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<document id="65712C9913332F17C9E29665A1153DE3" ID-DOI="10.1111/zoj.12362" ID-GBIF-Dataset="701ee46e-4e18-4f6e-a04f-28e1598f1dfd" ID-Zenodo-Dep="269748" IM.bibliography_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461051174125" checkinUser="plazi" docAuthor="Pablo M. Dellapé, María C. Melo &amp; Thomas J. Henry" docDate="2016" docId="6F64DB1B5E00FFECFF51FB07FE30F8D9" docLanguage="en" docName="Dellap-_et_al-2016-Zoological_Journal_of_the_Linnean_Society.pdf" docOrigin="Zoological Journal of the Linnean Society 177 (177)" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Heraeus mesoamericanus" docType="treatment" docVersion="8" lastPageNumber="96" masterDocId="935DA3635E40FFAFFFBBFFD2FFDAFFB7" masterDocTitle="A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species" masterLastPageNumber="134" masterPageNumber="29" pageNumber="93" updateTime="1698578884425" updateUser="plazi">
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<mods:title id="B1F957410156625215B73B88F35B3B30">A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species</mods:title>
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<mods:namePart id="F559321B69A548EBE7ACC7B71565CF34">Pablo M. Dellapé</mods:namePart>
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<mods:namePart id="9BC1A280C1F5E903BC324729E3D3CF43">María C. Melo</mods:namePart>
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<mods:namePart id="7D6020D2E747B14CD65596268FDAD8DA">Thomas J. Henry</mods:namePart>
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<mods:title id="F2F8C81ED240027CB0606FBF2CB55260">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="630FCC399EC30B0ADF34357977618AA4">2016</mods:date>
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<mods:number id="010E7300EF65DBFC6FB136D5117EF0FC">177</mods:number>
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<treatment id="6F64DB1B5E00FFECFF51FB07FE30F8D9" ID-DOI="http://doi.org/10.5281/zenodo.5611173" ID-GBIF-Taxon="121066346" ID-Zenodo-Dep="5611173" LSID="urn:lsid:plazi:treatment:6F64DB1B5E00FFECFF51FB07FE30F8D9" httpUri="http://treatment.plazi.org/id/6F64DB1B5E00FFECFF51FB07FE30F8D9" lastPageId="67" lastPageNumber="96" pageId="64" pageNumber="93">
<subSubSection id="AFD739865E00FFEFFF51FB07FD46FB5D" box="[234,668,1237,1258]" pageId="64" pageNumber="93" type="nomenclature">
<paragraph id="E7726A0D5E00FFEFFF51FB07FD46FB5D" blockId="64.[144,760,1237,1612]" box="[234,668,1237,1258]" pageId="64" pageNumber="93">
<heading id="BC3ADD615E00FFEFFF51FB07FD46FB5D" allCaps="true" bold="true" box="[234,668,1237,1258]" centered="true" fontSize="8" level="3" pageId="64" pageNumber="93" reason="5">
<taxonomicName id="20CD118E5E00FFEFFF51FB07FDE3FB5D" ID-CoL="3KXMH" box="[234,569,1237,1258]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="64" pageNumber="95" phylum="Arthropoda" rank="species" species="mesoamericanus" status="sp. nov.">
<emphasis id="D5B9B61F5E00FFEFFF51FB07FDE3FB5D" bold="true" box="[234,569,1237,1258]" italics="true" pageId="64" pageNumber="93">HERAEUS MESOAMERICANUS</emphasis>
</taxonomicName>
SP
<emphasis id="D5B9B61F5E00FFEFFDE1FB07FD46FB5D" bold="true" box="[602,668,1237,1258]" pageId="64" pageNumber="93">. NOV.</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AFD739865E00FFEFFEB8FB2EFD59FAA5" box="[259,643,1276,1298]" pageId="64" pageNumber="93" type="description">
<paragraph id="E7726A0D5E00FFEFFEB8FB2EFD59FAA5" blockId="64.[144,760,1237,1612]" box="[259,643,1276,1298]" pageId="64" pageNumber="93">
<heading id="BC3ADD615E00FFEFFEB8FB2EFD59FAA5" allCaps="true" box="[259,643,1276,1298]" centered="true" fontSize="9" level="6" pageId="64" pageNumber="93" reason="4">
(
<figureCitation id="7FF676885E00FFEFFEB2FB2EFEBEFAA6" box="[265,356,1276,1297]" captionStart="Figure 30" captionStartId="55.[164,240,1815,1834]" captionTargetBox="[168,1442,193,1783]" captionTargetId="figure@55.[164,1444,193,1786]" captionTargetPageId="55" captionText="Figure 30. Aedeagus. A, Heraeus bahiensis sp. nov.; B, Heraeus bolivianus sp. nov.; C, Heraeus brevirostris sp. nov.; D, Heraeus concolor Slater &amp; Baranowski, 1994; E, Heraeus ecuatorianus sp. nov.; F, Heraeus mesoamericanus sp. nov.; G, Heraeus mexicanus sp. nov.; H, Heraeus penai sp. nov." httpUri="https://zenodo.org/record/269778/files/figure.png" pageId="64" pageNumber="93">FIGS 30</figureCitation>
