488 lines
50 KiB
XML
488 lines
50 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zsE.95.36788" ID-GBIF-Dataset="bc2389bd-52b8-4411-9ef5-f37d12db4898" ID-Pensoft-Pub="1860-0743-2-507" ID-Pensoft-UUID="F3BFD206564F59C4967CE6267990A52E" ID-ZooBank="10.3897/zsE.95.36788" ModsDocID="1860-0743-2-507" checkinTime="1571900144293" checkinUser="pensoft" docAuthor="Brito, Pamella Silva de, Guimaraes, Erick Cristofore, Carvalho-Costa, Luis Fernando & Ottoni, Felipe Polivanov" docDate="2019" docId="00D38DF4C439548A9820A10076073CBC" docLanguage="en" docName="ZoosystEvol 95(2): 507-516" docOrigin="Zoosystematics and Evolution 95 (2)" docSource="http://dx.doi.org/" docTitle="Aphyocharax brevicaudatus Brito, Guimaraes, Carvalho-Costa & Ottoni, 2019, sp. nov." docType="treatment" docUuid="C5D86CB2-B51B-4B45-AFF7-6E483533B680" docUuidSource="ZooBank" docVersion="5" id="F3BFD206564F59C4967CE6267990A52E" lastPageNumber="507" masterDocId="F3BFD206564F59C4967CE6267990A52E" masterDocTitle="A new species of Aphyocharax Guenther, 1868 (Characiformes, Characidae) from the Maracacume river basin, eastern Amazon" masterLastPageNumber="516" masterPageNumber="507" pageNumber="507" updateTime="1684346104069" updateUser="felipe">
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<mods:titleInfo id="70C34DBCB0278992C33ADE8D92CE6642">
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<mods:title id="0F0E96F347B5F012C2D0E075CD6963D4">A new species of Aphyocharax Guenther, 1868 (Characiformes, Characidae) from the Maracacume river basin, eastern Amazon</mods:title>
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<mods:namePart id="7B18270537C70F511DC522943B136F68">Brito, Pamella Silva de</mods:namePart>
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<mods:namePart id="6CB2ADE7ADBD971A165AF03AB9D0B095">Guimaraes, Erick Cristofore</mods:namePart>
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<mods:namePart id="1A0662BF16D122F611A4DFA0839023D5">Carvalho-Costa, Luis Fernando</mods:namePart>
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<mods:name id="A0B1FB54C8AC9A15A9EAB8E8F7C4532D" type="personal">
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<mods:namePart id="922F5011B3AD33F41CC76B06E6D612A9">Ottoni, Felipe Polivanov</mods:namePart>
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<mods:typeOfResource id="BF527D9A2CCE2CF4E68817C24689AE47">text</mods:typeOfResource>
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<mods:title id="AB71DE75923707AC44D43484D32418BF">Zoosystematics and Evolution</mods:title>
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<mods:date id="E62F74F0B6C397947DD577FF677E7B6F">2019</mods:date>
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<mods:number id="9673A043345A1840E39F9ADDE266620E">95</mods:number>
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<mods:classification id="5E9E657AB2F0A643ECB11C71C8F6438C">journal article</mods:classification>
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<treatment id="00D38DF4C439548A9820A10076073CBC" ID-GBIF-Taxon="159834676" LSID="urn:lsid:zoobank.org:act:C5D86CB2-B51B-4B45-AFF7-6E483533B680" httpUri="http://treatment.plazi.org/id/00D38DF4C439548A9820A10076073CBC" lastPageNumber="507" pageId="0" pageNumber="507">
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<subSubSection id="A57C2284006F9C89461DC8112339910C" pageId="0" pageNumber="507" type="nomenclature">
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<paragraph id="8494002C4C4841B137849BFC390C4C06" pageId="0" pageNumber="507">
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<taxonomicName id="FD74FF10FAC918BF7A324C018AFBAC94" LSID="00D38DF4-C439-548A-9820-A10076073CBC" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax brevicaudatus" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus">Aphyocharax brevicaudatus</taxonomicName>
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<taxonomicNameLabel id="E91FFDE63A84C546268419CB712413BA" pageId="0" pageNumber="507">sp. nov.</taxonomicNameLabel>
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<figureCitation id="9F74653C034A608F6CD73769A3F949A3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Aphyocharax brevicaudatus sp. nov. a. CICCAA 02293, holotype (male), 35.9 mm SL; b. CICCAA 02294, paratype (female), 32.4 mm SL, Brazil: Maranhao state: Maracacume river basin. (Photographed by Erick Guimaraes)." figureDoi=".figure1" httpUri="https://binary.pensoft.net/fig/349754" pageId="0" pageNumber="507" tableDoi=".figure1">Figs 1</figureCitation>
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,
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<figureCitation id="2A746E8FC6FB897ED0D3C3E84D9AB0A3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Caudal-fin of Aphyocharax brevicaudatus, holotype, CICCAA 02293, (male)." figureDoi=".figure2" httpUri="https://binary.pensoft.net/fig/349755" pageId="0" pageNumber="507" tableDoi=".figure2">2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C78DE35C59435F60CEE713AE1D641A13" pageId="0" pageNumber="507" type="holotype">
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<paragraph id="3BB28BA17D661CB9754AE91B6D05757E" pageId="0" pageNumber="507">Holotype.</paragraph>
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<paragraph id="DE6BC11B8DE81ADFB08FCDEE1563FB10" pageId="0" pageNumber="507">
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CICCAA 02293, (male) 35.9 mm SL, Brazil,
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<normalizedToken id="D322C797608C22C1C80B1077EB60AD11" originalValue="Maranhão">Maranhao</normalizedToken>
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state,
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<normalizedToken id="377680C7D06A2E04E9D48F570B02926B" originalValue="Maracaçumé">Maracacume</normalizedToken>
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municipality,
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<normalizedToken id="9160EFEA6C5FFA0571A25D537BFE6AC7" originalValue="Maracaçumé">Maracacume</normalizedToken>
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River,
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<geoCoordinate id="79A9AB3FC3205C30B4EDBB6968A5B085" degrees="2" direction="south" minutes="3" orientation="latitude" precision="15" seconds="14" value="-2.0538888">2°3'14"S</geoCoordinate>
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,
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<geoCoordinate id="F0083FD158E0BC1DEA8E8997B455CFC5" degrees="45" direction="west" minutes="57" orientation="longitude" precision="15" seconds="16" value="-45.954445">45°57'16"W</geoCoordinate>
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; 29 Jun 2018, E.C.
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<normalizedToken id="5A1CF120BBC07D7700575E82BBEB563D" originalValue="Guimarães">Guimaraes</normalizedToken>
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and P.S. Brito.
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</paragraph>
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</subSubSection>
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<subSubSection id="EF4BB1E7B2D8B90C290EF0DB7902A2D6" pageId="0" pageNumber="507" type="paratypes">
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<paragraph id="78062F9D362FA1A394A62BA027DF19EC" pageId="0" pageNumber="507">Paratypes.</paragraph>
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<paragraph id="51DA5B2C485A058EB4187676228045B6" pageId="0" pageNumber="507">
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All from Brazil,
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<normalizedToken id="BACB731FC7E7251E4264685300CCA8E8" originalValue="Maranhão">Maranhao</normalizedToken>
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state: CICCAA 02294, 1 (female), 32.4 mm SL, CICCAA 02295, 35 (males), 20.9-31.7 mm SL,CICCAA 02296, 94 (females), 21-32.1 mm SL, CICCAA 02297, 30 (females) C&S, 22.2-30.8 mm SL, CICCAA 02312, 2 (males) C&S, 28.3-32.1 mm SL, UFRJ 11746, 10 (female), 24.2-30.2 mm SL; all collected with holotype.
