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<document id="006B7C453465A4E4400D7600FD8322CA" ID-CLB-Dataset="24286" ID-DOI="10.11646/zootaxa.4728.2.2" ID-GBIF-Dataset="a1c40b73-0c11-4863-a188-412c33523e02" ID-ISSN="1175-5326" ID-Zenodo-Dep="3621117" ID-ZooBank="AE4F8089-7D2E-4ABA-AA09-CD504705788A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1579677041620" checkinUser="plazi" docAuthor="Wagner, Rüdiger &amp; Withers, Phil" docDate="2020" docId="784D878B601F21008FF10570FCBF8D43" docLanguage="en" docName="zootaxa.4728.2.2.pdf" docOrigin="Zootaxa 4728 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tonnoiriella nigricauda" docType="treatment" docVersion="8" lastPageNumber="204" masterDocId="8474FFF3600C21158F66006DFFB28F20" masterDocTitle="The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae)" masterLastPageNumber="210" masterPageNumber="183" pageNumber="202" updateTime="1698778572713" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="0759CCE1A78E9D81C7BB104313FF1457" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="5B750183651F3A306A7A46B675491CDB">The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae)</mods:title>
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<mods:namePart id="27D01820C67432C807C75F0C6C61EFDE">Wagner, Rüdiger</mods:namePart>
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<mods:namePart id="855CF12E54544024132439542A397D04">Withers, Phil</mods:namePart>
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<mods:title id="91D1591EE6E6436993EFC8C2540AD31B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0B6BFAA715EFB5B8EB0FE3ABF8B76A9D">
<mods:date id="6AAD8E2832C9AB0E223E1AC7F005B1B8">2020</mods:date>
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<mods:number id="5F0A90115249FB7E0CBF0FC4C73825BA">2020-01-22</mods:number>
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<mods:number id="655DA172B30D2D47BC110C1D982BA7E2">4728</mods:number>
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<mods:number id="BBDEAAAA4BFDEF32B7DD7BF82B62F783">2</mods:number>
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<mods:start id="11348E4D0C45D7BA3B7E7068EF2159D4">183</mods:start>
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<mods:classification id="71FAC3E14D4B105788BF232051678D64">journal article</mods:classification>
<mods:identifier id="110A798C23027FA2F2C03D0A7BACD45E" type="CLB-Dataset">24286</mods:identifier>
<mods:identifier id="D80E51D9EC97BBBCA5AD09A5CD1CC2AE" type="DOI">10.11646/zootaxa.4728.2.2</mods:identifier>
<mods:identifier id="A1E67385461BF470160E78B50B19651A" type="GBIF-Dataset">a1c40b73-0c11-4863-a188-412c33523e02</mods:identifier>
<mods:identifier id="78BE7FD7F2B297EA727E610DDF732ADA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0C3BB831C42108021B9A3C08F18772D4" type="Zenodo-Dep">3621117</mods:identifier>
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<treatment id="784D878B601F21008FF10570FCBF8D43" ID-DOI="http://doi.org/10.5281/zenodo.5918611" ID-GBIF-Taxon="161827856" ID-Zenodo-Dep="5918611" LSID="urn:lsid:plazi:treatment:784D878B601F21008FF10570FCBF8D43" httpUri="http://treatment.plazi.org/id/784D878B601F21008FF10570FCBF8D43" lastPageId="21" lastPageNumber="204" pageId="19" pageNumber="202">
<subSubSection id="B8FE6516601F21068FF10570FDCF8A17" box="[151,637,1309,1336]" pageId="19" pageNumber="202" type="nomenclature">
<paragraph id="F05B369D601F21068FF10570FDCF8A17" blockId="19.[151,637,1309,1372]" box="[151,637,1309,1336]" pageId="19" pageNumber="202">