F, 33D, 34E–H, 36A, 37)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AFD739865E00FFEFFF2BFAF0FEA5F9FB" pageId="64" pageNumber="93" type="diagnosis">
<paragraph id="E7726A0D5E00FFEFFF2BFAF0FEDAFA80" blockId="64.[144,760,1237,1612]" box="[144,256,1314,1335]" pageId="64" pageNumber="93">
<heading id="BC3ADD615E00FFEFFF2BFAF0FEDAFA80" bold="true" box="[144,256,1314,1335]" fontSize="9" level="5" pageId="64" pageNumber="93" reason="6">
<emphasis id="D5B9B61F5E00FFEFFF2BFAF0FEDAFA80" bold="true" box="[144,256,1314,1335]" italics="true" pageId="64" pageNumber="93">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="E7726A0D5E00FFEFFF2BFA93FEA5F9FB" blockId="64.[144,760,1237,1612]" pageId="64" pageNumber="93">Labium extending to mesocoxae. Distiflagellomere with a sub-basal whitish band. Evaporative area short, distance from dorsal margin of auricle to dorsal margin of evaporative area shorter than distance from dorsal margin of evaporative area to dorsal margin of metapleura. Membrane brown, veins paler, without an apical pale spot. Meso- and metafemur with a subapical narrow brown band. Pygophore apex pointed dorsally in lateral view.</paragraph>
</subSubSection>
<subSubSection id="AFD739865E00FFEDFF2BF950FB32FAF2" lastPageId="66" lastPageNumber="95" pageId="64" pageNumber="93" type="description">
<paragraph id="E7726A0D5E00FFEFFF2BF950FE53F920" blockId="64.[144,758,1666,1902]" box="[144,393,1666,1687]" pageId="64" pageNumber="93">
<heading id="BC3ADD615E00FFEFFF2BF950FE53F920" bold="true" box="[144,393,1666,1687]" fontSize="9" level="5" pageId="64" pageNumber="93" reason="6">
<emphasis id="D5B9B61F5E00FFEFFF2BF950FE53F920" bold="true" box="[144,393,1666,1687]" italics="true" pageId="64" pageNumber="93">
Description (
<figureCitation id="7FF676885E00FFEFFE99F950FEAAF920" box="[290,368,1666,1687]" captionStart="Figure 36" captionStartId="65.[164,244,1794,1813]" captionTargetBox="[164,1442,193,1762]" captionTargetId="figure@65.[164,1444,193,1764]" captionTargetPageId="65" captionText="Figure 36. Habitus dorsal: A, Heraeus mesoamericanus sp. nov.; B, Heraeus mexicanus sp. nov.; C, Heraeus morganae sp. nov.; D, Heraeus nicaraguensis sp. nov.; E, Heraeus pacificus Barber, 1925; F, Heraeus pallidinervis sp. nov.; G, Heraeus penai sp. nov.; H, Heraeus plebejus Stål, 1874; I, Heraeus pulchellus Barber, 1954." httpUri="https://zenodo.org/record/269784/files/figure.png" pageId="64" pageNumber="93">Fig. 36</figureCitation>
A)
</emphasis>
</heading>
</paragraph>
<paragraph id="E7726A0D5E00FFEFFF2BF972FE87F901" blockId="64.[144,758,1666,1902]" box="[144,349,1696,1718]" pageId="64" pageNumber="93">Total length 5.61.</paragraph>
<paragraph id="E7726A0D5E00FFEFFF13F96CFBFFFA49" blockId="64.[144,758,1666,1902]" lastBlockId="64.[806,1422,1237,1902]" pageId="64" pageNumber="93">
<emphasis id="D5B9B61F5E00FFEFFF13F96CFF36F963" bold="true" box="[168,236,1726,1748]" italics="true" pageId="64" pageNumber="93">Head:</emphasis>
Brown, shiny, coriaceous, with short recumbent and long erect setae dorsally. Head length 1.18, width 0.85, postocular length 0.43. Eyes not surpassing dorsal margin of head in lateral view. Ocelli posterior of an imaginary line passing the posterior border of eyes. Interocular width 0.42, interocellar width 0.22. Labium pale brown with abundant erect setae, extending to mesocoxae. Labial segment lengths: I 0.72, II 0.82, III 0.60, and IV 0.24. Antennae pale brown, apical half of basiflagellomere and distiflagellomere darker, distiflagellomere with a subbasal whitish band; all segments with abundant short recumbent and sparse short erect setae. Antennal lengths: scape 0.46, pedicel 1.03, basiflagellomere 0.91, and distiflagellomere 1.03. Length of pale band on distiflagellomere 0.24.
