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<document id="D3455D7183B8C91CF1BF823F6190261C" ID-DOI="10.1080/00222930500087408" ID-ISSN="1464-5262" ID-Zenodo-Dep="5217766" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629301816659" checkinUser="felipe" docAuthor="Ohtsuka, Susumu, Nishida, Shuhei &amp; Machida, Ryuji J." docDate="2005" docId="03E0481BFFDFFF9CD7EA5340FEF42CE5" docLanguage="en" docName="JNatHist.39.27.2483-2514.pdf" docOrigin="Journal of Natural History 39 (27)" docSource="http://dx.doi.org/10.1080/00222930500087408" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Metacalanalis Ohtsuka &amp; Nishida &amp; Machida 2005, n. gen." docType="treatment" docVersion="3" lastPageNumber="2486" masterDocId="FFD93063FFDCFF99D63D5678FFAE2E01" masterDocTitle="Systematics and zoogeography of the deep-sea hyperbenthic family Arietellidae (Copepoda: Calanoida) collected from the Sulu Sea" masterLastPageNumber="2514" masterPageNumber="2483" pageNumber="2484" updateTime="1699067816347" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="223B1E186AEBEB9B9FE956B08C7FD4AA">Systematics and zoogeography of the deep-sea hyperbenthic family Arietellidae (Copepoda: Calanoida) collected from the Sulu Sea</mods:title>
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<mods:namePart id="557AF5405BEB56441D0A3327625BA244">Ohtsuka, Susumu</mods:namePart>
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<mods:namePart id="25D20D7BAC581B74FC263D9CFF2F10FF">Nishida, Shuhei</mods:namePart>
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<mods:namePart id="B36356669181F7BE740BA38A87142151">Machida, Ryuji J.</mods:namePart>
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<mods:title id="38E85463AD5FDD609148B41C000CFBF0">Journal of Natural History</mods:title>
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<mods:date id="6DAC645DCB2FAAAE401D3431C43AC845">2005</mods:date>
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<subSubSection id="C353AA86FFDFFF9AD7EA5340FCEE2B51" box="[471,832,1336,1360]" pageId="3" pageNumber="2484" type="nomenclature">
<paragraph id="8BF6F90DFFDFFF9AD7EA5340FCEE2B51" blockId="3.[421,877,1303,1360]" box="[471,832,1336,1360]" pageId="3" pageNumber="2484">
<heading id="D0BE4E61FFDFFF9AD7EA5340FCEE2B51" box="[471,832,1336,1360]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="2484" reason="2">
<emphasis id="B93D251FFFDFFF9AD7EA5340FD482B51" bold="true" box="[471,742,1336,1360]" pageId="3" pageNumber="2484">
Genus
<taxonomicName id="4C49828EFFDFFF9AD40C5340FD482B51" authority="Ohtsuka &amp; Nishida &amp; Machida, 2005" authorityName="Ohtsuka &amp; Nishida &amp; Machida" authorityYear="2005" box="[561,742,1336,1360]" class="Hexanauplia" family="Arietellidae" genus="Metacalanalis" kingdom="Animalia" order="Calanoida" pageId="3" pageNumber="2484" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93D251FFFDFFF9AD40C5340FD482B51" bold="true" box="[561,742,1336,1360]" italics="true" pageId="3" pageNumber="2484">Metacalanalis</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A20E9864FFDFFF9AD4D25340FCEE2B51" box="[751,832,1336,1360]" pageId="3" pageNumber="2484" rank="genus">n. gen.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C353AA86FFDFFF9CD64053F1FEF42CE5" lastPageId="5" lastPageNumber="2486" pageId="3" pageNumber="2484" type="diagnosis">
<paragraph id="8BF6F90DFFDFFF9AD64053F1FF482BA0" blockId="3.[125,230,1417,1441]" box="[125,230,1417,1441]" pageId="3" pageNumber="2484">
<emphasis id="B93D251FFFDFFF9AD64053F1FF482BA0" box="[125,230,1417,1441]" italics="true" pageId="3" pageNumber="2484">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BF6F90DFFDFFF9AD64053C2FE292895" blockId="3.[125,1172,1466,1684]" pageId="3" pageNumber="2484">
<emphasis id="B93D251FFFDFFF9AD64053C2FF7B2BD3" box="[125,213,1466,1490]" italics="true" pageId="3" pageNumber="2484">Female.</emphasis>
Body compact. Prosome plump, about 2.6 times as long as urosome; cephalosome asymmetrical anteriorly, with curved rostrum; prosomal ends almost symmetrical, without dorsolateral and ventrolateral processes. Urosome with genital double-somite wider than long; paired gonopores and copulatory pores located at mid-length of genital doublesomite; each copulatory pore slit-like, located at inner corner of gonopore; seminal receptacles located ventrally, massive; anal operculum lacking; caudal rami symmetrical, nearly as long as wide.
