treatments-xml/data/4A/41/CC/4A41CC3B077E5BC1905DF44748B003F9.xml

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<mods:title>Systematic revision of the genus Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae)</mods:title>
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<mods:namePart>Dayrat, Benoit</mods:namePart>
<mods:affiliation>Department of Biology, Pennsylvania State University, University Park, PA 16802, USA</mods:affiliation>
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<mods:namePart>Apte, Deepak</mods:namePart>
<mods:affiliation>Bombay Natural History Society, Hornbill House, Opp. Lion Gate, Shaheed Bhagat Singh Road, Mumbai 400 001, Maharashtra, India</mods:affiliation>
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<mods:namePart>Aslam, Sadar</mods:namePart>
<mods:affiliation>Centre of Excellence in Marine Biology, University of Karachi, Karachi 75270, Pakistan</mods:affiliation>
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<mods:namePart>Bourke, Adam</mods:namePart>
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<mods:namePart>Comendador, Joseph</mods:namePart>
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<mods:namePart>Khalil, Munawar</mods:namePart>
<mods:affiliation>Department of Marine Science, Universitas Malikussaleh, Reuleut Main Campus, Kecamatan Muara Batu, North Aceh, Aceh, 24355, Indonesia</mods:affiliation>
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<mods:namePart>Ngo, Xuan Qu ảng</mods:namePart>
<mods:affiliation>Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart>Tan, Siong Kiat</mods:namePart>
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<mods:namePart>Tan, Shau Hwai</mods:namePart>
<mods:affiliation>Centre for Marine and Coastal Studies, Universiti Sains Malaysia, 11800, Minden Penang, Malaysia &amp; Marine Science Laboratory, School of Biological Sciences, Universiti Sains Malaysia, 11800, Minden Penang, Malaysia</mods:affiliation>
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<treatment ID-GBIF-Taxon="168209563" LSID="urn:lsid:plazi:treatment:4A41CC3B077E5BC1905DF44748B003F9" httpUri="http://treatment.plazi.org/id/4A41CC3B077E5BC1905DF44748B003F9" lastPageNumber="1" pageId="0" pageNumber="1">
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Genus 
<taxonomicName LSID="4A41CC3B-077E-5BC1-905D-F44748B003F9" authority="Fleming, 1822 a" authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Peronia Fleming, 1822a</taxonomicName>
</paragraph>
</subSubSection>
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<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Onchis</taxonomicName>
<normalizedToken originalValue="Férussac">Ferussac</normalizedToken>
, 1822: xxxi. Nomen oblitum.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Peronia</taxonomicName>
Fleming, 1822a: 574; 
<bibRefCitation author="Fleming, J" journalOrPublisher="Archibald Constable, Edinburgh" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28230157" refId="B73" refString="Fleming, J, 1822b. The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2). Archibald Constable, Edinburgh, https://www.biodiversitylibrary.org/page/28230157" title="The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2)." url="https://www.biodiversitylibrary.org/page/28230157" year="1822 b">Fleming 1822b</bibRefCitation>
: 463. Nomen protectum.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Peronia</taxonomicName>
Blainville, 1824: 280 [junior homonym of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Peronia</taxonomicName>
Fleming, not a reference of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Peronia</taxonomicName>
Fleming].
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Gistel" authorityYear="1848" class="Gastropoda" family="Onchidiidae" genus="Eudrastus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eudrastus" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Eudrastus</taxonomicName>
Gistel, 1848: x.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Paraperonia</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a: 196.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Scaphis</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a: 203.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Aves" family="Onchidiidae" genus="Lessonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Chordata" rank="genus">Lessonia</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a: 213 [junior homonym of 
<taxonomicName class="Aves" family="Onchidiidae" genus="Lessonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Chordata" rank="genus">Lessonia</taxonomicName>
Swainson, 1832, replaced by 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Lessonina</taxonomicName>
Starobogatov, 1976].
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Quoya</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a: 216.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Lessonina</taxonomicName>
Starobogatov, 1976: 211.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Starobogatov" authorityYear="1976" class="Gastropoda" family="Onchidiidae" genus="Quoyella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Quoyella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Quoyella</taxonomicName>
Starobogatov, 1976: 211 [unnecessary replacement name for 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Quoya</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type species">
<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
: 
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
Cuvier, 1804, by monotypy.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
Fleming: 
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
Cuvier, 1804, by monotypy.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
Blainville: 
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
Blainville, 1824, by original designation.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Gistel" authorityYear="1848" class="Gastropoda" family="Onchidiidae" genus="Eudrastus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eudrastus" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Eudrastus</emphasis>
</taxonomicName>
: 
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
Quoy &amp; Gaimard, 1832, by subsequent designation (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.10914" author="Baker, HB" journalOrPublisher="The Nautilus" pageId="0" pageNumber="1" pagination="85 - 88" refId="B13" refString="Baker, HB, 1938. Nomenclature of Onchidiidae. The Nautilus 51: 85 - 88, DOI: https://doi.org/10.5962/bhl.part.10914" title="Nomenclature of Onchidiidae." url="https://doi.org/10.5962/bhl.part.10914" volume="51" year="1938">Baker 1938</bibRefCitation>
: 86).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia</emphasis>
</taxonomicName>
: 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia gondwanae" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gondwanae">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia gondwanae</emphasis>
</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a, by subsequent designation (
<bibRefCitation author="Starobogatov, YI" journalOrPublisher="Soviet Journal of Marine Biology" pageId="0" pageNumber="1" pagination="206 - 212" refId="B171" refString="Starobogatov, YI, 1976. Composition and taxonomic position of marine pulmonate mollusks. Soviet Journal of Marine Biology 4: 206 - 212" title="Composition and taxonomic position of marine pulmonate mollusks." volume="4" year="1976">Starobogatov 1976</bibRefCitation>
: 211).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Scaphis</emphasis>
</taxonomicName>
: 
<taxonomicName baseAuthorityName="Labbe" baseAuthorityYear="1934" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium astridae" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="astridae">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium astridae</emphasis>
</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934b, by subsequent designation (
<bibRefCitation author="Starobogatov, YI" journalOrPublisher="Soviet Journal of Marine Biology" pageId="0" pageNumber="1" pagination="206 - 212" refId="B171" refString="Starobogatov, YI, 1976. Composition and taxonomic position of marine pulmonate mollusks. Soviet Journal of Marine Biology 4: 206 - 212" title="Composition and taxonomic position of marine pulmonate mollusks." volume="4" year="1976">Starobogatov 1976</bibRefCitation>
: 211).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
: 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya indica" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya indica</emphasis>
</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a, by monotypy.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina</emphasis>
</taxonomicName>
: 
<taxonomicName authorityName="Lesson" authorityYear="1831" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium ferrugineum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ferrugineum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium ferrugineum</emphasis>
</taxonomicName>
Lesson, 1831a, by monotypy.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Starobogatov" authorityYear="1976" class="Gastropoda" family="Onchidiidae" genus="Quoyella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Quoyella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoyella</emphasis>
</taxonomicName>
: 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya indica" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya indica</emphasis>
</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a, by monotypy.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
: After the Greek 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="ὁ">ὁ</normalizedToken>
<normalizedToken originalValue="ὂγκος">ὂγκος</normalizedToken>
</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="1">oncos</emphasis>
, which means mass, or tumor.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
: After 
<normalizedToken originalValue="François">Francois</normalizedToken>
<normalizedToken originalValue="Péron">Peron</normalizedToken>
[1775-1810], zoologist of the Baudin expedition between 1800 and 1803, during which he collected the two slugs (from Mauritius and Timor) which Cuvier described as 
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
in 1804.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Gistel" authorityYear="1848" class="Gastropoda" family="Onchidiidae" genus="Eudrastus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eudrastus" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Eudrastus</emphasis>
</taxonomicName>
: Likely, although for unclear reasons, from the Greek 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="εὖ">εὖ</normalizedToken>
</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="1">eu</emphasis>
, for true, and 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="δραστέoς">δραστέoς</normalizedToken>
</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="1">drasteos</emphasis>
, a verbal adjective which means to be done.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia</emphasis>
</taxonomicName>
: From the Greek 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="παρα">παρα</normalizedToken>
</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="1">para</emphasis>
, meaning beside, and 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Scaphis</emphasis>
</taxonomicName>
: After the Greek 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="ἡ">ἡ</normalizedToken>
<normalizedToken originalValue="σκᾰφίs">σκᾰφίs</normalizedToken>
</emphasis>
, which means small boat (
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a: 202).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
: After the French naturalist Jean 
<normalizedToken originalValue="René">Rene</normalizedToken>
Constant Quoy [1790-1869], a member of two circumnavigations from 1817 to 1820 with captain Freycinet and from 1826 to 1829 with captain Dumont 
<normalizedToken originalValue="d’Urville">d'Urville</normalizedToken>
. Quoy and Joseph Paul Gaimard [1793-1858] described several species of onchidiids based on their collections in the southern seas. 
<taxonomicName authorityName="Starobogatov" authorityYear="1976" class="Gastropoda" family="Onchidiidae" genus="Quoyella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Quoyella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoyella</emphasis>
</taxonomicName>
has the same etymology.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina</emphasis>
</taxonomicName>
: After the French naturalist 
<normalizedToken originalValue="René">Rene</normalizedToken>
<normalizedToken originalValue="Primevère">Primevere</normalizedToken>
Lesson [1794-1849], a member of a circumnavigation from 1822 to 1825 with captain Duperrey. Lesson described several species of onchidiids based on his collections in the southern seas, such as the type species of 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Lesson" authorityYear="1831" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium ferrugineum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ferrugineum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium ferrugineum</emphasis>
</taxonomicName>
, which he collected in West Papua, Indonesia. 
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
invalid name 
<taxonomicName class="Aves" family="Tyrannidae" genus="Lessonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonia" order="Passeriformes" pageId="0" pageNumber="1" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonia</emphasis>
</taxonomicName>
was also dedicated to Lesson.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="gender">
<paragraph pageId="0" pageNumber="1">Gender.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
: Masculine. 
<normalizedToken originalValue="Férussac">Ferussac</normalizedToken>
did not specify the gender of 
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
which he did not combine with any specific name, and even the binomen 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis peronii</emphasis>
</taxonomicName>
, which 
<normalizedToken originalValue="Férussac">Ferussac</normalizedToken>
did not use 
<emphasis italics="true" pageId="0" pageNumber="1">per se</emphasis>
, would not help in that respect. Because 
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
is derived from the masculine Greek noun 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="ὁ">ὁ</normalizedToken>
<normalizedToken originalValue="ὂγκος">ὂγκος</normalizedToken>
</emphasis>
, it is considered to be of masculine gender.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
: Feminine. No gender was specified by Fleming, and the combination 
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
does not help to determine it. Because no gender was originally specified or indicated and because 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
ends in - 
<emphasis italics="true" pageId="0" pageNumber="1">a</emphasis>
, it is treated as a name of feminine gender (
<bibRefCitation author="ICZN" journalOrPublisher="Bulletin of Zoological Nomenclature" pageId="0" pageNumber="1" refId="B97" refString="ICZN, 1999. International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." title="International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." year="1999">ICZN 1999</bibRefCitation>
: Article 30.2.4). Indeed, 
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
, an early combination used by 
<bibRefCitation author="Blainville, HMD de" journalOrPublisher="Levrault, Strasbourg" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/25299228" refId="B23" refString="Blainville, HMD de, 1824. Mollusques. In: Cuvier F (Ed) Dictionnaire des Sciences Naturelles (Vol. 32). F. G. Levrault, Strasbourg, https://www.biodiversitylibrary.org/page/25299228" title="Mollusques. In: Cuvier F (Ed) Dictionnaire des Sciences Naturelles (Vol. 32). F. G." url="https://www.biodiversitylibrary.org/page/25299228" year="1824">Blainville (1824</bibRefCitation>
: 281), shows that 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
has always been treated as a name of feminine gender.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Gistel" authorityYear="1848" class="Gastropoda" family="Onchidiidae" genus="Eudrastus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eudrastus" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Eudrastus</emphasis>
</taxonomicName>
: Masculine. No gender was originally specified or indicated. 
<taxonomicName authorityName="Gistel" authorityYear="1848" class="Gastropoda" family="Onchidiidae" genus="Eudrastus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eudrastus" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Eudrastus</emphasis>
</taxonomicName>
ends in a word derived from a word of variable gender (a verbal adjective) and should be treated as masculine (
<bibRefCitation author="ICZN" journalOrPublisher="Bulletin of Zoological Nomenclature" pageId="0" pageNumber="1" refId="B97" refString="ICZN, 1999. International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." title="International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." year="1999">ICZN 1999</bibRefCitation>
: Article 30.1.4.2).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia</emphasis>
</taxonomicName>
: Feminine. Gender of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Scaphis</emphasis>
</taxonomicName>
: Feminine. The gender was not specified by 
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, but his original combinations 
<taxonomicName lsidName="S. atra" pageId="0" pageNumber="1" rank="species" species="atra">
<emphasis italics="true" pageId="0" pageNumber="1">S. atra</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="S. carbonaria" pageId="0" pageNumber="1" rank="species" species="carbonaria">
<emphasis italics="true" pageId="0" pageNumber="1">S. carbonaria</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="S. lata" pageId="0" pageNumber="1" rank="species" species="lata">
<emphasis italics="true" pageId="0" pageNumber="1">S. lata</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="S. punctata" pageId="0" pageNumber="1" rank="species" species="punctata">
<emphasis italics="true" pageId="0" pageNumber="1">S. punctata</emphasis>
</taxonomicName>
indicate that he treated 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Scaphis</emphasis>
</taxonomicName>
as a name of feminine gender, which is correct since 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Scaphis</emphasis>
</taxonomicName>
is derived from the feminine Greek noun 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="ἡ">ἡ</normalizedToken>
<normalizedToken originalValue="σκᾰφίs">σκᾰφίs</normalizedToken>
</emphasis>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
: Feminine. The gender was not specified by 
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, but his original combination 
<taxonomicName lsidName="Q. indica" pageId="0" pageNumber="1" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="1">Q. indica</emphasis>
</taxonomicName>
indicates that he treated 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
as a name of feminine gender, which is assumed to be the gender of 
<taxonomicName authorityName="Starobogatov" authorityYear="1976" class="Gastropoda" family="Onchidiidae" genus="Quoyella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Quoyella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoyella</emphasis>
</taxonomicName>
as well.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina</emphasis>
</taxonomicName>
: Feminine. The gender was not specified by Starobogatov, and no gender was specified for 
<taxonomicName class="Aves" family="Tyrannidae" genus="Lessonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonia" order="Passeriformes" pageId="0" pageNumber="1" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonia</emphasis>
</taxonomicName>
by 
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
. 
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
original combination 
<taxonomicName class="Aves" family="Tyrannidae" genus="Lessonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonia ferruginea" order="Passeriformes" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="ferruginea">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonia ferruginea</emphasis>
</taxonomicName>
indicates that he treated 
<taxonomicName class="Aves" family="Tyrannidae" genus="Lessonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonia" order="Passeriformes" pageId="0" pageNumber="1" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonia</emphasis>
</taxonomicName>
as a name of feminine gender, which is assumed to be the gender of 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina</emphasis>
</taxonomicName>
as well.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Body not flattened. Dorsal gills present. Dorsal eyes present. No retractable, central papilla present. Eyes at tip of short ocular tentacles. Male opening below right ocular tentacle and to its left. Foot wide. Pneumostome median, on ventral hyponotum. Intestinal loops of types I or V. Rectal gland absent. Accessory penial gland present, with muscular sac. Penis with hooks.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Phylogenetic analyses show that all species of slugs with dorsal gills belong to the same clade (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogenetic relationships between Peronia species based on concatenated mitochondrial COI and 16 S DNA sequences. Numbers by the branches are the bootstrap values (maximum likelihood analysis) and the posterior probabilities (Bayesian analysis). Only the values&gt; 50 % (ML) and&gt; 0.9 (Bayesian) are indicated. Numbers for each individual correspond to unique identifiers for DNA extraction. Information on specimens can be found in the lists of material examined and in Table 2. The color used for each species or mitochondrial unit is the same as the color used in Figs 3 - 6." figureDoi="10.3897/zookeys.972.52853.figure2" httpUri="https://binary.pensoft.net/fig/458300" pageId="0" pageNumber="1">2</figureCitation>
- 
<figureCitation captionStart="Figure 4" captionStartId="F6" captionText="Figure 4. Maximum parsimony consensus tree within Peronia based on concatenated nuclear ITS 2 and 28 S sequences. Numbers by the branches are the bootstrap values. Only the values&gt; 50 % are indicated. Numbers for each individual correspond to unique identifiers for DNA extraction. Information on specimens can be found in the lists of material examined and in Table 2. The color used for each species or mitochondrial unit is the same as the color used in Figs 2, 3, 5, 6." figureDoi="10.3897/zookeys.972.52853.figure4" httpUri="https://binary.pensoft.net/fig/458304" pageId="0" pageNumber="1">4</figureCitation>
). Seven generic names apply to that clade (excluding spelling mistakes, unjustified emendations, replaced names, and 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
Blainville, 1824, a junior homonym of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
Fleming, 1822a). Note that the species name of a type species can be valid (such as 
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
), synonymous (such as 
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
, junior synonym of 
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
), or even a 
<emphasis italics="true" pageId="0" pageNumber="1">nomen dubium</emphasis>
(such as 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya indica" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya indica</emphasis>
</taxonomicName>
). Remarks on the nomenclatural history of the genus 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
follow a chronological order.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">Cuvier (1804)</bibRefCitation>
described the first 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species as 
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
but did not mention the presence of dorsal gills. Nor did he illustrate them. He only described a mantle covered by small warts subdivided in even smaller warts. Dorsal gills are actually present on the dorsum of the type specimen of 
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
from Timor, but they are retracted, as most often seen in preserved specimens. 
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">Cuvier (1804</bibRefCitation>
: 41) also confessed that he would have believed 
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
to be terrestrial, due to its pulmonary cavity &quot;similar to that of reptiles&quot;, but that he regarded it as marine because 
<normalizedToken originalValue="Péron">Peron</normalizedToken>
was certain to have collected it in seawater. But, 
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">Cuvier (1804</bibRefCitation>
: 41) adds: &quot;I think at least that it comes to the surface to open its [pulmonary] hole, and naturally take air to breathe, as do our 
<emphasis italics="true" pageId="0" pageNumber="1">bulines</emphasis>
[
<taxonomicName class="Gastropoda" family="Bulinidae" genus="Bulinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulinus" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Bulinus</emphasis>
</taxonomicName>
] and our 
<emphasis italics="true" pageId="0" pageNumber="1">planorbes</emphasis>
[
<taxonomicName class="Gastropoda" family="Planorbidae" genus="Planorbis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Planorbis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Planorbis</emphasis>
</taxonomicName>
] which, although aquatic, breathe only air.&quot; Actually, 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
slugs hide in crevices at high tide and only come out at low tide.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">Cuvier (1804</bibRefCitation>
: 38) decided to classify his new species in 
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1800.tb00584.x" author="Buchannan, F" journalOrPublisher="Transactions of the Linnean Society of London" pageId="0" pageNumber="1" pagination="132 - 134" refId="B28" refString="Buchannan, F, 1800. An account of the Onchidium, a new genus of the class of vermes, found in Bengal. Transactions of the Linnean Society of London 5: 132 - 134, DOI: https://doi.org/10.1111/j.1096-3642.1800.tb00584.x" title="An account of the Onchidium, a new genus of the class of vermes, found in Bengal." url="https://doi.org/10.1111/j.1096-3642.1800.tb00584.x" volume="5" year="1800">
<normalizedToken originalValue="Buchannan’s">Buchannan's</normalizedToken>
(1800)
</bibRefCitation>
genus 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
because of the &quot;extreme external resemblance&quot; between 
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
and 
<normalizedToken originalValue="Buchannan’s">Buchannan's</normalizedToken>
<taxonomicName lsidName="O. typhae" pageId="0" pageNumber="1" rank="species" species="typhae">
<emphasis italics="true" pageId="0" pageNumber="1">O. typhae</emphasis>
</taxonomicName>
, despite the fact that, according to Buchannan, sexes are separate in 
<taxonomicName lsidName="O. typhae" pageId="0" pageNumber="1" rank="species" species="typhae">
<emphasis italics="true" pageId="0" pageNumber="1">O. typhae</emphasis>
</taxonomicName>
, while 
<normalizedToken originalValue="Cuvier’s">Cuvier's</normalizedToken>
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
is hermaphroditic. In his description of 
<taxonomicName lsidName="O. typhae" pageId="0" pageNumber="1" rank="species" species="typhae">
<emphasis italics="true" pageId="0" pageNumber="1">O. typhae</emphasis>
</taxonomicName>
, 
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1800.tb00584.x" author="Buchannan, F" journalOrPublisher="Transactions of the Linnean Society of London" pageId="0" pageNumber="1" pagination="132 - 134" refId="B28" refString="Buchannan, F, 1800. An account of the Onchidium, a new genus of the class of vermes, found in Bengal. Transactions of the Linnean Society of London 5: 132 - 134, DOI: https://doi.org/10.1111/j.1096-3642.1800.tb00584.x" title="An account of the Onchidium, a new genus of the class of vermes, found in Bengal." url="https://doi.org/10.1111/j.1096-3642.1800.tb00584.x" volume="5" year="1800">Buchannan (1800)</bibRefCitation>
wrote that slugs live in Bengal on leaves of 
<taxonomicName class="Liliopsida" family="Typhaceae" genus="Typha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Typha" order="Poales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Typha</emphasis>
</taxonomicName>
reeds and are &quot;very nearly allied&quot; to 
<taxonomicName class="Gastropoda" family="Limacidae" genus="Limax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limax" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Limax</emphasis>
</taxonomicName>
, suggesting that they are terrestrial, although he did not mention the presence of a pulmonary cavity and did not clearly state whether the slugs were terrestrial or not. At any rate, authors considered that 
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1800.tb00584.x" author="Buchannan, F" journalOrPublisher="Transactions of the Linnean Society of London" pageId="0" pageNumber="1" pagination="132 - 134" refId="B28" refString="Buchannan, F, 1800. An account of the Onchidium, a new genus of the class of vermes, found in Bengal. Transactions of the Linnean Society of London 5: 132 - 134, DOI: https://doi.org/10.1111/j.1096-3642.1800.tb00584.x" title="An account of the Onchidium, a new genus of the class of vermes, found in Bengal." url="https://doi.org/10.1111/j.1096-3642.1800.tb00584.x" volume="5" year="1800">
<normalizedToken originalValue="Buchannan’s">Buchannan's</normalizedToken>
(1800)
</bibRefCitation>
<taxonomicName lsidName="O. typhae" pageId="0" pageNumber="1" rank="species" species="typhae">
<emphasis italics="true" pageId="0" pageNumber="1">O. typhae</emphasis>
</taxonomicName>
was not a marine species and 
<bibRefCitation author="Blainville, HMD de" journalOrPublisher="Journal de Physique, de Chimie, d'Histoire Naturelle et des Arts" pageId="0" pageNumber="1" pagination="437 - 444" refId="B22" refString="Blainville, HMD de, 1817. Memoire sur quelques mollusques pulmobranches. Journal de Physique, de Chimie, d'Histoire Naturelle et des Arts 85: 437 - 444" title="Memoire sur quelques mollusques pulmobranches." volume="85" year="1817">Blainville (1817</bibRefCitation>
: 440) argued that 
<normalizedToken originalValue="Buchannan’s">Buchannan's</normalizedToken>
<taxonomicName lsidName="O. typhae" pageId="0" pageNumber="1" rank="species" species="typhae">
<emphasis italics="true" pageId="0" pageNumber="1">O. typhae</emphasis>
</taxonomicName>
was 
<normalizedToken originalValue="“generically”">&quot;generically&quot;</normalizedToken>
different from 
<normalizedToken originalValue="Cuvier’s">Cuvier's</normalizedToken>
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
, and that 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
should be restricted to 
<taxonomicName lsidName="O. typhae" pageId="0" pageNumber="1" rank="species" species="typhae">
<emphasis italics="true" pageId="0" pageNumber="1">O. typhae</emphasis>
</taxonomicName>
. However, 
<bibRefCitation author="Blainville, HMD de" journalOrPublisher="Journal de Physique, de Chimie, d'Histoire Naturelle et des Arts" pageId="0" pageNumber="1" pagination="437 - 444" refId="B22" refString="Blainville, HMD de, 1817. Memoire sur quelques mollusques pulmobranches. Journal de Physique, de Chimie, d'Histoire Naturelle et des Arts 85: 437 - 444" title="Memoire sur quelques mollusques pulmobranches." volume="85" year="1817">Blainville (1817)</bibRefCitation>
did not propose any new generic name for 
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">
<normalizedToken originalValue="Cuvier’s">Cuvier's</normalizedToken>
(1804)
</bibRefCitation>
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
In his 
<emphasis italics="true" pageId="0" pageNumber="1">
Histoire naturelle 
<normalizedToken originalValue="générale">generale</normalizedToken>
et 
<normalizedToken originalValue="particulière">particuliere</normalizedToken>
des mollusques terrestres et fluviatiles
</emphasis>
, 
<normalizedToken originalValue="Férussac">Ferussac</normalizedToken>
(1819: 80-82) agreed with 
<bibRefCitation author="Blainville, HMD de" journalOrPublisher="Journal de Physique, de Chimie, d'Histoire Naturelle et des Arts" pageId="0" pageNumber="1" pagination="437 - 444" refId="B22" refString="Blainville, HMD de, 1817. Memoire sur quelques mollusques pulmobranches. Journal de Physique, de Chimie, d'Histoire Naturelle et des Arts 85: 437 - 444" title="Memoire sur quelques mollusques pulmobranches." volume="85" year="1817">Blainville (1817)</bibRefCitation>
that 
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">
<normalizedToken originalValue="Cuvier’s">Cuvier's</normalizedToken>
(1804)
</bibRefCitation>
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
was distinct from 
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1800.tb00584.x" author="Buchannan, F" journalOrPublisher="Transactions of the Linnean Society of London" pageId="0" pageNumber="1" pagination="132 - 134" refId="B28" refString="Buchannan, F, 1800. An account of the Onchidium, a new genus of the class of vermes, found in Bengal. Transactions of the Linnean Society of London 5: 132 - 134, DOI: https://doi.org/10.1111/j.1096-3642.1800.tb00584.x" title="An account of the Onchidium, a new genus of the class of vermes, found in Bengal." url="https://doi.org/10.1111/j.1096-3642.1800.tb00584.x" volume="5" year="1800">
<normalizedToken originalValue="Buchannan’s">Buchannan's</normalizedToken>
(1800)
</bibRefCitation>
<taxonomicName lsidName="O. typhae" pageId="0" pageNumber="1" rank="species" species="typhae">
<emphasis italics="true" pageId="0" pageNumber="1">O. typhae</emphasis>
</taxonomicName>
, but, like Blainville, he refrained from creating a new generic name for 
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
. A year later, 
<bibRefCitation author="Fleming, J" editor="Brewster, D" journalOrPublisher="14). William Blackwood, Edinburgh" pageId="0" pageNumber="1" pagination="598 - 635" publicationUrl="https://archive.org/details/edinburghencyclo14edinuoft/page/n6/mode/2up" refId="B71" refString="Fleming, J, 1820. Mollusca. In: Brewster, D, Ed., The Edinburgh Encyclopaedia (Vol. 14). William Blackwood, Edinburgh: 598 - 635, https://archive.org/details/edinburghencyclo14edinuoft/page/n6/mode/2up" title="Mollusca." url="https://archive.org/details/edinburghencyclo14edinuoft/page/n6/mode/2up" volumeTitle="The Edinburgh Encyclopaedia (Vol." year="1820">Fleming (1820</bibRefCitation>
: 616) stated that he thought 
<normalizedToken originalValue="Cuvier’s">Cuvier's</normalizedToken>
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
should probably be classified in a different genus from 
<normalizedToken originalValue="Buchannan’s">Buchannan's</normalizedToken>
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
: &quot;This species [
<taxonomicName authorityName="Buchannan" authorityYear="1800" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium typhae" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="typhae">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium typhae</emphasis>
</taxonomicName>
Buchannan], however, if the description be accurate, differs essentially from the one described by Cuvier [
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
], and would lead us to infer that a new genus would be necessary for the reception of the species of the last-mentioned naturalist.&quot;
</paragraph>
<paragraph pageId="0" pageNumber="1">
The two generic names 
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
were independently created in 1822 for 
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
, respectively by 
<normalizedToken originalValue="Férussac">Ferussac</normalizedToken>
(1822) and 
<bibRefCitation author="Fleming, J" editor="Napier, M" journalOrPublisher="5). Archibald Constable, Edinburgh" pageId="0" pageNumber="1" pagination="567 - 584" publicationUrl="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" refId="B72" refString="Fleming, J, 1822a. Mollusca. In: Napier, M, Ed., Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol. 5). Archibald Constable, Edinburgh: 567 - 584, https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" title="Mollusca." url="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" volumeTitle="Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol." year="1822 a">Fleming (1822a</bibRefCitation>
, 
<bibRefCitation author="Fleming, J" journalOrPublisher="Archibald Constable, Edinburgh" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28230157" refId="B73" refString="Fleming, J, 1822b. The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2). Archibald Constable, Edinburgh, https://www.biodiversitylibrary.org/page/28230157" title="The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2)." url="https://www.biodiversitylibrary.org/page/28230157" year="1822 b">b</bibRefCitation>
) who both follow 
<bibRefCitation author="Blainville, HMD de" journalOrPublisher="Journal de Physique, de Chimie, d'Histoire Naturelle et des Arts" pageId="0" pageNumber="1" pagination="437 - 444" refId="B22" refString="Blainville, HMD de, 1817. Memoire sur quelques mollusques pulmobranches. Journal de Physique, de Chimie, d'Histoire Naturelle et des Arts 85: 437 - 444" title="Memoire sur quelques mollusques pulmobranches." volume="85" year="1817">
<normalizedToken originalValue="Blainville’s">Blainville's</normalizedToken>
(1817)
</bibRefCitation>
argument according to which a marine and hermaphroditic species (
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
) cannot be classified in the same genus as a terrestrial species with separate sexes (
<taxonomicName lsidName="O. typhae" pageId="0" pageNumber="1" rank="species" species="typhae">
<emphasis italics="true" pageId="0" pageNumber="1">O. typhae</emphasis>
</taxonomicName>
). Interestingly, neither 
<normalizedToken originalValue="Férussac">Ferussac</normalizedToken>
(1822) nor 
<bibRefCitation author="Fleming, J" editor="Napier, M" journalOrPublisher="5). Archibald Constable, Edinburgh" pageId="0" pageNumber="1" pagination="567 - 584" publicationUrl="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" refId="B72" refString="Fleming, J, 1822a. Mollusca. In: Napier, M, Ed., Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol. 5). Archibald Constable, Edinburgh: 567 - 584, https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" title="Mollusca." url="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" volumeTitle="Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol." year="1822 a">Fleming (1822a</bibRefCitation>
, 
<bibRefCitation author="Fleming, J" journalOrPublisher="Archibald Constable, Edinburgh" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28230157" refId="B73" refString="Fleming, J, 1822b. The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2). Archibald Constable, Edinburgh, https://www.biodiversitylibrary.org/page/28230157" title="The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2)." url="https://www.biodiversitylibrary.org/page/28230157" year="1822 b">b</bibRefCitation>
) mention dorsal gills (which, again, Cuvier did not mention in the original description of 
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
). Dorsal gills were first illustrated by 
<bibRefCitation author="Savigny, JC" journalOrPublisher="Marine Biodiversity" pageId="0" pageNumber="1" publicationUrl="http://gallica.bnf.fr/ark:/12148/btv1b2300300m" refId="B157" refString="Savigny, JC, 1817. Description de l'Egypte ou Recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'Armee francaise, publie par ordre du Gouvernement. Histoire Naturelle, Planches, Tome Deuxieme. Imprimerie Royale, Paris. [105 pls; gastropods on pls 1-3] http://gallica.bnf.fr/ark:/12148/btv1b2300300m" title="Description de l'Egypte ou Recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'Armee francaise, publie par ordre du Gouvernement. Histoire Naturelle, Planches, Tome Deuxieme. Imprimerie Royale, Paris. [105 pls; gastropods on pls 1 - 3]" url="http://gallica.bnf.fr/ark:/12148/btv1b2300300m" year="1817">Savigny (1817</bibRefCitation>
: pl. 2, fig. 3.5) in the 
<emphasis italics="true" pageId="0" pageNumber="1">
Description de 
<normalizedToken originalValue="l’Egypte">l'Egypte</normalizedToken>
</emphasis>
for slugs from the Red Sea; for a collation, see 
<bibRefCitation author="Baring, T" journalOrPublisher="London Institution, London" pageId="0" pageNumber="1" publicationUrl="https://archive.org/details/b2227411x/page/n2" refId="B14" refString="Baring, T, 1838. A Bibliographical Account and Collation of La description de l'Egypte: presented to the library of the London Institution. London Institution, London, https://archive.org/details/b2227411x/page/n2" title="A Bibliographical Account and Collation of La description de l'Egypte: presented to the library of the London Institution." url="https://archive.org/details/b2227411x/page/n2" year="1838">Baring (1838)</bibRefCitation>
and 
<bibRefCitation author="Sherborn, CD" journalOrPublisher="Wissenschaftliche Resultate. Dritter Band. Landmollusken. CW Kreidel's Verlag, Wiesbaden, 251 - 264 [1880]" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/30987440" refId="B160" refString="Sherborn, CD, 1897. On the dates of the Natural History portion of Savigny's 'Description de l'Egypte'. Proceedings of the Zoological Society of London [1897]: 285-288. https://www.biodiversitylibrary.org/page/30987440" title="On the dates of the Natural History portion of Savigny's ' Description de l'Egypte'. Proceedings of the Zoological Society of London [1897]: 285 - 288." url="https://www.biodiversitylibrary.org/page/30987440" year="1897">Sherborn (1897)</bibRefCitation>
. However, gills remained completely unnoticed because the explanation of 
<normalizedToken originalValue="Savigny’s">Savigny's</normalizedToken>
plate was published nearly ten years later by 
<bibRefCitation author="Audouin, V" editor="Audouin, V" journalOrPublisher="Histoire naturelle. Volume 1, partie 4. Imprimerie Imperiale, Paris" pageId="0" pageNumber="1" pagination="7 - 56" publicationUrl="https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" refId="B9" refString="Audouin, V, 1826. Explication sommaire des planches de mollusques de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny. In: Audouin, V, Ed., Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee francaise. Histoire naturelle. Volume 1, partie 4. Imprimerie Imperiale, Paris: 7 - 56, https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" title="Explication sommaire des planches de mollusques de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny." url="https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" volumeTitle="Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee francaise." year="1826">Audouin (1826</bibRefCitation>
: 18-20).
</paragraph>
<paragraph pageId="0" pageNumber="1">
The exact date of publication of 
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
is 13 April 1822 (when the pages xxv-xlvii were published); a collation for 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.10558" author="Ferussac, AEJPJF d'Audebard de" journalOrPublisher="Natural Product Reports" pageId="0" pageNumber="1" refId="B68" refString="Ferussac, AEJPJF d'Audebard de, 1821-1822. Tableaux systematiques des animaux mollusques classes en familles naturelles, dans lesquels on a etabli la concordance de tous les systemes; suivis d'un Prodrome general pour tous les mollusques terrestres ou fluviatiles, vivants ou fossiles. Arthus Bertrand &amp; Sowerby, Paris &amp; London, 114 + xlviii + 111 + 27 pp. [for a collation see Coan &amp; Kabat (2019)] https://doi.org/10.5962/bhl.title.10558" title="Tableaux systematiques des animaux mollusques classes en familles naturelles, dans lesquels on a etabli la concordance de tous les systemes; suivis d'un Prodrome general pour tous les mollusques terrestres ou fluviatiles, vivants ou fossiles. Arthus Bertrand &amp; Sowerby, Paris &amp; London, 114 + xlviii + 111 + 27 pp. [for a collation see Coan &amp; Kabat (2019)]" url="https://doi.org/10.5962/bhl.title.10558" year="1821 - 1822">
<normalizedToken originalValue="Férussac’s">Ferussac's</normalizedToken>
(1821-1822)
</bibRefCitation>
<taxonomicName genus="Tableaux" lsidName="Tableaux" pageId="0" pageNumber="1" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Tableaux</emphasis>
</taxonomicName>
can be found in 
<bibRefCitation author="Coan, EV" journalOrPublisher="The Veliger" pageId="0" pageNumber="1" publicationUrl="https://ams.wildapricot.org/2400-Years-of-Malacology" refId="B34" refString="Coan, EV, Kabat, AR, 2019. 2,400 years of malacology (16th ed.). American Malacological Society, 1712 + 111 pp. [Annex 1 - Book Collations] + 65 pp. [Annex 2 - Kuester Collation] + 62 pp. [Annex 3 - Journal Collations] https://ams.wildapricot.org/2400-Years-of-Malacology" title="2,400 years of malacology (16 th ed.). American Malacological Society, 1712 + 111 pp. [Annex 1 - Book Collations] + 65 pp. [Annex 2 - Kuester Collation] + 62 pp. [Annex 3 - Journal Collations]" url="https://ams.wildapricot.org/2400-Years-of-Malacology" year="2019">Coan and Kabat (2019)</bibRefCitation>
. 
<normalizedToken originalValue="Férussac">Ferussac</normalizedToken>
(1822: xxxi) clearly distinguishes two genera of onchidiids: &quot;Genre I. Onchide, 
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
; 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Onchidium</taxonomicName>
, Cuvier, Ocken. (Marin.)&quot; and &quot;Genre II. Onchidie, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
, Buchannan, Ocken.&quot; Note that 
<bibRefCitation author="Oken, L" journalOrPublisher="Transactions of the Zoological Society of London" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/59378221" refId="B138" refString="Oken, L, 1815. Lehrbuch der Naturgeschichte. Dritter Theil. Zoologie (Vol. 3). Jena, Bersasser, 850 + xviii pp. https://www.biodiversitylibrary.org/page/59378221" title="Lehrbuch der Naturgeschichte. Dritter Theil. Zoologie (Vol. 3). Jena, Bersasser, 850 + xviii pp." url="https://www.biodiversitylibrary.org/page/59378221" year="1815">Oken (1815</bibRefCitation>
: 307), to which 
<normalizedToken originalValue="Férussac">Ferussac</normalizedToken>
(1822) refers, merely listed 
<taxonomicName authorityName="Buchannan" authorityYear="1800" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium typhae" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="typhae">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium typhae</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
. According to 
<normalizedToken originalValue="Férussac">Ferussac</normalizedToken>
, 
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
clearly refers to 
<normalizedToken originalValue="Cuvier’s">Cuvier's</normalizedToken>
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
, supposedly marine and living underwater, and 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
is restricted to 
<normalizedToken originalValue="Buchannan’s">Buchannan's</normalizedToken>
<taxonomicName authorityName="Buchannan" authorityYear="1800" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium typhae" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="typhae">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium typhae</emphasis>
</taxonomicName>
, thought not to live underwater. In the 
<emphasis italics="true" pageId="0" pageNumber="1">
Tableau 
<normalizedToken originalValue="systématique">systematique</normalizedToken>
de la famille de limaces
</emphasis>
(part of the 16th livraison published on 13 July 1822), 
<normalizedToken originalValue="Férussac">Ferussac</normalizedToken>
(1822: 8) also considered two genera: the &quot;premier genre,&quot; i.e., the first genus ever described, 
<normalizedToken originalValue="Buchannan’s">Buchannan's</normalizedToken>
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
, and another genus, unnamed, with 
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
as type.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
is not etymologically rigorous. The latinization of 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="ὂγκος">ὂγκος</normalizedToken>
</emphasis>
is 
<emphasis italics="true" pageId="0" pageNumber="1">oncos</emphasis>
or 
<emphasis italics="true" pageId="0" pageNumber="1">oncus</emphasis>
, as in the English word oncology. The Greek letter 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="κ">κ</normalizedToken>
</emphasis>
is 
<normalizedToken originalValue="“c”">&quot;c&quot;</normalizedToken>
in Latin, while 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="χ">x</normalizedToken>
</emphasis>
becomes 
<normalizedToken originalValue="“ch.”">&quot;ch.&quot;</normalizedToken>
That 
<normalizedToken originalValue="Férussac">Ferussac</normalizedToken>
used 
<emphasis italics="true" pageId="0" pageNumber="1">onchis</emphasis>
instead of 
<emphasis italics="true" pageId="0" pageNumber="1">oncos</emphasis>
is not surprising, as naturalists often took liberties with the latinization of Greek words. A famous example being the word taxonomy, created as 
<emphasis italics="true" pageId="0" pageNumber="1">taxonomie</emphasis>
by 
<bibRefCitation author="De Candolle, AP" journalOrPublisher="Deterville, Paris" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/27673488" refId="B59" refString="De Candolle, AP, 1813. Theorie elementaire de la Botanique. Deterville, Paris, https://www.biodiversitylibrary.org/page/27673488" title="Theorie elementaire de la Botanique." url="https://www.biodiversitylibrary.org/page/27673488" year="1813">De Candolle (1813</bibRefCitation>
: 19) from the Greek words 
<emphasis italics="true" pageId="0" pageNumber="1">taxis</emphasis>
(arrangement, order) and 
<emphasis italics="true" pageId="0" pageNumber="1">nomos</emphasis>
(law, rule): 
<emphasis italics="true" pageId="0" pageNumber="1">taxis</emphasis>
should have stayed as 
<emphasis italics="true" pageId="0" pageNumber="1">taxi</emphasis>
- to form 
<emphasis italics="true" pageId="0" pageNumber="1">taxinomie</emphasis>
, taxinomy, exactly like in the English word taxidermy (from 
<emphasis italics="true" pageId="0" pageNumber="1">taxis</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="1">dermis</emphasis>
, skin). However, the Code does not require taxon names to be etymologically correct. Therefore, the intentional spelling change of 
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
to 
<taxonomicName class="Liliopsida" family="Dioscoreaceae" genus="Oncus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Oncus" order="Dioscoreales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Oncus</emphasis>
</taxonomicName>
by 
<bibRefCitation author="Agassiz, L" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/33664307" refId="B2" refString="Agassiz, L, 1846. Nomenclatoris Zoologici Index Universalis. Jent et Gassman, Soloduri, viii + 393 pp. [this version of the Index Universalis with two columns of names per page was first published with a printed date of 1846 on the title page although it may have been published only in 1847] https://biodiversitylibrary.org/page/33664307" title="Nomenclatoris Zoologici Index Universalis. Jent et Gassman, Soloduri, viii + 393 pp. [this version of the Index Universalis with two columns of names per page was first published with a printed date of 1846 on the title page although it may have been published only in 1847]" url="https://biodiversitylibrary.org/page/33664307" year="1846">Agassiz (1846</bibRefCitation>
: 259; 
<bibRefCitation author="Agassiz, L" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/1382903" refId="B3" refString="Agassiz, L, 1848. Nomenclatoris Zoologici Index Universalis, Jent et Gassman, Soloduri, x + 1135 pp. [this version of the Index Universalis with one column of names per page was published with a printed date of 1848] https://biodiversitylibrary.org/page/1382903" title="Nomenclatoris Zoologici Index Universalis, Jent et Gassman, Soloduri, x + 1135 pp. [this version of the Index Universalis with one column of names per page was published with a printed date of 1848]" url="https://biodiversitylibrary.org/page/1382903" year="1848">1848</bibRefCitation>
: 748) is an unjustified emendation because 
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
is not the result of &quot;inadvertent error, such as a lapsus calami or a 
<normalizedToken originalValue="copyist’s">copyist's</normalizedToken>
or 
<normalizedToken originalValue="printer’s">printer's</normalizedToken>
error&quot; (
<bibRefCitation author="ICZN" journalOrPublisher="Bulletin of Zoological Nomenclature" pageId="0" pageNumber="1" refId="B97" refString="ICZN, 1999. International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." title="International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." year="1999">ICZN 1999</bibRefCitation>
: Article 32.5.1) and therefore 
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
must not be corrected. The emendation of 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
into 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Oncidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oncidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Oncidium</emphasis>
</taxonomicName>
by 
<bibRefCitation author="Agassiz, L" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/33664307" refId="B2" refString="Agassiz, L, 1846. Nomenclatoris Zoologici Index Universalis. Jent et Gassman, Soloduri, viii + 393 pp. [this version of the Index Universalis with two columns of names per page was first published with a printed date of 1846 on the title page although it may have been published only in 1847] https://biodiversitylibrary.org/page/33664307" title="Nomenclatoris Zoologici Index Universalis. Jent et Gassman, Soloduri, viii + 393 pp. [this version of the Index Universalis with two columns of names per page was first published with a printed date of 1846 on the title page although it may have been published only in 1847]" url="https://biodiversitylibrary.org/page/33664307" year="1846">Agassiz (1846</bibRefCitation>
: 259; 
<bibRefCitation author="Agassiz, L" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/1382903" refId="B3" refString="Agassiz, L, 1848. Nomenclatoris Zoologici Index Universalis, Jent et Gassman, Soloduri, x + 1135 pp. [this version of the Index Universalis with one column of names per page was published with a printed date of 1848] https://biodiversitylibrary.org/page/1382903" title="Nomenclatoris Zoologici Index Universalis, Jent et Gassman, Soloduri, x + 1135 pp. [this version of the Index Universalis with one column of names per page was published with a printed date of 1848]" url="https://biodiversitylibrary.org/page/1382903" year="1848">1848</bibRefCitation>
: 748) also is unjustified for the same reason.
</paragraph>
<paragraph pageId="0" pageNumber="1">
The generic name 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
first appeared in two different venues, both published by Fleming (1822). One venue is 
<bibRefCitation author="Fleming, J" editor="Napier, M" journalOrPublisher="5). Archibald Constable, Edinburgh" pageId="0" pageNumber="1" pagination="567 - 584" publicationUrl="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" refId="B72" refString="Fleming, J, 1822a. Mollusca. In: Napier, M, Ed., Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol. 5). Archibald Constable, Edinburgh: 567 - 584, https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" title="Mollusca." url="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" volumeTitle="Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol." year="1822 a">
<normalizedToken originalValue="Fleming’s">Fleming's</normalizedToken>
(1822a
</bibRefCitation>
: 574) article &quot; 
<taxonomicName authorityName="Fleming" authorityYear="1822" lsidName="" pageId="0" pageNumber="1" phylum="Mollusca" rank="phylum">Mollusca</taxonomicName>
&quot; in the fifth volume (second part) of the 
<emphasis italics="true" pageId="0" pageNumber="1">Supplement to the fourth</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="1">fifth</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="1">
and sixth editions of the 
<normalizedToken originalValue="Encyclopædia">Encyclopaedia</normalizedToken>
Britannica
</emphasis>
published in May 1822 (as clearly indicated in a memorandum at the end of the sixth volume of the 
<emphasis italics="true" pageId="0" pageNumber="1">Supplement</emphasis>
), even though the 
<emphasis italics="true" pageId="0" pageNumber="1">Supplement</emphasis>
was only completed in 1824 (date on the title page). The other venue is 
<bibRefCitation author="Fleming, J" journalOrPublisher="Archibald Constable, Edinburgh" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28230157" refId="B73" refString="Fleming, J, 1822b. The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2). Archibald Constable, Edinburgh, https://www.biodiversitylibrary.org/page/28230157" title="The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2)." url="https://www.biodiversitylibrary.org/page/28230157" year="1822 b">
<normalizedToken originalValue="Fleming’s">Fleming's</normalizedToken>
(1822b
</bibRefCitation>
: 463) 
<emphasis italics="true" pageId="0" pageNumber="1">Philosophy of Zoology</emphasis>
which, according to 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.25768" author="Feuer, RC" journalOrPublisher="Great Basin Naturalist" pageId="0" pageNumber="1" pagination="55 - 60" refId="B69" refString="Feuer, RC, Smith, HM, 1972. The contributions of the 1822 works of Jarocki and Fleming to herpetological nomenclature. Great Basin Naturalist 32: 55 - 60, DOI: https://doi.org/10.5962/bhl.part.25768" title="The contributions of the 1822 works of Jarocki and Fleming to herpetological nomenclature." url="https://doi.org/10.5962/bhl.part.25768" volume="32" year="1972">Feuer and Smith (1972</bibRefCitation>
: 55), was not published earlier than May 1822 but no later than June 1822. The mention of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
in the 
<emphasis italics="true" pageId="0" pageNumber="1">Supplement</emphasis>
is considered here to be the earliest one because it was published in May 1822.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
Fleming, 1822a is an objective junior synonym of 
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Férussac">Ferussac</normalizedToken>
, 1822, because 
<normalizedToken originalValue="Férussac’s">Ferussac's</normalizedToken>
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
was published prior to 
<normalizedToken originalValue="Fleming’s">Fleming's</normalizedToken>
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
and both generic names share the same type species (
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
). However, to the best of our knowledge, 
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
has only been used twice in a binomen, and both times before 1899: by 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.51444" author="Stimpson, W" journalOrPublisher="Proceedings of the Academy of Natural Sciences, Philadelphia" pageId="0" pageNumber="1" pagination="375 - 384" refId="B172" refString="Stimpson, W, 1855. Descriptions of some of the new Marine Invertebrata from the Chinese and Japanese Seas. Proceedings of the Academy of Natural Sciences, Philadelphia 7: 375 - 384, DOI: https://doi.org/10.5962/bhl.title.51444" title="Descriptions of some of the new Marine Invertebrata from the Chinese and Japanese Seas." url="https://doi.org/10.5962/bhl.title.51444" volume="7" year="1855">Stimpson (1855)</bibRefCitation>
for 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis fruticosa" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fruticosa">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis fruticosa</emphasis>
</taxonomicName>
, a species name that has remained unnoticed until now, and by 
<bibRefCitation author="Moerch, OAL" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="21 - 43" publicationUrl="http://biodiversitylibrary.org/page/15338052" refId="B129" refString="Moerch, OAL, 1863. Contributions la faune malacologique des Antilles danoises. Journal de Conchyliologie 11: 21 - 43, http://biodiversitylibrary.org/page/15338052" title="Contributions la faune malacologique des Antilles danoises." url="http://biodiversitylibrary.org/page/15338052" volume="11" year="1863">
<normalizedToken originalValue="Mörch">Moerch</normalizedToken>
(1863)
</bibRefCitation>
for 
<taxonomicName authorityName="Morch" authorityYear="1863" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis (Peronella) armadilla" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="armadilla" subGenus="Peronella">Onchis (Peronella) armadilla</taxonomicName>
<normalizedToken originalValue="Mörch">Moerch</normalizedToken>
, 1863, i.e., 
<taxonomicName baseAuthorityName="Mörch" baseAuthorityYear="1863" class="Gastropoda" family="Onchidiidae" genus="Onchidella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidella armadilla" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="armadilla">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidella armadilla</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Mörch">Moerch</normalizedToken>
, 1863). Reversal of precedence applies here (
<bibRefCitation author="ICZN" journalOrPublisher="Bulletin of Zoological Nomenclature" pageId="0" pageNumber="1" refId="B97" refString="ICZN, 1999. International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." title="International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." year="1999">ICZN 1999</bibRefCitation>
: Article 23.9). 
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
, the senior synonym, &quot;has not been used as a valid name after 1899&quot; (
<bibRefCitation author="ICZN" journalOrPublisher="Bulletin of Zoological Nomenclature" pageId="0" pageNumber="1" refId="B97" refString="ICZN, 1999. International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." title="International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." year="1999">ICZN 1999</bibRefCitation>
: Article 23.9.1.1) and 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
, the junior synonym, &quot;has been used for a particular taxon, as its presumed valid name, in at least 25 works, published by at least 10 authors in the immediately preceding 50 years and encompassing a span of not less than 10 years.&quot; (
<bibRefCitation author="ICZN" journalOrPublisher="Bulletin of Zoological Nomenclature" pageId="0" pageNumber="1" refId="B97" refString="ICZN, 1999. International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." title="International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." year="1999">ICZN 1999</bibRefCitation>
: Article 23.9.1.2) A chronological list of 25 works meeting the criteria of ICZN Article 23.9.1.2 is provided here, all of which mentioning 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
, 
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia verruculata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia verruculata</emphasis>
</taxonomicName>
, or 
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
as valid names: 
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, 
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, 
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, 
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, 
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, 
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, 
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, 
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, 
<bibRefCitation author="Solanki, D" journalOrPublisher="The Journal of Zoology Studies" pageId="0" pageNumber="1" pagination="24 - 28" publicationUrl="https://www.journalofzoology.com/archives/volume4/issue3/PartA/6.1.pdf" refId="B168" refString="Solanki, D, Kanejiya, J, Gohil, B, 2017. Studies on ecological status, nutritive values and exploitation of Peronia verruculata, Cuvier, 1830 (Gastropoda: Onchidiidae) from Gulf of Khambhat, India. The Journal of Zoology Studies 4: 24 - 28, https://www.journalofzoology.com/archives/volume4/issue3/PartA/6.1.pdf" title="Studies on ecological status, nutritive values and exploitation of Peronia verruculata, Cuvier, 1830 (Gastropoda: Onchidiidae) from Gulf of Khambhat, India." url="https://www.journalofzoology.com/archives/volume4/issue3/PartA/6.1.pdf" volume="4" year="2017">Solanki et al. (2017)</bibRefCitation>
, 
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. 
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Férussac">Ferussac</normalizedToken>
, 1822, objective senior synonym, is regarded as a 
<emphasis italics="true" pageId="0" pageNumber="1">nomen oblitum</emphasis>
, and 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
Fleming, 1822a, objective junior synonym, is regarded as a 
<emphasis italics="true" pageId="0" pageNumber="1">nomen protectum</emphasis>
(
<bibRefCitation author="ICZN" journalOrPublisher="Bulletin of Zoological Nomenclature" pageId="0" pageNumber="1" refId="B97" refString="ICZN, 1999. International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." title="International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." year="1999">ICZN 1999</bibRefCitation>
: Article 23.9.1.2).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Fleming, J" editor="Napier, M" journalOrPublisher="5). Archibald Constable, Edinburgh" pageId="0" pageNumber="1" pagination="567 - 584" publicationUrl="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" refId="B72" refString="Fleming, J, 1822a. Mollusca. In: Napier, M, Ed., Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol. 5). Archibald Constable, Edinburgh: 567 - 584, https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" title="Mollusca." url="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" volumeTitle="Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol." year="1822 a">Fleming (1822a</bibRefCitation>
: 571, 574) classified 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
(with only the type species 
<taxonomicName lsidName="O. typhae" pageId="0" pageNumber="1" rank="species" species="typhae">
<emphasis italics="true" pageId="0" pageNumber="1">O. typhae</emphasis>
</taxonomicName>
) in a group of slugs that &quot;reside constantly on the land,&quot; and transferred 
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
to 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
, a genus for marine slugs that have &quot;their residence constantly in water&quot; and look like 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
. However, 
<bibRefCitation author="Fleming, J" editor="Napier, M" journalOrPublisher="5). Archibald Constable, Edinburgh" pageId="0" pageNumber="1" pagination="567 - 584" publicationUrl="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" refId="B72" refString="Fleming, J, 1822a. Mollusca. In: Napier, M, Ed., Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol. 5). Archibald Constable, Edinburgh: 567 - 584, https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" title="Mollusca." url="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" volumeTitle="Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol." year="1822 a">Fleming (1822a</bibRefCitation>
: 574) expressed doubts that 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
slugs are air-breathing, as 
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">Cuvier (1804)</bibRefCitation>
claimed in the original description of 
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
:
</paragraph>
<paragraph pageId="0" pageNumber="1">
&quot;This genus, which we have named in honor of M. Peron, was referred by Cuvier to the 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Onchidium</taxonomicName>
of Buchanan (
<normalizedToken originalValue="…">...</normalizedToken>
) and the species termed 
<taxonomicName lsidName="O. Peronii" pageId="0" pageNumber="1" rank="species" species="Peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. Peronii</emphasis>
</taxonomicName>
. It was found creeping upon marine rocks, under water, at the Mauritius, by M. Peron. M. Cuvier conjectures that it breathes free air, and has accordingly inserted it among the 
<taxonomicName genus="Pulmones" lsidName="Pulmones aquatique" pageId="0" pageNumber="1" rank="species" species="aquatique">
<emphasis italics="true" pageId="0" pageNumber="1">Pulmones aquatique</emphasis>
</taxonomicName>
[
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="Pulmonés">Pulmones</normalizedToken>
aquatiques
</emphasis>
, i.e., aquatic pulmonates]. Some doubts, however, may reasonably be entertained about the truth of this supposition. It would certainly be an unexpected occurrence to find a marine gasteropodous 
<taxonomicName lsidName="" pageId="0" pageNumber="1" phylum="Mollusca" rank="phylum">mollusca</taxonomicName>
obliged to come to the surface at intervals to respire. It will probably be found that it is truly branchiferous.&quot;
</paragraph>
<paragraph pageId="0" pageNumber="1">
It was 
<bibRefCitation author="Audouin, V" editor="Audouin, V" journalOrPublisher="Histoire naturelle. Volume 1, partie 4. Imprimerie Imperiale, Paris" pageId="0" pageNumber="1" pagination="7 - 56" publicationUrl="https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" refId="B9" refString="Audouin, V, 1826. Explication sommaire des planches de mollusques de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny. In: Audouin, V, Ed., Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee francaise. Histoire naturelle. Volume 1, partie 4. Imprimerie Imperiale, Paris: 7 - 56, https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" title="Explication sommaire des planches de mollusques de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny." url="https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" volumeTitle="Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee francaise." year="1826">Audouin (1826)</bibRefCitation>
who demonstrated later that both Cuvier and Fleming were correct because 
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
can breathe through both its pulmonary cavity and dorsal gills.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Blainville, HMD de" journalOrPublisher="Levrault, Strasbourg" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/25299228" refId="B23" refString="Blainville, HMD de, 1824. Mollusques. In: Cuvier F (Ed) Dictionnaire des Sciences Naturelles (Vol. 32). F. G. Levrault, Strasbourg, https://www.biodiversitylibrary.org/page/25299228" title="Mollusques. In: Cuvier F (Ed) Dictionnaire des Sciences Naturelles (Vol. 32). F. G." url="https://www.biodiversitylibrary.org/page/25299228" year="1824">Blainville (1824</bibRefCitation>
: 280) created the generic name 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
without being aware that 
<bibRefCitation author="Fleming, J" editor="Napier, M" journalOrPublisher="5). Archibald Constable, Edinburgh" pageId="0" pageNumber="1" pagination="567 - 584" publicationUrl="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" refId="B72" refString="Fleming, J, 1822a. Mollusca. In: Napier, M, Ed., Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol. 5). Archibald Constable, Edinburgh: 567 - 584, https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" title="Mollusca." url="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" volumeTitle="Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol." year="1822 a">Fleming (1822a</bibRefCitation>
, 
<bibRefCitation author="Fleming, J" journalOrPublisher="Archibald Constable, Edinburgh" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28230157" refId="B73" refString="Fleming, J, 1822b. The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2). Archibald Constable, Edinburgh, https://www.biodiversitylibrary.org/page/28230157" title="The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2)." url="https://www.biodiversitylibrary.org/page/28230157" year="1822 b">b</bibRefCitation>
) had already created exactly the same name two years before. Indeed, that 
<bibRefCitation author="Blainville, HMD de" journalOrPublisher="Levrault, Strasbourg" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/25299228" refId="B23" refString="Blainville, HMD de, 1824. Mollusques. In: Cuvier F (Ed) Dictionnaire des Sciences Naturelles (Vol. 32). F. G. Levrault, Strasbourg, https://www.biodiversitylibrary.org/page/25299228" title="Mollusques. In: Cuvier F (Ed) Dictionnaire des Sciences Naturelles (Vol. 32). F. G." url="https://www.biodiversitylibrary.org/page/25299228" year="1824">Blainville (1824</bibRefCitation>
: 258) wrote &quot;our genus 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="Péronie">Peronie</normalizedToken>
</emphasis>
&quot; clearly suggests that he thought he was the author of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
. Also, most past authors attributed the authorship of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
to Blainville instead of Fleming (e.g., 
<bibRefCitation author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" pagination="86 - 111" publicationUrl="https://biodiversitylibrary.org/page/41279271" refId="B173" refString="Stoliczka, F, 1869. The malacology of Lower Bengal. Journal of the Asiatic Society of Bengal 38: 86 - 111, https://biodiversitylibrary.org/page/41279271" title="The malacology of Lower Bengal." url="https://biodiversitylibrary.org/page/41279271" volume="38" year="1869">Stoliczka 1869</bibRefCitation>
: 100; 
<bibRefCitation author="Plate, L" journalOrPublisher="Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere" pageId="0" pageNumber="1" pagination="93 - 234" publicationUrl="https://biodiversitylibrary.org/page/11175213" refId="B146" refString="Plate, L, 1893. Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere 7: 93 - 234, https://biodiversitylibrary.org/page/11175213" title="Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden." url="https://biodiversitylibrary.org/page/11175213" volume="7" year="1893">Plate 1893</bibRefCitation>
: 102; 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
1934a
</bibRefCitation>
: 189). 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
Blainville, 1824 is a junior homonym of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
Fleming, 1822a and thus cannot be used as a valid name (
<bibRefCitation author="ICZN" journalOrPublisher="Bulletin of Zoological Nomenclature" pageId="0" pageNumber="1" refId="B97" refString="ICZN, 1999. International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." title="International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." year="1999">ICZN 1999</bibRefCitation>
: Article 52.2). However, 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
Blainville is also a junior objective synonym of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
Fleming, because they &quot;both denote nominal taxa with name-bearing types whose own names are themselves objectively synonymous.&quot; (
<bibRefCitation author="ICZN" journalOrPublisher="Bulletin of Zoological Nomenclature" pageId="0" pageNumber="1" refId="B97" refString="ICZN, 1999. International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." title="International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." year="1999">ICZN 1999</bibRefCitation>
: &quot;objective synonym&quot; in the glossary) Indeed, 
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
, the type species of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
Fleming, and 
<taxonomicName lsidName="P. mauritiana" pageId="0" pageNumber="1" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">P. mauritiana</emphasis>
</taxonomicName>
, the type species of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
Blainville are objective synonyms because they share the same lectotype, i.e., the specimen from Mauritius which 
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">Cuvier (1804</bibRefCitation>
: pl. 6) illustrated (see below, the comments on the type material of 
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="P. mauritiana" pageId="0" pageNumber="1" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">P. mauritiana</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
When he created the generic name 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
, 
<bibRefCitation author="Blainville, HMD de" journalOrPublisher="Levrault, Strasbourg" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/25299228" refId="B23" refString="Blainville, HMD de, 1824. Mollusques. In: Cuvier F (Ed) Dictionnaire des Sciences Naturelles (Vol. 32). F. G. Levrault, Strasbourg, https://www.biodiversitylibrary.org/page/25299228" title="Mollusques. In: Cuvier F (Ed) Dictionnaire des Sciences Naturelles (Vol. 32). F. G." url="https://www.biodiversitylibrary.org/page/25299228" year="1824">Blainville (1824</bibRefCitation>
: 280, 281) cited only one species name, 
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
, a junior objective synonym of 
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
. 
<bibRefCitation author="Blainville, HMD de" journalOrPublisher="Levrault, Strasbourg" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/25299228" refId="B23" refString="Blainville, HMD de, 1824. Mollusques. In: Cuvier F (Ed) Dictionnaire des Sciences Naturelles (Vol. 32). F. G. Levrault, Strasbourg, https://www.biodiversitylibrary.org/page/25299228" title="Mollusques. In: Cuvier F (Ed) Dictionnaire des Sciences Naturelles (Vol. 32). F. G." url="https://www.biodiversitylibrary.org/page/25299228" year="1824">Blainville (1824</bibRefCitation>
: 281) also claimed that he knew four or five other species of marine onchidiids from the southern hemisphere, without naming them, but 
<bibRefCitation author="Blainville, HMD de" editor="Cuvier, F" journalOrPublisher="38). F. G. Levrault, Strasbourg" pageId="0" pageNumber="1" pagination="519 - 524" publicationUrl="https://www.biodiversitylibrary.org/page/25302418" refId="B24" refString="Blainville, HMD de, 1826. Peronie. In: Cuvier, F, Ed., Dictionnaire des sciences naturelles (Vol. 38). F. G. Levrault, Strasbourg: 519 - 524, https://www.biodiversitylibrary.org/page/25302418" title="Peronie." url="https://www.biodiversitylibrary.org/page/25302418" volumeTitle="Dictionnaire des sciences naturelles (Vol." year="1826">Blainville (1826</bibRefCitation>
: 523) listed them two years later (Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Alphabetic list of the 51 existing species-group names of which the nomenclatural status is addressed in the present work. Details can be found in the text: comments on the four valid Peronia species names, their synonyms, and the junior homonym are in the species remarks; comments on the fifteen nomina dubia and the ten names that must be classified in other genera are in the general discussion." httpUri="http://table.plazi.org/id/F0EDB18E2E41A7D8B1806BBE53F2616C" pageId="0" pageNumber="1" tableUuid="F0EDB18E2E41A7D8B1806BBE53F2616C">1</tableCitation>
): 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia laevis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="laevis">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia laevis</emphasis>
</taxonomicName>
, a junior objective synonym of 
<taxonomicName baseAuthorityName="Quoy &amp; Gaimard" baseAuthorityYear="1825" class="Gastropoda" family="Onchidiidae" genus="Marmaronchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Marmaronchis vaigiensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="vaigiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Marmaronchis vaigiensis</emphasis>
</taxonomicName>
; 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia semituberculata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="semituberculata">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia semituberculata</emphasis>
</taxonomicName>
, a junior objective synonym of 
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium planatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="planatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium planatum</emphasis>
</taxonomicName>
, itself a 
<emphasis italics="true" pageId="0" pageNumber="1">nomen dubium</emphasis>
which may or may not refer to an onchidiid species; 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia oniscoides" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="oniscoides">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia oniscoides</emphasis>
</taxonomicName>
, which all authors ignored except for 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 243) and which clearly does not refer to a 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species (see general discussion). In addition, 
<bibRefCitation author="Blainville, HMD de" editor="Cuvier, F" journalOrPublisher="38). F. G. Levrault, Strasbourg" pageId="0" pageNumber="1" pagination="519 - 524" publicationUrl="https://www.biodiversitylibrary.org/page/25302418" refId="B24" refString="Blainville, HMD de, 1826. Peronie. In: Cuvier, F, Ed., Dictionnaire des sciences naturelles (Vol. 38). F. G. Levrault, Strasbourg: 519 - 524, https://www.biodiversitylibrary.org/page/25302418" title="Peronie." url="https://www.biodiversitylibrary.org/page/25302418" volumeTitle="Dictionnaire des sciences naturelles (Vol." year="1826">Blainville (1826</bibRefCitation>
: 523) also pointed out that 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium celticum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="celticum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium celticum</emphasis>
</taxonomicName>
, a name which Cuvier used for small marine slugs from the coast of Brittany, France, could also refer to a 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
; 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium celticum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="celticum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium celticum</emphasis>
</taxonomicName>
remained a 
<emphasis italics="true" pageId="0" pageNumber="1">nomen nudum</emphasis>
until 1832, when it was described by 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.43796" author="Audouin, V" journalOrPublisher="Crochard, Paris" pageId="0" pageNumber="1" refId="B10" refString="Audouin, V, Milne-Edwards, H, 1832. Recherches pour servir a l'histoire naturelle du littoral de la France. Introduction. (Vol. 1). Crochard, Paris, DOI: https://doi.org/10.5962/bhl.title.43796" title="Recherches pour servir a l'histoire naturelle du littoral de la France. Introduction. (Vol. 1)." url="https://doi.org/10.5962/bhl.title.43796" year="1832">Audouin and Milne-Edwards (1832</bibRefCitation>
: 118).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Like 
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">Cuvier (1804)</bibRefCitation>
, 
<normalizedToken originalValue="Férussac">Ferussac</normalizedToken>
(1822), and 
<bibRefCitation author="Fleming, J" editor="Napier, M" journalOrPublisher="5). Archibald Constable, Edinburgh" pageId="0" pageNumber="1" pagination="567 - 584" publicationUrl="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" refId="B72" refString="Fleming, J, 1822a. Mollusca. In: Napier, M, Ed., Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol. 5). Archibald Constable, Edinburgh: 567 - 584, https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" title="Mollusca." url="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" volumeTitle="Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol." year="1822 a">Fleming (1822a</bibRefCitation>
, 
<bibRefCitation author="Fleming, J" journalOrPublisher="Archibald Constable, Edinburgh" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28230157" refId="B73" refString="Fleming, J, 1822b. The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2). Archibald Constable, Edinburgh, https://www.biodiversitylibrary.org/page/28230157" title="The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2)." url="https://www.biodiversitylibrary.org/page/28230157" year="1822 b">b</bibRefCitation>
), 
<bibRefCitation author="Blainville, HMD de" journalOrPublisher="Levrault, Strasbourg" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/25299228" refId="B23" refString="Blainville, HMD de, 1824. Mollusques. In: Cuvier F (Ed) Dictionnaire des Sciences Naturelles (Vol. 32). F. G. Levrault, Strasbourg, https://www.biodiversitylibrary.org/page/25299228" title="Mollusques. In: Cuvier F (Ed) Dictionnaire des Sciences Naturelles (Vol. 32). F. G." url="https://www.biodiversitylibrary.org/page/25299228" year="1824">Blainville (1824)</bibRefCitation>
did not mention the existence of dorsal gills. Dorsal gills were first described by 
<bibRefCitation author="Audouin, V" editor="Audouin, V" journalOrPublisher="Histoire naturelle. Volume 1, partie 4. Imprimerie Imperiale, Paris" pageId="0" pageNumber="1" pagination="7 - 56" publicationUrl="https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" refId="B9" refString="Audouin, V, 1826. Explication sommaire des planches de mollusques de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny. In: Audouin, V, Ed., Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee francaise. Histoire naturelle. Volume 1, partie 4. Imprimerie Imperiale, Paris: 7 - 56, https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" title="Explication sommaire des planches de mollusques de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny." url="https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" volumeTitle="Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee francaise." year="1826">Audouin (1826</bibRefCitation>
: 18-20) in the explanation of a plate by 
<bibRefCitation author="Savigny, JC" journalOrPublisher="Marine Biodiversity" pageId="0" pageNumber="1" publicationUrl="http://gallica.bnf.fr/ark:/12148/btv1b2300300m" refId="B157" refString="Savigny, JC, 1817. Description de l'Egypte ou Recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'Armee francaise, publie par ordre du Gouvernement. Histoire Naturelle, Planches, Tome Deuxieme. Imprimerie Royale, Paris. [105 pls; gastropods on pls 1-3] http://gallica.bnf.fr/ark:/12148/btv1b2300300m" title="Description de l'Egypte ou Recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'Armee francaise, publie par ordre du Gouvernement. Histoire Naturelle, Planches, Tome Deuxieme. Imprimerie Royale, Paris. [105 pls; gastropods on pls 1 - 3]" url="http://gallica.bnf.fr/ark:/12148/btv1b2300300m" year="1817">Savigny (1817</bibRefCitation>
: pl. 2) from the 
<emphasis italics="true" pageId="0" pageNumber="1">
Description de 
<normalizedToken originalValue="l’Egypte">l'Egypte</normalizedToken>
</emphasis>
. 
<bibRefCitation author="Savigny, JC" journalOrPublisher="Marine Biodiversity" pageId="0" pageNumber="1" publicationUrl="http://gallica.bnf.fr/ark:/12148/btv1b2300300m" refId="B157" refString="Savigny, JC, 1817. Description de l'Egypte ou Recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'Armee francaise, publie par ordre du Gouvernement. Histoire Naturelle, Planches, Tome Deuxieme. Imprimerie Royale, Paris. [105 pls; gastropods on pls 1-3] http://gallica.bnf.fr/ark:/12148/btv1b2300300m" title="Description de l'Egypte ou Recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'Armee francaise, publie par ordre du Gouvernement. Histoire Naturelle, Planches, Tome Deuxieme. Imprimerie Royale, Paris. [105 pls; gastropods on pls 1 - 3]" url="http://gallica.bnf.fr/ark:/12148/btv1b2300300m" year="1817">
<normalizedToken originalValue="Savigny’s">Savigny's</normalizedToken>
(1817
</bibRefCitation>
: pl. 2, figs 3.1-3.8) plate displays eight drawings for two onchidiid slugs from the Red Sea, with one of them clearly representing a dorsal gill (
<bibRefCitation author="Savigny, JC" journalOrPublisher="Marine Biodiversity" pageId="0" pageNumber="1" publicationUrl="http://gallica.bnf.fr/ark:/12148/btv1b2300300m" refId="B157" refString="Savigny, JC, 1817. Description de l'Egypte ou Recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'Armee francaise, publie par ordre du Gouvernement. Histoire Naturelle, Planches, Tome Deuxieme. Imprimerie Royale, Paris. [105 pls; gastropods on pls 1-3] http://gallica.bnf.fr/ark:/12148/btv1b2300300m" title="Description de l'Egypte ou Recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'Armee francaise, publie par ordre du Gouvernement. Histoire Naturelle, Planches, Tome Deuxieme. Imprimerie Royale, Paris. [105 pls; gastropods on pls 1 - 3]" url="http://gallica.bnf.fr/ark:/12148/btv1b2300300m" year="1817">Savigny 1817</bibRefCitation>
: pl. 2, fig. 3.5). According to 
<bibRefCitation author="Audouin, V" editor="Audouin, V" journalOrPublisher="Histoire naturelle. Volume 1, partie 4. Imprimerie Imperiale, Paris" pageId="0" pageNumber="1" pagination="7 - 56" publicationUrl="https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" refId="B9" refString="Audouin, V, 1826. Explication sommaire des planches de mollusques de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny. In: Audouin, V, Ed., Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee francaise. Histoire naturelle. Volume 1, partie 4. Imprimerie Imperiale, Paris: 7 - 56, https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" title="Explication sommaire des planches de mollusques de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny." url="https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" volumeTitle="Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee francaise." year="1826">Audouin (1826</bibRefCitation>
: 19), it was Cuvier himself who identified those two slugs as 
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
, although 
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/33372853" refId="B45" refString="Cuvier, G, 1830. Le regne animal distribue d'apres son organisation (Vol. 3). Deterville, Paris, xvi + 504 pp. [pls I-XX] https://www.biodiversitylibrary.org/page/33372853" title="Le regne animal distribue d'apres son organisation (Vol. 3). Deterville, Paris, xvi + 504 pp. [pls I-XX]" url="https://www.biodiversitylibrary.org/page/33372853" year="1830">Cuvier (1830)</bibRefCitation>
later changed his mind and created the new name 
<taxonomicName authorityName="Cuvier" authorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium verruculatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium verruculatum</emphasis>
</taxonomicName>
for them. More importantly, 
<bibRefCitation author="Audouin, V" editor="Audouin, V" journalOrPublisher="Histoire naturelle. Volume 1, partie 4. Imprimerie Imperiale, Paris" pageId="0" pageNumber="1" pagination="7 - 56" publicationUrl="https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" refId="B9" refString="Audouin, V, 1826. Explication sommaire des planches de mollusques de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny. In: Audouin, V, Ed., Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee francaise. Histoire naturelle. Volume 1, partie 4. Imprimerie Imperiale, Paris: 7 - 56, https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" title="Explication sommaire des planches de mollusques de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny." url="https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" volumeTitle="Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee francaise." year="1826">Audouin (1826</bibRefCitation>
: 19) described in great detail the &quot;small vascular branches&quot; at the posterior end of the dorsum, or 
<normalizedToken originalValue="“tubercles”">&quot;tubercles&quot;</normalizedToken>
that work as &quot;true gills.&quot; And 
<bibRefCitation author="Audouin, V" editor="Audouin, V" journalOrPublisher="Histoire naturelle. Volume 1, partie 4. Imprimerie Imperiale, Paris" pageId="0" pageNumber="1" pagination="7 - 56" publicationUrl="https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" refId="B9" refString="Audouin, V, 1826. Explication sommaire des planches de mollusques de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny. In: Audouin, V, Ed., Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee francaise. Histoire naturelle. Volume 1, partie 4. Imprimerie Imperiale, Paris: 7 - 56, https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" title="Explication sommaire des planches de mollusques de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny." url="https://digi.ub.uni-heidelberg.de/diglit/jomard1809bd5_1_1/0548" volumeTitle="Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee francaise." year="1826">Audouin (1826</bibRefCitation>
: 19) even made this clever statement:
</paragraph>
<paragraph pageId="0" pageNumber="1">
&quot;The Onchidie thus would have at the same time a pulmonary apparatus and a branchial apparatus; and that structure is in perfect agreement with what we know of the habits of that mollusk: 
<normalizedToken originalValue="Péron">Peron</normalizedToken>
says that it is aquatic; on the contrary M. Cuvier, without the authority of this observer, would have believed it to be terrestrial. (...) We think that the Onchidie, at least the species illustrated here, enjoys the capacity to breathe under water thanks to the help of those ramified tubercles which cover the posterior end of its body, without the necessity of coming up to the surface; which is relatively difficult for an animal that slowly crawls at the bottom underwater. As for the pulmonary opening, it indicates that the onchidie breathes air as well; and we must suppose that several times in its life it finds itself in the condition to do so.&quot;
</paragraph>
<paragraph pageId="0" pageNumber="1">Audouin supposedly assumed that those slugs were truly aquatic.</paragraph>
<paragraph pageId="0" pageNumber="1">
Because 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
was originally used as a genus for all marine onchidiids by both 
<bibRefCitation author="Fleming, J" editor="Napier, M" journalOrPublisher="5). Archibald Constable, Edinburgh" pageId="0" pageNumber="1" pagination="567 - 584" publicationUrl="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" refId="B72" refString="Fleming, J, 1822a. Mollusca. In: Napier, M, Ed., Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol. 5). Archibald Constable, Edinburgh: 567 - 584, https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" title="Mollusca." url="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" volumeTitle="Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol." year="1822 a">Fleming (1822a</bibRefCitation>
, 
<bibRefCitation author="Fleming, J" journalOrPublisher="Archibald Constable, Edinburgh" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28230157" refId="B73" refString="Fleming, J, 1822b. The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2). Archibald Constable, Edinburgh, https://www.biodiversitylibrary.org/page/28230157" title="The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2)." url="https://www.biodiversitylibrary.org/page/28230157" year="1822 b">b</bibRefCitation>
) and 
<bibRefCitation author="Blainville, HMD de" journalOrPublisher="Levrault, Strasbourg" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/25299228" refId="B23" refString="Blainville, HMD de, 1824. Mollusques. In: Cuvier F (Ed) Dictionnaire des Sciences Naturelles (Vol. 32). F. G. Levrault, Strasbourg, https://www.biodiversitylibrary.org/page/25299228" title="Mollusques. In: Cuvier F (Ed) Dictionnaire des Sciences Naturelles (Vol. 32). F. G." url="https://www.biodiversitylibrary.org/page/25299228" year="1824">Blainville (1824</bibRefCitation>
, 
<bibRefCitation author="Blainville, HMD de" editor="Cuvier, F" journalOrPublisher="38). F. G. Levrault, Strasbourg" pageId="0" pageNumber="1" pagination="519 - 524" publicationUrl="https://www.biodiversitylibrary.org/page/25302418" refId="B24" refString="Blainville, HMD de, 1826. Peronie. In: Cuvier, F, Ed., Dictionnaire des sciences naturelles (Vol. 38). F. G. Levrault, Strasbourg: 519 - 524, https://www.biodiversitylibrary.org/page/25302418" title="Peronie." url="https://www.biodiversitylibrary.org/page/25302418" volumeTitle="Dictionnaire des sciences naturelles (Vol." year="1826">1826</bibRefCitation>
), several 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species names already existed by 1830: 
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="P. oniscoides" pageId="0" pageNumber="1" rank="species" species="oniscoides">
<emphasis italics="true" pageId="0" pageNumber="1">P. oniscoides</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="P. semituberculata" pageId="0" pageNumber="1" rank="species" species="semituberculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. semituberculata</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="P. laevis" pageId="0" pageNumber="1" rank="species" species="laevis">
<emphasis italics="true" pageId="0" pageNumber="1">P. laevis</emphasis>
</taxonomicName>
(see above). Of those names, only the two objective synonyms 
<taxonomicName lsidName="P. mauritiana" pageId="0" pageNumber="1" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">P. mauritiana</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
refer to true 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
slugs, i.e., slugs with dorsal gills (Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Alphabetic list of the 51 existing species-group names of which the nomenclatural status is addressed in the present work. Details can be found in the text: comments on the four valid Peronia species names, their synonyms, and the junior homonym are in the species remarks; comments on the fifteen nomina dubia and the ten names that must be classified in other genera are in the general discussion." httpUri="http://table.plazi.org/id/F0EDB18E2E41A7D8B1806BBE53F2616C" pageId="0" pageNumber="1" tableUuid="F0EDB18E2E41A7D8B1806BBE53F2616C">1</tableCitation>
). 
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/33372853" refId="B45" refString="Cuvier, G, 1830. Le regne animal distribue d'apres son organisation (Vol. 3). Deterville, Paris, xvi + 504 pp. [pls I-XX] https://www.biodiversitylibrary.org/page/33372853" title="Le regne animal distribue d'apres son organisation (Vol. 3). Deterville, Paris, xvi + 504 pp. [pls I-XX]" url="https://www.biodiversitylibrary.org/page/33372853" year="1830">Cuvier (1830</bibRefCitation>
: 46) did not see the need for a genus assignment for marine onchidiid species and still only recognized 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
, but other naturalists started transferring species names from 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
to 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
. Lesson (1833: pl. 19) transferred his own 
<taxonomicName authorityName="Lesson" authorityYear="1831" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium ferrugineum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ferrugineum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium ferrugineum</emphasis>
</taxonomicName>
Lesson, 1831a to 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
, and clearly specified that he agreed with Blainville that marine onchidiids should be classified in a distinct genus. Dorsal gills are very clearly described by 
<bibRefCitation author="Lesson, RP" journalOrPublisher="Bulletin des Sciences Naturelles et de Geologie" pageId="0" pageNumber="1" pagination="128 - 130" publicationUrl="https://www.biodiversitylibrary.org/page/4441590" refId="B113" refString="Lesson, RP, 1831a. Note sur deux Onchidies nouvelles. Bulletin des Sciences Naturelles et de Geologie 25: 128 - 130, https://www.biodiversitylibrary.org/page/4441590" title="Note sur deux Onchidies nouvelles." url="https://www.biodiversitylibrary.org/page/4441590" volume="25" year="1831 a">Lesson (1831a</bibRefCitation>
: 128-130; 1831b: 300-302; 
<bibRefCitation author="Lesson, RP" journalOrPublisher="Memorial encyclopedique et progressif des connaissances humaines" pageId="0" pageNumber="1" pagination="36 - 37" refId="B115" refString="Lesson, RP, 1832. Mollusques: Nouvelle espece d'Onchidie, Onchidium ferrugineum. Memorial encyclopedique et progressif des connaissances humaines 2: 36 - 37" title="Mollusques: Nouvelle espece d'Onchidie, Onchidium ferrugineum." volume="2" year="1832">1832</bibRefCitation>
: 36-37, fig. 32; 1833: pl. 19) in 
<taxonomicName lsidName="O. ferrugineum" pageId="0" pageNumber="1" rank="species" species="ferrugineum">
<emphasis italics="true" pageId="0" pageNumber="1">O. ferrugineum</emphasis>
</taxonomicName>
, but they were not the reason why he transferred it from 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
to 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
. Shortly after that, 
<bibRefCitation author="Oken, L" journalOrPublisher="Transactions of the Zoological Society of London" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/27509779" refId="B139" refString="Oken, L, 1834a. Voyage de la Corvette l'Astrolabe sous le Commandement de Dumont Durville. Isis von Oken [1834]: 283-310. https://biodiversitylibrary.org/page/27509779" title="Voyage de la Corvette l'Astrolabe sous le Commandement de Dumont Durville. Isis von Oken [1834]: 283 - 310." url="https://biodiversitylibrary.org/page/27509779" year="1834 a">Oken (1834a)</bibRefCitation>
also transferred six 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
species names by 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.2132" author="Quoy, JRC" journalOrPublisher="Tastu, Paris" pageId="0" pageNumber="1" refId="B148" refString="Quoy, JRC, Gaimard, JP, 1832-1833. Zoologie. Tome Second. Voyage de decouvertes de l'Astrolabe Execute par ordre du Roi, pendant les annees 1826-1827-1828-1829, sous le commandement de M. J. Dumont d'Urville. J. Tastu, Paris, DOI: https://doi.org/10.5962/bhl.title.2132" title="Zoologie. Tome Second. Voyage de decouvertes de l'Astrolabe Execute par ordre du Roi, pendant les annees 1826 - 1827 - 1828 - 1829, sous le commandement de M. J. Dumont d'Urville. J." url="https://doi.org/10.5962/bhl.title.2132" year="1832 - 1833">Quoy and Gaimard (1832-1833)</bibRefCitation>
to 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
(
<taxonomicName lsidName="P. cinerea" pageId="0" pageNumber="1" rank="species" species="cinerea">
<emphasis italics="true" pageId="0" pageNumber="1">P. cinerea</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="P. incisa" pageId="0" pageNumber="1" rank="species" species="incisa">
<emphasis italics="true" pageId="0" pageNumber="1">P. incisa</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="P. nigricans" pageId="0" pageNumber="1" rank="species" species="nigricans">
<emphasis italics="true" pageId="0" pageNumber="1">P. nigricans</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="P. patelloides" pageId="0" pageNumber="1" rank="species" species="patelloides">
<emphasis italics="true" pageId="0" pageNumber="1">P. patelloides</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="P. punctata" pageId="0" pageNumber="1" rank="species" species="punctata">
<emphasis italics="true" pageId="0" pageNumber="1">P. punctata</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="P. tongana" pageId="0" pageNumber="1" rank="species" species="tongana">
<emphasis italics="true" pageId="0" pageNumber="1">P. tongana</emphasis>
</taxonomicName>
), with no justification but most likely because he also adopted the idea that marine onchidiids should not be classified in 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
The name 
<taxonomicName authorityName="Gistel" authorityYear="1848" class="Gastropoda" family="Onchidiidae" genus="Eudrastus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eudrastus" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Eudrastus</emphasis>
</taxonomicName>
was created by 
<bibRefCitation author="Gistel, J" journalOrPublisher="Annual Reviews of Ecology, Evolution and Systematics" pageId="0" pageNumber="1" refId="B77" refString="Gistel, J, 1848. Naturgeschichte des Thierreichs. Fuer hoehere Schulen. Hoffmannische Verlags-Buchhandlung, Stuttgart, xvi + 216 pp. [32 pls]" title="Naturgeschichte des Thierreichs. Fuer hoehere Schulen. Hoffmannische Verlags-Buchhandlung, Stuttgart, xvi + 216 pp. [32 pls]" year="1848">Gistel (1848</bibRefCitation>
: x), as a replacement name for &quot; 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Peronia</taxonomicName>
(Quoy, Isis 1834. 287.).&quot; Gistel refers here to a report (
<bibRefCitation author="Oken, L" journalOrPublisher="Transactions of the Zoological Society of London" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/27509779" refId="B139" refString="Oken, L, 1834a. Voyage de la Corvette l'Astrolabe sous le Commandement de Dumont Durville. Isis von Oken [1834]: 283-310. https://biodiversitylibrary.org/page/27509779" title="Voyage de la Corvette l'Astrolabe sous le Commandement de Dumont Durville. Isis von Oken [1834]: 283 - 310." url="https://biodiversitylibrary.org/page/27509779" year="1834 a">Oken 1834a</bibRefCitation>
: 283-310) on 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.2132" author="Quoy, JRC" journalOrPublisher="Tastu, Paris" pageId="0" pageNumber="1" refId="B148" refString="Quoy, JRC, Gaimard, JP, 1832-1833. Zoologie. Tome Second. Voyage de decouvertes de l'Astrolabe Execute par ordre du Roi, pendant les annees 1826-1827-1828-1829, sous le commandement de M. J. Dumont d'Urville. J. Tastu, Paris, DOI: https://doi.org/10.5962/bhl.title.2132" title="Zoologie. Tome Second. Voyage de decouvertes de l'Astrolabe Execute par ordre du Roi, pendant les annees 1826 - 1827 - 1828 - 1829, sous le commandement de M. J. Dumont d'Urville. J." url="https://doi.org/10.5962/bhl.title.2132" year="1832 - 1833">
Quoy and 
<normalizedToken originalValue="Gaimard’s">Gaimard's</normalizedToken>
(1832-1833)
</bibRefCitation>
contribution to the 
<emphasis italics="true" pageId="0" pageNumber="1">
Voyage de 
<normalizedToken originalValue="découvertes">decouvertes</normalizedToken>
de 
<normalizedToken originalValue="l’Astrolabe">l'Astrolabe</normalizedToken>
</emphasis>
published in 
<emphasis italics="true" pageId="0" pageNumber="1">Isis</emphasis>
, the encyclopedic journal edited by Lorenz Oken from 1817 to 1848. This report was most likely written by Oken himself, as was often the case (
<bibRefCitation DOI="https://doi.org/10.1086/354208" author="Kertesz, GA" journalOrPublisher="Isis" pageId="0" pageNumber="1" pagination="497 - 503" refId="B104" refString="Kertesz, GA, 1986. Notes on Isis von Oken, 1817-1848. Isis 77: 497 - 503, DOI: https://doi.org/10.1086/354208" title="Notes on Isis von Oken, 1817 - 1848." url="https://doi.org/10.1086/354208" volume="77" year="1986">Kertesz 1986</bibRefCitation>
), which would explain that the six onchidiid specific names mentioned (
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="tongana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tongana">
<emphasis italics="true" pageId="0" pageNumber="1">tongana</emphasis>
</taxonomicName>
, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="incisa" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="incisa">
<emphasis italics="true" pageId="0" pageNumber="1">incisa</emphasis>
</taxonomicName>
, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="patelloides" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="patelloides">
<emphasis italics="true" pageId="0" pageNumber="1">patelloides</emphasis>
</taxonomicName>
, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="nigricans" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="nigricans">
<emphasis italics="true" pageId="0" pageNumber="1">nigricans</emphasis>
</taxonomicName>
, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="punctata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctata">
<emphasis italics="true" pageId="0" pageNumber="1">punctata</emphasis>
</taxonomicName>
, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="cinerea" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cinerea">
<emphasis italics="true" pageId="0" pageNumber="1">cinerea</emphasis>
</taxonomicName>
) are combined with 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
instead of 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
, the generic name originally used by 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.2132" author="Quoy, JRC" journalOrPublisher="Tastu, Paris" pageId="0" pageNumber="1" refId="B148" refString="Quoy, JRC, Gaimard, JP, 1832-1833. Zoologie. Tome Second. Voyage de decouvertes de l'Astrolabe Execute par ordre du Roi, pendant les annees 1826-1827-1828-1829, sous le commandement de M. J. Dumont d'Urville. J. Tastu, Paris, DOI: https://doi.org/10.5962/bhl.title.2132" title="Zoologie. Tome Second. Voyage de decouvertes de l'Astrolabe Execute par ordre du Roi, pendant les annees 1826 - 1827 - 1828 - 1829, sous le commandement de M. J. Dumont d'Urville. J." url="https://doi.org/10.5962/bhl.title.2132" year="1832 - 1833">Quoy and Gaimard (1832-1833)</bibRefCitation>
. Regardless of who authored that 
<emphasis italics="true" pageId="0" pageNumber="1">Isis</emphasis>
report, 
<bibRefCitation author="Gistel, J" journalOrPublisher="Annual Reviews of Ecology, Evolution and Systematics" pageId="0" pageNumber="1" refId="B77" refString="Gistel, J, 1848. Naturgeschichte des Thierreichs. Fuer hoehere Schulen. Hoffmannische Verlags-Buchhandlung, Stuttgart, xvi + 216 pp. [32 pls]" title="Naturgeschichte des Thierreichs. Fuer hoehere Schulen. Hoffmannische Verlags-Buchhandlung, Stuttgart, xvi + 216 pp. [32 pls]" year="1848">Gistel (1848)</bibRefCitation>
did create the new generic name 
<taxonomicName authorityName="Gistel" authorityYear="1848" class="Gastropoda" family="Onchidiidae" genus="Eudrastus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eudrastus" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Eudrastus</emphasis>
</taxonomicName>
for those six species. 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.10914" author="Baker, HB" journalOrPublisher="The Nautilus" pageId="0" pageNumber="1" pagination="85 - 88" refId="B13" refString="Baker, HB, 1938. Nomenclature of Onchidiidae. The Nautilus 51: 85 - 88, DOI: https://doi.org/10.5962/bhl.part.10914" title="Nomenclature of Onchidiidae." url="https://doi.org/10.5962/bhl.part.10914" volume="51" year="1938">Baker (1938</bibRefCitation>
: 86) subsequently designated 
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
Quoy &amp; Gaimard, 1832 (
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia tongana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tongana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia tongana</emphasis>
</taxonomicName>
in 
<emphasis italics="true" pageId="0" pageNumber="1">Isis</emphasis>
), as the type species of 
<taxonomicName authorityName="Gistel" authorityYear="1848" class="Gastropoda" family="Onchidiidae" genus="Eudrastus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eudrastus" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Eudrastus</emphasis>
</taxonomicName>
. 
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
is regarded here as a junior subjective synonym of 
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
, so 
<taxonomicName authorityName="Gistel" authorityYear="1848" class="Gastropoda" family="Onchidiidae" genus="Eudrastus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eudrastus" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Eudrastus</emphasis>
</taxonomicName>
is a junior subjective synonym of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
. 
<bibRefCitation DOI="https://doi.org/10.1093/oxfordjournals.mollus.a065863" author="Britton, KM" journalOrPublisher="Journal of Molluscan Studies" pageId="0" pageNumber="1" pagination="179 - 191" refId="B27" refString="Britton, KM, 1984. The Onchidiacea (Gastropoda, Pulmonata) of Hong Kong with a worldwide review of the genera. Journal of Molluscan Studies 50: 179 - 191, DOI: https://doi.org/10.1093/oxfordjournals.mollus.a065863" title="The Onchidiacea (Gastropoda, Pulmonata) of Hong Kong with a worldwide review of the genera." url="https://doi.org/10.1093/oxfordjournals.mollus.a065863" volume="50" year="1984">Britton (1984</bibRefCitation>
: 182-183) suggested that 
<taxonomicName authorityName="Gistel" authorityYear="1848" class="Gastropoda" family="Onchidiidae" genus="Eudrastus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eudrastus" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Eudrastus</emphasis>
</taxonomicName>
should be regarded as a junior synonym of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
because it seemed to be based on &quot;unimportant characters.&quot;
</paragraph>
<paragraph pageId="0" pageNumber="1">
John Edwards 
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="129 - 219" publicationUrl="https://www.biodiversitylibrary.org/page/12862913" refId="B85" refString="Gray, JE, 1847. A list of the genera of recent Mollusca, their synonyms and types. Proceedings of the Zoological Society of London 15: 129 - 219, https://www.biodiversitylibrary.org/page/12862913" title="A list of the genera of recent Mollusca, their synonyms and types." url="https://www.biodiversitylibrary.org/page/12862913" volume="15" year="1847">Gray (1847</bibRefCitation>
: 179) attributed the authorship of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
to Blainville (with an erroneous date of 1825) but, most importantly, gave its modern definition to 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
by restricting it to six species of slugs with &quot;radiating processes&quot; on the back (JE 
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">Gray 1850</bibRefCitation>
: 117): 
<taxonomicName lsidName="P. alderi" pageId="0" pageNumber="1" rank="species" species="alderi">
<emphasis italics="true" pageId="0" pageNumber="1">P. alderi</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="P. ferruginea" pageId="0" pageNumber="1" rank="species" species="ferruginea">
<emphasis italics="true" pageId="0" pageNumber="1">P. ferruginea</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="P. mauritiana" pageId="0" pageNumber="1" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">P. mauritiana</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="P. punctata" pageId="0" pageNumber="1" rank="species" species="punctata">
<emphasis italics="true" pageId="0" pageNumber="1">P. punctata</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="P. tongana" pageId="0" pageNumber="1" rank="species" species="tongana">
<emphasis italics="true" pageId="0" pageNumber="1">P. tongana</emphasis>
</taxonomicName>
. All those names refer to true 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
slugs with dorsal gills. JE 
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">Gray (1850</bibRefCitation>
: 117) restricted 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
to 
<normalizedToken originalValue="Buchannan’s">Buchannan's</normalizedToken>
<taxonomicName lsidName="O. typhae" pageId="0" pageNumber="1" rank="species" species="typhae">
<emphasis italics="true" pageId="0" pageNumber="1">O. typhae</emphasis>
</taxonomicName>
and included all the other marine species without dorsal gills in a new genus 
<taxonomicName authorityName="J.E. Gray" authorityYear="1850" class="Gastropoda" family="Onchidiidae" genus="Onchidella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidella</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
JE 
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">
<normalizedToken originalValue="Gray’s">Gray's</normalizedToken>
(1850)
</bibRefCitation>
clarity only lasted for a few years. Indeed, Adams and Adams (1855: 234) pointed out that 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
slugs differ from 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="J.E. Gray" authorityYear="1850" class="Gastropoda" family="Onchidiidae" genus="Onchidella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidella</emphasis>
</taxonomicName>
because of &quot;arbusculiform and other appendages of the mantle, which have sometimes been mistaken for gills.&quot; Because they did not believe that gills were distinct from other dorsal papillae, Adams and Adams (1855: 234) classified in 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
some names that belong to both 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
(
<taxonomicName lsidName="P. ferruginea" pageId="0" pageNumber="1" rank="species" species="ferruginea">
<emphasis italics="true" pageId="0" pageNumber="1">P. ferruginea</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="P. mauritiana" pageId="0" pageNumber="1" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">P. mauritiana</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="P. punctata" pageId="0" pageNumber="1" rank="species" species="punctata">
<emphasis italics="true" pageId="0" pageNumber="1">P. punctata</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="P. tongana" pageId="0" pageNumber="1" rank="species" species="tongana">
<emphasis italics="true" pageId="0" pageNumber="1">P. tongana</emphasis>
</taxonomicName>
) and to 
<taxonomicName authorityName="J.E. Gray" authorityYear="1850" class="Gastropoda" family="Onchidiidae" genus="Onchidella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidella</emphasis>
</taxonomicName>
(
<taxonomicName lsidName="O. celtica" pageId="0" pageNumber="1" rank="species" species="celtica">
<emphasis italics="true" pageId="0" pageNumber="1">O. celtica</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="O. indolens" pageId="0" pageNumber="1" rank="species" species="indolens">
<emphasis italics="true" pageId="0" pageNumber="1">O. indolens</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="O. marginata" pageId="0" pageNumber="1" rank="species" species="marginata">
<emphasis italics="true" pageId="0" pageNumber="1">O. marginata</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="O. parthenopeia" pageId="0" pageNumber="1" rank="species" species="parthenopeia">
<emphasis italics="true" pageId="0" pageNumber="1">O. parthenopeia</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
JE 
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">
<normalizedToken originalValue="Gray’s">Gray's</normalizedToken>
(1850)
</bibRefCitation>
classification was adopted by 
<bibRefCitation author="Keferstein, WM" editor="Bronn, HG" journalOrPublisher="Bd. 3 (2). Winter, Leipzig &amp; Heidelberg" pageId="0" pageNumber="1" pagination="1160 - 1306" publicationUrl="https://www.biodiversitylibrary.org/page/35742549" refId="B102" refString="Keferstein, WM, 1865a. Lungenschnecken: Pulmonata. In: Bronn, HG, Ed., Klassen und Ordnungen der Weichthiere (Malacozoa), wissenschaftlich dargestellt in Wort &amp; Bild. Bd. 3(2). Winter, Leipzig &amp; Heidelberg: 1160 - 1306, https://www.biodiversitylibrary.org/page/35742549" title="Lungenschnecken: Pulmonata." url="https://www.biodiversitylibrary.org/page/35742549" volumeTitle="Klassen und Ordnungen der Weichthiere (Malacozoa), wissenschaftlich dargestellt in Wort &amp; Bild." year="1865 a">Keferstein (1865a)</bibRefCitation>
but, until 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
(1934a)
</bibRefCitation>
work, all authors have ignored the genus 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
and simply used the genus 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
for slugs with and without dorsal gills (
<bibRefCitation author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" pagination="86 - 111" publicationUrl="https://biodiversitylibrary.org/page/41279271" refId="B173" refString="Stoliczka, F, 1869. The malacology of Lower Bengal. Journal of the Asiatic Society of Bengal 38: 86 - 111, https://biodiversitylibrary.org/page/41279271" title="The malacology of Lower Bengal." url="https://biodiversitylibrary.org/page/41279271" volume="38" year="1869">Stoliczka 1869</bibRefCitation>
; 
<bibRefCitation author="Semper, C" editor="Semper, C" journalOrPublisher="Wissenschaftliche Resultate. Dritter Band. Landmollusken. CW Kreidel's Verlag, Wiesbaden, 251 - 264 [1880]" pageId="0" pageNumber="1" pagination="265 - 290" publicationUrl="https://biodiversitylibrary.org/" refId="B159" refString="Semper, C, 1880-1885. Dritte Familie, Onchididae [for Onchidiidae]. In: Semper, C, Ed., Reisen im Archipel der Philippinen, Zweiter Theil. Wissenschaftliche Resultate. Dritter Band. Landmollusken. CW Kreidel's Verlag, Wiesbaden, 251-264 [1880]: 265 - 290, https://biodiversitylibrary.org/" title="Dritte Familie, Onchididae [for Onchidiidae]." url="https://biodiversitylibrary.org/" volumeTitle="Reisen im Archipel der Philippinen, Zweiter Theil." year="1880 - 1885">Semper 1880-1885</bibRefCitation>
; 
<bibRefCitation author="Plate, L" journalOrPublisher="Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere" pageId="0" pageNumber="1" pagination="93 - 234" publicationUrl="https://biodiversitylibrary.org/page/11175213" refId="B146" refString="Plate, L, 1893. Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere 7: 93 - 234, https://biodiversitylibrary.org/page/11175213" title="Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden." url="https://biodiversitylibrary.org/page/11175213" volume="7" year="1893">Plate 1893</bibRefCitation>
; 
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.12.1919.888" author="Bretnall, W" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="303 - 328" refId="B26" refString="Bretnall, W, 1919. Onchidiidae from Australia and the South-Western Pacific Islands. Records of the Australian Museum 12: 303 - 328, DOI: https://doi.org/10.3853/j.0067-1975.12.1919.888" title="Onchidiidae from Australia and the South-Western Pacific Islands." url="https://doi.org/10.3853/j.0067-1975.12.1919.888" volume="12" year="1919">Bretnall 1919</bibRefCitation>
; 
<bibRefCitation author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher" pageId="0" pageNumber="1" pagination="29 - 118" refId="B92" refString="Hoffmann, H, 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbuecher 55: 29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann 1928</bibRefCitation>
). 
<bibRefCitation author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" pagination="86 - 111" publicationUrl="https://biodiversitylibrary.org/page/41279271" refId="B173" refString="Stoliczka, F, 1869. The malacology of Lower Bengal. Journal of the Asiatic Society of Bengal 38: 86 - 111, https://biodiversitylibrary.org/page/41279271" title="The malacology of Lower Bengal." url="https://biodiversitylibrary.org/page/41279271" volume="38" year="1869">Stoliczka (1869</bibRefCitation>
: 100-102), who was the first one to re-examine live slugs of 
<taxonomicName lsidName="O. typhae" pageId="0" pageNumber="1" rank="species" species="typhae">
<emphasis italics="true" pageId="0" pageNumber="1">O. typhae</emphasis>
</taxonomicName>
since 
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1800.tb00584.x" author="Buchannan, F" journalOrPublisher="Transactions of the Linnean Society of London" pageId="0" pageNumber="1" pagination="132 - 134" refId="B28" refString="Buchannan, F, 1800. An account of the Onchidium, a new genus of the class of vermes, found in Bengal. Transactions of the Linnean Society of London 5: 132 - 134, DOI: https://doi.org/10.1111/j.1096-3642.1800.tb00584.x" title="An account of the Onchidium, a new genus of the class of vermes, found in Bengal." url="https://doi.org/10.1111/j.1096-3642.1800.tb00584.x" volume="5" year="1800">Buchannan (1800)</bibRefCitation>
, firmly argued that slugs with &quot;dorsal tufts&quot; were anatomically so similar to 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="J.E. Gray" authorityYear="1850" class="Gastropoda" family="Onchidiidae" genus="Onchidella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidella</emphasis>
</taxonomicName>
that only one name, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
, was needed. 
<bibRefCitation author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" pagination="86 - 111" publicationUrl="https://biodiversitylibrary.org/page/41279271" refId="B173" refString="Stoliczka, F, 1869. The malacology of Lower Bengal. Journal of the Asiatic Society of Bengal 38: 86 - 111, https://biodiversitylibrary.org/page/41279271" title="The malacology of Lower Bengal." url="https://biodiversitylibrary.org/page/41279271" volume="38" year="1869">Stoliczka (1869</bibRefCitation>
: 98) also clarified that 
<taxonomicName lsidName="O. typhae" pageId="0" pageNumber="1" rank="species" species="typhae">
<emphasis italics="true" pageId="0" pageNumber="1">O. typhae</emphasis>
</taxonomicName>
is not a terrestrial species but that, instead, it lives in &quot;damp places, generally close to tanks or ditches, especially those which are supplied during high tide with brackish water.&quot;
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" pagination="86 - 111" publicationUrl="https://biodiversitylibrary.org/page/41279271" refId="B173" refString="Stoliczka, F, 1869. The malacology of Lower Bengal. Journal of the Asiatic Society of Bengal 38: 86 - 111, https://biodiversitylibrary.org/page/41279271" title="The malacology of Lower Bengal." url="https://biodiversitylibrary.org/page/41279271" volume="38" year="1869">
<normalizedToken originalValue="Stoliczka’s">Stoliczka's</normalizedToken>
(1869)
</bibRefCitation>
strong influence can be seen in 
<bibRefCitation author="Semper, C" editor="Semper, C" journalOrPublisher="Wissenschaftliche Resultate. Dritter Band. Landmollusken. CW Kreidel's Verlag, Wiesbaden, 251 - 264 [1880]" pageId="0" pageNumber="1" pagination="265 - 290" publicationUrl="https://biodiversitylibrary.org/" refId="B159" refString="Semper, C, 1880-1885. Dritte Familie, Onchididae [for Onchidiidae]. In: Semper, C, Ed., Reisen im Archipel der Philippinen, Zweiter Theil. Wissenschaftliche Resultate. Dritter Band. Landmollusken. CW Kreidel's Verlag, Wiesbaden, 251-264 [1880]: 265 - 290, https://biodiversitylibrary.org/" title="Dritte Familie, Onchididae [for Onchidiidae]." url="https://biodiversitylibrary.org/" volumeTitle="Reisen im Archipel der Philippinen, Zweiter Theil." year="1880 - 1885">
<normalizedToken originalValue="Semper’s">Semper's</normalizedToken>
(1880-1885)
</bibRefCitation>
study of the onchidiids from the Philippines (and other parts of the Indo-West Pacific) in which all onchidiids are in 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
, with the exception of a single species in his new genus 
<taxonomicName authorityName="C.Semper" authorityYear="1882" class="Gastropoda" family="Onchidiidae" genus="Onchidina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidina</emphasis>
</taxonomicName>
Semper, 1882; for a collation of 
<normalizedToken originalValue="Semper’s">Semper's</normalizedToken>
work, see 
<bibRefCitation DOI="https://doi.org/10.3366/jsbnh.1969.5.2.144" author="Johnson, RI" journalOrPublisher="Journal of the Society for the Bibliography of Natural History" pageId="0" pageNumber="1" pagination="144 - 147" refId="B99" refString="Johnson, RI, 1969. Semper's Reisen im Archipel der Philippinen, wissenshaftliche Resultate, 1867-1916. A complete collation. Journal of the Society for the Bibliography of Natural History 5: 144 - 147, DOI: https://doi.org/10.3366/jsbnh.1969.5.2.144" title="Semper's Reisen im Archipel der Philippinen, wissenshaftliche Resultate, 1867 - 1916. A complete collation." url="https://doi.org/10.3366/jsbnh.1969.5.2.144" volume="5" year="1969">Johnson (1969)</bibRefCitation>
. 
<bibRefCitation author="Plate, L" journalOrPublisher="Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere" pageId="0" pageNumber="1" pagination="93 - 234" publicationUrl="https://biodiversitylibrary.org/page/11175213" refId="B146" refString="Plate, L, 1893. Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere 7: 93 - 234, https://biodiversitylibrary.org/page/11175213" title="Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden." url="https://biodiversitylibrary.org/page/11175213" volume="7" year="1893">Plate (1893)</bibRefCitation>
adopted a classification with five genera, but the four species of slugs with dorsal gills recognized by Plate are classified in 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
with thirteen species of slugs without dorsal gills. 
<bibRefCitation author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher" pageId="0" pageNumber="1" pagination="29 - 118" refId="B92" refString="Hoffmann, H, 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbuecher 55: 29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann (1928)</bibRefCitation>
adopted a classification with six genera, six species of slugs with dorsal gills being classified in 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
with 34 species without dorsal gills.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Then, suddenly, in 1934, the number of onchidiid taxon names for slugs with dorsal gills dramatically increased. Based on the onchidiid collection at the Paris Museum, 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a)
</bibRefCitation>
created fourteen new species-group names for slugs with dorsal gills (all but one name are species names) and four new generic names: 
<taxonomicName class="Aves" family="Tyrannidae" genus="Lessonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonia" order="Passeriformes" pageId="0" pageNumber="1" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonia</emphasis>
</taxonomicName>
(later replaced by 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina</emphasis>
</taxonomicName>
), 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia</emphasis>
</taxonomicName>
, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
, and 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Scaphis</emphasis>
</taxonomicName>
. Below, the nomenclatural status of 
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
generic names is justified first (they all are junior synonyms of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
), followed by opinions in the secondary literature.
</paragraph>
<paragraph pageId="0" pageNumber="1">
The generic name 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia</emphasis>
</taxonomicName>
was created by 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 196) for four species similar to 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
but with intestinal loops of type V (instead of type I). The type species is 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia gondwanae" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gondwanae">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia gondwanae</emphasis>
</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a, by subsequent designation (
<bibRefCitation author="Starobogatov, YI" journalOrPublisher="Soviet Journal of Marine Biology" pageId="0" pageNumber="1" pagination="206 - 212" refId="B171" refString="Starobogatov, YI, 1976. Composition and taxonomic position of marine pulmonate mollusks. Soviet Journal of Marine Biology 4: 206 - 212" title="Composition and taxonomic position of marine pulmonate mollusks." volume="4" year="1976">Starobogatov 1976</bibRefCitation>
: 211). 
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
description of 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia gondwanae" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gondwanae">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia gondwanae</emphasis>
</taxonomicName>
was based on 38 individuals with intestinal loops of types I and V which belong to different species. The application of the name 
<taxonomicName lsidName="P. gondwanae" pageId="0" pageNumber="1" rank="species" species="gondwanae">
<emphasis italics="true" pageId="0" pageNumber="1">P. gondwanae</emphasis>
</taxonomicName>
is clarified through the designation of a lectotype (see 
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
): 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia gondwanae" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gondwanae">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia gondwanae</emphasis>
</taxonomicName>
is a junior synonym of 
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia verruculata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia verruculata</emphasis>
</taxonomicName>
, and 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia</emphasis>
</taxonomicName>
is a junior synonym of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
The generic name 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Scaphis</emphasis>
</taxonomicName>
was created by 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 203) for nine species similar to 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
but supposedly with an oblique, almost vertical hyponotum. The type species is 
<taxonomicName baseAuthorityName="Labbe" baseAuthorityYear="1934" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium astridae" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="astridae">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium astridae</emphasis>
</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934b, by subsequent designation (
<bibRefCitation author="Starobogatov, YI" journalOrPublisher="Soviet Journal of Marine Biology" pageId="0" pageNumber="1" pagination="206 - 212" refId="B171" refString="Starobogatov, YI, 1976. Composition and taxonomic position of marine pulmonate mollusks. Soviet Journal of Marine Biology 4: 206 - 212" title="Composition and taxonomic position of marine pulmonate mollusks." volume="4" year="1976">Starobogatov 1976</bibRefCitation>
: 211). 
<taxonomicName baseAuthorityName="Labbe" baseAuthorityYear="1934" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium astridae" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="astridae">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium astridae</emphasis>
</taxonomicName>
is a junior synonym of 
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia verruculata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia verruculata</emphasis>
</taxonomicName>
, and 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Scaphis</emphasis>
</taxonomicName>
is a junior synonym of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Aves" family="Tyrannidae" genus="Lessonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonia" order="Passeriformes" pageId="0" pageNumber="1" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonia</emphasis>
</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a is objectively invalid because it is the junior homonym of 
<taxonomicName class="Aves" family="Tyrannidae" genus="Lessonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonia" order="Passeriformes" pageId="0" pageNumber="1" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonia</emphasis>
</taxonomicName>
Swainson, 1832 [
<taxonomicName class="Aves" lsidName="" pageId="0" pageNumber="1" rank="class">Aves</taxonomicName>
]. 
<bibRefCitation author="Starobogatov, YI" journalOrPublisher="Soviet Journal of Marine Biology" pageId="0" pageNumber="1" pagination="206 - 212" refId="B171" refString="Starobogatov, YI, 1976. Composition and taxonomic position of marine pulmonate mollusks. Soviet Journal of Marine Biology 4: 206 - 212" title="Composition and taxonomic position of marine pulmonate mollusks." volume="4" year="1976">Starobogatov (1976</bibRefCitation>
: 211) replaced it by 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina</emphasis>
</taxonomicName>
. 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 213-216, figs 48-50) described 
<taxonomicName class="Aves" family="Tyrannidae" genus="Lessonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonia" order="Passeriformes" pageId="0" pageNumber="1" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonia</emphasis>
</taxonomicName>
based on a single species, 
<taxonomicName authorityName="Lesson" authorityYear="1831" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium ferrugineum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ferrugineum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium ferrugineum</emphasis>
</taxonomicName>
Lesson, 1831a, of which he examined no other material than the four syntypes (MNHN-IM-2000-22951). The examination of the three remaining syntypes (one syntype was lost by or after 
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
) revealed that the lectotype (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.763.21252" author="Goulding, TC" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 111" refId="B82" refString="Goulding, TC, Khalil, M, Tan, SH, Dayrat, B, 2018b. Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda: Pulmonata: Onchidiidae). ZooKeys 763: 1 - 111, DOI: https://doi.org/10.3897/zookeys.763.21252" title="Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda: Pulmonata: Onchidiidae)." url="https://doi.org/10.3897/zookeys.763.21252" volume="763" year="2018 b">Goulding et al. 2018b</bibRefCitation>
: 75) belongs to a 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species and that the two paralectotypes belong to 
<taxonomicName baseAuthorityName="Lesson" baseAuthorityYear="1831" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">
<emphasis italics="true" pageId="0" pageNumber="1">Wallaconchis ater</emphasis>
</taxonomicName>
(Lesson, 1831a). Both 
<normalizedToken originalValue="Lesson’s">Lesson's</normalizedToken>
original description of 
<taxonomicName authorityName="Lesson" authorityYear="1831" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium ferrugineum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ferrugineum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium ferrugineum</emphasis>
</taxonomicName>
and 
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
re-description of 
<taxonomicName baseAuthorityName="Lesson" baseAuthorityYear="1831" class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina ferruginea" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ferruginea">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina ferruginea</emphasis>
</taxonomicName>
are a confusing combination of traits that characterize species from two distinct genera. For instance, the dorsal gills mentioned by both authors, are characteristic of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
, while the absence of an accessory penial gland mentioned by 
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(even though there is a penial gland in the lectotype) is characteristic of 
<taxonomicName authorityName="Goulding &amp; Dayrat" authorityYear="2018" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Wallaconchis</emphasis>
</taxonomicName>
. Thanks to the designation of a lectotype with dorsal gills, the name 
<taxonomicName authorityName="Lesson" authorityYear="1831" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium ferrugineum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ferrugineum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium ferrugineum</emphasis>
</taxonomicName>
clearly applies to a 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species and 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina</emphasis>
</taxonomicName>
becomes a junior synonym of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Starobogatov, YI" journalOrPublisher="Soviet Journal of Marine Biology" pageId="0" pageNumber="1" pagination="206 - 212" refId="B171" refString="Starobogatov, YI, 1976. Composition and taxonomic position of marine pulmonate mollusks. Soviet Journal of Marine Biology 4: 206 - 212" title="Composition and taxonomic position of marine pulmonate mollusks." volume="4" year="1976">Starobogatov (1976</bibRefCitation>
: 211) created 
<taxonomicName authorityName="Starobogatov" authorityYear="1976" class="Gastropoda" family="Onchidiidae" genus="Quoyella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Quoyella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoyella</emphasis>
</taxonomicName>
as a replacement name of 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a, which he treated as a junior homonym of &quot; 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
Deshayes, 1843&quot; [
<taxonomicName lsidName="" pageId="0" pageNumber="1" phylum="Mollusca" rank="phylum">Mollusca</taxonomicName>
, 
<taxonomicName class="Gastropoda" lsidName="" pageId="0" pageNumber="1" rank="class">Gastropoda</taxonomicName>
, 
<taxonomicName family="Planaxidae" lsidName="" pageId="0" pageNumber="1" rank="family">Planaxidae</taxonomicName>
]. In the second edition of 
<normalizedToken originalValue="Lamarck’s">Lamarck's</normalizedToken>
<emphasis italics="true" pageId="0" pageNumber="1">
Histoire naturelle des animaux sans 
<normalizedToken originalValue="vertèbres">vertebres</normalizedToken>
</emphasis>
, Deshayes indicates that he originally thought of creating a new genus 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
but that, after all, he decided not to (
<bibRefCitation author="Deshayes, GP" journalOrPublisher="Bailliere, Paris" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/41660756" refId="B64" refString="Deshayes, GP, Milne-Edwards, H, 1845. Histoire Naturelle des Animaux sans Vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent, par J.B.P.A. de Lamarck. Deuxieme edition, Tome neuvieme. Histoire des Mollusques. J.-B. Bailliere, Paris, https://www.biodiversitylibrary.org/page/41660756" title="Histoire Naturelle des Animaux sans Vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent, par J. B. P. A. de Lamarck. Deuxieme edition, Tome neuvieme. Histoire des Mollusques. J. - B." url="https://www.biodiversitylibrary.org/page/41660756" year="1845">Deshayes and Milne-Edwards 1845</bibRefCitation>
: 236). Deshayes still used the binomen &quot; 
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1833" class="Gastropoda" family="Planaxidae" genus="Planaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Planaxis decollata" order="Neogastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="decollata">
<emphasis italics="true" pageId="0" pageNumber="1">Planaxis decollata</emphasis>
</taxonomicName>
Quoy&quot; (
<bibRefCitation author="Deshayes, GP" journalOrPublisher="Bailliere, Paris" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/41660756" refId="B64" refString="Deshayes, GP, Milne-Edwards, H, 1845. Histoire Naturelle des Animaux sans Vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent, par J.B.P.A. de Lamarck. Deuxieme edition, Tome neuvieme. Histoire des Mollusques. J.-B. Bailliere, Paris, https://www.biodiversitylibrary.org/page/41660756" title="Histoire Naturelle des Animaux sans Vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent, par J. B. P. A. de Lamarck. Deuxieme edition, Tome neuvieme. Histoire des Mollusques. J. - B." url="https://www.biodiversitylibrary.org/page/41660756" year="1845">Deshayes and Milne-Edwards 1845</bibRefCitation>
: 238). However, in the 
<emphasis italics="true" pageId="0" pageNumber="1">Explication des planches</emphasis>
of his 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="Traité">Traite</normalizedToken>
<normalizedToken originalValue="élémentaire">elementaire</normalizedToken>
de conchyliologie
</emphasis>
, Deshayes (1853: 50) used 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
for two valid species names: 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya decollata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="decollata">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya decollata</emphasis>
</taxonomicName>
and 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya grateloupi" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya grateloupi</emphasis>
</taxonomicName>
. Regardless, according to 
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="129 - 219" publicationUrl="https://www.biodiversitylibrary.org/page/12862913" refId="B85" refString="Gray, JE, 1847. A list of the genera of recent Mollusca, their synonyms and types. Proceedings of the Zoological Society of London 15: 129 - 219, https://www.biodiversitylibrary.org/page/12862913" title="A list of the genera of recent Mollusca, their synonyms and types." url="https://www.biodiversitylibrary.org/page/12862913" volume="15" year="1847">Gray (1847</bibRefCitation>
: 138), the generic name 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
by Deshayes is an incorrect subsequent spelling of his 
<taxonomicName authorityName="Gray" authorityYear="1839" class="Gastropoda" family="Planaxidae" genus="Quoyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoyia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoyia</emphasis>
</taxonomicName>
JE Gray, 1839. As an incorrect subsequent spelling, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
Deshayes is not available (
<bibRefCitation author="ICZN" journalOrPublisher="Bulletin of Zoological Nomenclature" pageId="0" pageNumber="1" refId="B97" refString="ICZN, 1999. International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." title="International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." year="1999">ICZN 1999</bibRefCitation>
: Article 33.3) and, as a result, 
<taxonomicName authorityName="Starobogatov" authorityYear="1976" class="Gastropoda" family="Onchidiidae" genus="Quoyella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Quoyella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoyella</emphasis>
</taxonomicName>
is an unnecessary replacement name. Ironically, 
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="129 - 219" publicationUrl="https://www.biodiversitylibrary.org/page/12862913" refId="B85" refString="Gray, JE, 1847. A list of the genera of recent Mollusca, their synonyms and types. Proceedings of the Zoological Society of London 15: 129 - 219, https://www.biodiversitylibrary.org/page/12862913" title="A list of the genera of recent Mollusca, their synonyms and types." url="https://www.biodiversitylibrary.org/page/12862913" volume="15" year="1847">Gray (1847</bibRefCitation>
: 138) indicated that he originally found the generic name 
<taxonomicName authorityName="Gray" authorityYear="1839" class="Gastropoda" family="Planaxidae" genus="Quoyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoyia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoyia</emphasis>
</taxonomicName>
in a manuscript by Deshayes in 1830 (&quot; 
<taxonomicName authorityName="Gray" authorityYear="1839" class="Gastropoda" family="Planaxidae" genus="Quoyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoyia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Quoyia</taxonomicName>
, 
<emphasis italics="true" pageId="0" pageNumber="1">Desh</emphasis>
. MSS. 1830; 
<emphasis italics="true" pageId="0" pageNumber="1">Gray</emphasis>
, 1839 (...) 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Quoya</taxonomicName>
, 
<emphasis italics="true" pageId="0" pageNumber="1">Desh</emphasis>
. 1843&quot;). According to 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.10914" author="Baker, HB" journalOrPublisher="The Nautilus" pageId="0" pageNumber="1" pagination="85 - 88" refId="B13" refString="Baker, HB, 1938. Nomenclature of Onchidiidae. The Nautilus 51: 85 - 88, DOI: https://doi.org/10.5962/bhl.part.10914" title="Nomenclature of Onchidiidae." url="https://doi.org/10.5962/bhl.part.10914" volume="51" year="1938">Baker (1938</bibRefCitation>
: 87), 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
Agassiz, 1862 [
<taxonomicName lsidName="" pageId="0" pageNumber="1" phylum="Coelenterata" rank="phylum">Coelenterata</taxonomicName>
] is another homonym of 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1834a. However, the spelling of that generic name is not 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
but 
<taxonomicName authorityName="Gray" authorityYear="1839" class="Gastropoda" family="Planaxidae" genus="Quoyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoyia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoyia</emphasis>
</taxonomicName>
(Agassiz, 1862: 173). So, 
<taxonomicName authorityName="Gray" authorityYear="1839" class="Gastropoda" family="Planaxidae" genus="Quoyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoyia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoyia</emphasis>
</taxonomicName>
Agassiz, 1862 is a junior homonym of 
<taxonomicName authorityName="Gray" authorityYear="1839" class="Gastropoda" family="Planaxidae" genus="Quoyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoyia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoyia</emphasis>
</taxonomicName>
Gray, 1839, but 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a is not a junior homonym of 
<taxonomicName authorityName="Gray" authorityYear="1839" class="Gastropoda" family="Planaxidae" genus="Quoyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoyia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoyia</emphasis>
</taxonomicName>
. 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya indica" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya indica</emphasis>
</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a, type species of 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
by monotypy, is regarded here as a 
<emphasis italics="true" pageId="0" pageNumber="1">nomen dubium</emphasis>
even though it applies to a species with dorsal gills and thus belongs to 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
(see general discussion).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Nothing is ever simple in onchidiid taxonomy. Indeed, 
<bibRefCitation author="Labbe, A" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="0" pageNumber="1" pagination="312 - 317" refId="B110" refString="Labbe, A, 1935a. Sur une forme nouvelle de Silicoderme, Elophilus ajuthiae nov. Gen. nov. sp. Bulletin de la Societe Zoologique de France 60: 312 - 317" title="Sur une forme nouvelle de Silicoderme, Elophilus ajuthiae nov. Gen. nov. sp." volume="60" year="1935 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1935a
</bibRefCitation>
, 
<bibRefCitation author="Labbe, A" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="0" pageNumber="1" pagination="318 - 320" refId="B111" refString="Labbe, A, 1935b. Les branchies des Silicodermes. Bulletin de la Societe Zoologique de France 60: 318 - 320" title="Les branchies des Silicodermes." volume="60" year="1935 b">b</bibRefCitation>
) also described what he called 
<normalizedToken originalValue="“microgills”">&quot;microgills&quot;</normalizedToken>
in 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Elophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elophilus" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Elophilus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1935a, a name preoccupied by 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Elophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elophilus" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Elophilus</emphasis>
</taxonomicName>
Meigen, 1803 (
<taxonomicName lsidName="" order="Diptera" pageId="0" pageNumber="1" rank="order">Diptera</taxonomicName>
) and replaced by 
<taxonomicName authorityName="Starobogatov" authorityYear="1970" class="Gastropoda" family="Onchidiidae" genus="Labbella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labbella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Labbella</emphasis>
</taxonomicName>
Starobogatov, 1970. 
<bibRefCitation author="Labbe, A" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="0" pageNumber="1" pagination="312 - 317" refId="B110" refString="Labbe, A, 1935a. Sur une forme nouvelle de Silicoderme, Elophilus ajuthiae nov. Gen. nov. sp. Bulletin de la Societe Zoologique de France 60: 312 - 317" title="Sur une forme nouvelle de Silicoderme, Elophilus ajuthiae nov. Gen. nov. sp." volume="60" year="1935 a">
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
(1935a
</bibRefCitation>
, 
<bibRefCitation author="Labbe, A" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="0" pageNumber="1" pagination="318 - 320" refId="B111" refString="Labbe, A, 1935b. Les branchies des Silicodermes. Bulletin de la Societe Zoologique de France 60: 318 - 320" title="Les branchies des Silicodermes." volume="60" year="1935 b">b</bibRefCitation>
) microgills consolidated the old idea of a gradual continuum between regular dorsal papillae and dorsal gills. So, for instance, 
<bibRefCitation author="Marcus, Er" journalOrPublisher="Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse" pageId="0" pageNumber="1" pagination="873 - 934" refId="B122" refString="Marcus, Er, Marcus, Ev, 1960. Opisthobranchia aus dem Roten Meer und von den Malediven. Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse 12: 873 - 934" title="Opisthobranchia aus dem Roten Meer und von den Malediven." volume="12" year="1960">Marcus and Marcus (1960</bibRefCitation>
: 875) argued that one cannot say for sure whether a papilla is a dorsal gill or not. However, 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.636.8879" author="Dayrat, B" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 40" refId="B52" refString="Dayrat, B, Goulding, TC, Apte, D, Bhave, V, Comendador, J, Ngo, XQ, Tan, SK, Tan, SH, 2016. Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae). ZooKeys 636: 1 - 40, DOI: https://doi.org/10.3897/zookeys.636.8879" title="Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae)." url="https://doi.org/10.3897/zookeys.636.8879" volume="636" year="2016">Dayrat et al. (2016</bibRefCitation>
, 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.892.39524" author="Dayrat, B" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="27 - 58" refId="B58" refString="Dayrat, B, Goulding, TC, Khalil, M, Apte, D, Tan, SH, 2019d. A new species and new records of Onchidium slugs (Gastropoda, Euthyneura, Pulmonata, Onchidiidae) in South-East Asia. ZooKeys 892: 27 - 58, DOI: https://doi.org/10.3897/zookeys.892.39524" title="A new species and new records of Onchidium slugs (Gastropoda, Euthyneura, Pulmonata, Onchidiidae) in South-East Asia." url="https://doi.org/10.3897/zookeys.892.39524" volume="892" year="2019 d">2019d</bibRefCitation>
) demonstrated that there are no gills at all (not even microgills) on the notum of the type material of the type species of 
<taxonomicName authorityName="Starobogatov" authorityYear="1970" class="Gastropoda" family="Onchidiidae" genus="Labbella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labbella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Labbella</emphasis>
</taxonomicName>
which actually belongs to 
<taxonomicName baseAuthorityName="Westerlund" baseAuthorityYear="1883" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium stuxbergi" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="stuxbergi">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium stuxbergi</emphasis>
</taxonomicName>
(Westerlund, 1883). 
<taxonomicName authorityName="Starobogatov" authorityYear="1970" class="Gastropoda" family="Onchidiidae" genus="Labbella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labbella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Labbella</emphasis>
</taxonomicName>
is a junior synonym of 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
. Contrary to regular papillae, dorsal gills are distinctively branched, which is striking if specimens are fully relaxed before preservation but otherwise difficult to see. Finally, note that 
<bibRefCitation author="Labbe, A" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="0" pageNumber="1" pagination="318 - 320" refId="B111" refString="Labbe, A, 1935b. Les branchies des Silicodermes. Bulletin de la Societe Zoologique de France 60: 318 - 320" title="Les branchies des Silicodermes." volume="60" year="1935 b">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1935b
</bibRefCitation>
: 320) claimed that he observed rudimentary eyes on dorsal gills, which, to our knowledge, has never been confirmed.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 187, 188) rightly recognized the importance of dorsal gills for classification and he separated all five genera of slugs with dorsal gills from all other onchidiids. According to 
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, onchidiids deserved their own order, the Silicodermatae, composed of two suborders: Dendrobranchiatae (onchidiids with dorsal gills) and Abranchiatae (onchidiids without dorsal gills). Our phylogenetic analyses clearly demonstrate that all species of slugs with dorsal gills belong to a single clade, and that only one generic name (
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
) is necessary (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogenetic relationships between Peronia species based on concatenated mitochondrial COI and 16 S DNA sequences. Numbers by the branches are the bootstrap values (maximum likelihood analysis) and the posterior probabilities (Bayesian analysis). Only the values&gt; 50 % (ML) and&gt; 0.9 (Bayesian) are indicated. Numbers for each individual correspond to unique identifiers for DNA extraction. Information on specimens can be found in the lists of material examined and in Table 2. The color used for each species or mitochondrial unit is the same as the color used in Figs 3 - 6." figureDoi="10.3897/zookeys.972.52853.figure2" httpUri="https://binary.pensoft.net/fig/458300" pageId="0" pageNumber="1">2</figureCitation>
- 
<figureCitation captionStart="Figure 4" captionStartId="F6" captionText="Figure 4. Maximum parsimony consensus tree within Peronia based on concatenated nuclear ITS 2 and 28 S sequences. Numbers by the branches are the bootstrap values. Only the values&gt; 50 % are indicated. Numbers for each individual correspond to unique identifiers for DNA extraction. Information on specimens can be found in the lists of material examined and in Table 2. The color used for each species or mitochondrial unit is the same as the color used in Figs 2, 3, 5, 6." figureDoi="10.3897/zookeys.972.52853.figure4" httpUri="https://binary.pensoft.net/fig/458304" pageId="0" pageNumber="1">4</figureCitation>
). However, the species of slugs with no dorsal gills do not form a natural group (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogenetic relationships between Peronia species based on concatenated mitochondrial COI and 16 S DNA sequences. Numbers by the branches are the bootstrap values (maximum likelihood analysis) and the posterior probabilities (Bayesian analysis). Only the values&gt; 50 % (ML) and&gt; 0.9 (Bayesian) are indicated. Numbers for each individual correspond to unique identifiers for DNA extraction. Information on specimens can be found in the lists of material examined and in Table 2. The color used for each species or mitochondrial unit is the same as the color used in Figs 3 - 6." figureDoi="10.3897/zookeys.972.52853.figure2" httpUri="https://binary.pensoft.net/fig/458300" pageId="0" pageNumber="1">2</figureCitation>
- 
<figureCitation captionStart="Figure 4" captionStartId="F6" captionText="Figure 4. Maximum parsimony consensus tree within Peronia based on concatenated nuclear ITS 2 and 28 S sequences. Numbers by the branches are the bootstrap values. Only the values&gt; 50 % are indicated. Numbers for each individual correspond to unique identifiers for DNA extraction. Information on specimens can be found in the lists of material examined and in Table 2. The color used for each species or mitochondrial unit is the same as the color used in Figs 2, 3, 5, 6." figureDoi="10.3897/zookeys.972.52853.figure4" httpUri="https://binary.pensoft.net/fig/458304" pageId="0" pageNumber="1">4</figureCitation>
). In other words, the absence of dorsal gills is a plesiomorphic trait for the onchidiids and the presence of dorsal gills is a synapomorphy for the genus 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
(1934a
</bibRefCitation>
: 187) distinction between the tribes 
<taxonomicName authorityName="Keferstein" authorityYear="1865" family="Peroniidae" lsidName="" pageId="0" pageNumber="1" rank="family">Peroniidae</taxonomicName>
(
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
and 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia</emphasis>
</taxonomicName>
) and 
<taxonomicName authorityName="Labbe" authorityYear="1934" family="Scaphidae" lsidName="" pageId="0" pageNumber="1" rank="family">Scaphidae</taxonomicName>
(
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Scaphis</emphasis>
</taxonomicName>
, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina</emphasis>
</taxonomicName>
, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
) based on the orientation of the hyponotum (horizontal versus oblique) is meaningless. This trait obviously varies depending on preservation, and 
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
exclusively studied preserved material from the collections of the MNHN without access to live animals.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
(1934a
</bibRefCitation>
: 187) distinction between 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
and 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia</emphasis>
</taxonomicName>
based on the intestinal types (type I in 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
and type V in 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia</emphasis>
</taxonomicName>
) is unwarranted because 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species with intestinal loops of type V are not more closely-related to each other (Table 
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of traits that can help identify Peronia species, based exclusively on specimens examined for the present revision. Species are arranged in a phylogenetic order (Figs 2 - 4). Traits are described in detail in species descriptions." httpUri="http://table.plazi.org/id/8BDBED7FCCE9A8E1C67A716B088DDD7D" pageId="0" pageNumber="1" tableUuid="8BDBED7FCCE9A8E1C67A716B088DDD7D">4</tableCitation>
, Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogenetic relationships between Peronia species based on concatenated mitochondrial COI and 16 S DNA sequences. Numbers by the branches are the bootstrap values (maximum likelihood analysis) and the posterior probabilities (Bayesian analysis). Only the values&gt; 50 % (ML) and&gt; 0.9 (Bayesian) are indicated. Numbers for each individual correspond to unique identifiers for DNA extraction. Information on specimens can be found in the lists of material examined and in Table 2. The color used for each species or mitochondrial unit is the same as the color used in Figs 3 - 6." figureDoi="10.3897/zookeys.972.52853.figure2" httpUri="https://binary.pensoft.net/fig/458300" pageId="0" pageNumber="1">2</figureCitation>
- 
<figureCitation captionStart="Figure 4" captionStartId="F6" captionText="Figure 4. Maximum parsimony consensus tree within Peronia based on concatenated nuclear ITS 2 and 28 S sequences. Numbers by the branches are the bootstrap values. Only the values&gt; 50 % are indicated. Numbers for each individual correspond to unique identifiers for DNA extraction. Information on specimens can be found in the lists of material examined and in Table 2. The color used for each species or mitochondrial unit is the same as the color used in Figs 2, 3, 5, 6." figureDoi="10.3897/zookeys.972.52853.figure4" httpUri="https://binary.pensoft.net/fig/458304" pageId="0" pageNumber="1">4</figureCitation>
). Also, 
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
often made mistakes with respect to intestinal types: for instance, the type material of 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia gondwanae" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gondwanae">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia gondwanae</emphasis>
</taxonomicName>
includes individuals with loops of both types I and V, even though 
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
described it as a species with loops of type V. 
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
asserted that the position of the pneumostome and the size of the muscular sac differ between 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
and 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia</emphasis>
</taxonomicName>
. However, the position of the pneumostome varies between individuals and is not consistently on the right side of the median axis in species he classified as 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
(1934a
</bibRefCitation>
: 187) distinction between 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Scaphis</emphasis>
</taxonomicName>
, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
, and 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina</emphasis>
</taxonomicName>
, is also unwarranted. Again, the position of the pneumostome (on the right of a median line in 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Scaphis</emphasis>
</taxonomicName>
according to 
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
) varies between individuals. 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
(1934a)
</bibRefCitation>
re-description of 
<taxonomicName baseAuthorityName="Lesson" baseAuthorityYear="1831" class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina ferruginea" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ferruginea">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina ferruginea</emphasis>
</taxonomicName>
(the type species of 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina</emphasis>
</taxonomicName>
, by monotypy) was based on individuals of two different species (see above). The male opening of the lectotype, which bears dorsal gills, is on the left of the right ocular tentacle, exactly as in all 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species, while the male opening of the two paralectotypes, which belong to 
<taxonomicName baseAuthorityName="Lesson" baseAuthorityYear="1831" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">
<emphasis italics="true" pageId="0" pageNumber="1">Wallaconchis ater</emphasis>
</taxonomicName>
, is under the right ocular tentacle (Goulding et al. 2018: 75). 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 216, fig. 51) described a double male opening in 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya indica" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="indica">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya indica</emphasis>
</taxonomicName>
(the openings of the penis and of the accessory penial gland being supposedly separated), but this could not be confirmed in the type material. Regardless, male openings occasionally appear separated due to preservation (when the vestibule is everted) and that is by no means a trait of generic value.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Authors completely rejected 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
(1934a)
</bibRefCitation>
idea that the presence or absence of dorsal gills could be of any use in onchidiid classification (e.g., 
<bibRefCitation author="Marcus, Er" journalOrPublisher="Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse" pageId="0" pageNumber="1" pagination="873 - 934" refId="B122" refString="Marcus, Er, Marcus, Ev, 1960. Opisthobranchia aus dem Roten Meer und von den Malediven. Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse 12: 873 - 934" title="Opisthobranchia aus dem Roten Meer und von den Malediven." volume="12" year="1960">Marcus and Marcus 1960</bibRefCitation>
; 
<bibRefCitation author="Starobogatov, YI" journalOrPublisher="Soviet Journal of Marine Biology" pageId="0" pageNumber="1" pagination="206 - 212" refId="B171" refString="Starobogatov, YI, 1976. Composition and taxonomic position of marine pulmonate mollusks. Soviet Journal of Marine Biology 4: 206 - 212" title="Composition and taxonomic position of marine pulmonate mollusks." volume="4" year="1976">Starobogatov 1976</bibRefCitation>
). 
<bibRefCitation DOI="https://doi.org/10.1093/oxfordjournals.mollus.a065863" author="Britton, KM" journalOrPublisher="Journal of Molluscan Studies" pageId="0" pageNumber="1" pagination="179 - 191" refId="B27" refString="Britton, KM, 1984. The Onchidiacea (Gastropoda, Pulmonata) of Hong Kong with a worldwide review of the genera. Journal of Molluscan Studies 50: 179 - 191, DOI: https://doi.org/10.1093/oxfordjournals.mollus.a065863" title="The Onchidiacea (Gastropoda, Pulmonata) of Hong Kong with a worldwide review of the genera." url="https://doi.org/10.1093/oxfordjournals.mollus.a065863" volume="50" year="1984">Britton (1984</bibRefCitation>
: 180) even asserted that &quot;the division of the group into two subordinate taxa based on this character is no longer admissible.&quot; As for the status of 
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé’s">Labbe's</normalizedToken>
(1934a
</bibRefCitation>
, 
<bibRefCitation author="Labbe, A" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="0" pageNumber="1" pagination="312 - 317" refId="B110" refString="Labbe, A, 1935a. Sur une forme nouvelle de Silicoderme, Elophilus ajuthiae nov. Gen. nov. sp. Bulletin de la Societe Zoologique de France 60: 312 - 317" title="Sur une forme nouvelle de Silicoderme, Elophilus ajuthiae nov. Gen. nov. sp." volume="60" year="1935 a">1935a</bibRefCitation>
) generic names for slugs with dorsal gills, authors were not in agreement. 
<bibRefCitation author="Marcus, Ev" journalOrPublisher="Malacologia" pageId="0" pageNumber="1" pagination="181 - 223" publicationUrl="https://www.biodiversitylibrary.org/page/13147904" refId="B123" refString="Marcus, Ev, Marcus, Er, 1970. Some gastropods from Madagascar and west Mexico. Malacologia 10: 181 - 223, https://www.biodiversitylibrary.org/page/13147904" title="Some gastropods from Madagascar and west Mexico." url="https://www.biodiversitylibrary.org/page/13147904" volume="10" year="1970">Marcus and Marcus (1970</bibRefCitation>
: 213) regarded 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
and 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia</emphasis>
</taxonomicName>
&quot;at most as subgenera.&quot; 
<bibRefCitation author="Starobogatov, YI" journalOrPublisher="Soviet Journal of Marine Biology" pageId="0" pageNumber="1" pagination="206 - 212" refId="B171" refString="Starobogatov, YI, 1976. Composition and taxonomic position of marine pulmonate mollusks. Soviet Journal of Marine Biology 4: 206 - 212" title="Composition and taxonomic position of marine pulmonate mollusks." volume="4" year="1976">Starobogatov (1976)</bibRefCitation>
regarded all names as valid: 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina</emphasis>
</taxonomicName>
, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Starobogatov" authorityYear="1976" class="Gastropoda" family="Onchidiidae" genus="Quoyella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Quoyella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoyella</emphasis>
</taxonomicName>
(unnecessary replacement name for 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
), 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Scaphis</emphasis>
</taxonomicName>
, and 
<taxonomicName authorityName="Starobogatov" authorityYear="1970" class="Gastropoda" family="Onchidiidae" genus="Labbella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labbella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Labbella</emphasis>
</taxonomicName>
(supposedly with micro-gills). 
<bibRefCitation DOI="https://doi.org/10.1093/oxfordjournals.mollus.a065863" author="Britton, KM" journalOrPublisher="Journal of Molluscan Studies" pageId="0" pageNumber="1" pagination="179 - 191" refId="B27" refString="Britton, KM, 1984. The Onchidiacea (Gastropoda, Pulmonata) of Hong Kong with a worldwide review of the genera. Journal of Molluscan Studies 50: 179 - 191, DOI: https://doi.org/10.1093/oxfordjournals.mollus.a065863" title="The Onchidiacea (Gastropoda, Pulmonata) of Hong Kong with a worldwide review of the genera." url="https://doi.org/10.1093/oxfordjournals.mollus.a065863" volume="50" year="1984">Britton (1984</bibRefCitation>
: 182-183) suggested that 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Gistel" authorityYear="1848" class="Gastropoda" family="Onchidiidae" genus="Eudrastus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eudrastus" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Eudrastus</emphasis>
</taxonomicName>
and 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Scaphis</emphasis>
</taxonomicName>
should be regarded as junior synonyms of 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
because they seemed to be based on &quot;unimportant characters,&quot; but treated 
<taxonomicName authorityName="Starobogatov" authorityYear="1970" class="Gastropoda" family="Onchidiidae" genus="Labbella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labbella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Labbella</emphasis>
</taxonomicName>
(supposedly with micro-gills), 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina</emphasis>
</taxonomicName>
, and 
<taxonomicName authorityName="Starobogatov" authorityYear="1976" class="Gastropoda" family="Onchidiidae" genus="Quoyella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Quoyella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoyella</emphasis>
</taxonomicName>
(for 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
) as valid. In a recent review of the application of onchidiid generic names, 
<bibRefCitation DOI="https://doi.org/10.1080/00222933.2017.1347297" author="Dayrat, B" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="1" pagination="1851 - 1897" refId="B53" refString="Dayrat, B, Goulding, TC, Apte, D, Bhave, V, Ngo, XQ, 2017. A new genus and four new species of onchidiid slugs from South-East Asia (Mollusca: Gastropoda: Pulmonata: Onchidiidae). Journal of Natural History 51: 1851 - 1897, DOI: https://doi.org/10.1080/00222933.2017.1347297" title="A new genus and four new species of onchidiid slugs from South-East Asia (Mollusca: Gastropoda: Pulmonata: Onchidiidae)." url="https://doi.org/10.1080/00222933.2017.1347297" volume="51" year="2017">Dayrat et al. (2017</bibRefCitation>
: 1861) made it clear that all slugs with dorsal gills belong to one clade and that 
<taxonomicName authorityName="Gistel" authorityYear="1848" class="Gastropoda" family="Onchidiidae" genus="Eudrastus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eudrastus" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Eudrastus</emphasis>
</taxonomicName>
, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchis</emphasis>
</taxonomicName>
, 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Starobogatov" authorityYear="1976" class="Gastropoda" family="Onchidiidae" genus="Quoyella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Quoyella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoyella</emphasis>
</taxonomicName>
(for 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Quoya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quoya" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Quoya</emphasis>
</taxonomicName>
), and 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Scaphis</emphasis>
</taxonomicName>
all refer to that clade. Note that the application of 
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lessonina</emphasis>
</taxonomicName>
was fully clarified when a lectotype was designated for its type species 
<taxonomicName authorityName="Lesson" authorityYear="1831" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium ferrugineum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ferrugineum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium ferrugineum</emphasis>
</taxonomicName>
(Goulding et al. 2018: 75).
</paragraph>
</subSubSection>
</treatment>
</document>