</paragraph>
<paragraph id="E7726A0D5E00FFEDFC84F9D5FE21F9EF" blockId="64.[806,1422,1237,1902]" lastBlockId="66.[144,760,1266,1900]" lastPageId="66" lastPageNumber="95" pageId="64" pageNumber="93">
<emphasis id="D5B9B61F5E00FFEFFC84F9D5FC4FF9AA" bold="true" box="[831,917,1543,1565]" italics="true" pageId="64" pageNumber="93">Thorax:</emphasis>
Anterior pronotal lobe brown, collar and posterior lobe paler, with four longitudinal pale stripes, humeral angles yellowish; collar delimited posteriorly by a punctate sulcus; anterior and posterior lobes with short recumbent and long erect setae, longer on anterior lobe. Pronotum pruinose, punctate, punctures larger on posterior lobe. Collar length 0.08, anterior lobe length 0.51, posterior lobe length 0.54; anterior lobe width 0.89, posterior lobe width 1.42. Pleurae brown, acetabular areas paler, punctate, with short recumbent setae. Evaporative area short. Scutellum brown, with a longitudinal median dark stripe, apex paler; pruinose, punctate; with short recumbent and long erect setae, as on anterior pronotal lobe. Hemelytra pale brown, irregularly pigmented. Costal margin pale on proximal three-quarters, apex of corium darker with a subapical whitish spot and an small pale inner spot; setae short and recumbent; membrane brown with veins paler (
<figureCitation id="7FF676885E02FFEDFF62FA7BFEFCFA08" box="[217,294,1449,1471]" captionStart="Figure 36" captionStartId="65.[164,244,1794,1813]" captionTargetBox="[164,1442,193,1762]" captionTargetId="figure@65.[164,1444,193,1764]" captionTargetPageId="65" captionText="Figure 36. Habitus dorsal: A, Heraeus mesoamericanus sp. nov.; B, Heraeus mexicanus sp. nov.; C, Heraeus morganae sp. nov.; D, Heraeus nicaraguensis sp. nov.; E, Heraeus pacificus Barber, 1925; F, Heraeus pallidinervis sp. nov.; G, Heraeus penai sp. nov.; H, Heraeus plebejus Stål, 1874; I, Heraeus pulchellus Barber, 1954." httpUri="https://zenodo.org/record/269784/files/figure.png" pageId="66" pageNumber="95">Fig. 36</figureCitation>
A). Legs: Coxae, protrochanter, profemur, except distally, and a subapical narrow band on meso- and metafemur brown, remainder of legs pale brown, apex of tibiae, tarsi, and pretarsi darker (
<figureCitation id="7FF676885E02FFEDFD3DF9D7FD0FF9AC" box="[646,725,1541,1563]" captionStart="Figure 33" captionStartId="60.[144,223,1062,1081]" captionTargetBox="[305,1262,196,1030]" captionTargetId="figure@60.[303,1263,195,1033]" captionTargetPageId="60" captionText="Figure 33. Legs: A, Heraeus concolor Slater &amp; Baranowski, 1994; B, Heraeus ecuatorianus sp. nov.; C, Heraeus loja sp. nov.; D, Heraeus mesoamericanus sp. nov.; E, Heraeus mexicanus sp. nov.; F, Heraeus morganae sp. nov." httpUri="https://zenodo.org/record/269781/files/figure.png" pageId="66" pageNumber="95">Fig. 33</figureCitation>
D); setae semi-erect, longest on profemur. Profemur with spines restricted to apical half.
</paragraph>
<caption id="B3B23A855E01FFEEFF1FF8D0FA78F8F8" httpUri="https://zenodo.org/record/269784/files/figure.png" pageId="65" pageNumber="94" targetBox="[164,1442,193,1762]" targetPageId="65">
<paragraph id="E7726A0D5E01FFEEFF1FF8D0FA78F8F8" blockId="65.[164,1442,1794,1872]" pageId="65" pageNumber="94">
<emphasis id="D5B9B61F5E01FFEEFF1FF8D0FEC6F8A2" bold="true" box="[164,284,1794,1813]" pageId="65" pageNumber="94">Figure 36.</emphasis>
Habitus dorsal: A,
<emphasis id="D5B9B61F5E01FFEEFE47F8D0FC54F8A2" bold="true" box="[508,910,1794,1813]" pageId="65" pageNumber="94">
<taxonomicName id="20CD118E5E01FFEEFE47F8D0FCF5F8A2" box="[508,815,1794,1813]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="65" pageNumber="94" phylum="Arthropoda" rank="species" species="mesoamericanus" status="sp. nov.">
<emphasis id="D5B9B61F5E01FFEEFE47F8D0FCF5F8A2" bold="true" box="[508,815,1794,1813]" italics="true" pageId="65" pageNumber="94">Heraeus mesoamericanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CE8A0B645E01FFEEFC8DF8D0FC54F8A2" box="[822,910,1794,1813]" pageId="65" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; B,
<emphasis id="D5B9B61F5E01FFEEFC79F8D0FAD6F8A2" bold="true" box="[962,1292,1794,1813]" pageId="65" pageNumber="94">
<taxonomicName id="20CD118E5E01FFEEFC79F8D0FB77F8A2" box="[962,1197,1794,1813]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="65" pageNumber="94" phylum="Arthropoda" rank="species" species="mexicanus" status="sp. nov.">