</paragraph>
<paragraph id="8BF6F90DFFD8FF9DD69856F1FEB12C47" blockId="4.[138,1185,137,680]" pageId="4" pageNumber="2485">
Antennules asymmetrical, 21-segmented, left longer than right and reaching to pediger 3; posterior margin lacking long setules; first (IIV) and last (XXIVXXVIII) segments compound; segments II, IV, VI, VIII, and X lacking aesthetasc. Antenna with indistinctly three-segmented endopod; first endopodal segment bearing one inner seta, second and third segments bearing three and five setae, respectively; exopod indistinctly 10-segmented. Mandibular gnathobase without patch of long setules; four teeth present, dorsalmost tricuspid at tip; endopod represented by rudimentary segment with two unequal setae terminally; seta on first to fifth exopodal segments not reduced. Maxillule with one short and four long spines on praecoxal arthrite; coxal endite bearing single seta; coxal epipodite with six setae; endopod one-segmented, with two setae terminally. Maxilla with two setae and one vestigial element on first praecoxal endite; basal spine with short row of three or four minute spinules at mid-length. Maxilliped with innermost seta on fourth and fifth exopodal segments not reduced; setae
<emphasis id="B93D251FFFD8FF9DD4775476FDF62C24" box="[586,600,526,549]" italics="true" pageId="4" pageNumber="2485">a</emphasis>
and
<emphasis id="B93D251FFFD8FF9DD4AE5475FD312C24" box="[659,671,525,549]" italics="true" pageId="4" pageNumber="2485">b</emphasis>
(see
<bibRefCitation id="EFD884FCFFD8FF9DD4E05475FC622C24" author="Ohtsuka S &amp; Boxshall GA &amp; Roe HSJ" box="[733,972,525,549]" pageId="4" pageNumber="2485" pagination="105 - 172" refId="ref13031" refString="Ohtsuka S, Boxshall GA, Roe HSJ. 1994. Phylogenetic relationships between arietellid genera (Copepoda: Calanoida), with the establishment of three new genera. Bulletin of the Natural History Museum (Zoology Series) 60: 105 - 172." type="journal article" year="1994">Ohtsuka et al., 1994</bibRefCitation>
) on sixth segment not reduced.
