241 lines
46 KiB
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241 lines
46 KiB
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<document id="B327929E4BC6416665714D600D306BDC" ID-DOI="10.5852/ejt.2020.626" ID-GBIF-Dataset="c906d9cd-cfcb-43db-a24c-b57b39d9184b" ID-Zenodo-Dep="3754989" ID-ZooBank="D91881D6-55D8-48FC-A383-069BC643A91E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587066773978" checkinUser="valdenar" docAuthor="Enghoff, Henrik" docDate="2020" docId="5C11879EFFB5FFDEFD84EF7DFDF610CB" docLanguage="en" docName="ejt-626_enghoff.pdf.imf" docOrigin="European Journal of Taxonomy 626" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Aquattuor Frederiksen 2013" docType="treatment" docVersion="6" lastPageNumber="5" masterDocId="A028FFE6FFB6FFDAFFC1ED40FFC7143C" masterDocTitle="A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="32" masterPageNumber="1" pageNumber="4" updateTime="1698738618004" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A3DF22C917D83E511DD5980FFBBD1B40">A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<mods:namePart id="D7E9F9241BDB377545D49ECE4C723072">Enghoff, Henrik</mods:namePart>
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<mods:title id="46ECBF6E8400E0935CB5B76CFDC45FAA">European Journal of Taxonomy</mods:title>
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<mods:date id="C4E14E53111AB981188023D37BB2A6D3">2020</mods:date>
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<mods:number id="73D88F32B58A8A45080DDD867FE21C47">2020-04-14</mods:number>
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<treatment id="5C11879EFFB5FFDEFD84EF7DFDF610CB" ID-DOI="http://doi.org/10.5281/zenodo.3808346" ID-GBIF-Taxon="163642158" ID-Zenodo-Dep="3808346" LSID="urn:lsid:plazi:treatment:5C11879EFFB5FFDEFD84EF7DFDF610CB" httpUri="http://treatment.plazi.org/id/5C11879EFFB5FFDEFD84EF7DFDF610CB" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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<subSubSection id="9CA26503FFB5FFD9FD84EF7DFC28166B" box="[581,1007,573,599]" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="D4073688FFB5FFD9FD84EF7DFC28166B" blockId="3.[581,1007,573,599]" box="[581,1007,573,599]" pageId="3" pageNumber="4">
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Genus
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<taxonomicName id="13B84D0BFFB5FFD9FD56EF7DFC28166B" ID-CoL="7NQQQ" authority="Frederiksen, 2013" authorityName="Frederiksen" authorityYear="2013" box="[663,1007,573,599]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="E6CCEA9AFFB5FFD9FD56EF7DFCD4166B" bold="true" box="[663,787,573,599]" italics="true" pageId="3" pageNumber="4">Aquattuor</emphasis>
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<bibRefCitation id="B0294B79FFB5FFD9FCDAEF7DFC28166B" author="Frederiksen S. B." box="[795,1007,573,599]" pageId="3" pageNumber="4" pagination="59 - 66" refId="ref12038" refString="Frederiksen S. B. 2013. East African odontopygid millipedes 3: Two new genera, Lamelloramus and Aquattuor proposed to contain three new species (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3694: 59 - 66. https: // doi. org / 10.11646 / zootaxa. 3694.1.4" type="journal article" year="2013">Frederiksen, 2013</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="9CA26503FFB5FFDEFF7CEF3EFDF610CB" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph id="D4073688FFB5FFD9FF7CEF3EFC8816A5" blockId="3.[189,847,638,665]" box="[189,847,638,665]" pageId="3" pageNumber="4">
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<emphasis id="E6CCEA9AFFB5FFD9FF7CEF3EFEF116A4" bold="true" box="[189,310,638,664]" pageId="3" pageNumber="4">Diagnosis</emphasis>
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(modified after
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<bibRefCitation id="B0294B79FFB5FFD9FE32EF3EFC8116A5" author="Enghoff H. & Frederiksen S. B." box="[499,838,638,665]" pageId="3" pageNumber="4" pagination="1 - 25" refId="ref11852" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff & Frederiksen 2015</bibRefCitation>
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)
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</paragraph>
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<paragraph id="D4073688FFB5FFD9FF7CEFEDFC7B1768" blockId="3.[189,1399,685,852]" pageId="3" pageNumber="4">
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Differs from all other
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<taxonomicName id="13B84D0BFFB5FFD9FE7AEFEDFDB716FB" authorityName="Attems" authorityYear="1909" box="[443,624,685,711]" class="Diplopoda" family="Odontopygidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Odontopygidae</taxonomicName>
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by the structure of the limbus, which consists of large, rectangular, easily detachable flaps. Gonopods of the
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<taxonomicName id="13B84D0BFFB5FFD9FD61EF90FC8116D6" authorityName="Hoffman" authorityYear="1991" box="[672,838,720,746]" class="Diplopoda" family="Odontopygidae" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Prionopetalini">Prionopetalini</taxonomicName>
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<typeStatus id="0B03882AFFB5FFD9FC8FEF91FC4416D7" box="[846,899,721,747]" pageId="3" pageNumber="4">type</typeStatus>
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, i.e., basomere of telopodite with a torsate flexible zone followed by a posttorsal narrowing; solenomere elongated. Gonopodal proplica apically expanded into a distal palette; metaplica with an oblique flange; solenomere simple, thin and whip-like; telomere terminating in a long, curved, gutter-like to tubular part.