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</paragraph>
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</subSubSection>
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<subSubSection id="7200EB4DEEE6024BEB83889F21589460" pageId="0" pageNumber="507" type="diagnosis">
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<paragraph id="0E818BDF46F85F782E7239C0AFC64BF6" pageId="0" pageNumber="507">Diagnosis (PAA).</paragraph>
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<paragraph id="1657CB081B5B953ED4FCD8A59581DADC" pageId="0" pageNumber="507">
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<taxonomicName id="A415FE05C561A2E4AAD958DC1569215F" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax brevicaudatus" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus">
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<emphasis id="29DF9D1F056BEEDA79067FD5267887B7" italics="true" pageId="0" pageNumber="507">Aphyocharax brevicaudatus</emphasis>
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</taxonomicName>
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sp. nov. differs from all its congeners by possessing the upper lobe of the caudal fin longer than the lower lobe in mature males (vs upper and lower lobes similar in length, see
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<figureCitation id="B6799F49E1B2B2707F9937388A363E1D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Aphyocharax brevicaudatus sp. nov. a. CICCAA 02293, holotype (male), 35.9 mm SL; b. CICCAA 02294, paratype (female), 32.4 mm SL, Brazil: Maranhao state: Maracacume river basin. (Photographed by Erick Guimaraes)." figureDoi=".figure1" httpUri="https://binary.pensoft.net/fig/349754" pageId="0" pageNumber="507" tableDoi=".figure1">Figs 1</figureCitation>
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,
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<figureCitation id="014EBF749AD0D8A0BC7374EBF77CBF78" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Caudal-fin of Aphyocharax brevicaudatus, holotype, CICCAA 02293, (male)." figureDoi=".figure2" httpUri="https://binary.pensoft.net/fig/349755" pageId="0" pageNumber="507" tableDoi=".figure2">2</figureCitation>
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;
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<bibRefCitation id="D8D6B9D471B7B776CADBFF8A545ED5C8" DOI="https://doi.org/10.1016/j.ympev.2012.04.007" author="Tagliacollo, VA" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="507" pagination="297 - 307" refId="B36" refString="Tagliacollo, VA, Souza-Lima, R, Benine, RC, Oliveira, C, 2012. Molecular phylogeny of Aphyocharacinae (Characiformes, Characidae) with morphological diagnoses for the subfamily and recognized genera. Molecular Phylogenetics and Evolution 64: 297 - 307" title="Molecular phylogeny of Aphyocharacinae (Characiformes, Characidae) with morphological diagnoses for the subfamily and recognized genera." url="https://doi.org/10.1016/j.ympev.2012.04.007" volume="64" year="2012">Tagliacollo et al. 2012</bibRefCitation>
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: fig.4). Additionally, the new species is distinguished from
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<taxonomicName id="9B575EF194A51A88CB8CFE526D73B5BF" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax avary" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="avary">
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<emphasis id="45163771AFF07488648834412A3DD389" italics="true" pageId="0" pageNumber="507">Aphyocharax avary</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="01E4ADFBFC3DB25FAFF439A6D1451396" lsidName="A. pusillus" pageId="0" pageNumber="507" rank="species" species="pusillus">
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<emphasis id="ED1486E6718DD5BA1AA27A2E17CDCE2F" italics="true" pageId="0" pageNumber="507">A. pusillus</emphasis>
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</taxonomicName>
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by having hyaline middle caudal-fin rays (vs black or dark brown middle caudal-fin rays,
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<bibRefCitation id="A1C560BEFBA74043E056CB900403F111" DOI="https://doi.org/10.3897/zse.94.28201" author="Brito, PS" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="507" pagination="393 - 399" refId="B3" refString="Brito, PS, Guimaraes, EC, Katz, AM, Piorski, NM, Ottoni, FP, 2018. Taxonomic status of Aphyocharaxavary Fowler 1913, Aphyocharaxpusillus Guenther 1868 and Chirodonalburnus Guenther 1869 (Characiformes: Characidae). Zoosystematics and Evolution 94: 393 - 399" title="Taxonomic status of Aphyocharaxavary Fowler 1913, Aphyocharaxpusillus Guenther 1868 and Chirodonalburnus Guenther 1869 (Characiformes: Characidae)." url="https://doi.org/10.3897/zse.94.28201" volume="94" year="2018">Brito et al. 2018</bibRefCitation>
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: fig. 3); from
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<taxonomicName id="A4A73EF5ADF7685C5EF9C26DA1BCAD20" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax colifax" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="colifax">
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<emphasis id="414C15C4CB8973E5EAB6590297032ACA" italics="true" pageId="0" pageNumber="507">Aphyocharax colifax</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="FEDF3D8472E081C80B2DE6B32F3F3D0A" lsidName="A. yekwanae" pageId="0" pageNumber="507" rank="species" species="yekwanae">
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<emphasis id="84C6735173C766BF870CB81500B1AA37" italics="true" pageId="0" pageNumber="507">A. yekwanae</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="6F4F38CDB79209B4904688BE41118EB9" lsidName="A. rathbuni" pageId="0" pageNumber="507" rank="species" species="rathbuni">
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<emphasis id="1E633C6939EB235BD9007EC5E48DB527" italics="true" pageId="0" pageNumber="507">A. rathbuni</emphasis>
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</taxonomicName>
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by having caudal-fin light red colouration never surpassing the vertical line of the adipose-fin (vs red colouration extending to the lateral midline of body,
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<bibRefCitation id="A194C8527D323E847699741383513605" author="Willink, PW" journalOrPublisher="Ichthyological Exploration of Freshwaters" pageId="0" pageNumber="507" pagination="1 - 8" refId="B41" refString="Willink, PW, Chernoff, B, Machado-Allison, A, Provenzano, F, Petry, P, 2003. Aphyocharaxyekwanae, a new species of bloodfin tetra (Teleostei: Characiformes: Characidae) from the Guyana Shield of Venezuela. Ichthyological Exploration of Freshwaters 14: 1 - 8" title="Aphyocharaxyekwanae, a new species of bloodfin tetra (Teleostei: Characiformes: Characidae) from the Guyana Shield of Venezuela." volume="14" year="2003">Willink et al. 2003</bibRefCitation>
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: fig. 1); from
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<taxonomicName id="01DEC1433CB3A6C64FCD3AA0AF81FA16" lsidName="A. gracilis" pageId="0" pageNumber="507" rank="species" species="gracilis">
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<emphasis id="465A0212C14DD842C591B46860C9979D" italics="true" pageId="0" pageNumber="507">A. gracilis</emphasis>
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</taxonomicName>
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by having a larger body depth at dorsal-fin origin (body depth), 24.5-29.2% SL (vs 20.1-20.6% SL); and from
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<taxonomicName id="D3C32A18B603290B7D52598B21229352" lsidName="A. pusillus" pageId="0" pageNumber="507" rank="species" species="pusillus">
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<emphasis id="8C3A66AF1265B44BDD8F2E6539473E09" italics="true" pageId="0" pageNumber="507">A. pusillus</emphasis>
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</taxonomicName>
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by having teeth along 2/3 of the maxillary extension (vs along proximal half of the bone,
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<bibRefCitation id="FCCAE11C941C01371468F3694430D22A" DOI="https://doi.org/10.3897/zse.94.28201" author="Brito, PS" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="507" pagination="393 - 399" refId="B3" refString="Brito, PS, Guimaraes, EC, Katz, AM, Piorski, NM, Ottoni, FP, 2018. Taxonomic status of Aphyocharaxavary Fowler 1913, Aphyocharaxpusillus Guenther 1868 and Chirodonalburnus Guenther 1869 (Characiformes: Characidae). Zoosystematics and Evolution 94: 393 - 399" title="Taxonomic status of Aphyocharaxavary Fowler 1913, Aphyocharaxpusillus Guenther 1868 and Chirodonalburnus Guenther 1869 (Characiformes: Characidae)." url="https://doi.org/10.3897/zse.94.28201" volume="94" year="2018">Brito et al. 2018</bibRefCitation>
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: fig. 4).