</paragraph>
<paragraph id="8BF6F90DFFD8FF9DD6985437FC722CA9" blockId="4.[138,1185,137,680]" pageId="4" pageNumber="2485">Leg 1 with third exopodal segment bearing only one outer spine. Some elements of legs 14 asymmetrical. Leg 5 symmetrical, with coxae and intercoxal sclerite separate; endopod lacking; exopod unisegmented, with two lateral and two terminal spines.</paragraph>
<paragraph id="8BF6F90DFFD8FF9DD6B754A8FEFA2CE9" blockId="4.[138,340,720,744]" box="[138,340,720,744]" pageId="4" pageNumber="2485">
<emphasis id="B93D251FFFD8FF9DD6B754A8FF652CE9" box="[138,203,720,744]" italics="true" pageId="4" pageNumber="2485">Male.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BF6F90DFFD8FF9DD6B75559FF472D38" blockId="4.[138,233,801,825]" box="[138,233,801,825]" pageId="4" pageNumber="2485">
<emphasis id="B93D251FFFD8FF9DD6B75559FF472D38" box="[138,233,801,825]" italics="true" pageId="4" pageNumber="2485">Remarks</emphasis>
</paragraph>
<paragraph id="8BF6F90DFFD8FF9DD6B7552AFCC82BB3" blockId="4.[138,1185,850,1685]" pageId="4" pageNumber="2485">
The new genus is similar to the genera
<taxonomicName id="4C49828EFFD8FF9DD47B552AFCCC2D6B" authority="Cleve, 1901" authorityName="Cleve" authorityYear="1901" box="[582,866,850,874]" class="Hexanauplia" family="Arietellidae" genus="Metacalanus" kingdom="Animalia" order="Calanoida" pageId="4" pageNumber="2485" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD8FF9DD47B552AFD7F2D6B" box="[582,721,850,874]" italics="true" pageId="4" pageNumber="2485">Metacalanus</emphasis>
Cleve, 1901
</taxonomicName>
and
<taxonomicName id="4C49828EFFD8FF9DD5A7552AFB322D6B" authority="Alvarez, 1985" authorityName="Alvarez" authorityYear="1985" box="[922,1180,850,874]" class="Hexanauplia" family="Arietellidae" genus="Pilarella" kingdom="Animalia" order="Calanoida" pageId="4" pageNumber="2485" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD8FF9DD5A7552AFC5A2D6B" box="[922,1012,850,874]" italics="true" pageId="4" pageNumber="2485">Pilarella</emphasis>
<bibRefCitation id="EFD884FCFFD8FF9DD5C7552AFB322D6B" author="Alvarez MPJ" box="[1018,1180,850,874]" pageId="4" pageNumber="2485" pagination="189 - 195" refId="ref12255" refString="Alvarez MPJ. 1985. A new arietellid copepod (Crustacea): Pilarella longicornis, gen. n., sp. n., from the Brazilian continental shelf. Revista Brasileira de Zoologia 28: 189 - 195." type="journal article" year="1985">Alvarez, 1985</bibRefCitation>
</taxonomicName>
, and has mixed features of these two genera. The discriminative features between these are as follows (cf.
<bibRefCitation id="EFD884FCFFD8FF9DD77955EBFE492DAA" author="Alvarez MPJ" box="[324,487,915,939]" pageId="4" pageNumber="2485" pagination="189 - 195" refId="ref12255" refString="Alvarez MPJ. 1985. A new arietellid copepod (Crustacea): Pilarella longicornis, gen. n., sp. n., from the Brazilian continental shelf. Revista Brasileira de Zoologia 28: 189 - 195." type="journal article" year="1985">Alvarez 1985</bibRefCitation>
;
<bibRefCitation id="EFD884FCFFD8FF9DD7C455EBFD5A2DAA" author="Ohtsuka S &amp; Boxshall GA &amp; Roe HSJ" box="[505,756,915,939]" pageId="4" pageNumber="2485" pagination="105 - 172" refId="ref13031" refString="Ohtsuka S, Boxshall GA, Roe HSJ. 1994. Phylogenetic relationships between arietellid genera (Copepoda: Calanoida), with the establishment of three new genera. Bulletin of the Natural History Museum (Zoology Series) 60: 105 - 172." type="journal article" year="1994">Ohtsuka et al. 1994</bibRefCitation>
) (
<taxonomicName id="4C49828EFFD8FF9DD52955EBFCC02DAA" authorityName="Alvarez" authorityYear="1985" box="[788,878,915,939]" class="Hexanauplia" family="Arietellidae" genus="Pilarella" kingdom="Animalia" order="Calanoida" pageId="4" pageNumber="2485" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD8FF9DD52955EBFCC02DAA" box="[788,878,915,939]" italics="true" pageId="4" pageNumber="2485">Pilarella</emphasis>
</taxonomicName>