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</paragraph>
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<paragraph id="D4073688FFB5FFD9FF7CEE3BFE5417A9" blockId="3.[189,403,891,917]" box="[189,403,891,917]" pageId="3" pageNumber="4">
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<heading id="8F4F81E4FFB5FFD9FF7CEE3BFE5417A9" bold="true" box="[189,403,891,917]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
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<emphasis id="E6CCEA9AFFB5FFD9FF7CEE3BFE5417A9" bold="true" box="[189,403,891,917]" pageId="3" pageNumber="4">Descriptive notes</emphasis>
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</heading>
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</paragraph>
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<paragraph id="D4073688FFB5FFD9FF7CEEEAFC3117F9" blockId="3.[189,1014,938,965]" box="[189,1014,938,965]" pageId="3" pageNumber="4">
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For characters not mentioned here, see
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<bibRefCitation id="B0294B79FFB5FFD9FD48EEEAFC3517F9" author="Enghoff H. & Frederiksen S. B." box="[649,1010,938,965]" pageId="3" pageNumber="4" pagination="1 - 25" refId="ref11852" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff & Frederiksen (2015)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="D4073688FFB5FFD9FF7CEEABFB721039" blockId="3.[189,1205,1003,1030]" box="[189,1205,1003,1030]" pageId="3" pageNumber="4">
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Mandibular stipes (
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<figureCitation id="4C832A0DFFB5FFD9FE61EEABFE2E1039" box="[416,489,1003,1029]" captionStart="Fig" captionStartId="3.[189,232,1913,1939]" captionTargetBox="[195,1389,1227,1874]" captionTargetId="figure@3.[189,1399,1215,1882]" captionTargetPageId="3" captionText="Fig. 2. Aquattuor udzungwensis Enghoff, 2015, specimens from Chita, heads, lateral view. A. Male (NHMD 621667). B. Female (NHMD 621667). Scale bars = 0.1 mm. The arrow points at the distoventral stipital expansion lobe in the male." figureDoi="http://doi.org/10.5281/zenodo.3754993" httpUri="https://zenodo.org/record/3754993/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
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): in males, but not in females with a disto-ventral expansion.
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</paragraph>
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<paragraph id="D4073688FFB5FFDEFF7CE96DFCC815EA" blockId="3.[189,1399,1069,1166]" lastBlockId="4.[189,1398,267,470]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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First pair of male legs: in the genus
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<taxonomicName id="13B84D0BFFB5FFD9FDBDE96DFD34107B" authorityName="Frederiksen" authorityYear="2013" box="[636,755,1069,1095]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="E6CCEA9AFFB5FFD9FDBDE96DFD34107B" box="[636,755,1069,1095]" italics="true" pageId="3" pageNumber="4">Aquattuor</emphasis>
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</taxonomicName>
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the first pair of legs is remarkably diverse.
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<figureCitation id="4C832A0DFFB5FFD9FACCE96DFAB1107B" box="[1293,1398,1069,1095]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="5.[189,232,1849,1875]" captionStartId-1="6.[189,232,1890,1916]" captionStartId-2="7.[189,232,1887,1913]" captionStartId-3="8.[189,232,1887,1913]" captionStartId-4="9.[189,232,1877,1903]" captionTargetBox-0="[205,1381,684,1788]" captionTargetBox-1="[226,1373,279,1795]" captionTargetBox-2="[234,1343,270,1842]" captionTargetBox-3="[200,1384,275,1839]" captionTargetBox-4="[206,1353,584,1811]" captionTargetId-0="figure@5.[189,1399,675,1801]" captionTargetId-1="figure@6.[189,1399,265,1821]" captionTargetId-2="figure@7.[229,1354,262,1848]" captionTargetId-3="figure@8.[189,1399,262,1849]" captionTargetId-4="figure@9.