<emphasis id="D5B9B61F5E01FFEEFC79F8D0FB77F8A2" bold="true" box="[962,1197,1794,1813]" italics="true" pageId="65" pageNumber="94">Heraeus mexicanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CE8A0B645E01FFEEFB0FF8D0FAD6F8A2" box="[1204,1292,1794,1813]" pageId="65" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; C,
<emphasis id="D5B9B61F5E01FFEEFA84F8D0FE50F885" bold="true" pageId="65" pageNumber="94">
<taxonomicName id="20CD118E5E01FFEEFA84F8D0FEF9F885" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="65" pageNumber="94" phylum="Arthropoda" rank="species" species="morganae" status="sp. nov.">
<emphasis id="D5B9B61F5E01FFEEFA84F8D0FEF9F885" bold="true" italics="true" pageId="65" pageNumber="94">Heraeus morganae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CE8A0B645E01FFEEFE91F8CDFE50F885" box="[298,394,1823,1842]" pageId="65" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; D,
<emphasis id="D5B9B61F5E01FFEEFE7FF8CDFC8CF885" bold="true" box="[452,854,1823,1842]" pageId="65" pageNumber="94">
<taxonomicName id="20CD118E5E01FFEEFE7FF8CDFD34F885" box="[452,750,1823,1842]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="65" pageNumber="94" phylum="Arthropoda" rank="species" species="nicaraguensis" status="sp. nov.">
<emphasis id="D5B9B61F5E01FFEEFE7FF8CDFD34F885" bold="true" box="[452,750,1823,1842]" italics="true" pageId="65" pageNumber="94">Heraeus nicaraguensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CE8A0B645E01FFEEFD4DF8CDFC8CF885" box="[758,854,1823,1842]" pageId="65" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; E,
<taxonomicName id="20CD118E5E01FFEEFC34F8CDFADDF885" authority="Barber, 1925" authorityName="Barber" authorityYear="1925" box="[911,1287,1823,1842]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="65" pageNumber="94" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="D5B9B61F5E01FFEEFC34F8CDFB84F885" bold="true" box="[911,1118,1823,1842]" italics="true" pageId="65" pageNumber="94">Heraeus pacificus</emphasis>
Barber, 1925
</taxonomicName>
; F,
<emphasis id="D5B9B61F5E01FFEEFA82F8CDFE4DF8E7" bold="true" pageId="65" pageNumber="94">
<taxonomicName id="20CD118E5E01FFEEFA82F8CDFEE1F8F8" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="65" pageNumber="94" phylum="Arthropoda" rank="species" species="pallidinervis" status="sp. nov.">
<emphasis id="D5B9B61F5E01FFEEFA82F8CDFEE1F8F8" bold="true" italics="true" pageId="65" pageNumber="94">Heraeus pallidinervis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CE8A0B645E01FFEEFEFAF8EFFE4DF8E7" box="[321,407,1853,1872]" pageId="65" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; G,
<emphasis id="D5B9B61F5E01FFEEFE7EF8EFFD11F8E7" bold="true" box="[453,715,1852,1872]" pageId="65" pageNumber="94">
<taxonomicName id="20CD118E5E01FFEEFE7EF8EFFDB5F8F8" box="[453,623,1852,1872]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="65" pageNumber="94" phylum="Arthropoda" rank="species" species="penai" status="sp. nov.">
<emphasis id="D5B9B61F5E01FFEEFE7EF8EFFDB5F8F8" bold="true" box="[453,623,1852,1872]" italics="true" pageId="65" pageNumber="94">Heraeus penai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CE8A0B645E01FFEEFDCEF8EFFD11F8E7" box="[629,715,1853,1872]" pageId="65" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; H,
<taxonomicName id="20CD118E5E01FFEEFD42F8EFFBFBF8E7" authority="Stal, 1874" authorityName="Stal" authorityYear="1874" box="[761,1057,1852,1872]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="65" pageNumber="94" phylum="Arthropoda" rank="species" species="plebejus">
<emphasis id="D5B9B61F5E01FFEEFD42F8EFFC71F8F8" bold="true" box="[761,939,1852,1872]" italics="true" pageId="65" pageNumber="94">Heraeus plebejus</emphasis>
Stål, 1874
</taxonomicName>
; I,
<taxonomicName id="20CD118E5E01FFEEFBF8F8EFFA47F8F8" authority="Barber, 1954" authorityName="Barber" authorityYear="1954" box="[1091,1437,1852,1872]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="65" pageNumber="94" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="D5B9B61F5E01FFEEFBF8F8EFFAD1F8F8" bold="true" box="[1091,1291,1852,1872]" italics="true" pageId="65" pageNumber="94">Heraeus pulchellus</emphasis>
Barber, 1954
</taxonomicName>
.
</paragraph>
</caption>
<caption id="B3B23A855E02FFEDFF2BFB5CFC53FB16" box="[144,905,1166,1185]" httpUri="https://zenodo.org/record/269785/files/figure.png" pageId="66" pageNumber="95" targetBox="[147,1420,198,1132]" targetPageId="66">
<paragraph id="E7726A0D5E02FFEDFF2BFB5CFC53FB16" blockId="66.[144,905,1166,1185]" box="[144,905,1166,1185]" pageId="66" pageNumber="95">
<emphasis id="D5B9B61F5E02FFEDFF2BFB5CFEDFFB16" bold="true" box="[144,261,1166,1185]" pageId="66" pageNumber="95">Figure 37.</emphasis>
Distributional map of
<taxonomicName id="20CD118E5E02FFEDFDBBFB5CFCF6FB16" box="[512,812,1166,1185]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="66" pageNumber="95" phylum="Arthropoda" rank="species" species="mesoamericanus" status="sp. nov.">
<emphasis id="D5B9B61F5E02FFEDFDBBFB5CFCF6FB16" bold="true" box="[512,812,1166,1185]" italics="true" pageId="66" pageNumber="95">Heraeus mesoamericanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CE8A0B645E02FFEDFC88FB5CFC53FB16" box="[819,905,1166,1185]" pageId="66" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="E7726A0D5E02FFEDFF13F9B3FD2DF8DB" blockId="66.[144,760,1266,1900]" pageId="66" pageNumber="95">
<emphasis id="D5B9B61F5E02FFEDFF13F9B3FEC0F9C0" bold="true" box="[168,282,1633,1655]" italics="true" pageId="66" pageNumber="95">Abdomen:</emphasis>
Brown, with abundant short recumbent setae. Male genitalia: Pygophore (
<figureCitation id="7FF676885E02FFEDFDB1F952FD8DF922" box="[522,599,1664,1686]" captionStart="Figure 34" captionStartId="63.[164,241,1714,1733]" captionTargetBox="[256,1348,194,1683]" captionTargetId="figure@63.[253,1354,192,1684]" captionTargetPageId="63" captionText="Figure 34. Male genitalia. Heraeus loja sp. nov.: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus mesoamericanus sp. nov.: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus mexicanus sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view. Heraeus morganae sp. nov.: M, pygophore, dorsal view; N, pygophore, lateral view; O, right paramere, inner view; P, right paramere, external view." httpUri="https://zenodo.org/record/269782/files/figure.png" pageId="66" pageNumber="95">Fig. 34</figureCitation>
E, F) rounded, elongate, with pointed posterior margin, apex with a dorsally directed protuberance, declivent posteriorly in lateral view; anterior margin of dorsal aperture rounded. Inner projections subquadrangular. Parameres:
<figureCitation id="7FF676885E02FFEDFF2BF8CBFF21F899" box="[144,251,1817,1839]" captionStart="Figure 34" captionStartId="63.[164,241,1714,1733]" captionTargetBox="[256,1348,194,1683]" captionTargetId="figure@63.[253,1354,192,1684]" captionTargetPageId="63" captionText="Figure 34. Male genitalia. Heraeus loja sp. nov.: A, pygophore, dorsal view; B, pygophore, lateral view; C, right paramere, inner view; D, right paramere, external view. Heraeus mesoamericanus sp. nov.: E, pygophore, dorsal view; F, pygophore, lateral view; G, right paramere, inner view; H, right paramere, external view. Heraeus mexicanus sp. nov.: I, pygophore, dorsal view; J, pygophore, lateral view; K, right paramere, inner view; L, right paramere, external view. Heraeus morganae sp. nov.: M, pygophore, dorsal view; N, pygophore, lateral view; O, right paramere, inner view; P, right paramere, external view." httpUri="https://zenodo.org/record/269782/files/figure.png" pageId="66" pageNumber="95">Figure 34</figureCitation>
(G, H). Aedeagus: conjunctiva and vesica with a few sparse spines laterally, vesica with two sclerotized lobes;
<emphasis id="D5B9B61F5E02FFEDFF62F884FE74F8DC" bold="true" box="[217,430,1878,1899]" italics="true" pageId="66" pageNumber="95">processus gonopori</emphasis>
long and slender (
<figureCitation id="7FF676885E02FFEDFD33F884FD00F8DB" box="[648,730,1878,1900]" captionStart="Figure 30" captionStartId="55.[164,240,1815,1834]" captionTargetBox="[168,1442,193,1783]" captionTargetId="figure@55.[164,1444,193,1786]" captionTargetPageId="55" captionText="Figure 30. Aedeagus. A, Heraeus bahiensis sp. nov.; B, Heraeus bolivianus sp. nov.; C, Heraeus brevirostris sp. nov.; D, Heraeus concolor Slater &amp; Baranowski, 1994; E, Heraeus ecuatorianus sp. nov.; F, Heraeus mesoamericanus sp. nov.; G, Heraeus mexicanus sp. nov.; H, Heraeus penai sp. nov." httpUri="https://zenodo.org/record/269778/files/figure.png" pageId="66" pageNumber="95">Fig. 30</figureCitation>
F).
</paragraph>
<paragraph id="E7726A0D5E02FFEDFC9DFB23FB32FAF2" blockId="66.[806,1421,1265,1349]" pageId="66" pageNumber="95">
<emphasis id="D5B9B61F5E02FFEDFC9DFB23FAEAFAB0" bold="true" box="[806,1328,1265,1287]" italics="true" pageId="66" pageNumber="95">Variability observed in other material studied</emphasis>
In the specimens from Belize, the labium is slightly longer, barely extending to metacoxae.
</paragraph>
</subSubSection>
<subSubSection id="AFD739865E02FFEDFC9DFAA0FC56FA72" pageId="66" pageNumber="95" type="distribution">
<paragraph id="E7726A0D5E02FFEDFC9DFAA0FC6BFA3F" blockId="66.[806,1421,1394,1477]" box="[806,945,1394,1416]" pageId="66" pageNumber="95">
<heading id="BC3ADD615E02FFEDFC9DFAA0FC6BFA3F" bold="true" box="[806,945,1394,1416]" fontSize="9" level="5" pageId="66" pageNumber="95" reason="6">
<emphasis id="D5B9B61F5E02FFEDFC9DFAA0FC6BFA3F" bold="true" box="[806,945,1394,1416]" italics="true" pageId="66" pageNumber="95">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="E7726A0D5E02FFEDFC9DFA43FC56FA72" blockId="66.[806,1421,1394,1477]" pageId="66" pageNumber="95">
Belize, Costa Rica, Ecuador, Honduras, and Panama (
<figureCitation id="7FF676885E02FFEDFC95FA7DFCA5FA72" box="[814,895,1455,1477]" captionStart="Figure 37" captionStartId="66.[144,223,1166,1185]" captionTargetBox="[147,1420,198,1132]" captionTargetId="figure@66.[144,1424,195,1136]" captionTargetPageId="66" captionText="Figure 37. Distributional map of Heraeus mesoamericanus sp. nov." httpUri="https://zenodo.org/record/269785/files/figure.png" pageId="66" pageNumber="95">Fig. 37</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="AFD739865E02FFEDFC9DFA20FB3BF9D3" pageId="66" pageNumber="95" type="etymology">
<paragraph id="E7726A0D5E02FFEDFC9DFA20FC47F9BF" blockId="66.[806,1421,1522,1636]" box="[806,925,1522,1544]" pageId="66" pageNumber="95">
<heading id="BC3ADD615E02FFEDFC9DFA20FC47F9BF" bold="true" box="[806,925,1522,1544]" fontSize="9" level="5" pageId="66" pageNumber="95" reason="6">
<emphasis id="D5B9B61F5E02FFEDFC9DFA20FC47F9BF" bold="true" box="[806,925,1522,1544]" italics="true" pageId="66" pageNumber="95">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="E7726A0D5E02FFEDFC9DF9C3FB3BF9D3" blockId="66.[806,1421,1522,1636]" pageId="66" pageNumber="95">
The specific epithet ‘
<taxonomicName id="20CD118E5E02FFEDFBA2F9C3FB0FF991" box="[1049,1237,1553,1574]" class="Insecta" family="Rhyparochromidae" genus="Heraeus" kingdom="Animalia" order="Hemiptera" pageId="66" pageNumber="95" phylum="Arthropoda" rank="species" species="mesoamericanus">
<emphasis id="D5B9B61F5E02FFEDFBA2F9C3FB0FF991" bold="true" box="[1049,1237,1553,1574]" italics="true" pageId="66" pageNumber="95">mesoamericanus</emphasis>
</taxonomicName>
’ is an adjective meaning Central American, and refers to the known geographic distribution of the species.
</paragraph>
</subSubSection>
<subSubSection id="AFD739865E02FFECFC9DF943FE30F8D9" lastPageId="67" lastPageNumber="96" pageId="66" pageNumber="95" type="materials_examined">
<paragraph id="E7726A0D5E02FFEDFC9DF943FC19F910" blockId="66.[806,1421,1681,1795]" box="[806,963,1681,1703]" pageId="66" pageNumber="95">
<heading id="BC3ADD615E02FFEDFC9DF943FC19F910" bold="true" box="[806,963,1681,1703]" fontSize="9" level="5" pageId="66" pageNumber="95" reason="6">
<emphasis id="D5B9B61F5E02FFEDFC9DF943FC19F910" bold="true" box="[806,963,1681,1703]" italics="true" pageId="66" pageNumber="95">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="E7726A0D5E02FFEDFC9DF97DFC57F8B4" blockId="66.[806,1421,1681,1795]" pageId="66" pageNumber="95">
<emphasis id="D5B9B61F5E02FFEDFC9DF97DFC4BF972" bold="true" box="[806,913,1711,1733]" italics="true" pageId="66" pageNumber="95">Holotype:</emphasis>
♂,
<emphasis id="D5B9B61F5E02FFEDFC79F97DFBB0F972" bold="true" box="[962,1130,1711,1733]" pageId="66" pageNumber="95">COSTA RICA</emphasis>
,
<emphasis id="D5B9B61F5E02FFEDFBC1F97DFB3BF972" bold="true" box="[1146,1249,1711,1733]" italics="true" pageId="66" pageNumber="95">San Jose</emphasis>
, San Isidro de Coronado, 550 ft, 11/12-VI-[19]67, BL trap, E.B. Fogar (USNM).
</paragraph>
<paragraph id="E7726A0D5E02FFECFC9DF8C9FE12FEE2" blockId="66.[806,1421,1819,1902]" lastBlockId="67.[188,779,196,341]" lastPageId="67" lastPageNumber="96" pageId="66" pageNumber="95">
<emphasis id="D5B9B61F5E02FFEDFC9DF8C9FC4DF887" bold="true" box="[806,919,1819,1840]" italics="true" pageId="66" pageNumber="95">Paratypes:</emphasis>
5♂, 11♀, same data as for holotype (USNM); 1♂, 2♀, San Isidro de Coronado, Finca Joseph Smids, 5500 ft, 31-V/2-VI-[19]67, blacklight trap, E.B. Fagan (USNM); 1♂, 1♀, 31-V/2-VI-[19]67, blacklight trap, E.B. Fagan (MLP);
<emphasis id="D5B9B61F5E03FFECFE1AFF36FDFEFF4E" bold="true" box="[417,548,228,249]" italics="true" pageId="67" pageNumber="96">Puntarenas</emphasis>
: 7♂, 7♀, finca Las Cruces, 6 km S San Vito de Java, 4200 ft, 28-IX/2- X-1986, J.E. Eger (USNM); 1♂, 1♀, 28-IX/2-X- 1986, J.E. Eger (MLP).
</paragraph>
<paragraph id="E7726A0D5E03FFECFF1FFE6BFE39FE78" blockId="67.[164,780,441,1902]" box="[164,483,441,463]" pageId="67" pageNumber="96">
<heading id="BC3ADD615E03FFECFF1FFE6BFE39FE78" bold="true" box="[164,483,441,463]" fontSize="9" level="5" pageId="67" pageNumber="96" reason="6">
<emphasis id="D5B9B61F5E03FFECFF1FFE6BFE39FE78" bold="true" box="[164,483,441,463]" italics="true" pageId="67" pageNumber="96">Additional material studied</emphasis>
</heading>
</paragraph>
<paragraph id="E7726A0D5E03FFECFF1FFE0AFCD0FE59" blockId="67.[164,780,441,1902]" box="[164,778,472,494]" pageId="67" pageNumber="96">
<emphasis id="D5B9B61F5E03FFECFF1FFE0AFEC8FE5A" bold="true" box="[164,274,472,493]" pageId="67" pageNumber="96">BELIZE:</emphasis>
2♂, 3♀, Orange Walk Dist., Chan Chich
</paragraph>
<paragraph id="E7726A0D5E03FFECFF1FFE25FCD1FDBB" blockId="67.[164,780,441,1902]" box="[164,779,502,525]" pageId="67" pageNumber="96">Lodge, 1-IV-[20]09, BLT, E. Flota (USNM); 1♂, Cayo</paragraph>
<paragraph id="E7726A0D5E03FFECFF1FFDC4FCD0FD9C" blockId="67.[164,780,441,1902]" box="[164,778,533,556]" pageId="67" pageNumber="96">Dist., Chiquibul Forest Res., 27-III-[20]08, BL Trap,</paragraph>
<paragraph id="E7726A0D5E03FFECFF1FFDE1FCD0FDFD" blockId="67.[164,780,441,1902]" box="[164,778,563,586]" pageId="67" pageNumber="96">27-III-[20]08, B. Miller (USNM); 1♂, Cayo Dist., Benque</paragraph>
<paragraph id="E7726A0D5E03FFECFF1FFD81FCD6FDDF" blockId="67.[164,780,441,1902]" box="[164,780,595,617]" pageId="67" pageNumber="96">Viejo del Carmen, Riverside N, Mopan</paragraph>
<paragraph id="E7726A0D5E03FFECFF1FFDA3FCD1FD30" blockId="67.[164,780,441,1902]" box="[164,779,625,647]" pageId="67" pageNumber="96">River Resort, 20-XI-[20]07, BL Trap, M. Chrysler</paragraph>
<paragraph id="E7726A0D5E03FFECFF1FFD42FCD1FD11" blockId="67.[164,780,441,1902]" box="[164,779,655,678]" pageId="67" pageNumber="96">(USNM); 1♂, 13-XII-[20]07, BL Trap, M. Chrysler</paragraph>
<paragraph id="E7726A0D5E03FFECFF1FFD7DFED1FD72" blockId="67.[164,780,441,1902]" box="[164,267,687,709]" pageId="67" pageNumber="96">(USNM).</paragraph>
<paragraph id="E7726A0D5E03FFECFF1FFD1FFEF9FC2C" blockId="67.[164,780,441,1902]" pageId="67" pageNumber="96">
<emphasis id="D5B9B61F5E03FFECFF1FFD1FFE8BFD54" bold="true" box="[164,337,717,739]" pageId="67" pageNumber="96">COSTA RICA:</emphasis>
1♂, Torrazu, 20-VIII-[19]36, on
<taxonomicName id="20CD118E5E03FFECFD73FD1FFE5DFCB5" authority="E. Marin" authorityName="E. Marin" class="Insecta" family="Rhyparochromidae" genus="Coffee" kingdom="Animalia" order="Hemiptera" pageId="67" pageNumber="96" phylum="Arthropoda" rank="species" species="arabica">
<emphasis id="D5B9B61F5E03FFECFD73FD1FFEC8FCB6" bold="true" italics="true" pageId="67" pageNumber="96">Coffee arabica</emphasis>
, E. Marin
</taxonomicName>
(USNM);
<emphasis id="D5B9B61F5E03FFECFE46FD3EFD82FCB6" bold="true" box="[509,600,748,769]" italics="true" pageId="67" pageNumber="96">Cartago</emphasis>
: 1♀, Turrialba, Int. Amer. Inst. Agr. Sci., blacklight trap, 18/20-VIII- [19]64, R.E. Woodruff (USNM);
<emphasis id="D5B9B61F5E03FFECFDC8FCFBFCDEFC89" bold="true" box="[627,772,809,830]" italics="true" pageId="67" pageNumber="96">Puntarenas</emphasis>
: 1♂, Turrialba, 27-I-1965, J.A. Slater &amp; N.T. Davis (AMNH); 1♀, on
<taxonomicName id="20CD118E5E03FFECFE0CFCB4FD5CFCCB" box="[439,646,870,892]" class="Magnoliopsida" family="Asteraceae" genus="Chrysanthemum" kingdom="Plantae" order="Asterales" pageId="67" pageNumber="96" phylum="Tracheophyta" rank="genus">
<emphasis id="D5B9B61F5E03FFECFE0CFCB4FD5CFCCB" bold="true" box="[439,646,870,892]" italics="true" pageId="67" pageNumber="96">Chrysanthemum</emphasis>
</taxonomicName>
, V-[19]87 (USNM).
</paragraph>
<paragraph id="E7726A0D5E03FFECFF1FFC71FE22FBA2" blockId="67.[164,780,441,1902]" pageId="67" pageNumber="96">
<emphasis id="D5B9B61F5E03FFECFF1FFC71FE14FC0E" bold="true" box="[164,462,931,953]" pageId="67" pageNumber="96">
ECUADOR:
<emphasis id="D5B9B61F5E03FFECFEFBFC71FE14FC0E" bold="true" box="[320,462,931,953]" italics="true" pageId="67" pageNumber="96">Tungurahua</emphasis>
</emphasis>
: 2♂, Baños, 6200 ft, 24-I- 1976, Spangler
<emphasis id="D5B9B61F5E03FFECFECCFC10FE7DFC60" bold="true" box="[375,423,961,983]" italics="true" pageId="67" pageNumber="96">et al</emphasis>
. (USNM); 2♂, 3♀, Finca Las Cruces, 6 km S San Vito de Java, 4200 ft, 28-IX/2- X-1986, J.E. Eger (USNM).
</paragraph>
<paragraph id="E7726A0D5E03FFECFF1FFBCFFE4DFB7A" blockId="67.[164,780,441,1902]" pageId="67" pageNumber="96">
<emphasis id="D5B9B61F5E03FFECFF1FFBCFFE9CFB84" bold="true" box="[164,326,1053,1075]" pageId="67" pageNumber="96">HONDURAS:</emphasis>
1♂, 1♀, Comayagua, Siguatepeque, 25- VIII-[19]66, mosquito trap, R.E. Woodruff (USNM); 1♂, 21 km NW Siguatepeque, Ch. 5 rd., 1-VIII- 1977, O’Brien &amp; Marshall (AMNH); 1♀, Sta. B., 13 km SE El Mochito, 22-VII-1977, C.W. &amp; L. O’Brien &amp; Marshall (AMNH).
</paragraph>
<paragraph id="E7726A0D5E03FFECFF1FFB07FE30F8D9" blockId="67.[164,780,441,1902]" pageId="67" pageNumber="96">
<emphasis id="D5B9B61F5E03FFECFF1FFB07FE4BFB5C" bold="true" box="[164,401,1237,1259]" pageId="67" pageNumber="96">
PANAMA:
<emphasis id="D5B9B61F5E03FFECFE95FB07FE4BFB5C" bold="true" box="[302,401,1237,1259]" italics="true" pageId="67" pageNumber="96">Chiriqui</emphasis>
</emphasis>
, 3♂, 15♀, Dist. Renacimiento, Santa Clara, 4000 ft, 28/29-V-[19]76, at light, Engleman &amp; Thurman (AMNH); 1♂, 1♀, 28/29-V- [19]76, at light, Engleman &amp; Thurman (MLP); 2♂, 1♀, 20/22-V-[19]77, Engleman (AMNH); 1♂, 3♀, 29- V-[19]76, D. Engleman (AMNH); 1♂, Rovira, Elvira Farm, 15-VIII-[19]64, 220 ft, blacklight trap, A.B. Broce (USNM); 3♂, Rovira, 5-VII-[19]64, 2500 ft, mosquito light trap, A. Bruse (USNM); 1♂, 9-V-[19]64, mosquito light trap, A. Bruse (USNM); 1♀, Rovira, 4-VII- [19]64, blacklight trap, A.B. Brose (USNM); 1♂, 1♀, Volcan Chiriqui, 3500 ft, 4-XI-[19]73, D. Engleman (AMNH); 1♂, 1♀, Volcan Aria, 4500 ft, 25-V- [19]73, lagos, D. Engleman (USNM); 1♂, 11-IV- [19]73, D. Engleman (USNM); 1♀, 2.5 mi. W El Volcan, 4400 ft, Las Lagunas, 9-VII-1974, O’Briens &amp; Marshall (AMNH); 1♂, 2♀, Res. For. La Fortuna, El Vivero, 20-VII-1995, UV-HG lights, L.B. O’Brien (USNM);
<emphasis id="D5B9B61F5E03FFECFE89F92EFE4FF8A6" bold="true" box="[306,405,1788,1809]" italics="true" pageId="67" pageNumber="96">Panama</emphasis>
, 1♂, Cerro Campana, 29-VII- 1995, C.W. &amp; L.B. O’Brien (USNM); 1♀, Interamerican Hwy. 3 km E of Ipeli, 3-V-1992, UV and HG vapour light, A. Gillogly (USNM).
</paragraph>
</subSubSection>
</treatment>
</document>
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