1527,1902]" box="[826,938,1604,1625]" pageId="35" pageNumber="64">
<heading id="BC3ADD615E63FF8CFC81F996FC70F9EE" bold="true" box="[826,938,1604,1625]" fontSize="9" level="5" pageId="35" pageNumber="64" reason="6">
<emphasis id="D5B9B61F5E63FF8CFC81F996FC70F9EE" bold="true" box="[826,938,1604,1625]" italics="true" pageId="35" pageNumber="64">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="E7726A0D5E63FF8CFC81F9B1FB72F886" blockId="35.[826,1442,1527,1902]" pageId="35" pageNumber="64">Strongly setose. Labium extending to mesocoxae. Basiflagellomere clavate. Anterior pronotal lobe dark brown, posterior lobe paler with humeral angles concolorous. Profemur pale brown, mottled with brown. Hemelytra uniformly pale brown, with a white subapical corial spot. Aedeagus spinose, with spines forming an arc on sclerotized vesical lobes.</paragraph>
<paragraph id="E7726A0D5E63FF8BFCE8F8E8FE6AFE24" blockId="35.[826,1442,1527,1902]" lastBlockId="36.[144,759,197,403]" lastPageId="36" lastPageNumber="65" pageId="35" pageNumber="64">
<taxonomicName id="20CD118E5E63FF8CFCE8F8E8FBF1F8F8" box="[851,1067,1849,1871]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="64" phylum="Arthropoda" rank="species" species="antennalis" status="sp. nov.">
<emphasis id="D5B9B61F5E63FF8CFCE8F8E8FBF1F8F8" bold="true" box="[851,1067,1849,1871]" italics="true" pageId="35" pageNumber="64">Heraeus antennalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CE8A0B645E63FF8CFB8AF8E8FB59F8F8" box="[1073,1155,1850,1871]" pageId="35" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="20CD118E5E63FF8CFB07F8E8FA93F8F8" box="[1212,1353,1849,1871]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="64" phylum="Arthropoda" rank="species" species="splendens" status="sp. nov.">
<emphasis id="D5B9B61F5E63FF8CFB07F8E8FA93F8F8" bold="true" box="[1212,1353,1849,1871]" italics="true" pageId="35" pageNumber="64">H. splendens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CE8A0B645E63FF8CFAEBF8E8FA78F8F8" box="[1360,1442,1850,1871]" pageId="35" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
are the only two species in the genus with the basiflagellomere clavate.
<taxonomicName id="20CD118E5E64FF8BFE10FF17FD59FF6C" box="[427,643,197,219]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="65" phylum="Arthropoda" rank="species" species="antennalis" status="sp. nov.">
<emphasis id="D5B9B61F5E64FF8BFE10FF17FD59FF6C" bold="true" box="[427,643,197,219]" italics="true" pageId="36" pageNumber="65">Heraeus antennalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CE8A0B645E64FF8BFD31FF17FD07FF6D" box="[650,733,197,218]" pageId="36" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
is strongly setose, with mottled profemora, and the posterior pronotal lobe uniformly pale brown, whereas
<taxonomicName id="20CD118E5E64FF8BFF2BFEF3FEC5FE81" box="[144,287,288,310]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="65" phylum="Arthropoda" rank="species" species="splendens" status="sp. nov.">
<emphasis id="D5B9B61F5E64FF8BFF2BFEF3FEC5FE81" bold="true" box="[144,287,288,310]" italics="true" pageId="36" pageNumber="65">H. splendens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CE8A0B645E64FF8BFE9DFEF3FEA0FE81" box="[294,378,289,310]" pageId="36" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
is less setose, with a pale invert- ed V-shaped mark on the posterior pronotal lobe, the profemora are dark but paler dorsally, and the outer margin of clavus is pale.