[189,1399,573,1842]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="Fig. 3. Aquattuor spp., first pair of male legs. A–C. A. fasciatus (Attems, 1896), non-type male from Zanzibar (NHMD 621641). D–F. A. claudiahempae Enghoff & Frederiksen, 2015, paratype from Kilimanjaro (NHMD 621638). G–I. A. claudiahempae, specimen from Udzungwa Mts (NHMD 621639). A, D, G. Anterior view. B, E, H. Sublateral view. C, F, I. Ventral view. Abbreviation: pfp = prefemoral process. Scale bars = 0.1 mm." captionText-1="Fig. 4. A–C. Aquattuor nguruensis sp. nov., holotype, ♂ (VMNH110617), first pair of legs. Notice mite larva (”hypopus”) on left femur. D–F. A. denticulatus Frederiksen, 2013, paratype, ♂ (NHMD 621640), left leg of first pair. A, D. Anterior view. B, E. Sublateral view. C, F. Sub-ventral view. Abbreviation: pfd = prefemoral depression. Scale bars = 0.1 mm." captionText-2="Fig. 5. Aquattuor spp., first pair of male legs. A–C. A. major Enghoff, 2015, paratype, ♂ (NHMD 621644), first pair of legs in situ. D–F. A. longipala Enghoff, 2015, male from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642). A, D. Anterior view. B, E. Sublateral view. C, F. Sub-ventral view. Abbreviations: dpl = distal prefemoral lobe; pfp = prefemoral process. Scale bars = 0.1 mm." captionText-3="Fig. 6. Aquattuor spp., first pair of male legs. A–C. A. mollilobus sp. nov., paratype, ♂ (NHMD 621647). D–F. A. stereosathe Enghoff, 2015, paratype, ♂ (NHMD 621650). A, D. Anterior view. B, E. (Sub)- lateral view. C, F. (Sub)-ventral view. Abbreviations: dpl = distal prefemoral lobe; pfp = prefemoral process. Scale bars = 0.1 mm." captionText-4="Fig. 7. Aquattuor spp., first pair of male legs. A–C. A. submajor Enghoff, 2015, paratype, ♂ (NHMD 621651). A. Anterior view. B. Sublateral view. C. Ventral view. D–E. A. udzungwensis Enghoff, 2015, male from Udzungwa Mts National Park, Kidatu (NHMD 621662). D. Anterior view. E. Ventral view. Scale bars = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3754995" figureDoi-1="http://doi.org/10.5281/zenodo.3754997" figureDoi-2="http://doi.org/10.5281/zenodo.3754999" figureDoi-3="http://doi.org/10.5281/zenodo.3755001" figureDoi-4="http://doi.org/10.5281/zenodo.3755003" httpUri-0="https://zenodo.org/record/3754995/files/figure.png" httpUri-1="https://zenodo.org/record/3754997/files/figure.png" httpUri-2="https://zenodo.org/record/3754999/files/figure.png" httpUri-3="https://zenodo.org/record/3755001/files/figure.png" httpUri-4="https://zenodo.org/record/3755003/files/figure.png" pageId="3" pageNumber="4">Figs 3–7</figureCitation>
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illustrate the first pair of legs in all known species of the genus, with a focus on the prefemur. In all of them, the prefemur carries a more or less finger-shaped process (
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<emphasis id="E6CCEA9AFFB5FFD9FC24E933FBCD10B1" box="[997,1034,1139,1165]" italics="true" pageId="3" pageNumber="4">pfp</emphasis>
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) which projects orad from the anterior surface of the prefemur (
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<figureCitation id="4C832A0DFFB2FFDEFD8DEC4BFD7D1519" box="[588,698,267,293]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="5.[189,232,1849,1875]" captionStartId-1="6.[189,232,1890,1916]" captionStartId-2="7.[189,232,1887,1913]" captionStartId-3="8.[189,232,1887,1913]" captionStartId-4="9.[189,232,1877,1903]" captionTargetBox-0="[205,1381,684,1788]" captionTargetBox-1="[226,1373,279,1795]" captionTargetBox-2="[234,1343,270,1842]" captionTargetBox-3="[200,1384,275,1839]" captionTargetBox-4="[206,1353,584,1811]" captionTargetId-0="figure@5.[189,1399,675,1801]" captionTargetId-1="figure@6.[189,1399,265,1821]" captionTargetId-2="figure@7.[229,1354,262,1848]" captionTargetId-3="figure@8.[189,1399,262,1849]" captionTargetId-4="figure@9.[189,1399,573,1842]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="Fig. 3. Aquattuor spp., first pair of male legs. A–C. A. fasciatus (Attems, 1896), non-type male from Zanzibar (NHMD 621641). D–F. A. claudiahempae Enghoff & Frederiksen, 2015, paratype from Kilimanjaro (NHMD 621638). G–I. A. claudiahempae, specimen from Udzungwa Mts (NHMD 621639). A, D, G. Anterior view. B, E, H. Sublateral view. C, F, I. Ventral view. Abbreviation: pfp = prefemoral process. Scale bars = 0.1 mm." captionText-1="Fig. 4. A–C. Aquattuor nguruensis sp. nov., holotype, ♂ (VMNH110617), first pair of legs. Notice mite larva (”hypopus”) on left femur. D–F. A. denticulatus Frederiksen, 2013, paratype, ♂ (NHMD 621640), left leg of first pair. A, D. Anterior view. B, E. Sublateral view. C, F. Sub-ventral view. Abbreviation: pfd = prefemoral depression. Scale bars = 0.1 mm." captionText-2="Fig. 5. Aquattuor spp., first pair of male legs. A–C. A. major Enghoff, 2015, paratype, ♂ (NHMD 621644), first pair of legs in situ. D–F. A. longipala Enghoff, 2015, male from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642). A, D. Anterior view. B, E. Sublateral view. C, F. Sub-ventral view. Abbreviations: dpl = distal prefemoral lobe; pfp = prefemoral process. Scale bars = 0.1 mm." captionText-3="Fig. 6. Aquattuor spp., first pair of male legs. A–C. A. mollilobus sp. nov., paratype, ♂ (NHMD 621647). D–F. A. stereosathe Enghoff, 2015, paratype, ♂ (NHMD 621650). A, D. Anterior view. B, E. (Sub)- lateral view. C, F. (Sub)-ventral view. Abbreviations: dpl = distal prefemoral lobe; pfp = prefemoral process. Scale bars = 0.1 mm." captionText-4="Fig. 7. Aquattuor spp., first pair of male legs. A–C. A. submajor Enghoff, 2015, paratype, ♂ (NHMD 621651). A. Anterior view. B. Sublateral view. C. Ventral view. D–E. A. udzungwensis Enghoff, 2015, male from Udzungwa Mts National Park, Kidatu (NHMD 621662). D. Anterior view. E. Ventral view. Scale bars = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3754995" figureDoi-1="http://doi.org/10.5281/zenodo.3754997" figureDoi-2="http://doi.org/10.5281/zenodo.3754999" figureDoi-3="http://doi.org/10.5281/zenodo.3755001" figureDoi-4="http://doi.org/10.5281/zenodo.3755003" httpUri-0="https://zenodo.org/record/3754995/files/figure.png" httpUri-1="https://zenodo.org/record/3754997/files/figure.png" httpUri-2="https://zenodo.org/record/3754999/files/figure.png" httpUri-3="https://zenodo.org/record/3755001/files/figure.png" httpUri-4="https://zenodo.org/record/3755003/files/figure.png" pageId="4" pageNumber="5">Figs 3–7</figureCitation>
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). The distal (ventral) surface of the process is smooth, the basal (dorsal) surface is more or less scaly-rugose. The shape of the process varies from slender fingershaped to relatively broad/triangular, and in one species (
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<taxonomicName id="13B84D0BFFB2FFDEFC40EC12FBE01550" authority="Enghoff, 2020" authorityName="Enghoff" authorityYear="2020" box="[897,1063,338,364]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="nguruensis" status="sp. nov.">
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<emphasis id="E6CCEA9AFFB2FFDEFC40EC12FBE01550" box="[897,1063,338,364]" italics="true" pageId="4" pageNumber="5">A. nguruensis</emphasis>
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</taxonomicName>
|
||
<taxonomicNameLabel id="FDFF57E1FFB2FFDEFBF2EC12FB541550" box="[1075,1171,338,364]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) there is a clearly delimited depression (
|
||
<emphasis id="E6CCEA9AFFB2FFDEFE03EC35FE2015B3" box="[450,487,373,399]" italics="true" pageId="4" pageNumber="5">pfd</emphasis>
|
||
) on the distal surface of the process (
|
||
<figureCitation id="4C832A0DFFB2FFDEFC60EC35FC2015B3" box="[929,999,373,399]" captionStart="Fig" captionStartId="6.[189,232,1890,1916]" captionTargetBox="[226,1373,279,1795]" captionTargetId="figure@6.[189,1399,265,1821]" captionTargetPageId="6" captionText="Fig. 4. A–C. Aquattuor nguruensis sp. nov., holotype, ♂ (VMNH110617), first pair of legs. Notice mite larva (”hypopus”) on left femur. D–F. A. denticulatus Frederiksen, 2013, paratype, ♂ (NHMD 621640), left leg of first pair. A, D. Anterior view. B, E. Sublateral view. C, F. Sub-ventral view. Abbreviation: pfd = prefemoral depression. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3754997" httpUri="https://zenodo.org/record/3754997/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
|
||
A–C). In
|
||
<taxonomicName id="13B84D0BFFB2FFDEFB96EC35FAB515AC" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[1111,1394,373,400]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="major">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFB96EC35FB7915B3" box="[1111,1214,373,399]" italics="true" pageId="4" pageNumber="5">A. major</emphasis>
|
||