versus
<taxonomicName id="4C49828EFFD8FF9DD5EF55EBFBF32DAA" authorityName="Cleve" authorityYear="1901" box="[978,1117,915,939]" class="Hexanauplia" family="Arietellidae" genus="Metacalanus" kingdom="Animalia" order="Calanoida" pageId="4" pageNumber="2485" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD8FF9DD5EF55EBFBF32DAA" box="[978,1117,915,939]" italics="true" pageId="4" pageNumber="2485">Metacalanus</emphasis>
</taxonomicName>
): (1) location and configuration of female gonopores and copulatory pores (medially located and paired versus posteriorly and paired or unpaired); (2) relative length of caudal rami (about five times longer than wide versus at most twice); (3) segmentation and relative length of antennules (21-segmented in both, left antennule reaching beyond urosomal end versus 16- or 18-segmented in left and 18- or 20-segmented in right, left antennule not reaching prosomal end); (4) setal number on the second exopodal segments of antenna (three medial setae on the second segment versus only two); (5) armature of the praecoxal arthrite, coxal endite and endopod of maxillule (formula with 6, 1, 2 versus 02, 01, 01); (6) armature and ornamentation of elements on the maxillary endites (two distinct setae + one rudimentary element on the first praecoxal endite versus one + one). The new genus exhibits symplesiomorphies in (1), (3), (4), (5), and (6) with
<taxonomicName id="4C49828EFFD8FF9DD59C528FFC552B0E" authorityName="Alvarez" authorityYear="1985" box="[929,1019,1271,1295]" class="Hexanauplia" family="Arietellidae" genus="Pilarella" kingdom="Animalia" order="Calanoida" pageId="4" pageNumber="2485" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD8FF9DD59C528FFC552B0E" box="[929,1019,1271,1295]" italics="true" pageId="4" pageNumber="2485">Pilarella</emphasis>
</taxonomicName>
. On the other hand, the relative shortness of the caudal rami and the antennules of the new genus are reminiscent of
<taxonomicName id="4C49828EFFD8FF9DD7715340FE792B51" authorityName="Cleve" authorityYear="1901" box="[332,471,1336,1360]" class="Hexanauplia" family="Arietellidae" genus="Metacalanus" kingdom="Animalia" order="Calanoida" pageId="4" pageNumber="2485" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD8FF9DD7715340FE792B51" box="[332,471,1336,1360]" italics="true" pageId="4" pageNumber="2485">Metacalanus</emphasis>
</taxonomicName>
. The fifth leg of female
<taxonomicName id="4C49828EFFD8FF9DD5125340FC662B51" authorityName="Ohtsuka &amp; Nishida &amp; Machida" authorityYear="2005" box="[815,968,1336,1360]" class="Hexanauplia" family="Arietellidae" genus="Metacalanalis" kingdom="Animalia" order="Calanoida" pageId="4" pageNumber="2485" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD8FF9DD5125340FC662B51" box="[815,968,1336,1360]" italics="true" pageId="4" pageNumber="2485">Metacalanalis</emphasis>
</taxonomicName>
exhibits a more plesiomorphic state than those of
<taxonomicName id="4C49828EFFD8FF9DD41D5321FD052B70" authorityName="Cleve" authorityYear="1901" box="[544,683,1369,1393]" class="Hexanauplia" family="Arietellidae" genus="Metacalanus" kingdom="Animalia" order="Calanoida" pageId="4" pageNumber="2485" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD8FF9DD41D5321FD052B70" box="[544,683,1369,1393]" italics="true" pageId="4" pageNumber="2485">Metacalanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C49828EFFD8FF9DD4D15321FCE82B70" authorityName="Alvarez" authorityYear="1985" box="[748,838,1369,1393]" class="Hexanauplia" family="Arietellidae" genus="Pilarella" kingdom="Animalia" order="Calanoida" pageId="4" pageNumber="2485" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD8FF9DD4D15321FCE82B70" box="[748,838,1369,1393]" italics="true" pageId="4" pageNumber="2485">Pilarella</emphasis>
</taxonomicName>
, and is rather similar to that of another closely related genus
<taxonomicName id="4C49828EFFD8FF9DD7C55301FC882B90" authority="Scott, 1897" authorityName="Scott" authorityYear="1897" box="[504,806,1401,1425]" class="Hexanauplia" family="Arietellidae" genus="Paramisophria" kingdom="Animalia" order="Calanoida" pageId="4" pageNumber="2485" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD8FF9DD7C55301FD392B90" box="[504,663,1401,1425]" italics="true" pageId="4" pageNumber="2485">Paramisophria</emphasis>
Scott, 1897
</taxonomicName>
, in particular
<taxonomicName id="4C49828EFFD8FF9DD5FE5302FE6C2BB3" authority="Ohtsuka, Fosshagen and Iliffe, 1993" authorityName="Ohtsuka, Fosshagen and Iliffe" authorityYear="1993" class="Hexanauplia" family="Arietellidae" genus="Paramisophria" kingdom="Animalia" order="Calanoida" pageId="4" pageNumber="2485" phylum="Arthropoda" rank="species" species="reducta">
<emphasis id="B93D251FFFD8FF9DD5FE5302FB9F2B90" box="[963,1073,1401,1425]" italics="true" pageId="4" pageNumber="2485">P. reducta</emphasis>
<bibRefCitation id="EFD884FCFFD8FF9DD2055301FE6C2BB3" author="Ohtsuka S &amp; Fosshagen A &amp; Iliffe TM" pageId="4" pageNumber="2485" pagination="57 - 67" refId="ref13107" refString="Ohtsuka S, Fosshagen A, Iliffe TM. 1993. Two new species of Paramisophria (Copepoda, Calanoida, Arietellidae) from anchialine caves on the Canary and Galapagos Islands. Sarsia 78: 57 - 67." type="journal article" year="1993">Ohtsuka, Fosshagen and Iliffe, 1993</bibRefCitation>
</taxonomicName>
except for the absence of endopod.
</paragraph>
<paragraph id="8BF6F90DFFD8FF9CD69853C3FD8F2EC1" blockId="4.[138,1185,850,1685]" lastBlockId="5.[125,1172,137,513]" lastPageId="5" lastPageNumber="2486" pageId="4" pageNumber="2485">
<taxonomicName id="4C49828EFFD8FF9DD69853C3FE902BD2" authorityName="Ohtsuka &amp; Nishida &amp; Machida" authorityYear="2005" box="[165,318,1467,1491]" class="Hexanauplia" family="Arietellidae" genus="Metacalanalis" kingdom="Animalia" order="Calanoida" pageId="4" pageNumber="2485" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD8FF9DD69853C3FE902BD2" box="[165,318,1467,1491]" italics="true" pageId="4" pageNumber="2485">Metacalanalis</emphasis>
</taxonomicName>
is distinguishable from
<taxonomicName id="4C49828EFFD8FF9DD46C53C3FD722BD2" authorityName="Cleve" authorityYear="1901" box="[593,732,1467,1491]" class="Hexanauplia" family="Arietellidae" genus="Metacalanus" kingdom="Animalia" order="Calanoida" pageId="4" pageNumber="2485" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD8FF9DD46C53C3FD722BD2" box="[593,732,1467,1491]" italics="true" pageId="4" pageNumber="2485">Metacalanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C49828EFFD8FF9DD52B53C3FCDE2BD2" authorityName="Alvarez" authorityYear="1985" box="[790,880,1467,1491]" class="Hexanauplia" family="Arietellidae" genus="Pilarella" kingdom="Animalia" order="Calanoida" pageId="4" pageNumber="2485" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD8FF9DD52B53C3FCDE2BD2" box="[790,880,1467,1491]" italics="true" pageId="4" pageNumber="2485">Pilarella</emphasis>
</taxonomicName>
by the combination of the following features: (1) caudal rami about 1.2 times as long as wide; (2) left antennule reaching at most to the posterior margin of pediger 3; (3) antenna showing the most primitive condition in the family
<taxonomicName id="4C49828EFFD8FF9DD4005064FD6B2835" authorityName="Sars" authorityYear="1902" box="[573,709,1564,1588]" class="Hexanauplia" family="Arietellidae" kingdom="Animalia" order="Calanoida" pageId="4" pageNumber="2485" phylum="Arthropoda" rank="family">Arietellidae</taxonomicName>
, with an indistinctly three-segmented endopod and an indistinctly 10-segmented exopod with setal formula 0, 0, 1, 1, 1, 1, 1, 1, 0, 2; (4) praecoxal arthrite of the maxillule with one short and four long spines; (5) coxal epipodite of the maxillule with six setae; (6) basal spine of the maxilla with a short row of minute spinules medially; (7) fifth leg of the female with a unisegmented exopod bearing two lateral and two terminal spines.
</paragraph>
<paragraph id="8BF6F90DFFD9FF9CD6A556B1FBE42F80" blockId="5.[125,1172,137,513]" pageId="5" pageNumber="2486">
The antenna of
<taxonomicName id="4C49828EFFD9FF9CD76556B1FE5F2EE0" authorityName="Ohtsuka &amp; Nishida &amp; Machida" authorityYear="2005" box="[344,497,201,225]" class="Hexanauplia" family="Arietellidae" genus="Metacalanalis" kingdom="Animalia" order="Calanoida" pageId="5" pageNumber="2486" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD9FF9CD76556B1FE5F2EE0" box="[344,497,201,225]" italics="true" pageId="5" pageNumber="2486">Metacalanalis</emphasis>
</taxonomicName>
exhibits a primitive condition, with an indistinctly threesegmented endopod and an indistinctly 10-segmented exopod. The endopod has a suture between the second and third segments clearly visible. Only the most primitive calanoid family
<taxonomicName id="4C49828EFFD9FF9CD6F75751FE372F40" authorityName="Giesbrecht" authorityYear="1893" box="[202,409,297,321]" class="Hexanauplia" family="Pseudocyclopidae" kingdom="Animalia" order="Calanoida" pageId="5" pageNumber="2486" phylum="Arthropoda" rank="family">Pseudocyclopidae</taxonomicName>
(monotypic,
<taxonomicName id="4C49828EFFD9FF9CD40C5751FCF42F40" authority="Brady, 1872" authorityName="Brady" authorityYear="1872" box="[561,858,297,321]" class="Hexanauplia" family="Pseudocyclopidae" genus="Pseudocyclops" kingdom="Animalia" order="Calanoida" pageId="5" pageNumber="2486" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD9FF9CD40C5751FD6B2F40" box="[561,709,297,321]" italics="true" pageId="5" pageNumber="2486">Pseudocyclops</emphasis>
Brady, 1872
</taxonomicName>
) is hitherto known to bear a distinctly three-segmented antennary endopod (
<bibRefCitation id="EFD884FCFFD9FF9CD49E5731FC6A2F60" author="Huys R &amp; Boxshall GA" box="[675,964,329,353]" pageId="5" pageNumber="2486" refId="ref12790" refString="Huys R, Boxshall GA. 1991. Copepod evolution. London: The Ray Society. 468 p." type="book" year="1991">Huys and Boxshall 1991</bibRefCitation>
). A 10-segmented exopod is also the most primitive condition in copepods (
<bibRefCitation id="EFD884FCFFD9FF9CD5255711FB932F80" author="Huys R &amp; Boxshall GA" box="[792,1085,361,385]" pageId="5" pageNumber="2486" refId="ref12790" refString="Huys R, Boxshall GA. 1991. Copepod evolution. London: The Ray Society. 468 p." type="book" year="1991">Huys and Boxshall 1991</bibRefCitation>
).
</paragraph>
<paragraph id="8BF6F90DFFD9FF9CD6A557F2FE702C00" blockId="5.[125,1172,137,513]" pageId="5" pageNumber="2486">
Asymmetry of legs 13 is unique to the new genus, although the asymmetrical antennules are common in the hyperbenthic arietellid genera
<taxonomicName id="4C49828EFFD9FF9CD4D057D2FCD62FC3" authorityName="Cleve" authorityYear="1901" box="[749,888,426,450]" class="Hexanauplia" family="Arietellidae" genus="Metacalanus" kingdom="Animalia" order="Calanoida" pageId="5" pageNumber="2486" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD9FF9CD4D057D2FCD62FC3" box="[749,888,426,450]" italics="true" pageId="5" pageNumber="2486">Metacalanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C49828EFFD9FF9CD5FD57D2FBF12FC3" authorityName="Scott" authorityYear="1897" box="[960,1119,426,450]" class="Hexanauplia" family="Arietellidae" genus="Paramisophria" kingdom="Animalia" order="Calanoida" pageId="5" pageNumber="2486" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD9FF9CD5FD57D2FBF12FC3" box="[960,1119,426,450]" italics="true" pageId="5" pageNumber="2486">Paramisophria</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFD884FCFFD9FF9CD64057B2FECD2FE3" author="Ohtsuka S &amp; Boxshall GA &amp; Roe HSJ" box="[125,355,458,482]" pageId="5" pageNumber="2486" pagination="105 - 172" refId="ref13031" refString="Ohtsuka S, Boxshall GA, Roe HSJ. 1994. Phylogenetic relationships between arietellid genera (Copepoda: Calanoida), with the establishment of three new genera. Bulletin of the Natural History Museum (Zoology Series) 60: 105 - 172." type="journal article" year="1994">Ohtsuka et al. 1994</bibRefCitation>
). These are either an adaptation to its hyperbenthic life or aberrant (see Remarks for the new species).
</paragraph>
<paragraph id="8BF6F90DFFD9FF9CD6405453FCA32C43" blockId="5.[125,781,554,578]" box="[125,781,554,578]" pageId="5" pageNumber="2486">
<emphasis id="B93D251FFFD9FF9CD6405453FD8C2C43" box="[125,546,554,578]" italics="true" pageId="5" pageNumber="2486">
<typeStatus id="54F247AFFFD9FF9CD6405453FF1D2C43" box="[125,179,555,578]" pageId="5" pageNumber="2486">Type</typeStatus>
species.
<taxonomicName id="4C49828EFFD9FF9CD72B5452FD8C2C43" authority="Ohtsuka &amp; Nishida &amp; Machida, 2005" authorityName="Ohtsuka &amp; Nishida &amp; Machida" authorityYear="2005" box="[278,546,554,578]" class="Hexanauplia" family="Arietellidae" genus="Metacalanalis" kingdom="Animalia" order="Calanoida" pageId="5" pageNumber="2486" phylum="Arthropoda" rank="species" species="hakuhoae" status="sp. nov.">Metacalanalis hakuhoae</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A20E9864FFD9FF9CD4165452FDC22C43" box="[555,620,554,578]" pageId="5" pageNumber="2486" rank="species">n. sp.</taxonomicNameLabel>
(monotypic).
</paragraph>
<paragraph id="8BF6F90DFFD9FF9CD6405403FF402C92" blockId="5.[125,238,635,659]" box="[125,238,635,659]" pageId="5" pageNumber="2486">
<emphasis id="B93D251FFFD9FF9CD6405403FF402C92" box="[125,238,635,659]" italics="true" pageId="5" pageNumber="2486">Etymology</emphasis>
</paragraph>
<paragraph id="8BF6F90DFFD9FF9CD64054D4FEF42CE5" blockId="5.[125,1172,684,740]" pageId="5" pageNumber="2486">
The generic name is derived from the closely related genus
<taxonomicName id="4C49828EFFD9FF9CD55E54D4FB3E2CC5" authority="Cleve, 1901" authorityName="Cleve" authorityYear="1901" box="[867,1168,684,708]" class="Hexanauplia" family="Arietellidae" genus="Metacalanus" kingdom="Animalia" order="Calanoida" pageId="5" pageNumber="2486" phylum="Arthropoda" rank="genus">
<emphasis id="B93D251FFFD9FF9CD55E54D4FC402CC5" box="[867,1006,684,708]" italics="true" pageId="5" pageNumber="2486">Metacalanus</emphasis>
Cleve, 1901
</taxonomicName>
. Gender masculine.
</paragraph>
</subSubSection>
</treatment>
</document>