</paragraph>
</subSubSection>
<subSubSection id="AFD739865E64FF8BFF2BFE0AFD59F944" pageId="36" pageNumber="65" type="description">
<paragraph id="E7726A0D5E64FF8BFF2BFE0AFE51FE5A" blockId="36.[144,760,472,1779]" box="[144,395,472,493]" pageId="36" pageNumber="65">
<heading id="BC3ADD615E64FF8BFF2BFE0AFE51FE5A" bold="true" box="[144,395,472,493]" fontSize="9" level="5" pageId="36" pageNumber="65" reason="6">
<emphasis id="D5B9B61F5E64FF8BFF2BFE0AFE51FE5A" bold="true" box="[144,395,472,493]" italics="true" pageId="36" pageNumber="65">
Description (
<figureCitation id="7FF676885E64FF8BFE99FE0AFEAAFE5A" box="[290,368,472,493]" captionStart="Figure 15" captionStartId="32.[144,229,1610,1629]" captionTargetBox="[144,1422,192,1580]" captionTargetId="figure@32.[144,1424,192,1580]" captionTargetPageId="32" captionText="Figure 15. Habitus dorsal: A, Heraeus annulatus sp. nov.; B, Heraeus antennalis sp. nov.; C, Heraeus chamamecinus sp. nov.; D, Heraeus illitus Distant, 1882; E, Heraeus inca sp. nov.; F, Heraeus panamaensis sp. nov.; G, Heraeus similis sp. nov.; H, Heraeus spinosus sp. nov. I, Heraeus splendens sp. nov.; J, Heraeus tiputini sp. nov." httpUri="https://zenodo.org/record/269763/files/figure.png" pageId="36" pageNumber="65">Fig. 15</figureCitation>
B)
</emphasis>
</heading>
</paragraph>
<paragraph id="E7726A0D5E64FF8BFF2BFE25FDC4FDBB" blockId="36.[144,760,472,1779]" box="[144,542,503,525]" pageId="36" pageNumber="65">Total length 6.36. Strongly setose.</paragraph>
<paragraph id="E7726A0D5E64FF8BFF13FDC7FD94FC60" blockId="36.[144,760,472,1779]" pageId="36" pageNumber="65">
<emphasis id="D5B9B61F5E64FF8BFF13FDC7FF36FD9C" bold="true" box="[168,236,533,555]" italics="true" pageId="36" pageNumber="65">Head:</emphasis>
Reddish brown, with an elongate neck and short recumbent and long erect setae. Head length 1.46, width 0.99. Postocular length 0.62. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli placed at level of an imaginary line passing the posterior border of eyes. Interocular width 0.53, interocellar width 0.27. Labium brown, with erect setae, extending to mesocoxae. Labial segment lengths: I 0.66, II 0.80, III 0.62, and IV 0.32. Antenna brown, pedicel and base of basiflagellomere paler, with short recumbent and sparse short erect setae, distiflagellomere with a narrow sub-basal whitish band. Basiflagellomere clavate. Antennal lengths: scape 0.43, pedicel 1.23, basiflagellomere 0.96, and distiflagellomere 1.06. Length of pale band on distiflagellomere 0.32.
</paragraph>
<paragraph id="E7726A0D5E64FF8BFF13FC32FF08F98C" blockId="36.[144,760,472,1779]" pageId="36" pageNumber="65">
<emphasis id="D5B9B61F5E64FF8BFF13FC32FEDAFC41" bold="true" box="[168,256,992,1014]" italics="true" pageId="36" pageNumber="65">Thorax:</emphasis>
Anterior pronotal lobe dark brown, posteri- or lobe paler, humeral angles concolorous; collar impunctate, delimited posteriorly by a sulcus; anteri- or and posterior lobes with long erect setae, posterior lobe with short recumbent setae. Pronotum pruinose, posterior lobe punctate. Collar length 0.10, anterior lobe length 0.55, posterior lobe length 0.55; anterior lobe width 0.94, posterior lobe width 1.47. Pleurae dark brown, acetabular areas paler, punctate, with short recumbent setae. Evaporative area extended. Scutellum brown darker at base and along a longitudinal line to the apex, with long erect setae. Hemelytra pale brown, pattern as in