<bibRefCitation id="B0294B79FFB2FFDEFB04EC35FAB515AC" author="Enghoff H. & Frederiksen S. B." box="[1221,1394,373,400]" pageId="4" pageNumber="5" pagination="1 - 25" refId="ref11852" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13B84D0BFFB2FFDEFF7CECD9FDC5158F" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[189,514,408,435]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="longipala">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFF7CECD9FE89158E" box="[189,334,408,435]" italics="true" pageId="4" pageNumber="5">A. longipala</emphasis>
|
||
<bibRefCitation id="B0294B79FFB2FFDEFE96ECD8FDC5158F" author="Enghoff H. & Frederiksen S. B." box="[343,514,408,435]" pageId="4" pageNumber="5" pagination="1 - 25" refId="ref11852" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="13B84D0BFFB2FFDEFDFCECD9FD1D158E" authority="Enghoff, 2020" authorityName="Enghoff" authorityYear="2020" box="[573,730,408,435]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mollilobus" status="sp. nov.">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFDFCECD9FD1D158E" box="[573,730,408,435]" italics="true" pageId="4" pageNumber="5">A. mollilobus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FDFF57E1FFB2FFDEFD22ECD9FCF8158F" box="[739,831,409,435]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
there is in addition a second, more distal, more irregularly shaped prefemoral lobe (
|
||
<emphasis id="E6CCEA9AFFB2FFDEFDA5ECFBFD4E15E9" box="[612,649,443,469]" italics="true" pageId="4" pageNumber="5">dpl</emphasis>
|
||
,
|
||
<figureCitation id="4C832A0DFFB2FFDEFD59ECFBFCC415EA" box="[664,771,443,470]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="7.[189,232,1887,1913]" captionStartId-1="8.[189,232,1887,1913]" captionTargetBox-0="[234,1343,270,1842]" captionTargetBox-1="[200,1384,275,1839]" captionTargetId-0="figure@7.[229,1354,262,1848]" captionTargetId-1="figure@8.[189,1399,262,1849]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Fig. 5. Aquattuor spp., first pair of male legs. A–C. A. major Enghoff, 2015, paratype, ♂ (NHMD 621644), first pair of legs in situ. D–F. A. longipala Enghoff, 2015, male from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642). A, D. Anterior view. B, E. Sublateral view. C, F. Sub-ventral view. Abbreviations: dpl = distal prefemoral lobe; pfp = prefemoral process. Scale bars = 0.1 mm." captionText-1="Fig. 6. Aquattuor spp., first pair of male legs. A–C. A. mollilobus sp. nov., paratype, ♂ (NHMD 621647). D–F. A. stereosathe Enghoff, 2015, paratype, ♂ (NHMD 621650). A, D. Anterior view. B, E. (Sub)- lateral view. C, F. (Sub)-ventral view. Abbreviations: dpl = distal prefemoral lobe; pfp = prefemoral process. Scale bars = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3754999" figureDoi-1="http://doi.org/10.5281/zenodo.3755001" httpUri-0="https://zenodo.org/record/3754999/files/figure.png" httpUri-1="https://zenodo.org/record/3755001/files/figure.png" pageId="4" pageNumber="5">Figs 5–6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="80C76600FFB5FFD9FF7CEA39FD6813E6" ID-DOI="http://doi.org/10.5281/zenodo.3754993" ID-Zenodo-Dep="3754993" httpUri="https://zenodo.org/record/3754993/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1913,1939]" targetBox="[195,1389,1227,1874]" targetPageId="3">
|
||
<paragraph id="D4073688FFB5FFD9FF7CEA39FD6813E6" blockId="3.[189,1399,1913,2010]" pageId="3" pageNumber="4">
|
||
<emphasis id="E6CCEA9AFFB5FFD9FF7CEA39FEC913AF" bold="true" box="[189,270,1913,1939]" pageId="3" pageNumber="4">Fig. 2.</emphasis>
|
||
<taxonomicName id="13B84D0BFFB5FFD9FED8EA3AFD3F13A8" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[281,760,1913,1940]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="udzungwensis">
|
||
<emphasis id="E6CCEA9AFFB5FFD9FED8EA3AFDFA13AF" box="[281,573,1913,1940]" italics="true" pageId="3" pageNumber="4">Aquattuor udzungwensis</emphasis>
|
||
<bibRefCitation id="B0294B79FFB5FFD9FD89EA39FD3F13A8" author="Enghoff H. & Frederiksen S. B." box="[584,760,1913,1940]" pageId="3" pageNumber="4" pagination="1 - 25" refId="ref11852" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
, specimens from Chita, heads, lateral view.
|
||
<emphasis id="E6CCEA9AFFB5FFD9FAD2EA39FAEF13AF" bold="true" box="[1299,1320,1913,1939]" pageId="3" pageNumber="4">A</emphasis>
|
||
. Male (NHMD 621667).