<heading id="AB1381F1601F21068FF10570FDCF8A17" bold="true" box="[151,637,1309,1336]" fontSize="11" level="1" pageId="19" pageNumber="202" reason="1">
<taxonomicName id="37E44D1E601F21068FF10570FDCF8A17" ID-CoL="8FGGB" authority="(Tonnoir, 1919)" baseAuthorityName="Tonnoir" baseAuthorityYear="1919" box="[151,637,1309,1336]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="202" phylum="Arthropoda" rank="species" species="nigricauda">
<emphasis id="C290EA8F601F21068FF10570FDCF8A17" bold="true" box="[151,637,1309,1336]" pageId="19" pageNumber="202">
<emphasis id="C290EA8F601F21068FF10570FE058A18" bold="true" box="[151,439,1309,1336]" italics="true" pageId="19" pageNumber="202">Tonnoiriella nigricauda</emphasis>
(
<bibRefCitation id="94754B6C601F21068EA00570FDC68A17" author="Tonnoir, A. L." box="[454,628,1309,1335]" pageId="19" pageNumber="202" pagination="8 - 17" refId="ref20540" refString="Tonnoir, A. L. (1919) Contribution a l`etude des Psychodidae de Belgique I. Annales de la Societe entomologique de Belgique, 59, 8 - 17." type="journal article" year="1919">Tonnoir, 1919</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601F21068FF1052CFE9C8A7C" box="[151,302,1345,1372]" pageId="19" pageNumber="202" type="description">
<paragraph id="F05B369D601F21068FF1052CFE9C8A7C" blockId="19.[151,637,1309,1372]" box="[151,302,1345,1372]" pageId="19" pageNumber="202">
(
<figureCitation id="68DF2A18601F21068FF9052CFE948A7C" box="[159,294,1345,1372]" captionStart="FIGURES 43–49" captionStartId="20.[151,264,1910,1935]" captionTargetBox="[231,1376,218,1841]" captionTargetId="figure@20.[191,1397,181,1884]" captionTargetPageId="20" captionText="FIGURES 43–49: 43–45 T. nigricauda 43 hypandrium, gonopods, aedeagus, 44 aedeagus ventral view; 45 variability of the ventral distiphallus sclerite in England, Poland, North- and South Germany (left to right?); 46 T. goncalvesi genitalia ventral view; 47 T. andradei hypandrium, gonopods, aedeagus; 48–49 T. arcuata (holotype) 48 hypandrium, gonopods, aedeagus, 49 epandrium and epandrial processes, oblique view." figureDoi="http://doi.org/10.5281/zenodo.3621129" httpUri="https://zenodo.org/record/3621129/files/figure.png" pageId="19" pageNumber="202">Figs. 43–45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601F21068FF105EBFBFA8A9C" pageId="19" pageNumber="202" type="reference_group">
<paragraph id="F05B369D601F21068FF105EBFBE68ABE" blockId="19.[151,1108,1413,1469]" box="[151,1108,1413,1438]" pageId="19" pageNumber="202">
<taxonomicName id="37E44D1E601F21068FF105EBFDB58ABE" ID-CoL="89NMQ" authority="Tonnoir, 1919" authorityName="Tonnoir" authorityYear="1919" box="[151,519,1413,1438]" class="Insecta" family="Psychodidae" genus="Pericoma" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="202" phylum="Arthropoda" rank="species" species="nigricauda">
<emphasis id="C290EA8F601F21068FF105EBFEDD8ABE" box="[151,367,1414,1438]" italics="true" pageId="19" pageNumber="202">Pericoma nigricauda</emphasis>
<bibRefCitation id="94754B6C601F21068E1305E8FDB58ABE" author="Tonnoir, A. L." box="[373,519,1413,1438]" pageId="19" pageNumber="202" pagination="8 - 17" refId="ref20540" refString="Tonnoir, A. L. (1919) Contribution a l`etude des Psychodidae de Belgique I. Annales de la Societe entomologique de Belgique, 59, 8 - 17." type="journal article" year="1919">Tonnoir, 1919</bibRefCitation>
</taxonomicName>
: Annales de la Société entomologique de
<collectingCountry id="88F3760D601F21068CCB05E8FBBA8ABE" box="[941,1032,1413,1438]" name="Belgium" pageId="19" pageNumber="202">Belgique</collectingCountry>
59: 11.
</paragraph>
<paragraph id="F05B369D601F21068FF105C9FBFA8A9C" blockId="19.[151,1108,1413,1469]" box="[151,1096,1444,1469]" pageId="19" pageNumber="202">
<taxonomicName id="37E44D1E601F21068FF105C9FE3B8A9C" ID-CoL="8FGGB" baseAuthorityName="Tonnoir" baseAuthorityYear="1919" box="[151,393,1444,1468]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="202" phylum="Arthropoda" rank="species" species="nigricauda">
<emphasis id="C290EA8F601F21068FF105C9FE3B8A9C" box="[151,393,1444,1468]" italics="true" pageId="19" pageNumber="202">Tonnoiriella nigricauda</emphasis>
</taxonomicName>
;
<bibRefCitation id="94754B6C601F21068EF005C9FDB98A9C" author="Jezek, J." box="[406,523,1444,1469]" pageId="19" pageNumber="202" pagination="7 - 18" refId="ref20095" refString="Jezek, J. (1999) Rare and new Palaearctic Tonnoiriella - species (Diptera, Psychodidae). Casopis Narodniho muzea Rada prirodovedna, 168 (1 - 4), 7 - 18." type="journal article" year="1999">Ježek 1999</bibRefCitation>
: Časopis Národniho muzea Řada přirodovědná, 168: 17.
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601F21068FF10587FAAB8951" pageId="19" pageNumber="202" type="materials_examined">
<paragraph id="F05B369D601F21068FF10587FAAB8951" blockId="19.[151,1437,1514,2045]" pageId="19" pageNumber="202">
<emphasis id="C290EA8F601F21068FF10587FF4D8925" bold="true" box="[151,255,1514,1541]" pageId="19" pageNumber="202">Material</emphasis>
:
<typeStatus id="2F5F883F601F21068E760586FEFA8925" box="[272,328,1515,1541]" pageId="19" pageNumber="202">Type</typeStatus>
material not inspected.
<materialsCitation id="408C3CC0601F21068D3A0586FE228909" ID-GBIF-Occurrence="2556505418" collectingDate="1987-05-15" collectorName="Wiedienska" country="Poland" county="Roztocze" location="Sopol river" municipality="Fryszerka" pageId="19" pageNumber="202" specimenCount="1" specimenCount-male="1">
Further specimens
<specimenCount id="E6E2FD14601F21068C530586FCEA8925" box="[821,856,1515,1541]" pageId="19" pageNumber="202" type="male">1♂</specimenCount>
<collectingCountry id="88F3760D601F21068C070586FC018925" box="[865,947,1515,1541]" name="Poland" pageId="19" pageNumber="202">Poland</collectingCountry>
:
<collectingCounty id="193A4E11601F21068CD90586FB988925" box="[959,1066,1515,1541]" pageId="19" pageNumber="202">Roztocze</collectingCounty>
,
<collectingMunicipality id="103FACE7601F21068B510586FB1A8925" box="[1079,1192,1515,1541]" pageId="19" pageNumber="202">Fryszerka</collectingMunicipality>
,
<location id="F53B6046601F21068BD00586FA858925" LSID="urn:lsid:plazi:treatment:784D878B601F21008FF10570FCBF8D43:F53B6046601F21068BD00586FA858925" box="[1206,1335,1515,1541]" country="Poland" county="Roztocze" municipality="Fryszerka" name="Sopol river" pageId="19" pageNumber="202">Sopol river</location>
,
<date id="845A105D601F21068A220587FF7D8909" pageId="19" pageNumber="202" value="1987-05-15">
<collectingDate id="941EE9B5601F21068A220587FF7D8909" pageId="19" pageNumber="202" value="1987-05-15">15 May 1987</collectingDate>
</date>
leg.