<figureCitation id="7FF676885E64FF8BFDDBFA82FD14FAD1" box="[608,718,1360,1382]" captionStart="Figure 15" captionStartId="32.[144,229,1610,1629]" captionTargetBox="[144,1422,192,1580]" captionTargetId="figure@32.[144,1424,192,1580]" captionTargetPageId="32" captionText="Figure 15. Habitus dorsal: A, Heraeus annulatus sp. nov.; B, Heraeus antennalis sp. nov.; C, Heraeus chamamecinus sp. nov.; D, Heraeus illitus Distant, 1882; E, Heraeus inca sp. nov.; F, Heraeus panamaensis sp. nov.; G, Heraeus similis sp. nov.; H, Heraeus spinosus sp. nov. I, Heraeus splendens sp. nov.; J, Heraeus tiputini sp. nov." httpUri="https://zenodo.org/record/269763/files/figure.png" pageId="36" pageNumber="65">Figure 15</figureCitation>
(B); inner corial spot diffuse, subapical corial spot present. Membrane brown, with veins paler and with a pale apical spot. Legs: Pale brown except mottled profemur, metafemur with a subapical dark band; and meso- and metatibia with a subproximal dark band, apex of tibiae darker (
<figureCitation id="7FF676885E64FF8BFEECF9D5FE72F9AA" box="[343,424,1543,1565]" captionStart="Figure 16" captionStartId="33.[164,240,1062,1081]" captionTargetBox="[325,1282,195,1030]" captionTargetId="figure@33.[323,1283,193,1032]" captionTargetPageId="33" captionText="Figure 16. Legs: A, Heraeus annulatus sp. nov.; B, Heraeus antennalis sp. nov.; C, Heraeus chamamecinus sp. nov.; D, Heraeus illitus Distant, 1882; E, Heraeus inca sp. nov.; F, Heraeus panamaensis sp. nov." httpUri="https://zenodo.org/record/269764/files/figure.png" pageId="36" pageNumber="65">Fig. 16</figureCitation>
B). Profemur with long erect setae.
</paragraph>
<paragraph id="E7726A0D5E64FF8BFF13F996FD59F944" blockId="36.[144,760,472,1779]" pageId="36" pageNumber="65">
<emphasis id="D5B9B61F5E64FF8BFF13F996FEC2F9ED" bold="true" box="[168,280,1604,1626]" italics="true" pageId="36" pageNumber="65">Abdomen:</emphasis>
Dark brown to reddish brown, with abundant short recumbent setae. Male genitalia: Pygophore (
<figureCitation id="7FF676885E64FF8BFF23F950FF3DF920" box="[152,231,1666,1688]" captionStart="Figure 17" captionStartId="34.[144,222,1714,1733]" captionTargetBox="[241,1332,196,1676]" captionTargetId="figure@34.[229,1338,192,1684]" captionTargetPageId="34" captionText="Figure 17. Male genitalia. Heraeus annulatus sp. nov.: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus antennalis sp. nov.: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus chamamecinus sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view. Heraeus illitus Distant, 1882: M, pygophore, dorsal view; N, pygophore, lateral view; O, right paramere, inner view; P, right paramere, external view." httpUri="https://zenodo.org/record/269765/files/figure.png" pageId="36" pageNumber="65">Fig. 17</figureCitation>
E, F) rounded, anterior margin of dorsal aperture rounded. Parameres:
<figureCitation id="7FF676885E64FF8BFE75F972FDE2F902" box="[462,568,1696,1718]" captionStart="Figure 17" captionStartId="34.[144,222,1714,1733]" captionTargetBox="[241,1332,196,1676]" captionTargetId="figure@34.[229,1338,192,1684]" captionTargetPageId="34" captionText="Figure 17. Male genitalia. Heraeus annulatus sp. nov.: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus antennalis sp. nov.: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus chamamecinus sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view. Heraeus illitus Distant, 1882: M, pygophore, dorsal view; N, pygophore, lateral view; O, right paramere, inner view; P, right paramere, external view." httpUri="https://zenodo.org/record/269765/files/figure.png" pageId="36" pageNumber="65">Figure 17</figureCitation>
(G, H). Aedeagus spinose, with spines forming an arc on sclerotized vesical lobes;
<emphasis id="D5B9B61F5E64FF8BFF61F90CFE6BF945" bold="true" box="[218,433,1757,1779]" italics="true" pageId="36" pageNumber="65">processus gonopori</emphasis>
long and slender.
</paragraph>
</subSubSection>
<subSubSection id="AFD739865E64FF8BFF2BF8EBFE84F8D9" box="[144,350,1849,1902]" pageId="36" pageNumber="65" type="distribution">
<paragraph id="E7726A0D5E64FF8BFF2BF8EBFEC1F8F8" blockId="36.[144,350,1849,1902]" box="[144,283,1849,1871]" pageId="36" pageNumber="65">
<heading id="BC3ADD615E64FF8BFF2BF8EBFEC1F8F8" bold="true" box="[144,283,1849,1871]" fontSize="9" level="5" pageId="36" pageNumber="65" reason="6">
<emphasis id="D5B9B61F5E64FF8BFF2BF8EBFEC1F8F8" bold="true" box="[144,283,1849,1871]" italics="true" pageId="36" pageNumber="65">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="E7726A0D5E64FF8BFF2BF88AFE84F8D9" blockId="36.[144,350,1849,1902]" box="[144,350,1880,1902]" pageId="36" pageNumber="65">
Ecuador (
<figureCitation id="7FF676885E64FF8BFEBBF88AFE8AF8D9" box="[256,336,1880,1902]" captionStart="Figure 18" captionStartId="35.[164,247,1160,1179]" captionTargetBox="[167,1441,197,1127]" captionTargetId="figure@35.[164,1444,194,1131]" captionTargetPageId="35" captionText="Figure 18. Distributional map of Heraeus antennalis sp. nov., Heraeus annulatus sp. nov., and Heraeus panamaensis sp. nov." httpUri="https://zenodo.org/record/269766/files/figure.png" pageId="36" pageNumber="65">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="AFD739865E64FF8BFC9DFF16FB47FEAF" pageId="36" pageNumber="65" type="etymology">
<paragraph id="E7726A0D5E64FF8BFC9DFF16FC47FF6D" blockId="36.[806,1420,196,280]" box="[806,925,196,218]" pageId="36" pageNumber="65">
<heading id="BC3ADD615E64FF8BFC9DFF16FC47FF6D" bold="true" box="[806,925,196,218]" fontSize="9" level="5" pageId="36" pageNumber="65" reason="6">
<emphasis id="D5B9B61F5E64FF8BFC9DFF16FC47FF6D" bold="true" box="[806,925,196,218]" italics="true" pageId="36" pageNumber="65">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="E7726A0D5E64FF8BFC9DFF31FB47FEAF" blockId="36.[806,1420,196,280]" pageId="36" pageNumber="65">The specific ephitet is a noun and refers to the clavate basiflagellomere of the antenna.</paragraph>
</subSubSection>
<subSubSection id="AFD739865E64FF8BFC9DFE92FB56FE24" pageId="36" pageNumber="65" type="materials_examined">
<paragraph id="E7726A0D5E64FF8BFC9DFE92FC19FEE2" blockId="36.[806,1421,319,403]" box="[806,963,319,341]" pageId="36" pageNumber="65">
<heading id="BC3ADD615E64FF8BFC9DFE92FC19FEE2" bold="true" box="[806,963,319,341]" fontSize="9" level="5" pageId="36" pageNumber="65" reason="6">
<emphasis id="D5B9B61F5E64FF8BFC9DFE92FC19FEE2" bold="true" box="[806,963,319,341]" italics="true" pageId="36" pageNumber="65">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="E7726A0D5E64FF8BFC9DFE8CFB56FE24" blockId="36.[806,1421,319,403]" pageId="36" pageNumber="65">
<emphasis id="D5B9B61F5E64FF8BFC9DFE8CFC4BFEC3" bold="true" box="[806,913,350,372]" italics="true" pageId="36" pageNumber="65">Holotype:</emphasis>
♂,
<emphasis id="D5B9B61F5E64FF8BFC79FE8CFB89FEC3" bold="true" box="[962,1107,350,372]" pageId="36" pageNumber="65">ECUADOR:</emphasis>
Yanayacu, 400 m a.s.l., IX/ X-1977, L.E. Peña G. (USNM).
</paragraph>
</subSubSection>
</treatment>
</document>
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