|
||
<emphasis id="E6CCEA9AFFB5FFD9FE5CEADDFE76138B" bold="true" box="[413,433,1949,1975]" pageId="3" pageNumber="4">B</emphasis>
|
||
. Female (NHMD 621667). Scale bars = 0.1 mm. The arrow points at the distoventral stipital expansion lobe in the male.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D4073688FFB2FFDEFF7CECBDFBA8169F" blockId="4.[189,1398,509,676]" pageId="4" pageNumber="5">
|
||
Another
|
||
<typeStatus id="0B03882AFFB2FFDEFEE5ECBDFE90162B" box="[292,343,509,535]" pageId="4" pageNumber="5">type</typeStatus>
|
||
of variation is seen in the setation of the telopodites of the first pair of legs (
|
||
<figureCitation id="4C832A0DFFB2FFDEFB19ECBDFAD8162B" box="[1240,1311,509,535]" captionStart="Fig" captionStartId="10.[189,232,1878,1904]" captionTargetBox="[205,1373,1111,1837]" captionTargetId="figure@10.[189,1399,1092,1845]" captionTargetPageId="10" captionText="Fig. 8. Aquattuor spp., males, left leg of first pair. A. A. stereosathe Enghoff, 2015, paratype, ♂ (NHMD 621650): setae same length as in normal walking legs. B. A. longipala Enghoff, 2015, specimen from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642): setae of femur shortened. C. A. mollilobus sp. nov., paratype, ♂ (NHMD 621646): all setae shortened. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3755005" httpUri="https://zenodo.org/record/3755005/files/figure.png" pageId="4" pageNumber="5">Fig. 8</figureCitation>
|
||
). Here,
|
||
<taxonomicName id="13B84D0BFFB2FFDEFF7CEF60FE9F1606" authority="Enghoff, 2020" authorityName="Enghoff" authorityYear="2020" box="[189,344,544,570]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mollilobus" status="sp. nov.">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFF7CEF60FE9F1606" box="[189,344,544,570]" italics="true" pageId="4" pageNumber="5">A. mollilobus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FDFF57E1FFB2FFDEFE9FEF60FE701606" box="[350,439,544,570]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="4C832A0DFFB2FFDEFE07EF60FDD91606" box="[454,542,544,570]" captionStart="Fig" captionStartId="10.[189,232,1878,1904]" captionTargetBox="[205,1373,1111,1837]" captionTargetId="figure@10.[189,1399,1092,1845]" captionTargetPageId="10" captionText="Fig. 8. Aquattuor spp., males, left leg of first pair. A. A. stereosathe Enghoff, 2015, paratype, ♂ (NHMD 621650): setae same length as in normal walking legs. B. A. longipala Enghoff, 2015, specimen from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642): setae of femur shortened. C. A. mollilobus sp. nov., paratype, ♂ (NHMD 621646): all setae shortened. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3755005" httpUri="https://zenodo.org/record/3755005/files/figure.png" pageId="4" pageNumber="5">Fig. 8C</figureCitation>
|
||
) stands out by having very short setae on all podomeres.
|
||
<taxonomicName id="13B84D0BFFB2FFDEFB72EF60FAB11606" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[1203,1398,544,570]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="major">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFB72EF60FAB11606" box="[1203,1398,544,570]" italics="true" pageId="4" pageNumber="5">Aquattuor major</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="13B84D0BFFB2FFDEFF2CEF04FEBB1661" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[237,380,579,606]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="longipala">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFF2CEF04FEBB1661" box="[237,380,579,606]" italics="true" pageId="4" pageNumber="5">A. longipala</emphasis>
|
||
</taxonomicName>
|
||
have long setae on the tarsus, but very short ones on the femur (
|
||
<figureCitation id="4C832A0DFFB2FFDEFB9CEF03FB7F1661" box="[1117,1208,579,605]" captionStart="Fig" captionStartId="10.[189,232,1878,1904]" captionTargetBox="[205,1373,1111,1837]" captionTargetId="figure@10.[189,1399,1092,1845]" captionTargetPageId="10" captionText="Fig. 8. Aquattuor spp., males, left leg of first pair. A. A. stereosathe Enghoff, 2015, paratype, ♂ (NHMD 621650): setae same length as in normal walking legs. B. A. longipala Enghoff, 2015, specimen from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642): setae of femur shortened. C. A. mollilobus sp. nov., paratype, ♂ (NHMD 621646): all setae shortened. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3755005" httpUri="https://zenodo.org/record/3755005/files/figure.png" pageId="4" pageNumber="5">Fig. 8B</figureCitation>
|
||
). The remaining species have long setae (same length as on the following leg-pairs) on all podomeres (
|
||
<figureCitation id="4C832A0DFFB2FFDEFB7AEF26FADC16BC" box="[1211,1307,614,640]" captionStart="Fig" captionStartId="10.[189,232,1878,1904]" captionTargetBox="[205,1373,1111,1837]" captionTargetId="figure@10.[189,1399,1092,1845]" captionTargetPageId="10" captionText="Fig. 8. Aquattuor spp., males, left leg of first pair. A. A. stereosathe Enghoff, 2015, paratype, ♂ (NHMD 621650): setae same length as in normal walking legs. B. A. longipala Enghoff, 2015, specimen from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642): setae of femur shortened. C. A. mollilobus sp. nov., paratype, ♂ (NHMD 621646): all setae shortened. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3755005" httpUri="https://zenodo.org/record/3755005/files/figure.png" pageId="4" pageNumber="5">Fig. 8A</figureCitation>
|
||
). These differences in setation do not seem to reflect the setation of normal walking legs.
|
||
</paragraph>
|
||
<paragraph id="D4073688FFB2FFDEFF7CEF8CFC5B1735" blockId="4.[189,1398,715,777]" pageId="4" pageNumber="5">
|
||
In one species (
|
||
<taxonomicName id="13B84D0BFFB2FFDEFEB6EF8BFDFC16D9" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[375,571,715,741]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="udzungwensis">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFEB6EF8BFDFC16D9" box="[375,571,715,741]" italics="true" pageId="4" pageNumber="5">A. udzungwensis</emphasis>
|
||
</taxonomicName>
|
||
), the podomeres distal to the prefemur of the first leg pair seem to be invariably missing (several specimens examined) (
|
||
<figureCitation id="4C832A0DFFB2FFDEFCCEEFAEFC931735" box="[783,852,750,777]" captionStart="Fig" captionStartId="9.[189,232,1877,1903]" captionTargetBox="[206,1353,584,1811]" captionTargetId="figure@9.[189,1399,573,1842]" captionTargetPageId="9" captionText="Fig. 7. Aquattuor spp., first pair of male legs. A–C. A. submajor Enghoff, 2015, paratype, ♂ (NHMD 621651). A. Anterior view. B. Sublateral view. C. Ventral view. D–E. A. udzungwensis Enghoff, 2015, male from Udzungwa Mts National Park, Kidatu (NHMD 621662). D. Anterior view. E. Ventral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3755003" httpUri="https://zenodo.org/record/3755003/files/figure.png" pageId="4" pageNumber="5">Fig. 7</figureCitation>
|
||
D–E).
|
||
</paragraph>
|
||
<paragraph id="D4073688FFB2FFDEFF7CEE70FD681776" blockId="4.[189,687,816,842]" box="[189,687,816,842]" pageId="4" pageNumber="5">Adhesive pads on walking legs are absent.</paragraph>
|
||
<paragraph id="D4073688FFB2FFDEFF7CEE31FD8E17B0" blockId="4.[189,585,881,908]" box="[189,585,881,908]" pageId="4" pageNumber="5">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFF7CEE31FE4E17B7" bold="true" box="[189,393,881,907]" pageId="4" pageNumber="5">Included species</emphasis>
|
||
(alphabetically)
|
||
</paragraph>
|
||
<paragraph id="D4073688FFB2FFDEFF7CEEE0FC881786" blockId="4.[189,847,927,1271]" box="[189,847,927,954]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="13B84D0BFFB2FFDEFF7CEEE0FC881786" authority="Enghoff & Frederiksen, 2015" authorityName="Enghoff & Frederiksen" authorityYear="2015" box="[189,847,927,954]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="claudiahempae">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFF7CEEE0FE291786" box="[189,494,928,954]" italics="true" pageId="4" pageNumber="5">Aquattuor claudiahempae</emphasis>
|
||
<bibRefCitation id="B0294B79FFB2FFDEFE34EEE0FC881786" author="Enghoff H. & Frederiksen S. B." box="[501,847,927,954]" pageId="4" pageNumber="5" pagination="1 - 25" refId="ref11852" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff & Frederiksen, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="D4073688FFB2FFDEFF7CEE83FD3617E2" blockId="4.[189,847,927,1271]" box="[189,753,963,990]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="13B84D0BFFB2FFDEFF7CEE83FD8F17E1" authority="Frederiksen, 2013" authorityName="Frederiksen" authorityYear="2013" box="[189,584,963,989]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="denticulatus">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFF7CEE83FEAB17E1" box="[189,364,963,989]" italics="true" pageId="4" pageNumber="5">A. denticulatus</emphasis>
|
||
<bibRefCitation id="B0294B79FFB2FFDEFEB5EE83FD8F17E1" author="Frederiksen S. B." box="[372,584,963,989]" pageId="4" pageNumber="5" pagination="59 - 66" refId="ref12038" refString="Frederiksen S. B. 2013. East African odontopygid millipedes 3: Two new genera, Lamelloramus and Aquattuor proposed to contain three new species (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3694: 59 - 66. https: // doi. org / 10.11646 / zootaxa. 3694.1.4" type="journal article" year="2013">Frederiksen, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="0B03882AFFB2FFDEFD96EE83FD4C17E1" box="[599,651,963,989]" pageId="4" pageNumber="5">type</typeStatus>
|
||
species)
|
||
</paragraph>
|
||
<paragraph id="D4073688FFB2FFDEFF7CEEA6FE39103D" blockId="4.[189,847,927,1271]" box="[189,510,998,1025]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="13B84D0BFFB2FFDEFF7CEEA6FE39103D" authority="(Attems, 1896)" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[189,510,998,1025]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="fasciatus">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFF7CEEA6FE82103C" box="[189,325,998,1024]" italics="true" pageId="4" pageNumber="5">A. fasciatus</emphasis>
|
||
(
|
||