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</paragraph>
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<caption id="2B53E4BCBFBFB79E48E326EBB78BB9B4" doi=".figure1" httpUri="https://binary.pensoft.net/fig/349754" pageId="0" pageNumber="507" start="Figure 1" startId="F1">
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<paragraph id="C9D61CAAC26F9A843E1002AACE86416E" pageId="0" pageNumber="507">
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<emphasis id="D501900C8F618FAF6BC88CCC242633ED" bold="true" pageId="0" pageNumber="507">Figure 1.</emphasis>
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<taxonomicName id="C8085F651B6CF0CF8EDC227C694A8871" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax brevicaudatus" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus">
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<emphasis id="B04CF2F0B57EBDCDDB4EF04ABC65E7FA" italics="true" pageId="0" pageNumber="507">Aphyocharax brevicaudatus</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis id="F136A7E3FF37540400EB731E55821230" bold="true" pageId="0" pageNumber="507">a.</emphasis>
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CICCAA 02293, holotype (male), 35.9 mm SL;
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<emphasis id="E3FB18FF2652F85536A346F603672405" bold="true" pageId="0" pageNumber="507">b.</emphasis>
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CICCAA 02294, paratype (female), 32.4 mm SL, Brazil:
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<normalizedToken id="9BF9388C959E4586A69B962782ED6FC3" originalValue="Maranhão">Maranhao</normalizedToken>
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state:
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<normalizedToken id="9974E0BEC4CFDD1A70CD0F84D605436E" originalValue="Maracaçumé">Maracacume</normalizedToken>
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river basin. (Photographed by Erick
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<normalizedToken id="89A7F7B94C9DF304C884FEA1F1C41C3E" originalValue="Guimarães">Guimaraes</normalizedToken>
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).
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</paragraph>
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</caption>
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<caption id="A400B703D2C5CDA673EBFD5B51C82890" doi=".figure2" httpUri="https://binary.pensoft.net/fig/349755" pageId="0" pageNumber="507" start="Figure 2" startId="F2">
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<paragraph id="8F85D203CD449C03833775A6A831C7C5" pageId="0" pageNumber="507">
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<emphasis id="BEFCC1CE9C72B9D756CAD5613852FE30" bold="true" pageId="0" pageNumber="507">Figure 2.</emphasis>
|
|
Caudal-fin of
|
|
<taxonomicName id="F6CA3986DA4E526CB70E432AEC2B6B6A" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax brevicaudatus" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus">
|
|
<emphasis id="B706BAB8FE72FD61EF16ABDE60EDEFA8" italics="true" pageId="0" pageNumber="507">Aphyocharax brevicaudatus</emphasis>
|
|
</taxonomicName>
|
|
, holotype, CICCAA 02293, (male).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="7D46339F9431662F2A552035AEC1CFDC" pageId="0" pageNumber="507" type="description">
|
|
<paragraph id="61CC8AF3230E098F20D21CA53F5FDD53" pageId="0" pageNumber="507">Description.</paragraph>
|
|
<paragraph id="979552420CC52862BCF7E2120909BF7A" pageId="0" pageNumber="507">
|
|
Morphometric data is presented in Table
|
|
<tableCitation id="539B61AB92A38437335562E7960EA53F" captionStartId="T2" httpUri="http://table.plazi.org/id/3EFCC811FE6F0397D03F2972FBFDBF6D" pageId="0" pageNumber="507" tableUuid="3EFCC811FE6F0397D03F2972FBFDBF6D">2</tableCitation>
|
|
. Body shape is generally fusiform, slightly elongate, greatest body depth slightly anterior to dorsal-fin base; dorsal body profile straight or slightly convex from snout to vertical through anterior nostrils; straight or slightly convex from posterior nostrils to tip of supraoccipital bone; straight or slightly convex from this point to dorsal-fin origin; slightly convex along dorsal-fin base; postdorsal profile straight from base of last dorsal-fin ray to adipose-fin origin; slightly concave from adipose-fin to end of caudal peduncle; ventral profile convex from snout to pelvic-fin insertion; straight or slightly convex from this point to anal-fin origin; straight along anal-fin base; long snout, with its length larger than orbital diameter; five infraorbital bones; fourth infraorbital absent and sixth infraorbital reduced; posterior border of maxilla rounded, extending vertically through anterior margin of orbit, not reaching third infraorbital.
|
|
</paragraph>
|
|
<caption id="3EFCC811FE6F0397D03F2972FBFDBF6D" ID-Table-UUID="3EFCC811FE6F0397D03F2972FBFDBF6D" httpUri="http://table.plazi.org/id/3EFCC811FE6F0397D03F2972FBFDBF6D" pageId="0" pageNumber="507" start="Table 2" startId="T2">
|
|
<paragraph id="81A0ECCF6127BDFFF59344931E521C5C" pageId="0" pageNumber="507">
|
|
<emphasis id="36909077EC16B231A71F0C0320BEEFB0" bold="true" pageId="0" pageNumber="507">Table 2.</emphasis>
|
|
Morphometric data (
|
|
<emphasis id="B169C40EA406262A1234934B52D68A08" italics="true" pageId="0" pageNumber="507">N</emphasis>
|
|
= 141) of the holotype and paratypes of
|
|
<taxonomicName id="65EBCDB3A635376DCB43C8A9FC6E6406" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax brevicaudatus" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus">
|
|
<emphasis id="56D10EB63130B12478C9A28DE6570B4C" italics="true" pageId="0" pageNumber="507">Aphyocharax brevicaudatus</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. from the
|
|
<normalizedToken id="F04CA18725A62268CA73CDAF03B01DB9" originalValue="Maracaçumé">Maracacume</normalizedToken>
|
|
river basin. SD: Standard deviation.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="5D768F5380907A1B93581D1228879D6D" pageId="0" pageNumber="507">