<collectorName id="5D11534B601F21068E6F0662FE228909" box="[265,400,1551,1577]" pageId="19" pageNumber="202">Wiedienska</collectorName>
</materialsCitation>
;
<materialsCitation id="408C3CC0601F21068EFB0662FF68896D" ID-GBIF-Occurrence="2556505442" collectingDate="1986-09-08" collectorName="Holm" country="Germany" location="Schierenseebach" pageId="19" pageNumber="202" specimenCount="1" specimenCount-male="1" stateProvince="Schleswig-Holstein">
<specimenCount id="E6E2FD14601F21068EFB0662FE0D8909" box="[413,447,1551,1577]" pageId="19" pageNumber="202" type="male">1♂</specimenCount>
<collectingCountry id="88F3760D601F21068EAE0662FD878909" box="[456,565,1551,1577]" name="Germany" pageId="19" pageNumber="202">Germany</collectingCountry>
:
<collectingRegion id="3220F87F601F21068D270662FCAD8909" box="[577,799,1551,1577]" country="Germany" name="Schleswig-Holstein" pageId="19" pageNumber="202">Schleswig-Holstein</collectingRegion>
,
<location id="F53B6046601F21068C4A0662FC558909" LSID="urn:lsid:plazi:treatment:784D878B601F21008FF10570FCBF8D43:F53B6046601F21068C4A0662FC558909" box="[812,999,1551,1577]" country="Germany" name="Schierenseebach" pageId="19" pageNumber="202" stateProvince="Schleswig-Holstein">Schierenseebach</location>
(lower course)
<date id="845A105D601F21068BFD0662FAD88909" box="[1179,1386,1551,1577]" pageId="19" pageNumber="202" value="1986-09-08">
<collectingDate id="941EE9B5601F21068BFD0662FAD88909" box="[1179,1386,1551,1577]" pageId="19" pageNumber="202" value="1986-09-08">8 September 1986</collectingDate>
</date>
leg.
<collectorName id="5D11534B601F21068FF1065EFF68896D" box="[151,218,1587,1613]" pageId="19" pageNumber="202">Holm</collectorName>
</materialsCitation>
;
<materialsCitation id="408C3CC0601F21068F8E065EFEB28951" ID-GBIF-Occurrence="2556505423" collectingDate="1978-07-27" collectorName="Halvelka" country="Germany" county="Baden-Wurrtemberg" location="Oberhausen-Rheinhausen" municipality="Taubergiessen" pageId="19" pageNumber="202" specimenCount="1" specimenCount-male="1">
<specimenCount id="E6E2FD14601F21068F8E065EFEB9896D" box="[232,267,1587,1613]" pageId="19" pageNumber="202" type="male">1♂</specimenCount>
<collectingCountry id="88F3760D601F21068E72065EFE33896D" box="[276,385,1587,1613]" name="Germany" pageId="19" pageNumber="202">Germany</collectingCountry>
:
<collectingCounty id="193A4E11601F21068EEB065EFDC5896D" box="[397,631,1587,1613]" pageId="19" pageNumber="202">Baden-Würrtemberg</collectingCounty>
, NSG
<collectingMunicipality id="103FACE7601F21068DA3065EFCD2896D" box="[709,864,1587,1613]" pageId="19" pageNumber="202">Taubergiessen</collectingMunicipality>
, near
<location id="F53B6046601F21068CC3065EFB7A896D" LSID="urn:lsid:plazi:treatment:784D878B601F21008FF10570FCBF8D43:F53B6046601F21068CC3065EFB7A896D" box="[933,1224,1587,1613]" country="Germany" county="Baden-Wurrtemberg" municipality="Taubergiessen" name="Oberhausen-Rheinhausen" pageId="19" pageNumber="202">Oberhausen-Rheinhausen</location>
,
<date id="845A105D601F21068BB3065EFAD8896D" box="[1237,1386,1587,1613]" pageId="19" pageNumber="202" value="1978-07-27">
<collectingDate id="941EE9B5601F21068BB3065EFAD8896D" box="[1237,1386,1587,1613]" pageId="19" pageNumber="202" value="1978-07-27">27 July 1978</collectingDate>
</date>
leg.
<collectorName id="5D11534B601F21068FF1063AFEB28951" box="[151,256,1623,1649]" pageId="19" pageNumber="202">Halvelka</collectorName>
</materialsCitation>
;
<materialsCitation id="408C3CC0601F21068E6D063AFAAB8951" ID-GBIF-Occurrence="2556505421" box="[267,1305,1622,1649]" collectingDate="1985-05-19" collectingDateMax="1985-05-26" collectingDateMin="1985-05-19" collectorName="Irwin &amp; Urwin." country="United Kingdom" location="Upton Broad" pageId="19" pageNumber="202" specimenCount="1" specimenCount-male="1" stateProvince="Norfolk">
<specimenCount id="E6E2FD14601F21068E6D063AFE9C8951" box="[267,302,1623,1649]" pageId="19" pageNumber="202" type="male">1♂</specimenCount>
<collectingCountry id="88F3760D601F21068E53063AFE7E8951" box="[309,460,1623,1649]" name="United Kingdom" pageId="19" pageNumber="202">Great Britain</collectingCountry>
:
<collectingRegion id="3220F87F601F21068EB1063AFD818951" box="[471,563,1623,1649]" country="United Kingdom" name="Norfolk" pageId="19" pageNumber="202">Norfolk</collectingRegion>
,
<location id="F53B6046601F21068D5B063AFD638951" LSID="urn:lsid:plazi:treatment:784D878B601F21008FF10570FCBF8D43:F53B6046601F21068D5B063AFD638951" box="[573,721,1623,1649]" country="United Kingdom" name="Upton Broad" pageId="19" pageNumber="202" stateProvince="Norfolk">Upton Broad</location>
,
<collectingMethod id="29A54E8A601F21068DBB063AFCD58951" box="[733,871,1623,1649]" pageId="19" pageNumber="202">malaise trap</collectingMethod>
,
<date id="845A105D601F21068C15063AFB808951" box="[883,1074,1622,1649]" pageId="19" pageNumber="202" value="1985-05-19" valueMax="1985-05-26" valueMin="1985-05-19">
<collectingDate id="941EE9B5601F21068C15063AFB808951" box="[883,1074,1622,1649]" pageId="19" pageNumber="202" value="1985-05-19" valueMax="1985-05-26" valueMin="1985-05-19">19–26 May 1985</collectingDate>
</date>
leg.
<collectorName id="5D11534B601F21068B0C063AFB158951" box="[1130,1191,1623,1649]" pageId="19" pageNumber="202">Irwin</collectorName>
&amp;
<collectorName id="5D11534B601F21068BAC063AFAAB8951" box="[1226,1305,1623,1649]" pageId="19" pageNumber="202">Urwin.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601F21008FA10617FE118EAB" lastPageId="21" lastPageNumber="204" pageId="19" pageNumber="202" type="description">
<paragraph id="F05B369D601F21068FA10617FEB78869" blockId="19.[151,1437,1514,2045]" pageId="19" pageNumber="202">
<emphasis id="C290EA8F601F21068FA10617FEE389B5" bold="true" box="[199,337,1658,1685]" pageId="19" pageNumber="202">Description</emphasis>
(Polish specimen): Head with eyebridge of 3 facet rows; distance between eyes 4.5–5 facet diameters; a single row postocular bristles. Antenna with scape, pedicel and 14 flagellomeres; absolute length of antennomeres: 0.048-0.048-0.054-0.052-0.048-0.048-0.048-0.048-0.048-0.046-0.042-0.032-0.030-0.024-
<quantity id="371C9B78601F21068BBC06AFFA2A89FD" box="[1242,1432,1730,1757]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" metricValueMax="4.8" metricValueMin="2.1999999999999997" pageId="19" pageNumber="202" unit="mm" value="0.035" valueMax="0.048" valueMin="0.022">0.022-0.048 mm</quantity>
; relative length 24-24-27-26-24-24-24-24-24-23-
<date id="845A105D601F21068DA3068BFCFF8821" box="[709,845,1766,1793]" pageId="19" pageNumber="202" value="1921-12-16" valueMax="1921-12-15" valueMin="1921-12-16">21-16-15-12</date>
-11-13. Flagellomeres 5–11 with a pair of digitiform curved ascoids in the distal third. Palpus: absolute segment length 0.056-0.070-
<quantity id="371C9B78601F21068B6E0767FB748805" box="[1032,1222,1802,1829]" metricMagnitude="-4" metricUnit="m" metricValue="1.1199999999999999" metricValueMax="1.44" metricValueMin="0.8" pageId="19" pageNumber="202" unit="mm" value="0.11199999999999999" valueMax="0.144" valueMin="0.08">0.080-0.144 mm</quantity>
; relative length 28- 35-40-72.
</paragraph>
<paragraph id="F05B369D601F21068FA1073EFC70884D" blockId="19.[151,1437,1514,2045]" box="[199,962,1875,1901]" pageId="19" pageNumber="202">
Wing 3.06 times longer than wide; length
<quantity id="371C9B78601F21068DF9073EFCB7884D" box="[671,773,1875,1901]" metricMagnitude="-3" metricUnit="m" metricValue="1.89" pageId="19" pageNumber="202" unit="mm" value="1.89">1.89 mm</quantity>
, width
<quantity id="371C9B78601F21068C31073EFC0C884D" box="[855,958,1875,1901]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="19" pageNumber="202" unit="mm" value="0.62">0.62 mm</quantity>
.
</paragraph>
<paragraph id="F05B369D601F21068FA1071AFCC988DD" blockId="19.[151,1437,1514,2045]" pageId="19" pageNumber="202">
Male terminalia with hypandrium thin with short median extension (
<figureCitation id="68DF2A18601F21068CA9071AFB9688B1" box="[975,1060,1911,1937]" captionStart="FIGURES 43–49" captionStartId="20.[151,264,1910,1935]" captionTargetBox="[231,1376,218,1841]" captionTargetId="figure@20.[191,1397,181,1884]" captionTargetPageId="20" captionText="FIGURES 43–49: 43–45 T. nigricauda 43 hypandrium, gonopods, aedeagus, 44 aedeagus ventral view; 45 variability of the ventral distiphallus sclerite in England, Poland, North- and South Germany (left to right?); 46 T. goncalvesi genitalia ventral view; 47 T. andradei hypandrium, gonopods, aedeagus; 48–49 T. arcuata (holotype) 48 hypandrium, gonopods, aedeagus, 49 epandrium and epandrial processes, oblique view." figureDoi="http://doi.org/10.5281/zenodo.3621129" httpUri="https://zenodo.org/record/3621129/files/figure.png" pageId="19" pageNumber="202">Fig. 43</figureCitation>
). Gonocoxites
<date id="845A105D601F21068BA8071AFB5888B1" box="[1230,1258,1911,1937]" pageId="19" pageNumber="202">2x</date>
longer than the apical width, 2 short setae in the basal half, a longer one at about middle, the longest near the apical end of the gonocoxite is 3-4 times longer than the more basal setae. Condyles shorter than gonocoxites, almost rectangular. Gonostylus more than
<date id="845A105D601F21068EF0078EFE0388DD" box="[406,433,2019,2045]" pageId="19" pageNumber="202">2x</date>
longer than the gonocoxite, evenly bent.
</paragraph>
<caption id="A49B6615601821018FF1071BFD2288DB" ID-DOI="http://doi.org/10.5281/zenodo.3621129" ID-Zenodo-Dep="3621129" httpUri="https://zenodo.org/record/3621129/files/figure.png" pageId="20" pageNumber="203" startId="20.[151,264,1910,1935]" targetBox="[231,1376,218,1841]" targetPageId="20">
<paragraph id="F05B369D601821018FF1071BFD2288DB" blockId="20.[151,1436,1910,2043]" pageId="20" pageNumber="203">
<emphasis id="C290EA8F601821018FF1071BFEEA88AF" bold="true" box="[151,344,1910,1935]" pageId="20" pageNumber="203">FIGURES 43–49:</emphasis>
43–45
<taxonomicName id="37E44D1E601821018EC00715FD9D88AF" baseAuthorityName="Tonnoir" baseAuthorityYear="1919" box="[422,559,1911,1935]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="203" phylum="Arthropoda" rank="species" species="nigricauda">
<emphasis id="C290EA8F601821018EC00715FD9D88AF" box="[422,559,1911,1935]" italics="true" pageId="20" pageNumber="203">T. nigricauda</emphasis>
</taxonomicName>
43 hypandrium, gonopods, aedeagus, 44 aedeagus ventral view; 45 variability of the ventral distiphallus sclerite in England, Poland, North- and South Germany (left to right?); 46
<taxonomicName id="37E44D1E601821018B0E07F1FB5D8893" authorityName="Wagner &amp; Withers" authorityYear="2020" box="[1128,1263,1947,1971]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="203" phylum="Arthropoda" rank="species" species="goncalvesi">
<emphasis id="C290EA8F601821018B0E07F1FB5D8893" box="[1128,1263,1947,1971]" italics="true" pageId="20" pageNumber="203">T. goncalvesi</emphasis>
</taxonomicName>
genitalia ventral view; 47
<taxonomicName id="37E44D1E601821018F9E07ADFEDF88F7" authorityName="Wagner &amp; Withers" authorityYear="2020" box="[248,365,1983,2007]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="203" phylum="Arthropoda" rank="species" species="andradei">
<emphasis id="C290EA8F601821018F9E07ADFEDF88F7" box="[248,365,1983,2007]" italics="true" pageId="20" pageNumber="203">T. andradei</emphasis>
</taxonomicName>
hypandrium, gonopods, aedeagus; 48–49
<taxonomicName id="37E44D1E601821018C7907ADFC3588F7" authorityName="Jezek" authorityYear="1997" box="[799,903,1984,2007]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="203" phylum="Arthropoda" rank="species" species="arcuata">
<emphasis id="C290EA8F601821018C7907ADFC3588F7" box="[799,903,1984,2007]" italics="true" pageId="20" pageNumber="203">T. arcuata</emphasis>
</taxonomicName>
(holotype) 48 hypandrium, gonopods, aedeagus, 49 epandrium and epandrial processes, oblique view.
</paragraph>
</caption>
<paragraph id="F05B369D601921008FA100F4FD548E63" blockId="21.[151,1437,153,611]" pageId="21" pageNumber="204">
Ventral bridge of parameres with large oval lobes adnate in the midst, extending to about middle of gonocoxite. The median inner prolongation is not clearly identifyable. Aedeagus dorsoventrally flat, basiphallus widened, medially with stronger sclerotization. Ventral lobe of the basiphallus split, slightly asymmetrical with one arm connected to the distiphallus, the other arm with a short tip. The dorsal lobe of the basiphallus straight broad and symmetric; without an apical cross sclerite (
<figureCitation id="68DF2A18601921008DE30144FD6B8E63" box="[645,729,297,323]" captionStart="FIGURES 43–49" captionStartId="20.[151,264,1910,1935]" captionTargetBox="[231,1376,218,1841]" captionTargetId="figure@20.[191,1397,181,1884]" captionTargetPageId="20" captionText="FIGURES 43–49: 43–45 T. nigricauda 43 hypandrium, gonopods, aedeagus, 44 aedeagus ventral view; 45 variability of the ventral distiphallus sclerite in England, Poland, North- and South Germany (left to right?); 46 T. goncalvesi genitalia ventral view; 47 T. andradei hypandrium, gonopods, aedeagus; 48–49 T. arcuata (holotype) 48 hypandrium, gonopods, aedeagus, 49 epandrium and epandrial processes, oblique view." figureDoi="http://doi.org/10.5281/zenodo.3621129" httpUri="https://zenodo.org/record/3621129/files/figure.png" pageId="21" pageNumber="204">Fig. 44</figureCitation>
).
</paragraph>
<paragraph id="F05B369D601921008FA10120FE118EAB" blockId="21.[151,1437,153,611]" pageId="21" pageNumber="204">Epandrium elongate rhomboid; epandrial processes almost straight with 8 distally frazzled retinacula. Hypoproct oval, microsetose.</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601921008FA101F9FA8A8E8F" box="[199,1336,404,431]" pageId="21" pageNumber="204" type="distribution">
<paragraph id="F05B369D601921008FA101F9FA8A8E8F" blockId="21.[151,1437,153,611]" box="[199,1336,404,431]" pageId="21" pageNumber="204">
<emphasis id="C290EA8F601921008FA101F9FEEB8E8F" bold="true" box="[199,345,404,431]" pageId="21" pageNumber="204">Distribution</emphasis>
: Specimens collected along lowland rivers and streams in Northern and Central Europe.
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601921008FA101D4FCBF8D43" pageId="21" pageNumber="204" type="discussion">
<paragraph id="F05B369D601921008FA101D4FCBF8D43" blockId="21.[151,1437,153,611]" pageId="21" pageNumber="204">
<emphasis id="C290EA8F601921008FA101D4FE868EF3" bold="true" box="[199,308,441,467]" pageId="21" pageNumber="204">Remarks</emphasis>
: Eyebridge of Polish specimen with 2 cross lines of 4 facet rows; wing length by width 2.98–3.20; distance between eyes 3.5–4 facet diameters. Wing length 1.65 (Taubergiessen) to
<quantity id="371C9B78601921008B5F01B0FB2D8ED7" box="[1081,1183,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="2.07" pageId="21" pageNumber="204" unit="mm" value="2.07">2.07 mm</quantity>
(GB
<collectingRegion id="3220F87F601921008BB901B0FA8F8ED8" box="[1247,1341,477,504]" country="United Kingdom" name="Norfolk" pageId="21" pageNumber="204">Norfolk</collectingRegion>
). British specimen with 9 retinacula (3 short, 6 long), specimen from the Taubergiessen with 8 retinacula (3 short, 5 long). Variability of genitalia is minor. There is some variation in the shape of the ventral distiphallus sclerite (
<figureCitation id="68DF2A18601921008A7F0248FAD88D1F" box="[1305,1386,548,575]" captionStart="FIGURES 43–49" captionStartId="20.[151,264,1910,1935]" captionTargetBox="[231,1376,218,1841]" captionTargetId="figure@20.[191,1397,181,1884]" captionTargetPageId="20" captionText="FIGURES 43–49: 43–45 T. nigricauda 43 hypandrium, gonopods, aedeagus, 44 aedeagus ventral view; 45 variability of the ventral distiphallus sclerite in England, Poland, North- and South Germany (left to right?); 46 T. goncalvesi genitalia ventral view; 47 T. andradei hypandrium, gonopods, aedeagus; 48–49 T. arcuata (holotype) 48 hypandrium, gonopods, aedeagus, 49 epandrium and epandrial processes, oblique view." figureDoi="http://doi.org/10.5281/zenodo.3621129" httpUri="https://zenodo.org/record/3621129/files/figure.png" pageId="21" pageNumber="204">Fig. 45</figureCitation>
) but differences are too small to distinguish between species.
</paragraph>
</subSubSection>
</treatment>
</document>
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