<bibRefCitation id="B0294B79FFB2FFDEFE94EEA6FE31103D" author="Attems C. G." box="[341,502,998,1025]" pageId="4" pageNumber="5" pagination="23 - 42" refId="ref11215" refString="Attems C. G. 1896. Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden. Jahrbuch der hamburgischen wissenschaftlichen Anstalten 13, Suppl.: 23 - 42. Available from: https: // biodiversitylibrary. org / page / 28745426 [accessed 1 Oct. 2019]." type="book chapter" year="1896">Attems, 1896</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="D4073688FFB2FFDEFF7CE94AFE391018" blockId="4.[189,847,927,1271]" box="[189,510,1033,1060]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="13B84D0BFFB2FFDEFF7CE94AFE391018" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[189,510,1033,1060]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="longipala">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFF7CE94AFE8A101F" box="[189,333,1033,1060]" italics="true" pageId="4" pageNumber="5">A. longipala</emphasis>
|
||
<bibRefCitation id="B0294B79FFB2FFDEFE94E949FE391018" author="Enghoff H. & Frederiksen S. B." box="[341,510,1033,1060]" pageId="4" pageNumber="5" pagination="1 - 25" refId="ref11852" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="D4073688FFB2FFDEFF7CE96DFE13107B" blockId="4.[189,847,927,1271]" box="[189,468,1068,1095]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="13B84D0BFFB2FFDEFF7CE96DFE13107B" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[189,468,1068,1095]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="major">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFF7CE96DFEE4107A" box="[189,291,1069,1095]" italics="true" pageId="4" pageNumber="5">A. major</emphasis>
|
||
<bibRefCitation id="B0294B79FFB2FFDEFEEAE96CFE13107B" author="Enghoff H. & Frederiksen S. B." box="[299,468,1068,1095]" pageId="4" pageNumber="5" pagination="1 - 25" refId="ref11852" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="D4073688FFB2FFDEFF7CE910FE7C1056" blockId="4.[189,847,927,1271]" box="[189,443,1104,1130]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="13B84D0BFFB2FFDEFF7CE910FE9E1056" authority="Enghoff, 2020" authorityName="Enghoff" authorityYear="2020" box="[189,345,1104,1130]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mollilobus" status="sp. nov.">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFF7CE910FE9E1056" box="[189,345,1104,1130]" italics="true" pageId="4" pageNumber="5">A. mollilobus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FDFF57E1FFB2FFDEFEA1E910FE7C1056" box="[352,443,1104,1130]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="D4073688FFB2FFDEFF7CE933FE0710B1" blockId="4.[189,847,927,1271]" box="[189,448,1139,1165]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="13B84D0BFFB2FFDEFF7CE933FE9910B1" authority="Enghoff, 2020" authorityName="Enghoff" authorityYear="2020" box="[189,350,1139,1165]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="nguruensis" status="sp. nov.">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFF7CE933FE9910B1" box="[189,350,1139,1165]" italics="true" pageId="4" pageNumber="5">A. nguruensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FDFF57E1FFB2FFDEFEA4E933FE0710B1" box="[357,448,1139,1165]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="D4073688FFB2FFDEFF7CE9D6FDD5108D" blockId="4.[189,847,927,1271]" box="[189,530,1174,1201]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="13B84D0BFFB2FFDEFF7CE9D6FDD5108D" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[189,530,1174,1201]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="stereosathe">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFF7CE9D6FEA5108C" box="[189,354,1174,1200]" italics="true" pageId="4" pageNumber="5">A. stereosathe</emphasis>
|
||
<bibRefCitation id="B0294B79FFB2FFDEFEA8E9D6FDD5108D" author="Enghoff H. & Frederiksen S. B." box="[361,530,1174,1201]" pageId="4" pageNumber="5" pagination="1 - 25" refId="ref11852" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="D4073688FFB2FFDEFF7CE9FAFE3A10E8" blockId="4.[189,847,927,1271]" box="[189,509,1209,1236]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="13B84D0BFFB2FFDEFF7CE9FAFE3A10E8" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[189,509,1209,1236]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="submajor">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFF7CE9FAFE8B10EF" box="[189,332,1209,1236]" italics="true" pageId="4" pageNumber="5">A. submajor</emphasis>
|
||
<bibRefCitation id="B0294B79FFB2FFDEFE92E9F9FE3A10E8" author="Enghoff H. & Frederiksen S. B." box="[339,509,1209,1236]" pageId="4" pageNumber="5" pagination="1 - 25" refId="ref11852" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="D4073688FFB2FFDEFF7CE99DFDF610CB" blockId="4.[189,847,927,1271]" box="[189,561,1244,1271]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="13B84D0BFFB2FFDEFF7CE99DFDF610CB" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff & Frederiksen" authorityYear="2015" box="[189,561,1244,1271]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="udzungwensis">
|
||
<emphasis id="E6CCEA9AFFB2FFDEFF7CE99DFE4710CA" box="[189,384,1244,1271]" italics="true" pageId="4" pageNumber="5">A. udzungwensis</emphasis>
|
||
<bibRefCitation id="B0294B79FFB2FFDEFE46E99CFDF610CB" author="Enghoff H. & Frederiksen S. B." box="[391,561,1244,1271]" pageId="4" pageNumber="5" pagination="1 - 25" refId="ref11852" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |