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<mods:title id="61104EF864342D0C3521244B4B20CE70">Review of the steatiticus - species group of the cuskeel genus Neobythites (Ophidiidae) from the Indo-Pacific, with description of two new species</mods:title>
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<heading id="B62ED46CFFF18F33FF69E11334C75078" bold="true" box="[151,458,1598,1624]" fontSize="11" level="1" pageId="1" pageNumber="158" reason="1">
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<taxonomicName id="2AD91883FFF18F33FF69E113348D5078" authority="Uiblein &amp; Nielsen, 2018" authorityName="Uiblein &amp; Nielsen" authorityYear="2018" box="[151,384,1598,1624]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="1" pageNumber="158" phylum="Chordata" rank="species" species="gloriae" status="sp. nov.">
<emphasis id="DFADBF12FFF18F33FF69E113348D5078" bold="true" box="[151,384,1598,1624]" italics="true" pageId="1" pageNumber="158">Neobythites gloriae</emphasis>
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<paragraph id="ED666300FFF18F33FF69E14C34BB505A" blockId="1.[151,458,1598,1658]" box="[151,438,1633,1658]" pageId="1" pageNumber="158">
(
<figureCitation id="75E27F85FFF18F33FF60E14C342B505A" box="[158,294,1633,1658]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1503,1525]" captionStartId-1="5.[151,250,876,898]" captionStartId-2="7.[151,250,1622,1644]" captionTargetBox-0="[163,1422,185,1492]" captionTargetBox-1="[172,1414,190,861]" captionTargetBox-2="[179,1412,223,1588]" captionTargetId-0="figure@3.[163,1423,185,1492]" captionTargetId-1="figure@5.[171,1415,189,862]" captionTargetId-2="figure@7.[151,1436,195,1601]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="7" captionText-0="FIGURE 1. Indo-Pacific Neobythites steatiticus species group. All photographs at left side and F show preserved fishes; the remaining five photographs at right side show fresh fishes, photographed on board immediately after capture. A) Neobythites gloriae n. sp. USNM 309008, HT, 158 mm SL, inner Gulf of Oman; B) N. lombokensis, ZMUC P 77744, HT, 93 mm SL, SE off Lombok, preserved (B1) and fresh (B2; side reversed); C) N. longipes, ZMUC P77740, 98 mm SL, SE off Lombok, preserved (C1) and fresh (C2); D) N. malayanus, ZMUC P77742, 138 mm SL, SE off Lombok, preserved (D1) and fresh (D2); E) N. malhaensis, ZMUC P77840, PT, 135 mm SL, Saya de Malha bank, preserved; F) N. meteori, ZMH 5621, HT, 102 mm SL, off Sokotra, preserved; G) N. steatiticus, ZMUC P771752, 162 mm SL, off Myanmar, preserved (G1) and fresh (G2); H1) N. stefanovi, ZMUC P77841, PT, 198 mm SL, Gulf of Aden, preserved; H2) N. stefanovi, SMF 26785, 111 mm SL, Red Sea, fresh. Photographers: Oddgeir Alvheim (G2), Thomas Gloerfelt-Tarp (B2, C2, D2), Marcus Krag (A, B1, C1, D1, E, G1, H1), Ralph Thiel (F), Michael Türkay (H2). The blue bars indicate 1 cm length for the respective specimen." captionText-1="FIGURE 2. Distribution of studied specimens of Neobythites, with numbers indicating multiple sampling events." captionText-2="FIGURE 3. Relationships between SL, morphometric body-shape characters, and number of dorsal-fin rays covered by ocellus spot in Neobythites gloriae n. sp., N. steatiticus, and two populations of N. stefanovi." httpUri-0="https://zenodo.org/record/1186738/files/figure.png" httpUri-1="https://zenodo.org/record/1186740/files/figure.png" httpUri-2="https://zenodo.org/record/1186742/files/figure.png" pageId="1" pageNumber="158">Figures 13</figureCitation>
,
<tableCitation id="A05B56BBFFF18F33FECDE14C34A0505A" box="[307,429,1633,1658]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStartId-0="9.[147,218,545,567]" captionStartId-1="11.[147,221,552,571]" captionStartId-2="13.[151,242,1453,1476]" captionText-0="TABLE 1. Meristic, morphometric, and quantitative ocellus and otolith characters in eight steatiticus-group species.Clear differences from Neobythites gloriae n. sp. are indicated by bold italics." captionText-1="TABLE 2. Meristic, morphometric, and quantitative ocellus characters in populations of Neobythites steatiticus and N. stefanovi." captionText-2="TABLE 3. Distribution of selected meristic characters in the steatiticus-species group, with data for the two populations of Neobythites stefanovi." pageId="1" pageNumber="158">Tables 13</tableCitation>
)
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<taxonomicName id="2AD91883FFF18F33FF69E18F37025098" ID-CoL="46C4M" authority=": Nielsen 1995" authorityName=": Nielsen" authorityYear="1995" box="[151,527,1698,1720]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="1" pageNumber="158" phylum="Chordata" rank="species" species="steatiticus">
<emphasis id="DFADBF12FFF18F33FF69E18F34765098" box="[151,379,1698,1720]" italics="true" pageId="1" pageNumber="158">Neobythites steatiticus</emphasis>
:
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(in part, Arabian Gulf, Gulf of
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<taxonomicName id="2AD91883FFF18F33FF69E1EC377D50F7" ID-CoL="46C4N" authority=": Nielsen &amp; Uiblein 1993" authorityName=": Nielsen &amp; Uiblein" authorityYear="1993" box="[151,624,1729,1751]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="1" pageNumber="158" phylum="Chordata" rank="species" species="stefanovi">
<emphasis id="DFADBF12FFF18F33FF69E1EC347D50F7" box="[151,368,1729,1751]" italics="true" pageId="1" pageNumber="158">Neobythites stefanovi</emphasis>
:
<bibRefCitation id="89481EF1FFF18F33FE83E1EC377D50F7" box="[381,624,1729,1751]" pageId="1" pageNumber="158" refId="ref12492" refString="Nielsen, J. G. &amp; Uiblein F. (1993) Tiefenwasser- und Tiefseefische aus dem Roten Meer. XVI. A new species of Neobythites from the NW Indian Ocean and the Red Sea. Senckenbergiana maritima, 23, 109 - 113." type="journal article">Nielsen &amp; Uiblein 1993</bibRefCitation>
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and
<bibRefCitation id="89481EF1FFF18F33FD5FE1EC362750F7" box="[673,810,1729,1751]" pageId="1" pageNumber="158" refId="ref12268" refString="Nielsen, J. G. (1995) A review of the species of the genus Neobythites (Pisces, Ophidiidae) from the western Indian Ocean with description of seven new species. Ichthyological Bulletin, 62, 1 - 19." type="journal article">Nielsen 1995</bibRefCitation>
,
<bibRefCitation id="89481EF1FFF18F33FCCAE1EC366A50F7" box="[820,871,1729,1751]" pageId="1" pageNumber="158" refId="ref12355" refString="Nielsen, J. G. (2002) Revision of the Indo-Pacific species of Neobythites (Teleostei, Ophidiidae) with 15 new species. Galathea Report, 19, 5 - 105." type="journal article">2002</bibRefCitation>
(in part, Gulf of
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,
<quantity id="2A21CEE5FFF18F33FC9BE04436F751A1" box="[869,1018,1897,1922]" metricMagnitude="-1" metricUnit="m" metricValue="1.385" metricValueMax="1.45" metricValueMin="1.3199999999999998" pageId="1" pageNumber="158" unit="mm" value="138.5" valueMax="145.0" valueMin="132.0">132145 mm</quantity>
SL),
<location id="E80635DBFFF18F33FBC6E04431D551A2" LSID="urn:lsid:plazi:treatment:6570D216FFF18F30FF69E113348451FC:E80635DBFFF18F33FBC6E04431D551A2" box="[1080,1240,1897,1922]" country="Oman" latitude="25.583334" longLatPrecision="1245" longitude="57.783333" name="Gulf of Oman" pageId="1" pageNumber="158">
Gulf of
<collectingCountry id="95CE2390FFF18F33FB6FE04731D551A2" box="[1169,1240,1898,1922]" name="Oman" pageId="1" pageNumber="158">Oman</collectingCountry>
</location>
,
<geoCoordinate id="88ED05C7FFF18F33FB1DE044304451A1" box="[1251,1353,1897,1922]" direction="north" orientation="latitude" pageId="1" pageNumber="158" precision="925" value="25.583334">25°35' N</geoCoordinate>
,
<geoCoordinate id="88ED05C7FFF18F33FAAAE04435A65185" direction="east" orientation="longitude" pageId="1" pageNumber="158" precision="925" value="57.783333">57°47' E</geoCoordinate>
,
<elevation id="66F48433FFF18F33FF45E0A3340B5185" box="[187,262,1934,1958]" metricMagnitude="2" metricUnit="m" metricValue="3.11" pageId="1" pageNumber="158" unit="m" value="311.0">
<quantity id="2A21CEE5FFF18F33FF45E0A3340B5185" box="[187,262,1934,1958]" metricMagnitude="2" metricUnit="m" metricValue="3.11" pageId="1" pageNumber="158" unit="m" value="311.0">311 m</quantity>
</elevation>
depth
</materialsCitation>
;
<materialsCitation id="5DB1695DFFF18F33FE9AE0A3348351EA" ID-GBIF-Occurrence="1826816964" collectionCode="BMHN" country="United Arab Emirates" elevation="26" location="Arabian Gulf" pageId="1" pageNumber="158" specimenCode="BMHN 1910.1" specimenCount="1" specimenCount-female="1" stateProvince="Dubai" typeStatus="paratype">
<specimenCode id="BD7FCB7BFFF18F33FE9AE0A3371F5186" box="[356,530,1934,1958]" collectionCode="BMHN" pageId="1" pageNumber="158">BMHN 1910.1</specimenCode>
.31.23 (
<specimenCount id="FBDFA889FFF18F33FD90E0A337D85186" box="[622,725,1933,1958]" pageId="1" pageNumber="158" type="female">1 female</specimenCount>
,
<quantity id="2A21CEE5FFF18F33FD18E0A336455185" box="[742,840,1934,1958]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="158" unit="mm" value="130.0">130 mm</quantity>
SL),
<location id="E80635DBFFF18F33FC6EE0A031205186" LSID="urn:lsid:plazi:treatment:6570D216FFF18F30FF69E113348451FC:E80635DBFFF18F33FC6EE0A031205186" box="[912,1069,1933,1958]" country="United Arab Emirates" name="Arabian Gulf" pageId="1" pageNumber="158" stateProvince="Dubai">Arabian Gulf</location>
, 10' W of
<collectingRegion id="2F1DADE2FFF18F33FB4AE0A031F65186" box="[1204,1275,1933,1958]" country="United Arab Emirates" name="Dubayy" pageId="1" pageNumber="158">Dubai</collectingRegion>
,
<collectingCountry id="95CE2390FFF18F33FAF2E0A035F351EA" name="United Arab Emirates" pageId="1" pageNumber="158">United Arab Emirates</collectingCountry>
,
<elevation id="66F48433FFF18F33FEF4E09C344951E9" box="[266,324,1969,1994]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="158" unit="m" value="26.0">
<quantity id="2A21CEE5FFF18F33FEF4E09C344951E9" box="[266,324,1969,1994]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="158" unit="m" value="26.0">26 m</quantity>
</elevation>
depth
</materialsCitation>
;
<materialsCitation id="5DB1695DFFF18F33FE64E09F34AF51CE" ID-GBIF-Occurrence="1826817285" collectionCode="ZMUC" country="United Arab Emirates" elevation="26" location="Arabian Gulf" pageId="1" pageNumber="158" specimenCode="USNM 440378" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Dubai" typeStatus="holotype">
<specimenCode id="BD7FCB7BFFF18F33FE64E09F374A51EA" box="[410,583,1970,1994]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="158">USNM 440378</specimenCode>
(
<specimenCount id="FBDFA889FFF18F33FDA4E09F37A551EA" box="[602,680,1969,1994]" pageId="1" pageNumber="158" type="male">1 male</specimenCount>
,
<quantity id="2A21CEE5FFF18F33FD4AE09F361F51E9" box="[692,786,1970,1994]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="158" unit="mm" value="140.0">140 mm</quantity>
SL), and
<collectionCode id="8BC8FBC5FFF18F33FC7AE09F36D851EA" box="[900,981,1970,1994]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="1" pageNumber="158">ZMUC</collectionCode>
P771754 (
<specimenCount id="FBDFA889FFF18F33FBAAE09F31BA51EA" box="[1108,1207,1969,1994]" pageId="1" pageNumber="158" type="female">1 female</specimenCount>
,
<quantity id="2A21CEE5FFF18F33FB3DE09F302F51E9" box="[1219,1314,1970,1994]" metricMagnitude="-1" metricUnit="m" metricValue="1.31" pageId="1" pageNumber="158" unit="mm" value="131.0">131 mm</quantity>
SL), same data as for the
<typeStatus id="3262DDA2FFF18F33FEC2E0F834AF51CE" box="[316,418,2005,2030]" pageId="1" pageNumber="158" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A5C3308BFFF28F30FF39E7BA308A578D" pageId="2" pageNumber="159" type="diagnosis">
<paragraph id="ED666300FFF28F30FF39E7BA308A578D" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="159">
<emphasis id="DFADBF12FFF28F30FF39E7BA344C5690" bold="true" box="[199,321,151,176]" pageId="2" pageNumber="159">Diagnosis.</emphasis>
No spines on hind margin of preopercle; dorsal fin-rays 9093; anal-fin rays 7276; pectoral finrays 2427; precaudal vertebrae 12; total vertebrae 5456; pseudobranchial filaments 34; long gill rakers on anterior arch 1113; head length 25.528.0 % SL; pelvic-fin length 13.517.0 % SL, pelvic fins not reaching beyond anus; orbit length 5.35.9 % SL, 19.523.5 % head length, and 2.02.2 times in upper-jaw length; longest gill filament 3.94.9 % SL and 15.519.0 % head length; ocellus spot placed well posterior to a vertical line through anus, the ocellus-spot distance being 49.051 % SL, and the spot covering 68 dorsal-fin rays; margins of posterior fifth to a quarter of dorsal fin and posterior half to two thirds of anal fin dark pigmented; no vertical bars on body; otolith length 4.85.0 % SL, sulcus length 3.43.7 % SL, and ostium height 21.026.5 % sulcus length.
</paragraph>
</subSubSection>
<subSubSection id="A5C3308BFFF28F30FF39E69A34395310" pageId="2" pageNumber="159" type="description">
<paragraph id="ED666300FFF28F30FF39E69A31C754D0" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="159">
<emphasis id="DFADBF12FFF28F30FF39E69A345F57F0" bold="true" box="[199,338,439,464]" pageId="2" pageNumber="159">Description</emphasis>
(based on
<typeStatus id="3262DDA2FFF28F30FE2EE69A373857F0" box="[464,565,439,464]" pageId="2" pageNumber="159" type="holotype">holotype</typeStatus>
, differences in
<typeStatus id="3262DDA2FFF28F30FD1BE695365D57F0" box="[741,848,440,464]" pageId="2" pageNumber="159" type="paratype">paratypes</typeStatus>
mentioned in round brackets). Rather elongate fish with complete lateral line in preserved specimens; on the anterior half of body, the lateral line runs with 810 scale rows to dorsal edge, decreasing posteriorad; head and body fully covered with cycloid scales; origin of dorsal fin above hind margin of opercle; origin of anal fin slightly in front of midpoint of fish; tip of pectoral fins ending just above origin of anal fin; pelvic fins reaching almost to origin of anal fin; blunt snout slightly shorter than diameter of eye; upper jaw ending well behind eye; anterior nostril ending in a short tube and the larger posterior nostril in a mere hole; no spines on preopercle; strong opercular spine; anterior gill raker with 12 (23) blunt and 34 (23) long rakers on upper branch; one long raker in angle and lower branch with eight long and 6 (7) blunt rakers; gill filaments 1.53.0 (14) times as long as long rakers; distinct pseudobranchial filaments 3 (34).
</paragraph>
<paragraph id="ED666300FFF28F30FF39E5D131A95518" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="159">
<emphasis id="DFADBF12FFF28F30FF39E5D1341B5534" box="[199,278,764,788]" italics="true" pageId="2" pageNumber="159">Otolith</emphasis>
. Otolith oval, its height 1.6 (1.51.7) times in its length; sulcus large, 1.3 (1.31.4) times in otolith length; ostium 1.5 times in sulcus length; ostium height 3.3 (2.63.3) times in ostium length.
</paragraph>
<paragraph id="ED666300FFF28F30FF39E468378955E8" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="159">
<emphasis id="DFADBF12FFF28F30FF39E4683435557C" box="[199,312,837,860]" italics="true" pageId="2" pageNumber="159">Dentition.</emphasis>
Premaxillaries and dentaries with small, pointed teeth in 34 irregular rows anteriorly, decreasing to 12 rows posteriorad. Teeth in outer rows somewhat longer. Vomer triangular with many small, close-set teeth. Palatines with several rows of small, close-set teeth. Two median basibranchial tooth patches, the anterior elongate and the posterior much smaller and circular.
</paragraph>
<paragraph id="ED666300FFF28F30FF39E4F936B45278" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="159">
<emphasis id="DFADBF12FFF28F30FF39E4F9346B55CC" box="[199,358,980,1004]" italics="true" pageId="2" pageNumber="159">Axial skeleton</emphasis>
(from radiographs). Precaudal vertebrae 12 and caudal vertebrae 42 (4245) and all neural and haemal spines pointed; first neural spine one third length of second spine, that is longer than the following spines; bases of neural spines more or less enlarged; parapophyses present on posterior 78 precaudal vertebrae; pleural ribs on vertebrae 311 (
<date id="996745C0FFF28F30FE5EE36D37345278" box="[416,569,1088,1112]" pageId="2" pageNumber="159" value="2012-04-03" valueMax="2012-10-03" valueMin="2012-04-03">34 to 1012</date>
); epipleural ribs on vertebrae 38.
</paragraph>
<paragraph id="ED666300FFF28F30FF39E34934395310" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="159">
<emphasis id="DFADBF12FFF28F30FF39E3493418525C" box="[199,277,1124,1148]" italics="true" pageId="2" pageNumber="159">Colour</emphasis>
. Preserved fish (
<typeStatus id="3262DDA2FFF28F30FE26E3493731525D" box="[472,572,1124,1149]" pageId="2" pageNumber="159" type="holotype">holotype</typeStatus>
[
<figureCitation id="75E27F85FFF28F30FDB2E34937C8525C" box="[588,709,1124,1149]" captionStart="FIGURE 1" captionStartId="3.[151,250,1503,1525]" captionTargetBox="[163,1422,185,1492]" captionTargetId="figure@3.[163,1423,185,1492]" captionTargetPageId="3" captionText="FIGURE 1. Indo-Pacific Neobythites steatiticus species group. All photographs at left side and F show preserved fishes; the remaining five photographs at right side show fresh fishes, photographed on board immediately after capture. A) Neobythites gloriae n. sp. USNM 309008, HT, 158 mm SL, inner Gulf of Oman; B) N. lombokensis, ZMUC P 77744, HT, 93 mm SL, SE off Lombok, preserved (B1) and fresh (B2; side reversed); C) N. longipes, ZMUC P77740, 98 mm SL, SE off Lombok, preserved (C1) and fresh (C2); D) N. malayanus, ZMUC P77742, 138 mm SL, SE off Lombok, preserved (D1) and fresh (D2); E) N. malhaensis, ZMUC P77840, PT, 135 mm SL, Saya de Malha bank, preserved; F) N. meteori, ZMH 5621, HT, 102 mm SL, off Sokotra, preserved; G) N. steatiticus, ZMUC P771752, 162 mm SL, off Myanmar, preserved (G1) and fresh (G2); H1) N. stefanovi, ZMUC P77841, PT, 198 mm SL, Gulf of Aden, preserved; H2) N. stefanovi, SMF 26785, 111 mm SL, Red Sea, fresh. Photographers: Oddgeir Alvheim (G2), Thomas Gloerfelt-Tarp (B2, C2, D2), Marcus Krag (A, B1, C1, D1, E, G1, H1), Ralph Thiel (F), Michael Türkay (H2). The blue bars indicate 1 cm length for the respective specimen." httpUri="https://zenodo.org/record/1186738/files/figure.png" pageId="2" pageNumber="159">Figure 1A</figureCitation>
] and
<typeStatus id="3262DDA2FFF28F30FCF6E3493675525C" box="[776,888,1124,1148]" pageId="2" pageNumber="159" type="paratype">paratypes</typeStatus>
). Body mottled brown, abdomen silvery bluish, lateral line pale brown, rather indistinct; head pale brown with brown preopercular area behind eye, gill cover transparent, eyes bluish; ocellus spot dark brown, its horizontal diameter about orbit length, placed well behind a vertical line through anal-fin origin; dorsal and anal fins with pale- to dark-brown distal margins along posterior fifth to quarter of dorsal fin and posterior half to two-thirds of anal fin; margins becoming darker towards dark caudal-fin tip.
</paragraph>
</subSubSection>
<subSubSection id="A5C3308BFFF28F30FF39E21131F35375" box="[199,1278,1340,1365]" pageId="2" pageNumber="159" type="etymology">
<paragraph id="ED666300FFF28F30FF39E21131F35375" blockId="2.[151,1437,151,2013]" box="[199,1278,1340,1365]" pageId="2" pageNumber="159">
<emphasis id="DFADBF12FFF28F30FF39E211344B5375" bold="true" box="[199,326,1340,1365]" pageId="2" pageNumber="159">Etymology</emphasis>
. The new species name honors the wife of the first author, Gloria Jansen Echevarria.
</paragraph>
</subSubSection>
<subSubSection id="A5C3308BFFF28F30FF39E2723684500C" pageId="2" pageNumber="159" type="distribution">
<paragraph id="ED666300FFF28F30FF39E27234E553BD" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="159">
<emphasis id="DFADBF12FFF28F30FF39E27234545358" bold="true" box="[199,345,1375,1400]" pageId="2" pageNumber="159">Distribution</emphasis>
. Inner Gulf of
<collectingCountry id="95CE2390FFF28F30FDFFE24D37455358" box="[513,584,1376,1400]" name="Oman" pageId="2" pageNumber="159">Oman</collectingCountry>
, NW Indian Ocean, at
<quantity id="2A21CEE5FFF28F30FCB9E24D36C85358" box="[839,965,1376,1400]" metricMagnitude="2" metricUnit="m" metricValue="2.96" metricValueMax="3.2" metricValueMin="2.7199999999999998" pageId="2" pageNumber="159" unit="m" value="296.0" valueMax="320.0" valueMin="272.0">272320 m</quantity>
depth, and southern Arabian Gulf,
<collectingCountry id="95CE2390FFF28F30FAAEE272343153BD" name="United Arab Emirates" pageId="2" pageNumber="159">United Arab Emirates</collectingCountry>
, at
<quantity id="2A21CEE5FFF28F30FE9DE2A9349653BC" box="[355,411,1412,1437]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="159" unit="m" value="26.0">26 m</quantity>
depth.
</paragraph>
<paragraph id="ED666300FFF28F30FF39E28A3684500C" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="159">
<emphasis id="DFADBF12FFF28F30FF39E28A346053E0" bold="true" box="[199,365,1447,1472]" pageId="2" pageNumber="159">Comparisons.</emphasis>
<taxonomicName id="2AD91883FFF28F30FE82E285376D53E0" authority="Uiblein &amp; Nielsen, 2018" authorityName="Uiblein &amp; Nielsen" authorityYear="2018" box="[380,608,1448,1472]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="2" pageNumber="159" phylum="Chordata" rank="species" species="gloriae" status="sp. nov.">
<emphasis id="DFADBF12FFF28F30FE82E285376D53E0" box="[380,608,1448,1472]" italics="true" pageId="2" pageNumber="159">Neobythites gloriae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C49E0269FFF28F30FD91E28A37BA53E0" box="[623,695,1447,1472]" pageId="2" pageNumber="159" rank="species">
<emphasis id="DFADBF12FFF28F30FD91E28A37A253E0" bold="true" box="[623,687,1447,1472]" pageId="2" pageNumber="159">n. sp</emphasis>
.
</taxonomicNameLabel>
differs from all congeners by the following combination of characters: a single ocellus placed on dorsal fin at mid-body, preopercular spines lacking, 1113 developed gill rakers, head length 25.528.0 % SL, gill filament length 3.94.9 % SL, orbit length 19.523.5 % head length and 4550 % upper-jaw length, and otolith sulcus length 3.43.7 % SL.
</paragraph>
</subSubSection>
<subSubSection id="A5C3308BFFF28F30FF39E115309F5124" pageId="2" pageNumber="159" type="description">
<paragraph id="ED666300FFF28F30FF39E115309F5124" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="159">
<taxonomicName id="2AD91883FFF28F30FF39E11534AE5070" authority="Uiblein &amp; Nielsen, 2018" authorityName="Uiblein &amp; Nielsen" authorityYear="2018" box="[199,419,1592,1616]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="2" pageNumber="159" phylum="Chordata" rank="species" species="gloriae" status="sp. nov.">
<emphasis id="DFADBF12FFF28F30FF39E11534AE5070" box="[199,419,1592,1616]" italics="true" pageId="2" pageNumber="159">Neobythites gloriae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C49E0269FFF28F30FE55E11A34E75070" box="[427,490,1591,1616]" pageId="2" pageNumber="159" rank="species">
<emphasis id="DFADBF12FFF28F30FE55E11A34EE5070" bold="true" box="[427,483,1591,1616]" pageId="2" pageNumber="159">n. sp</emphasis>
.
</taxonomicNameLabel>
is most similar to
<taxonomicName id="2AD91883FFF28F30FD42E114364C5070" authorityName=": Nielsen &amp; Uiblein" authorityYear="1993" box="[700,833,1592,1616]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="2" pageNumber="159" phylum="Chordata" rank="species" species="stefanovi">
<emphasis id="DFADBF12FFF28F30FD42E114364C5070" box="[700,833,1592,1616]" italics="true" pageId="2" pageNumber="159">N. stefanovi</emphasis>
</taxonomicName>
, sharing the following characters: dorsal fin with one ocellus, preopercular spine absent, pelvic fins ending anterior to anus, similar number of long rakers on anterior gill arch, eyes five times or less in head length, and similar otolith sulcus length.
<taxonomicName id="2AD91883FFF28F30FBF2E1AD31E550B8" authorityName="Uiblein &amp; Nielsen" authorityYear="2018" box="[1036,1256,1664,1688]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="2" pageNumber="159" phylum="Chordata" rank="species" species="gloriae">
<emphasis id="DFADBF12FFF28F30FBF2E1AD31E550B8" box="[1036,1256,1664,1688]" italics="true" pageId="2" pageNumber="159">Neobythites gloriae</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2AD91883FFF28F30FA7CE1AC35F1509C" authorityName=": Nielsen &amp; Uiblein" authorityYear="1993" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="2" pageNumber="159" phylum="Chordata" rank="species" species="stefanovi">
<emphasis id="DFADBF12FFF28F30FA7CE1AC35F1509C" italics="true" pageId="2" pageNumber="159">N. stefanovi</emphasis>
</taxonomicName>
having longer gill filaments (3.94.9
<emphasis id="DFADBF12FFF28F30FD58E18837C9509C" box="[678,708,1701,1724]" italics="true" pageId="2" pageNumber="159">vs.</emphasis>
1.43.7 % SL) and a slightly smaller ocellus spot (spot covering 68
<emphasis id="DFADBF12FFF28F30FF36E1E435D250C0" box="[200,223,1737,1760]" italics="true" pageId="2" pageNumber="159">vs</emphasis>
. 812 dorsal-fin rays). These two characters in combination with head length, orbit length, and ocellus-spot distance best distinguish
<taxonomicName id="2AD91883FFF28F30FE51E1C0372C5124" authorityName="Uiblein &amp; Nielsen" authorityYear="2018" box="[431,545,1772,1796]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="2" pageNumber="159" phylum="Chordata" rank="species" species="gloriae">
<emphasis id="DFADBF12FFF28F30FE51E1C0372C5124" box="[431,545,1772,1796]" italics="true" pageId="2" pageNumber="159">N. gloriae</emphasis>
</taxonomicName>
from
<taxonomicName id="2AD91883FFF28F30FD9BE1C037F55124" authorityName=": Nielsen" authorityYear="1995" box="[613,760,1773,1796]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="2" pageNumber="159" phylum="Chordata" rank="species" species="steatiticus">
<emphasis id="DFADBF12FFF28F30FD9BE1C037F55124" box="[613,760,1773,1796]" italics="true" pageId="2" pageNumber="159">N. steatiticus</emphasis>
</taxonomicName>
and
<taxonomicName id="2AD91883FFF28F30FCD0E1C036CC5124" authorityName=": Nielsen" authorityYear="1995" box="[814,961,1773,1796]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="2" pageNumber="159" phylum="Chordata" rank="species" species="steatiticus">
<emphasis id="DFADBF12FFF28F30FCD0E1C036CC5124" box="[814,961,1773,1796]" italics="true" pageId="2" pageNumber="159">N. steatiticus</emphasis>
</taxonomicName>
from
<taxonomicName id="2AD91883FFF28F30FBFBE1C031875124" authorityName=": Nielsen &amp; Uiblein" authorityYear="1993" box="[1029,1162,1772,1796]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="2" pageNumber="159" phylum="Chordata" rank="species" species="stefanovi">
<emphasis id="DFADBF12FFF28F30FBFBE1C031875124" box="[1029,1162,1772,1796]" italics="true" pageId="2" pageNumber="159">N. stefanovi</emphasis>
</taxonomicName>
(
<figureCitation id="75E27F85FFF28F30FB67E1C131F65124" box="[1177,1275,1772,1797]" captionStart="FIGURE 3" captionStartId="7.[151,250,1622,1644]" captionTargetBox="[179,1412,223,1588]" captionTargetId="figure@7.[151,1436,195,1601]" captionTargetPageId="7" captionText="FIGURE 3. Relationships between SL, morphometric body-shape characters, and number of dorsal-fin rays covered by ocellus spot in Neobythites gloriae n. sp., N. steatiticus, and two populations of N. stefanovi." httpUri="https://zenodo.org/record/1186742/files/figure.png" pageId="2" pageNumber="159">Figure 3</figureCitation>
;
<tableCitation id="A05B56BBFFF28F30FAF6E1C130895124" box="[1288,1412,1772,1797]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStartId-0="9.[147,218,545,567]" captionStartId-1="11.[147,221,552,571]" captionStartId-2="13.[151,242,1453,1476]" captionText-0="TABLE 1. Meristic, morphometric, and quantitative ocellus and otolith characters in eight steatiticus-group species.Clear differences from Neobythites gloriae n. sp. are indicated by bold italics." captionText-1="TABLE 2. Meristic, morphometric, and quantitative ocellus characters in populations of Neobythites steatiticus and N. stefanovi." captionText-2="TABLE 3. Distribution of selected meristic characters in the steatiticus-species group, with data for the two populations of Neobythites stefanovi." pageId="2" pageNumber="159">Tables 13</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A5C3308BFFF28F30FF39E022348451FC" pageId="2" pageNumber="159" type="discussion">
<paragraph id="ED666300FFF28F30FF39E022348451FC" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="159">
<emphasis id="DFADBF12FFF28F30FF39E02234395108" bold="true" box="[199,308,1807,1832]" pageId="2" pageNumber="159">Remarks</emphasis>
. The description of
<taxonomicName id="2AD91883FFF28F30FDE4E03D37F55108" authority="Uiblein &amp; Nielsen, 2018" authorityName="Uiblein &amp; Nielsen" authorityYear="2018" box="[538,760,1808,1832]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="2" pageNumber="159" phylum="Chordata" rank="species" species="gloriae" status="sp. nov.">
<emphasis id="DFADBF12FFF28F30FDE4E03D37F55108" box="[538,760,1808,1832]" italics="true" pageId="2" pageNumber="159">Neobythites gloriae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C49E0269FFF28F30FCFEE022364D5108" box="[768,832,1807,1832]" pageId="2" pageNumber="159" rank="species">
<emphasis id="DFADBF12FFF28F30FCFEE02236355108" bold="true" box="[768,824,1807,1832]" pageId="2" pageNumber="159">n. sp</emphasis>
.
</taxonomicNameLabel>
is based on three former
<taxonomicName id="2AD91883FFF28F30FB9DE03C31E45108" authorityName=": Nielsen &amp; Uiblein" authorityYear="1993" box="[1123,1257,1808,1832]" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="2" pageNumber="159" phylum="Chordata" rank="species" species="stefanovi">
<emphasis id="DFADBF12FFF28F30FB9DE03C31E45108" box="[1123,1257,1808,1832]" italics="true" pageId="2" pageNumber="159">N. stefanovi</emphasis>
</taxonomicName>
<typeStatus id="3262DDA2FFF28F30FB0CE03C30535109" box="[1266,1374,1809,1833]" pageId="2" pageNumber="159" type="paratype">paratypes</typeStatus>
from the inner Gulf of
<collectingCountry id="95CE2390FFF28F30FEA9E0193491516C" box="[343,412,1844,1868]" name="Oman" pageId="2" pageNumber="159">Oman</collectingCountry>
(
<bibRefCitation id="89481EF1FFF28F30FE54E01937B5516C" box="[426,696,1844,1869]" pageId="2" pageNumber="159" refId="ref12492" refString="Nielsen, J. G. &amp; Uiblein F. (1993) Tiefenwasser- und Tiefseefische aus dem Roten Meer. XVI. A new species of Neobythites from the NW Indian Ocean and the Red Sea. Senckenbergiana maritima, 23, 109 - 113." type="journal article">Nielsen &amp; Uiblein 1993</bibRefCitation>
) and six specimens previously identified as
<taxonomicName id="2AD91883FFF28F30FB5AE01835D65150" authority="(Nielsen 1995)" authorityName="Nielsen" authorityYear="1995" baseAuthorityName="Nielsen" baseAuthorityYear="1995" class="Actinopterygii" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="2" pageNumber="159" phylum="Chordata" rank="species" species="steatiticus">
<emphasis id="DFADBF12FFF28F30FB5AE018303B516C" box="[1188,1334,1845,1868]" italics="true" pageId="2" pageNumber="159">N. steatiticus</emphasis>
(
<bibRefCitation id="89481EF1FFF28F30FABBE01935DE5150" pageId="2" pageNumber="159" refId="ref12268" refString="Nielsen, J. G. (1995) A review of the species of the genus Neobythites (Pisces, Ophidiidae) from the western Indian Ocean with description of seven new species. Ichthyological Bulletin, 62, 1 - 19." type="journal article">Nielsen 1995</bibRefCitation>
)
</taxonomicName>
. Apart from this earlier misidentified material, no additional specimens and no photographs of fresh fish were available for our study. The depth of
<quantity id="2A21CEE5FFF28F30FD4EE05137FD51B4" box="[688,752,1916,1941]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="159" unit="m" value="26.0">26 m</quantity>
reported for the single Arabian Gulf specimen (BMHN 1910.1.31.23), indicated as questionable by
<bibRefCitation id="89481EF1FFF28F30FD6FE08D36305199" box="[657,829,1952,1977]" pageId="2" pageNumber="159" refId="ref12268" refString="Nielsen, J. G. (1995) A review of the species of the genus Neobythites (Pisces, Ophidiidae) from the western Indian Ocean with description of seven new species. Ichthyological Bulletin, 62, 1 - 19." type="journal article">Nielsen (1995)</bibRefCitation>
, is here confirmed based on original data recordings available from NHM.
</paragraph>
</subSubSection>
</treatment>
</document>