|
|
<table id="949628357E538E11D5ECEBC0BE564E39" pageId="0" pageNumber="507">
|
|
<tr id="B84EC7143D5E648F3B84B063756648FD" pageId="0" pageNumber="507">
|
|
<th id="1714897CEF053F9B15B839D33C528B4C" colspan="1" isEmpty="true" pageId="0" pageNumber="507" rowspan="1">-</th>
|
|
<th id="59AD5104114B30E9774F6A2B8A87D368" colspan="1" pageId="0" pageNumber="507" rowspan="1">Holotype (Male)</th>
|
|
<th id="D179455BF7E76CD253D23C648B53FFFB" colspan="1" pageId="0" pageNumber="507" rowspan="1">Paratypes (Male) N = 35</th>
|
|
<th id="0A99022D6CA9599FFA9AFD2A3E130C85" colspan="1" pageId="0" pageNumber="507" rowspan="1">Mean</th>
|
|
<th id="C6275D1D47E4E3A4181EC3A2DE9FB378" colspan="1" pageId="0" pageNumber="507" rowspan="1">SD</th>
|
|
<th id="EBE0D51741826910BB74CE0221FD5DA4" colspan="1" pageId="0" pageNumber="507" rowspan="1">Paratypes (Female) N = 105</th>
|
|
<th id="7B96AE70451EC1B662F13B909D339663" colspan="1" pageId="0" pageNumber="507" rowspan="1">Mean</th>
|
|
<th id="85EBED9CC4C75F62C6166FEDB180F833" colspan="1" pageId="0" pageNumber="507" rowspan="1">SD</th>
|
|
</tr>
|
|
<tr id="DB26187A9F36C395EE2A45D1C68089B0" pageId="0" pageNumber="507">
|
|
<td id="6E25E47C2F9C8FAE323C199F6FDCCF28" colspan="1" pageId="0" pageNumber="507" rowspan="1">Standard length (mm)</td>
|
|
<td id="60BCBE21066244E0D6619E5D599A692F" colspan="1" pageId="0" pageNumber="507" rowspan="1">35.9</td>
|
|
<td id="4E0C64979B6D6173617062157B31F4ED" colspan="1" pageId="0" pageNumber="507" rowspan="1">20.9-35.9</td>
|
|
<td id="AE93AA7403CBE93C378FE09C1EF20717" colspan="1" pageId="0" pageNumber="507" rowspan="1">26.6</td>
|
|
<td id="2106CACF55DA8F47AB4F0641C06586C6" colspan="1" pageId="0" pageNumber="507" rowspan="1">-</td>
|
|
<td id="FA43A842F045A70D822D0DECA1AF6197" colspan="1" pageId="0" pageNumber="507" rowspan="1">21.0-32.4</td>
|
|
<td id="BDDAB9874979BBD2DCBBB8A13D258ECC" colspan="1" pageId="0" pageNumber="507" rowspan="1">28.0</td>
|
|
<td id="07FE5DC7465B2CEA8DBEF68265939B67" colspan="1" pageId="0" pageNumber="507" rowspan="1">-</td>
|
|
</tr>
|
|
<tr id="00EAD0C7586D760A2F053BB1FCF2855B" pageId="0" pageNumber="507">
|
|
<td id="A8FBFD87CCABDAE4ADD5E5B6A6788061" colspan="8" pageId="0" pageNumber="507" rowspan="1">
|
|
<emphasis id="2C16FC0BC5CA05B59C0E484BBE4E5B0E" bold="true" pageId="0" pageNumber="507">Percentages of standard length</emphasis>
|
|
</td>
|
|
</tr>
|
|
<tr id="F2181D4D559CC8824E8C32AE2E5E9F20" pageId="0" pageNumber="507">
|
|
<td id="515D06AD0AD5992AB45100E16CEF343E" colspan="1" pageId="0" pageNumber="507" rowspan="1">Depth at dorsal-fin origin (body depth)</td>
|
|
<td id="9FB384451163908ADD71966F0F124096" colspan="1" pageId="0" pageNumber="507" rowspan="1">25.4</td>
|
|
<td id="DD2B67FBB517AD5D5BEFF2A4DE123F03" colspan="1" pageId="0" pageNumber="507" rowspan="1">24.5-28.7</td>
|
|
<td id="5BB30145AF5EDB8D03E368534C64C6BC" colspan="1" pageId="0" pageNumber="507" rowspan="1">25.9</td>
|
|
<td id="955D77E82A3BC92913A6E58C7BD74CB7" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.0</td>
|
|
<td id="DAC69A31CBC3E23177E521E4C3F03B30" colspan="1" pageId="0" pageNumber="507" rowspan="1">25.6-29.1</td>
|
|
<td id="A1FB7D6A6ECEE4E3B2BEFC3DCCBD1804" colspan="1" pageId="0" pageNumber="507" rowspan="1">26.3</td>
|
|
<td id="05D3A58C66A3604BE866CB15DF8516E3" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.8</td>
|
|
</tr>
|
|
<tr id="74A247774CC7172EB6E9E4DD0429606D" pageId="0" pageNumber="507">
|
|
<td id="A4A49F246140B1ACC9FF2FE5DF128BCC" colspan="1" pageId="0" pageNumber="507" rowspan="1">Snout to dorsal-fin origin</td>
|
|
<td id="A3A641A676EB60118F1B1BC409024464" colspan="1" pageId="0" pageNumber="507" rowspan="1">53.1</td>
|
|
<td id="3EB8C8E83EBA11C5091E0064C72FD998" colspan="1" pageId="0" pageNumber="507" rowspan="1">51.9-55.6</td>
|
|
<td id="0DE7357885FEAB57865381109F6782DA" colspan="1" pageId="0" pageNumber="507" rowspan="1">52.6</td>
|
|
<td id="275BE8DEA50A3700E6AC3C1E413C7662" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.1</td>
|
|
<td id="DC57293DECD2632D3356C227A2708D5E" colspan="1" pageId="0" pageNumber="507" rowspan="1">51.8-54.5</td>
|
|
<td id="E9E3B5435B5AC2E3842289E3FA101470" colspan="1" pageId="0" pageNumber="507" rowspan="1">52.1</td>
|
|
<td id="36370DA8401B2C099261580F7B0D93BA" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.7</td>
|
|
</tr>
|
|
<tr id="8B25BFC6C2A53FF7E41A5CFE4C39CDF0" pageId="0" pageNumber="507">
|
|
<td id="BD0A2AFC6F29A2CF2DD21BB740EEF20F" colspan="1" pageId="0" pageNumber="507" rowspan="1">Snout to pectoral-fin origin</td>
|
|
<td id="5523F0F105BAA676644508C742D7FFBF" colspan="1" pageId="0" pageNumber="507" rowspan="1">23.2</td>
|
|
<td id="25367EE6CA673642FABBB11B328CAA82" colspan="1" pageId="0" pageNumber="507" rowspan="1">23.0-27.7</td>
|
|
<td id="21EAE4AD7D83A0B2F5E7239A420A9C2F" colspan="1" pageId="0" pageNumber="507" rowspan="1">23.9</td>
|
|
<td id="805C7BF3F4B574B2CE764A4C7136D360" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.9</td>
|
|
<td id="5AA2B8F006AD085199BADA3AE1EBFED0" colspan="1" pageId="0" pageNumber="507" rowspan="1">22.6-25.2</td>
|
|
<td id="0E68DBB31C9666EF922083682E4422CE" colspan="1" pageId="0" pageNumber="507" rowspan="1">23.5</td>
|
|
<td id="1FD41F8F459E553AE44A148CC9288A44" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.6</td>
|
|
</tr>
|
|
<tr id="AA4E74761A30D18599125862B64C00FB" pageId="0" pageNumber="507">
|
|
<td id="4FE8C42E15BCF7578FC48CBF94259911" colspan="1" pageId="0" pageNumber="507" rowspan="1">Snout to pelvic-fin origin</td>
|
|
<td id="278043B74F1F7538BF4B75C3077DC9D5" colspan="1" pageId="0" pageNumber="507" rowspan="1">46.3</td>
|
|
<td id="60D05759A111D412603939D515BB8A76" colspan="1" pageId="0" pageNumber="507" rowspan="1">45.1-49.4</td>
|
|
<td id="35141E206523A7C034CB653EDFFC2740" colspan="1" pageId="0" pageNumber="507" rowspan="1">45.2</td>
|
|
<td id="41ECCFC4DD3B6092DF5AAE8F7E89F44A" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.6</td>
|
|
<td id="9BA5B6404897DDD61AE557337163ACD9" colspan="1" pageId="0" pageNumber="507" rowspan="1">44.2-47.1</td>
|
|
<td id="BB1FB2A56AB511CAB49A4DC137D542DA" colspan="1" pageId="0" pageNumber="507" rowspan="1">44.6</td>
|
|
<td id="EADEEAD5FBC5FF7358858A10264F1E31" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.9</td>
|
|
</tr>
|
|
<tr id="FE1A743D136D1B3D9ED675037F044FCB" pageId="0" pageNumber="507">
|
|
<td id="EBC28DC720099F654A68909926FE089B" colspan="1" pageId="0" pageNumber="507" rowspan="1">Snout to anal-fin origin</td>
|
|
<td id="5EDA417F65B8B5905466FECA0EA9B907" colspan="1" pageId="0" pageNumber="507" rowspan="1">67.4</td>
|
|
<td id="6CA114C151CE6F18344CF8148A0A996B" colspan="1" pageId="0" pageNumber="507" rowspan="1">63.9-68.6</td>
|
|
<td id="685CC3D9289365DDCE9FBCEB622200A1" colspan="1" pageId="0" pageNumber="507" rowspan="1">64.4</td>
|
|
<td id="3BC9E46EB45587B3980D0F565E2ED9CC" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.9</td>
|
|
<td id="C281B028A639FEC5C5D7CC7560CBBECB" colspan="1" pageId="0" pageNumber="507" rowspan="1">64.0-68.5</td>
|
|
<td id="65E1275E4A586168D7BFCFB7FA26B910" colspan="1" pageId="0" pageNumber="507" rowspan="1">64.2</td>
|
|
<td id="9E2A31CA0A0E545062CEE3FB7DC86EC6" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.7</td>
|
|
</tr>
|
|
<tr id="0D016187F8829217C7B428EA3FA28E81" pageId="0" pageNumber="507">
|
|
<td id="9C8382D74EDFDB896A94FD3A99629FDB" colspan="1" pageId="0" pageNumber="507" rowspan="1">Caudal peduncle depth</td>
|
|
<td id="6D7B31754EB4AC2A157B30B5BE49FBC0" colspan="1" pageId="0" pageNumber="507" rowspan="1">10.8</td>
|
|
<td id="668C427B7A74AEE069DEFB2630F29354" colspan="1" pageId="0" pageNumber="507" rowspan="1">10.1-12.5</td>
|
|
<td id="E552F0127521D6BDA8CFF794E3D558A0" colspan="1" pageId="0" pageNumber="507" rowspan="1">11.3</td>
|
|
<td id="69C75A792377B052A4D42D60AA6A9082" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.5</td>
|
|
<td id="1371201EDA44F8056381033AA6401188" colspan="1" pageId="0" pageNumber="507" rowspan="1">10.9-12.2</td>
|
|
<td id="6F2A84CF7EAFC253D050D203C86877B7" colspan="1" pageId="0" pageNumber="507" rowspan="1">11.3</td>
|
|
<td id="638ADE417E9F8AFDE54CAFFA5506A158" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.3</td>
|
|
</tr>
|
|
<tr id="0137C278708D507DD98FF6E593E85D58" pageId="0" pageNumber="507">
|
|
<td id="9BB1C2C5FB59795538A6FA0C7E7B0FD4" colspan="1" pageId="0" pageNumber="507" rowspan="1">Caudal peduncle length</td>
|
|
<td id="D65454636011A101415B5B2D487C35DF" colspan="1" pageId="0" pageNumber="507" rowspan="1">13.2</td>
|
|
<td id="40A9E5F8A0A692AA8D7F6FC516341D90" colspan="1" pageId="0" pageNumber="507" rowspan="1">12.2-17.2</td>
|
|
<td id="B44B0778C801A7E0CFA4325B1028DBF1" colspan="1" pageId="0" pageNumber="507" rowspan="1">14.0</td>
|
|
<td id="E5E2F829490A79FA66F74DA449868661" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.2</td>
|
|
<td id="820BE4D2C37354E2A492E9A89BC1F970" colspan="1" pageId="0" pageNumber="507" rowspan="1">12.2-14.9</td>
|
|
<td id="BE2E9E51BE6F2265E1090549AF160357" colspan="1" pageId="0" pageNumber="507" rowspan="1">13.1</td>
|
|
<td id="7C91971912A99559EC0550CFE4F39C90" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.7</td>
|
|
</tr>
|
|
<tr id="022B845FCE0D342AF327B054BE82F530" pageId="0" pageNumber="507">
|
|
<td id="C40D15F7554B82A1B890C73EA6C881F8" colspan="1" pageId="0" pageNumber="507" rowspan="1">Pectoral-fin length</td>
|
|
<td id="015DD6206826F2F0D4C93020BD17E05B" colspan="1" pageId="0" pageNumber="507" rowspan="1">20.4</td>
|
|
<td id="B63868BF76E77665CAE04EA559CF5CDA" colspan="1" pageId="0" pageNumber="507" rowspan="1">17.9-22.5</td>
|
|
<td id="F2FF0FD668967E47E8F7FB698CA12376" colspan="1" pageId="0" pageNumber="507" rowspan="1">19.7</td>
|
|
<td id="CADD912C8ACCFDAF4AA0A3E8C0D5A61E" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.3</td>
|
|
<td id="DB8A576EE53A857DD5BEDAD2094C9156" colspan="1" pageId="0" pageNumber="507" rowspan="1">18.6-21.1</td>
|
|
<td id="E79AEF4D943FE3AFB5C8BF7E5331BB84" colspan="1" pageId="0" pageNumber="507" rowspan="1">19.3</td>
|
|
<td id="1BE7CF571F29C7AAACA1154EE39DD790" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.6</td>
|
|
</tr>
|
|
<tr id="5FCFAF8EF7F8CD558B685E2911A157D4" pageId="0" pageNumber="507">
|
|
<td id="7F63940273BEF90548F7CD732375DA85" colspan="1" pageId="0" pageNumber="507" rowspan="1">Pelvic-fin length</td>
|
|
<td id="15C713EB93FA1EA276E2731B92BFAD0A" colspan="1" pageId="0" pageNumber="507" rowspan="1">15.9</td>
|
|
<td id="DD4A5506EA7DE67D494290AAC1366FF4" colspan="1" pageId="0" pageNumber="507" rowspan="1">14.6-20.6</td>
|
|
<td id="7210A940BAC022C50096A476540CA98B" colspan="1" pageId="0" pageNumber="507" rowspan="1">15.6</td>
|
|
<td id="A33DBAE13CC1F0954879594901EF2114" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.5</td>
|
|
<td id="260CB9C68EB9AFFB090C333A9C85315D" colspan="1" pageId="0" pageNumber="507" rowspan="1">14.0-17.1</td>
|
|
<td id="DD3A0A67744E36863C44848883A7DD25" colspan="1" pageId="0" pageNumber="507" rowspan="1">15.3</td>
|
|
<td id="EEB18DAB7B1CC02E9384EF927DDA5462" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.7</td>
|
|
</tr>
|
|
<tr id="9059B6CD7DA50AA21D2D7D6826B4C596" pageId="0" pageNumber="507">
|
|
<td id="CC804AD4C710B75DA5081A7FC12FCA99" colspan="1" pageId="0" pageNumber="507" rowspan="1">Dorsal-fin base length</td>
|
|
<td id="7D3CD35E4B3D88FD3DC61D7241277CA0" colspan="1" pageId="0" pageNumber="507" rowspan="1">11.6</td>
|
|
<td id="65B184C1802450C93CE755F5CC713F5D" colspan="1" pageId="0" pageNumber="507" rowspan="1">9.5-13.4</td>
|
|
<td id="9F3295CC667F695BF5C5045CC9519F72" colspan="1" pageId="0" pageNumber="507" rowspan="1">11.3</td>
|
|
<td id="878ECD9539226BE0ABA8044ECD4634B3" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.5</td>
|
|
<td id="FD115A300BA6655E860B6798645A54D5" colspan="1" pageId="0" pageNumber="507" rowspan="1">10.8-13.0</td>
|
|
<td id="FC08D9405A2D31F2CD76B82CB68B8790" colspan="1" pageId="0" pageNumber="507" rowspan="1">11.8</td>
|
|
<td id="A0C4D9A52D6ACBFD5AAB1AC385E5195A" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.5</td>
|
|
</tr>
|
|
<tr id="287C03D3FF3C4DA6ECF595B5DD67DE2D" pageId="0" pageNumber="507">
|
|
<td id="CA848489FD7E1B390FDCD1F30DD43211" colspan="1" pageId="0" pageNumber="507" rowspan="1">Dorsal-fin height</td>
|
|
<td id="2AAE4BF1A1E02D355396C63B42F78957" colspan="1" pageId="0" pageNumber="507" rowspan="1">23.1</td>
|
|
<td id="B4D83166D6E7B91FDEE7DEF411156EB8" colspan="1" pageId="0" pageNumber="507" rowspan="1">21.2-24.8</td>
|
|
<td id="68F812680B69886ACC595F47D781A1C9" colspan="1" pageId="0" pageNumber="507" rowspan="1">22.4</td>
|
|
<td id="7E309179A727250557FCEA61BA8C2478" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.5</td>
|
|
<td id="CAAA2FED4C54A3629E045E00444F6A85" colspan="1" pageId="0" pageNumber="507" rowspan="1">20.8-24.0</td>
|
|
<td id="0D4E1BE433343A0D8E0B69A171098A23" colspan="1" pageId="0" pageNumber="507" rowspan="1">22.3</td>
|
|
<td id="0DA131FDB6140715C1CC980209094138" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.7</td>
|
|
</tr>
|
|
<tr id="054C607AD5505BF21987D2CDAF5ADEF2" pageId="0" pageNumber="507">
|
|
<td id="EE15CFDE142BA89F5B7BC292F9B4A598" colspan="1" pageId="0" pageNumber="507" rowspan="1">Anal-fin base length</td>
|
|
<td id="1E66F33D90A7A711971ECF753B6804D0" colspan="1" pageId="0" pageNumber="507" rowspan="1">18.9</td>
|
|
<td id="1592E8CCA19CB84EB7FCF1326A3920CF" colspan="1" pageId="0" pageNumber="507" rowspan="1">16.7-21.1</td>
|
|
<td id="7873617097CC7CBB8767EF4EA970771E" colspan="1" pageId="0" pageNumber="507" rowspan="1">18.1</td>
|
|
<td id="062C9C4177AA917757662CA6948815E7" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.4</td>
|
|
<td id="BC0BE15BA30604CBB4849BBB584CC705" colspan="1" pageId="0" pageNumber="507" rowspan="1">16.8-20.7</td>
|
|
<td id="76E74D6572F1840BC777034F0649C438" colspan="1" pageId="0" pageNumber="507" rowspan="1">18.3</td>
|
|
<td id="1F70ADAAB36D7D7982B64D4953D30644" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.0</td>
|
|
</tr>
|
|
<tr id="D652A2A7CBD070F2538133F2232604DA" pageId="0" pageNumber="507">
|
|
<td id="A41C3D741FE47280D0E843FCCA45C989" colspan="1" pageId="0" pageNumber="507" rowspan="1">Eye to dorsal-fin origin</td>
|
|
<td id="E850D26DDF2B7227CED3D3E208A68C22" colspan="1" pageId="0" pageNumber="507" rowspan="1">42.6</td>
|
|
<td id="220AC9E708DE34734E2B439401D7AB48" colspan="1" pageId="0" pageNumber="507" rowspan="1">40.6-54.6</td>
|
|
<td id="B421CDF8E47807A0FE8A918B609DB817" colspan="1" pageId="0" pageNumber="507" rowspan="1">42.1</td>
|
|
<td id="F5B6D2908FFA296D692814A4FB497818" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.6</td>
|
|
<td id="37931B5F8E2A81F067DFF466EF3134F6" colspan="1" pageId="0" pageNumber="507" rowspan="1">41.4-52.4</td>
|
|
<td id="65C991557079CFF13E84CB992E966D9A" colspan="1" pageId="0" pageNumber="507" rowspan="1">41.8</td>
|
|
<td id="946F6FB6D16AA3B287E6BB4AE96D00B5" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.9</td>
|
|
</tr>
|
|
<tr id="157F8F06D57F8D1B2B3B16A10EB634D6" pageId="0" pageNumber="507">
|
|
<td id="AD195A535228B2BAB9C4BA2A61749688" colspan="1" pageId="0" pageNumber="507" rowspan="1">Dorsal-fin origin to caudal-fin base</td>
|
|
<td id="18C2C9988B2B9F0FE7EE1375416F812B" colspan="1" pageId="0" pageNumber="507" rowspan="1">47.6</td>
|
|
<td id="655D923D307A7206504FD82D553B4B71" colspan="1" pageId="0" pageNumber="507" rowspan="1">46.5-49.5</td>
|
|
<td id="7EF01D02E785CCAD52226E08B8467CBC" colspan="1" pageId="0" pageNumber="507" rowspan="1">46.5</td>
|
|
<td id="FC79B6BB3459BD74A8340F332CE024C9" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.7</td>
|
|
<td id="6962DFC30841AB8D4874C91B17796B5D" colspan="1" pageId="0" pageNumber="507" rowspan="1">46.4-49.4</td>
|
|
<td id="3559940794A4C03D377C8DB8B591B1DC" colspan="1" pageId="0" pageNumber="507" rowspan="1">46.5</td>
|
|
<td id="029DE403425095353EC9B1B518102D0F" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.7</td>
|
|
</tr>
|
|
<tr id="42013D27D684FF35DF955C1FE606B178" pageId="0" pageNumber="507">
|
|
<td id="EB394E63E04CA4A16F649E7AD20488F4" colspan="1" pageId="0" pageNumber="507" rowspan="1">Head length</td>
|
|
<td id="A7318BEA821EAFCB06C9265BF9938C90" colspan="1" pageId="0" pageNumber="507" rowspan="1">24.0</td>
|
|
<td id="F181F3638CB35504CDFABBEE1B7969D0" colspan="1" pageId="0" pageNumber="507" rowspan="1">22.3-26.6</td>
|
|
<td id="8D3ED807A99A14D13E270B05AECDEED5" colspan="1" pageId="0" pageNumber="507" rowspan="1">24.0</td>
|
|
<td id="28C12C7C04F99ACD21B986976A0D750A" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.7</td>
|
|
<td id="3318F6617F7BDA0878E690CEB6BD065F" colspan="1" pageId="0" pageNumber="507" rowspan="1">22.3-24.9</td>
|
|
<td id="B54955D2F2698FFDEB407404E337A52F" colspan="1" pageId="0" pageNumber="507" rowspan="1">23.1</td>
|
|
<td id="788BD16E7953A76C1A322359A1B23352" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.6</td>
|
|
</tr>
|
|
<tr id="E229658E05E50CACFB8A2BB888BDD100" pageId="0" pageNumber="507">
|
|
<td id="B4B12BC6BBC065465E7CB878D69B3088" colspan="8" pageId="0" pageNumber="507" rowspan="1">
|
|
<emphasis id="559A4DDEB36DC4E8E0D410F7AE4347FB" bold="true" pageId="0" pageNumber="507">Percentages of head length</emphasis>
|
|
</td>
|
|
</tr>
|
|
<tr id="4D965DD1CCC8EA08E52E7ECD61FFFE46" pageId="0" pageNumber="507">
|
|
<td id="79211A942A80C4C68E6E1E7E24E1ED71" colspan="1" pageId="0" pageNumber="507" rowspan="1">Horizontal eye diameter</td>
|
|
<td id="C6171F9BC97F0AE05FDCB6CFA98317E9" colspan="1" pageId="0" pageNumber="507" rowspan="1">30.2</td>
|
|
<td id="6A5FFB70685663C4E4885516C9DA7B17" colspan="1" pageId="0" pageNumber="507" rowspan="1">28.7 -36.0</td>
|
|
<td id="36ECAFCC9C0B481CABEB17335106BDA2" colspan="1" pageId="0" pageNumber="507" rowspan="1">31.4</td>
|
|
<td id="09A1AC0780E10863FCEB3E0E6562D423" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.5</td>
|
|
<td id="67ECE8C7EEF18E4BDBA45DB9DE505B2E" colspan="1" pageId="0" pageNumber="507" rowspan="1">29.5-34.8</td>
|
|
<td id="FB12EA527BB1551461AE42A0067391E8" colspan="1" pageId="0" pageNumber="507" rowspan="1">31.6</td>
|
|
<td id="FE8314E4DAC1C522F3C0ED5FE858549A" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.4</td>
|
|
</tr>
|
|
<tr id="AFB5ECE061A0975B92DBD92D48C232F8" pageId="0" pageNumber="507">
|
|
<td id="765D92DA8B0F252B626B9AF1DF1AC204" colspan="1" pageId="0" pageNumber="507" rowspan="1">Snout length</td>
|
|
<td id="CB0309AF585A923B8005C96931B995CA" colspan="1" pageId="0" pageNumber="507" rowspan="1">24.2</td>
|
|
<td id="D9449B4557C18D2C74B10909B903E361" colspan="1" pageId="0" pageNumber="507" rowspan="1">19.7 -28.8</td>
|
|
<td id="7A47FE9EE8B4FCE9DADB08669194A7E3" colspan="1" pageId="0" pageNumber="507" rowspan="1">23.5</td>
|
|
<td id="6B32B8500FB050624F6C346512F2554E" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.6</td>
|
|
<td id="92A78C95CCD84EC267A2ED7D15867283" colspan="1" pageId="0" pageNumber="507" rowspan="1">22.8-29.3</td>
|
|
<td id="BF96C08971A53672309A11AF8FCB4FD7" colspan="1" pageId="0" pageNumber="507" rowspan="1">25.4</td>
|
|
<td id="8AE85640E349B17EB0B1C12977F9EDD8" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.2</td>
|
|
</tr>
|
|
<tr id="ECD365D1109DA625FE2F5D9C849D7853" pageId="0" pageNumber="507">
|
|
<td id="A5DF337923B432B3C1D7E6BB7BDE43B2" colspan="1" pageId="0" pageNumber="507" rowspan="1">Least interorbital width</td>
|
|
<td id="D2E6A977206D33E9675900793E610F37" colspan="1" pageId="0" pageNumber="507" rowspan="1">36.8</td>
|
|
<td id="E29A6A4EB29C8D91D0BA21F2A0156ABD" colspan="1" pageId="0" pageNumber="507" rowspan="1">32.7 -38.9</td>
|
|
<td id="AA53C674E7B00D0D1D34A6156E4982F3" colspan="1" pageId="0" pageNumber="507" rowspan="1">34.1</td>
|
|
<td id="87407EB570D625E9F724B97EB7E54341" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.1</td>
|
|
<td id="503262C8D413F8D0E7F96B1689A2B0D0" colspan="1" pageId="0" pageNumber="507" rowspan="1">32.9 -37.0</td>
|
|
<td id="5B9B89D7A3D60D73644F7764D58D9E5F" colspan="1" pageId="0" pageNumber="507" rowspan="1">11.1</td>
|
|
<td id="047E19016DEBE8279D9BC0AB2A4D5CD7" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.1</td>
|
|
</tr>
|
|
<tr id="E8634A0D93DB2C5FAA18C83E33E0D8DA" pageId="0" pageNumber="507">
|
|
<td id="8F93AAE3A8274D493EDBE7A4BCCCA9B3" colspan="1" pageId="0" pageNumber="507" rowspan="1">Upper jaw length</td>
|
|
<td id="502F5C763CE2E61F0B337A700C01F260" colspan="1" pageId="0" pageNumber="507" rowspan="1">34.2</td>
|
|
<td id="03DE6E334993B407BFD0A3F508E099A4" colspan="1" pageId="0" pageNumber="507" rowspan="1">31.9 -37.3</td>
|
|
<td id="972951336F75D384EF44DE2F84284A57" colspan="1" pageId="0" pageNumber="507" rowspan="1">33.4</td>
|
|
<td id="90C7C4A76C6C53E040E0098E61BCE166" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.2</td>
|
|
<td id="6E688B50FE4B6B0EE1FC477724836B2E" colspan="1" pageId="0" pageNumber="507" rowspan="1">32.7-39.9</td>
|
|
<td id="1BC7AFFA514C94DDBD3383AA8AD1962D" colspan="1" pageId="0" pageNumber="507" rowspan="1">33.9</td>
|
|
<td id="D8081D43D5BFB08FC21E32E47F268D34" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.4</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<paragraph id="94BAD326026AC02A7393776F9E267F95" pageId="0" pageNumber="507">All teeth unicuspid or tricuspid and lateral cusps, when present, much smaller; premaxillary teeth in one rows with 6(9), 7(23) tricuspid teeth; maxilla with 11(3), 12(12), 13(14), or 14(3) unicuspid teeth; dentary with 6 (2) or 7 (30) larger tricuspid teeth followed by 6(26) or 7(6) smaller tricuspid teeth.</paragraph>
|
|
<paragraph id="38F2131A1A154507057D8E69F813B9F0" pageId="0" pageNumber="507">Scales cycloid and same size over entire body generally. Predorsal scales mostly regular, but sometimes irregular just posterior to supraoccipital and/or slightly anterior to dorsal-fin. Scales covering anterior third of caudal-fin, with up to two, three, or four scales beyond posterior margin of hypural plate. Lateral line interrupted; last scale on caudal-fin base, with 9+1(12),10+1(74), 11+1(50), or 12+1(5). Longitudinal scales series including lateral-line scales 35(3), 36(3), 37(56), 38(49), or 39(30). Longitudinal scales rows between dorsal-fin origin and lateral line 5(1), 6(93) or 7(47). Horizontal scale rows between lateral line and pelvic-fin origin 4 (141), Axillary scale present. Scales in median series between tip of supraoccipital spine and dorsal-fin origin 13+1(24),14+1(65), 15+1(26), or 16+1(26). Circumpeduncular scales 13(18), 14(115), or 15(8).</paragraph>
|
|
<paragraph id="3A6071BC842E55828EC52302814716A4" pageId="0" pageNumber="507">
|
|
Dorsal-fin rays i+10(99) or ii+10(42). Dorsal-fin origin situated posterior to vertical through pelvic-fin insertion, near middle of body. First dorsal-fin pterygiophore main body located of 8th and 9th vertebrae. Adipose-fin present. Anal-fin i+14(20), iii+15(18), ii+16(61), iii+16(24), ii+17(10), iii+17(5), ii+18 (3). Anteriormost anal-fin pterygiophore inserting at 14th and 15th vertebrae. Anterior anal-fin margin slightly convex, with anteriormost rays more elongate and slightly more thickened than remaining rays, forming a distinct lobe. Remaining rays smaller with straight distal margin. Pectoral-fin rays i+9(8), i+10(113), or i+11(20). Tip of pectoral-fin not reaching pelvic-fin origin, when adpressed. Pelvic-fin rays i+7(120) or ii+7(21). Tip of pelvic-fin not reaching anal-fin origin, when adpressed. Caudal-fin with a sexually dimorphic pattern, described below (
|
|
<figureCitation id="6BF9F0373733ACCC4B3EEF399FE00748" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Aphyocharax brevicaudatus sp. nov. a. CICCAA 02293, holotype (male), 35.9 mm SL; b. CICCAA 02294, paratype (female), 32.4 mm SL, Brazil: Maranhao state: Maracacume river basin. (Photographed by Erick Guimaraes)." figureDoi=".figure1" httpUri="https://binary.pensoft.net/fig/349754" pageId="0" pageNumber="507" tableDoi=".figure1">Fig. 1</figureCitation>
|
|
). Principal caudal-fin rays 10+9(130) or 10+10(11); dorsal procurrent rays 8(2), 9(3) or 10(27) and ventral procurrent rays 7(2), 8(3) or 9(27).
|
|
</paragraph>
|
|
<paragraph id="44133E6A9013BE4CD25499F3B5CE7AC5" pageId="0" pageNumber="507">Branchiostegal rays 4(32). Supraneurals 6(4) 7(27) or 8(1). Total vertebrae 31 (1), 32(30) or 33(1).</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="B4DEBF1A9F96F9F3132F7BE9271B2F7C" pageId="0" pageNumber="507" type="colour in alcohol">
|
|
<paragraph id="A5B1099A16E5578E50D000179275E5CE" pageId="0" pageNumber="507">Colour in alcohol.</paragraph>
|
|
<paragraph id="8B8F9BC4B50AF4F6E9BA7995CC239AD5" pageId="0" pageNumber="507">Ground colouration light brown to yellowish brown. Inconspicuous light brown to light gray stripe from humeral spot to caudal-fin base, more conspicuous on posterior half. Humeral region with one conspicuous dark brown to black humeral spot. Smaller dark brown or black chromatophores homogeneously scattered. Smaller dark brown or black chromatophores homogeneously scattered along body, except on chest. Head ground colouration similar to trunk, with dark brown chromatophores present on jaws, tip of snout, opercle, and dorsal portion of head. Dorsal, adipose, anal, caudal, pectoral, and pelvic fins hyaline to light brown.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="E2BBA13A07510B94708163BD31C56806" pageId="0" pageNumber="507" type="sexual dimorphism">
|
|
<paragraph id="C4674D279C50C5AD269BDB6F2DE51046" pageId="0" pageNumber="507">Sexual dimorphism.</paragraph>
|
|
<paragraph id="25A772AC15C6F468C38FD84BA355F533" pageId="0" pageNumber="507">
|
|
Caudal-fin of mature males with upper lobe longer (about 2/3 longer) than lower one, while both cauldal-fin lobes have similar leght in females (
|
|
<figureCitation id="60D998F0B0A4A4E562C5EEDBB875A2D3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Aphyocharax brevicaudatus sp. nov. a. CICCAA 02293, holotype (male), 35.9 mm SL; b. CICCAA 02294, paratype (female), 32.4 mm SL, Brazil: Maranhao state: Maracacume river basin. (Photographed by Erick Guimaraes)." figureDoi=".figure1" httpUri="https://binary.pensoft.net/fig/349754" pageId="0" pageNumber="507" tableDoi=".figure1">Fig. 1</figureCitation>
|
|
). Gill glands were found in all analyzed mature males of
|
|
<taxonomicName id="AEC4BAADB1CC43D7D3C5F2683E15C954" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax brevicaudatus" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus">
|
|
<emphasis id="1067BFD582ABC007789EB2442CA00BA5" italics="true" pageId="0" pageNumber="507">Aphyocharax brevicaudatus</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and were always absent in females. They were always located on anteriormost portion of lower branch of first gill arch, extending posteriorly through variable number of gill filaments.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="962407BAF6D848F50885C7B743B52ADD" pageId="0" pageNumber="507" type="etymology">
|
|
<paragraph id="54EA3AAFA0756AA69E97945D5D29862E" pageId="0" pageNumber="507">Etymology.</paragraph>
|
|
<paragraph id="23FFAA4F9F52989DBED935354F66A4CB" pageId="0" pageNumber="507">
|
|
The name
|
|
<taxonomicName id="C0E01867B95B148B6A8A5DE1AD521E8B" lsidName="brevicaudatus" pageId="0" pageNumber="507" rank="species" species="brevicaudatus">
|
|
<emphasis id="4F149630F50DCF124757CF0BAF0DDE93" italics="true" pageId="0" pageNumber="507">brevicaudatus</emphasis>
|
|
</taxonomicName>
|
|
is a contraction of the
|
|
<taxonomicName id="4725710DB797C89426DFC6246DCB2A77" genus="Latin" lsidName="Latin" pageId="0" pageNumber="507" rank="genus">
|
|
<emphasis id="6C1E9A4C51EAC1761797F6762C0119A4" italics="true" pageId="0" pageNumber="507">Latin</emphasis>
|
|
</taxonomicName>
|
|
words
|
|
<emphasis id="50EBAD29997167F683CC959B9AF59CA9" italics="true" pageId="0" pageNumber="507">brevis</emphasis>
|
|
meaning
|
|
<normalizedToken id="21F5ED1B757AF689608769DE336D4C49" originalValue="“short”">"short"</normalizedToken>
|
|
and
|
|
<emphasis id="CB73E1DD17BAA13C8DC4A3B2A19A3F66" italics="true" pageId="0" pageNumber="507">cauda</emphasis>
|
|
meaning
|
|
<normalizedToken id="5B224B3ED4C089E32BD26ACC300A8F3D" originalValue="“tail”">"tail"</normalizedToken>
|
|
, an allusion to the shorter caudal-fin lower lobe in the mature males of the new species.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="B3115941FAE04B031C74CEABE609FF46" pageId="0" pageNumber="507" type="distribution">
|
|
<paragraph id="C6AC7FA05E080D1C8EA5E9CDD6988007" pageId="0" pageNumber="507">Geographic distribution.</paragraph>
|
|
<paragraph id="BA94F5A806A42F3A5E3335A0C01B996E" pageId="0" pageNumber="507">
|
|
<taxonomicName id="5658E19C210AE54F63B2BDF8B5DFBB97" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax brevicaudatus" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus">
|
|
<emphasis id="910871E111F68B72965223604FFC4F59" italics="true" pageId="0" pageNumber="507">Aphyocharax brevicaudatus</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. is currently known only from a single locality, the
|
|
<normalizedToken id="13216ED74ED6878581EEC0AF2C90871D" originalValue="Maracaçumé">Maracacume</normalizedToken>
|
|
river basin, a small and isolated coastal river basin of the eastern Amazon region (
|
|
<figureCitation id="B1D4BAEA2695CE77AEE36B09732FEB63" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Type locality of Aphyocharax brevicaudatus sp. nov." figureDoi=".figure3" httpUri="https://binary.pensoft.net/fig/349756" pageId="0" pageNumber="507" tableDoi=".figure3">Fig. 3</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="FA485D3C129933051084F0EEB5FBC4F8" doi=".figure3" httpUri="https://binary.pensoft.net/fig/349756" pageId="0" pageNumber="507" start="Figure 3" startId="F3">
|
|
<paragraph id="40ACB8ED1BCB1C0D74583B76F64217A4" pageId="0" pageNumber="507">
|
|
<emphasis id="4660275D148162B4432253CA1E56DA2A" bold="true" pageId="0" pageNumber="507">Figure 3.</emphasis>
|
|
Type locality of
|
|
<taxonomicName id="EA353186EAC711CC592FFE4404F9C0BE" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax brevicaudatus" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus">
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<emphasis id="DBE3BDA32FFAD577AAB3682B0DB92AD8" italics="true" pageId="0" pageNumber="507">Aphyocharax brevicaudatus</emphasis>
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</taxonomicName>
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sp. nov.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |