268 lines
38 KiB
XML
268 lines
38 KiB
XML
<document id="F54A968A1A0CF7044F8CEFAEDC1B07BD" ID-DOI="10.5281/zenodo.5355486" ID-ISSN="2345-7600" ID-Zenodo-Dep="5355486" ID-ZooBank="C2FD963A-4DB8-43C9-B940-DA1F7FD333F1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630215542607" checkinUser="valdenar" docAuthor="Manuel-Santos, Marivene, Ng, Peter K. L. & Freitag, Hendrik" docDate="2016" docId="5C112C7B902DFFDA9BBFF831EA1C9CD8" docLanguage="en" docName="64rbz335-342.pdf" docOrigin="Raffles Bulletin of Zoology 64" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Geosesarma tagbanua Manuel-Santos & Ng & Freitag 2016, new species" docType="treatment" docVersion="3" lastPageNumber="341" masterDocId="A02854039028FFDC9B54FFACEB299A6D" masterDocTitle="Two new species of Geosesarma De Man, 1892 (Crustacea: Brachyura: Sesarmidae) from Palawan, the Philippines" masterLastPageNumber="342" masterPageNumber="335" pageNumber="340" updateTime="1699189379378" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
|
||
<mods:mods id="5939C3C7A29BD52AD13A7A595CB0D087" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="96468DBA1AC2E4BA69AAC0F7F9B09455">
|
||
<mods:title id="E6A7D003ECC8C4ED1BEFA19E0B03E12B">Two new species of Geosesarma De Man, 1892 (Crustacea: Brachyura: Sesarmidae) from Palawan, the Philippines</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="F5CA50E7AAAC00593779A041BFE7261A" type="personal">
|
||
<mods:role id="F5F2DBE595E035D546BEB10B42CECD4A">
|
||
<mods:roleTerm id="60FE9A95C4062F449502D658D3D1AC2A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E6883A9A8E86615677161A315A00B2EC">Manuel-Santos, Marivene</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="56E2D38729EEED76641BD34ECF5206A5" type="personal">
|
||
<mods:role id="0FA02BF019846566B4B455F4F611FBC1">
|
||
<mods:roleTerm id="8BB2F121B2773E5990392945F9EBB897">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="3E42B1180994177C28BE99D1FA22A25B">Ng, Peter K. L.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="CCD8EACC756DB639E8180A78FD6E18B9" type="personal">
|
||
<mods:role id="126D54E6465DE0B9992FB4F27B906EC5">
|
||
<mods:roleTerm id="127F8F5B74CC0D5DEA33A38F7BB08221">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="284AD0A2CDA73438DD9E8DFF41FBCB7E">Freitag, Hendrik</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="4BAFC37793EFEDC9E10DEB34AA2845CF">text</mods:typeOfResource>
|
||
<mods:relatedItem id="3A78B652A5C3707A4A52FAB497C413DE" type="host">
|
||
<mods:titleInfo id="B2007213F26CEC9EEC56BFE758855E1E">
|
||
<mods:title id="AA69898A76213CFC4EACEA3A6FF67CD8">Raffles Bulletin of Zoology</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="B1E6B7D7FFADE01ED3DE6D200737251E">
|
||
<mods:date id="1BD2EFC4BB21B4EEC3E81B4A26B0F924">2016</mods:date>
|
||
<mods:detail id="B486129FDC35035AC15FF0AAF2ED8ED1" type="pubDate">
|
||
<mods:number id="1034E138CA14F00FA931D70A60FAB9CC">2016-11-09</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="582F9DF6DD9F987C3E17994F3A04A843" type="volume">
|
||
<mods:number id="AF74033E1A76BAAF4A0469BAD10D2C20">64</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="E454A5914A4CFABE113DD7B880F05C93" unit="page">
|
||
<mods:start id="2A8A952689548DD10BEC5216E4179605">335</mods:start>
|
||
<mods:end id="062C16D6A1C850FBDC425AA1BA92C0DA">342</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="654098A98BE918746093608F9E6EB1B2">journal article</mods:classification>
|
||
<mods:identifier id="22D89A6306E251EC91EDCF43F98867F1" type="DOI">10.5281/zenodo.5355486</mods:identifier>
|
||
<mods:identifier id="D40B1ECA8678EAFE99223F342B33E9CD" type="ISSN">2345-7600</mods:identifier>
|
||
<mods:identifier id="CC275E99A07ACB052544DF0F6F6B7F27" type="Zenodo-Dep">5355486</mods:identifier>
|
||
<mods:identifier id="BAF3E248BC4B15CE97FC1B6EE47216ED" type="ZooBank">C2FD963A-4DB8-43C9-B940-DA1F7FD333F1</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="5C112C7B902DFFDA9BBFF831EA1C9CD8" LSID="urn:lsid:plazi:treatment:5C112C7B902DFFDA9BBFF831EA1C9CD8" httpUri="http://treatment.plazi.org/id/5C112C7B902DFFDA9BBFF831EA1C9CD8" lastPageId="6" lastPageNumber="341" pageId="5" pageNumber="340">
|
||
<subSubSection id="9CA2CEE6902DFFD99BBFF831E95E9DD8" box="[235,631,1949,1973]" pageId="5" pageNumber="340" type="nomenclature">
|
||
<paragraph id="D4079D6D902DFFD99BBFF831E95E9DD8" blockId="5.[235,631,1949,2005]" box="[235,631,1949,1973]" pageId="5" pageNumber="340">
|
||
<heading id="8F4F2A01902DFFD99BBFF831E95E9DD8" bold="true" box="[235,631,1949,1973]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="340" reason="2">
|
||
<emphasis id="E6CC417F902DFFD99BBFF831E95E9DD8" bold="true" box="[235,631,1949,1973]" pageId="5" pageNumber="340">
|
||
<taxonomicName id="13B8E6EE902DFFD99BBFF831EACB9DD8" ID-CoL="6KB9Z" authority="Manuel-Santos & Ng & Freitag, 2016" authorityName="Manuel-Santos & Ng & Freitag" authorityYear="2016" box="[235,482,1949,1973]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="340" phylum="Arthropoda" rank="species" species="tagbanua" status="sp. nov.">
|
||
<emphasis id="E6CC417F902DFFD99BBFF831EACB9DD8" bold="true" box="[235,482,1949,1973]" italics="true" pageId="5" pageNumber="340">Geosesarma tagbanua</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="FDFFFC04902DFFD99AA5F831E95E9DD8" box="[497,631,1949,1973]" pageId="5" pageNumber="340" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CA2CEE6902DFFD99A4EF811E96E9DB8" box="[282,583,1981,2005]" pageId="5" pageNumber="340" type="description">
|
||
<paragraph id="D4079D6D902DFFD99A4EF811E96E9DB8" blockId="5.[235,631,1949,2005]" box="[282,583,1981,2005]" pageId="5" pageNumber="340">
|
||
(
|
||
<figureCitation id="4C8381E8902DFFD99A76F811EA8C9DB8" box="[290,421,1981,2005]" captionStart="Fig" captionStartId="3.[113,148,1206,1228]" captionTargetBox="[115,1451,190,1191]" captionTargetPageId="3" captionText="Fig. 2. Geosesarma tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves. A, overall habitus; B, ventral view of cephalothorax; C, left chela and carpus, dorsal view; D, left chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.5355490" httpUri="https://zenodo.org/record/5355490/files/figure.png" pageId="5" pageNumber="340">Figs. 2A–D</figureCitation>
|
||
,
|
||
<figureCitation id="4C8381E8902DFFD99AE5F811EADA9DB8" box="[433,499,1981,2005]" captionStart="Fig" captionStartId="3.[812,847,1206,1228]" captionText="Fig. 3. Geosesarma tagbanua, new species, paratype male (21.2 × 21.7 mm) (NMCR-25052), Narra, Tuas, Estrella Creek. A, overall habitus; B, ventral view of cephalothorax; C, right chela, outer view." pageId="5" pageNumber="340">3A–C</figureCitation>
|
||
,
|
||
<figureCitation id="4C8381E8902DFFD99954F811E9179DB8" box="[512,574,1981,2005]" captionStart="Fig" captionStartId="4.[135,170,1184,1206]" captionTargetBox="[136,775,189,1168]" captionTargetId="figure-262@4.[135,775,189,1168]" captionTargetPageId="4" captionText="Fig. 4. A–D, Geosesarma batak, new species, holotype male (22.8 × 22.4 mm) (NMCR-40105), Concepcion, near Cleopatra Needle; E, F, G. batak, new species, paratype male (18.9 × 18.5 mm) (ZRC 2015.291), from Laptay/Napsan; G–J, G. tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves; K, L, G. tagbanua, new species, paratype male (21.2 × 21.7 mm) (NMCR-25052), Narra, Tuas, Estrella Creek. A–C, F, G–I, L, left G1 (from different angles); D, E, J, K, left G2." figureDoi="http://doi.org/10.5281/zenodo.5355494" httpUri="https://zenodo.org/record/5355494/files/figure.png" pageId="5" pageNumber="340">4G–L</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CA2CEE6902DFFD99B25F851EE859B18" pageId="5" pageNumber="340" type="materials_examined">
|
||
<paragraph id="D4079D6D902DFFD99B25F851E9D99258" blockId="5.[113,752,2045,2101]" pageId="5" pageNumber="340">
|
||
<emphasis id="E6CC417F902DFFD99B25F851EA749278" bold="true" box="[113,349,2045,2069]" pageId="5" pageNumber="340">Material examined.</emphasis>
|
||
<materialsCitation id="64D09730902DFFD99A33F851E9D99258" location="Tabon Caves" municipality="Lipuun Point" pageId="5" pageNumber="340" specimenCount="1" stateProvince="Quezon" typeStatus="holotype">
|
||
<typeStatus id="0B0323CF902DFFD99A33F851EAFA9278" box="[359,467,2045,2069]" pageId="5" pageNumber="340" type="holotype">Holotype</typeStatus>
|
||
: male (21.8 ×
|
||
<quantity id="13403088902DFFD999D5F851E9C09278" box="[641,745,2045,2069]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="5" pageNumber="340" unit="mm" value="21.8">21.8 mm</quantity>
|
||
) (NMCR-40107),
|
||
<collectingRegion id="167C538F902DFFD99A79F7B1EAAA9258" box="[301,387,2077,2101]" country="Philippines" name="Quezon" pageId="5" pageNumber="340">Quezon</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="34630717902DFFD99ADBF7B1E9369258" box="[399,543,2077,2101]" pageId="5" pageNumber="340">Lipuun Point</collectingMunicipality>
|
||
,
|
||
<location id="D167CBB6902DFFD99978F7B1E9959258" LSID="urn:lsid:plazi:treatment:5C112C7B902DFFDA9BBFF831EA1C9CD8:D167CBB6902DFFD99978F7B1E9959258" box="[556,700,2077,2101]" municipality="Lipuun Point" name="Tabon Caves" pageId="5" pageNumber="340" stateProvince="Quezon">Tabon Caves</location>
|
||
, c. 4
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="D4079D6D902DFFD99879FF11EE859B18" blockId="5.[811,1452,189,374]" pageId="5" pageNumber="340">
|
||
km NNW
|
||
<collectingRegion id="167C538F902DFFD998CCFF11E8C39AB8" box="[920,1002,189,213]" country="Philippines" name="Quezon" pageId="5" pageNumber="340">Quezon</collectingRegion>
|
||
town, c.
|
||
<geoCoordinate id="B18CFBAA902DFFD99F13FF11EFBC9AB8" box="[1095,1173,189,213]" degrees="9" direction="north" minutes="16" orientation="latitude" pageId="5" pageNumber="340" precision="925" value="9.266666">9°16’N</geoCoordinate>
|
||
<geoCoordinate id="B18CFBAA902DFFD99FCEFF11EE289AB8" box="[1178,1281,189,213]" degrees="117" direction="east" minutes="59" orientation="longitude" pageId="5" pageNumber="340" precision="925" value="117.98333">117°59’E</geoCoordinate>
|
||
, coll. J. Orcullo,
|
||
<date id="A006BBAD902DFFD9987FFF71E8869A98" box="[811,943,221,245]" pageId="5" pageNumber="340" value="2003-03">March 2003</date>
|
||
.
|
||
<materialsCitation id="64D09730902DFFD99895FF72EE289B78" collectionCode="ZRC" location="Tabon Caves" municipality="Lipuun Point" pageId="5" pageNumber="340" specimenCode="SMTD-S172" specimenCount="3" specimenCount-male="3" stateProvince="Quezon" typeStatus="paratype">
|
||
<typeStatus id="0B0323CF902DFFD99895FF72EF049A9B" box="[961,1069,222,246]" pageId="5" pageNumber="340" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="C2BE56E4902DFFD99F6DFF71EFA79A98" box="[1081,1166,221,245]" pageId="5" pageNumber="340" type="male">3 males</specimenCount>
|
||
(largest 20.0 ×
|
||
<quantity id="13403088902DFFD99E15FF71EE8D9A98" box="[1345,1444,221,245]" metricMagnitude="-2" metricUnit="m" metricValue="2.01" pageId="5" pageNumber="340" unit="mm" value="20.1">20.1 mm</quantity>
|
||
) (
|
||
<collectionCode id="B2A905A8902DFFD99860FF51E8459B78" box="[820,876,253,277]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="340">ZRC</collectionCode>
|
||
,
|
||
<specimenCode id="841E3516902DFFD99823FF51EF2C9B78" box="[887,1029,253,277]" collectionCode="SMTD-S" pageId="5" pageNumber="340">SMTD-S172</specimenCode>
|
||
), same data as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="64D09730902DFFD99E44FF51EE809B18" collectingDate="1990-05-24" collectorName="M. R. Manuel-Santos" country="Philippines" elevation="370" latitude="9.316667" location="Estrella Creek" longLatPrecision="1300" longitude="118.333336" pageId="5" pageNumber="340" specimenCount="1" specimenCount-male="1" stateProvince="Palawan" typeStatus="paratype">
|
||
<specimenCount id="C2BE56E4902DFFD99E44FF51EE719B78" box="[1296,1368,253,277]" pageId="5" pageNumber="340" type="male">1 male</specimenCount>
|
||
(21.2 ×
|
||
<quantity id="13403088902DFFD99879FEB1E8A69B58" box="[813,911,285,309]" metricMagnitude="-2" metricUnit="m" metricValue="2.17" pageId="5" pageNumber="340" unit="mm" value="21.7">21.7 mm</quantity>
|
||
) (NMCR-25052), Narra, Tuas,
|
||
<location id="D167CBB6902DFFD99FBEFEB1EEAE9B58" LSID="urn:lsid:plazi:treatment:5C112C7B902DFFDA9BBFF831EA1C9CD8:D167CBB6902DFFD99FBEFEB1EEAE9B58" box="[1258,1415,285,309]" country="Philippines" latitude="9.316667" longLatPrecision="1300" longitude="118.333336" name="Estrella Creek" pageId="5" pageNumber="340" stateProvince="Palawan">Estrella Creek</location>
|
||
, ca
|
||
<elevation id="5F957A5E902DFFD99879FE91E8B49B38" box="[813,925,317,341]" metricMagnitude="2" metricUnit="m" metricValue="3.7" pageId="5" pageNumber="340" unit="m" value="370.0">
|
||
<quantity id="13403088902DFFD99879FE91E85A9B38" box="[813,883,317,341]" metricMagnitude="2" metricUnit="m" metricValue="3.7" pageId="5" pageNumber="340" unit="m" value="370.0">370 m</quantity>
|
||
asl
|
||
</elevation>
|
||
,
|
||
<geoCoordinate id="B18CFBAA902DFFD998FFFE91E8D29B38" box="[939,1019,317,341]" degrees="9" direction="north" minutes="19" orientation="latitude" pageId="5" pageNumber="340" precision="925" value="9.316667">9°19’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B18CFBAA902DFFD99F5EFE91EF5A9B38" box="[1034,1139,317,341]" degrees="118" direction="east" minutes="20" orientation="longitude" pageId="5" pageNumber="340" precision="925" value="118.333336">118°20’E</geoCoordinate>
|
||
, coll.
|
||
<collectorName id="794DF8BB902DFFD99FEDFE92EE8E9B38" box="[1209,1447,317,341]" pageId="5" pageNumber="340">M. R. Manuel-Santos</collectorName>
|
||
,
|
||
<date id="A006BBAD902DFFD99879FEF1E8949B18" box="[813,957,349,374]" pageId="5" pageNumber="340" value="1990-05-24">
|
||
<collectingDate id="B0424245902DFFD99879FEF1E8949B18" box="[813,957,349,374]" pageId="5" pageNumber="340" value="1990-05-24">24 May 1990</collectingDate>
|
||
</date>
|
||
. All locations in
|
||
<collectingRegion id="167C538F902DFFD99F2FFEF1EFFC9B18" box="[1147,1237,349,373]" country="Philippines" name="Palawan" pageId="5" pageNumber="340">Palawan</collectingRegion>
|
||
Island,
|
||
<collectingCountry id="ACAFDDFD902DFFD99E79FEF1EE809B18" box="[1325,1449,349,373]" name="Philippines" pageId="5" pageNumber="340">Philippines</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CA2CEE6902DFFD99879FE31EFDD9978" pageId="5" pageNumber="340" type="diagnosis">
|
||
<paragraph id="D4079D6D902DFFD99879FE31EFDD9978" blockId="5.[811,1453,413,789]" pageId="5" pageNumber="340">
|
||
<emphasis id="E6CC417F902DFFD99879FE31E88B9BD8" bold="true" box="[813,930,413,437]" pageId="5" pageNumber="340">Diagnosis.</emphasis>
|
||
Carapace squarish; dorsal surfaces flat, rough, frontal margin strongly deflexed at 90° from horizontal view; postfrontal crest divided into 4 dentiform granulated lobes. Outer surface of chela densely covered with low rounded granules; dorsal margin with incomplete granular ridge. Ambulatory legs relatively longer, fourth merus 0.9 times as long as carapace width. Male abdominal somite 6 slightly shorter than telson; male telson relatively narrower, 1.2 times wider than long. G1 long, slender; shaft rather abruptly tapering subapically, forming angle, distal chitinous part bent laterad at 45°, moderately slender, long, broad, relatively straight except for curved apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CA2CEE6902DFFDA987BFC91EA9A9B98" lastPageId="6" lastPageNumber="341" pageId="5" pageNumber="340" type="description">
|
||
<paragraph id="D4079D6D902DFFD9987BFC91EE859FD8" blockId="5.[811,1452,829,1461]" pageId="5" pageNumber="340">
|
||
<emphasis id="E6CC417F902DFFD9987BFC91EF509938" bold="true" box="[815,1145,829,853]" pageId="5" pageNumber="340">
|
||
Description of
|
||
<typeStatus id="0B0323CF902DFFD99880FC91EF1F9938" box="[980,1078,829,853]" pageId="5" pageNumber="340" type="holotype">holotype</typeStatus>
|
||
male.
|
||
</emphasis>
|
||
Carapace (
|
||
<figureCitation id="4C8381E8902DFFD99FBFFC91EE179938" box="[1259,1342,829,853]" captionStart="Fig" captionStartId="3.[113,148,1206,1228]" captionTargetBox="[115,1451,190,1191]" captionTargetPageId="3" captionText="Fig. 2. Geosesarma tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves. A, overall habitus; B, ventral view of cephalothorax; C, left chela and carpus, dorsal view; D, left chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.5355490" httpUri="https://zenodo.org/record/5355490/files/figure.png" pageId="5" pageNumber="340">Fig. 2A</figureCitation>
|
||
) squarish, about as long as wide; lateral margins almost subparallel or slightly diverging towards posterior carapace margin; dorsal surfaces approximately flat, not convex, rough, with low granules; gastric and branchial regions swollen; hepatic region depressed; grooves deep, regions well defined. Postfrontal crest overall straight, with 4 dentiform postfrontal lobes, reaching or extending slightly beyond frontal margin in dorsal view; anterior margins granulate; median cleft connected with mediogastric groove; protogastric protuberances behind relatively inconspicuous lateral postfrontal lobes; front prominently deflexed from dorsal carapace at right angles; ventral front margin sinuous; sublateral areas distinctly lobiform. Orbital margin angularly bent abruptly, not sinuous. External orbital tooth broadly triangular, directed antero-laterad, entirely slightly upward directed; tip sharp, pointed; first anterolateral tooth smaller, broadly triangular, tip acute, pointing antero-laterad; second anterolateral tooth very low or inconspicuous; lateral margins finely granular, slightly cristate. Posterolateral regions with fine oblique striae.
|
||
</paragraph>
|
||
<paragraph id="D4079D6D902DFFD99879FA71EE7E9F98" blockId="5.[813,1367,1501,1525]" box="[813,1367,1501,1525]" pageId="5" pageNumber="340">
|
||
Third maxillipeds (
|
||
<figureCitation id="4C8381E8902DFFD998AAFA71EF789F98" box="[1022,1105,1501,1525]" captionStart="Fig" captionStartId="3.[113,148,1206,1228]" captionTargetBox="[115,1451,190,1191]" captionTargetPageId="3" captionText="Fig. 2. Geosesarma tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves. A, overall habitus; B, ventral view of cephalothorax; C, left chela and carpus, dorsal view; D, left chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.5355490" httpUri="https://zenodo.org/record/5355490/files/figure.png" pageId="5" pageNumber="340">Fig. 2B</figureCitation>
|
||
) as in previous species.
|
||
</paragraph>
|
||
<paragraph id="D4079D6D902DFFD99879F9B1EF379C98" blockId="5.[811,1452,1565,1781]" pageId="5" pageNumber="340">
|
||
Thoracic sternites (
|
||
<figureCitation id="4C8381E8902DFFD998A2F9B1EF6F9C58" box="[1014,1094,1565,1589]" captionStart="Fig" captionStartId="3.[113,148,1206,1228]" captionTargetBox="[115,1451,190,1191]" captionTargetPageId="3" captionText="Fig. 2. Geosesarma tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves. A, overall habitus; B, ventral view of cephalothorax; C, left chela and carpus, dorsal view; D, left chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.5355490" httpUri="https://zenodo.org/record/5355490/files/figure.png" pageId="5" pageNumber="340">Fig. 2B</figureCitation>
|
||
) as in previous species. Abdomen (
|
||
<figureCitation id="4C8381E8902DFFD99861F991E8A19C38" box="[821,904,1597,1621]" captionStart="Fig" captionStartId="3.[113,148,1206,1228]" captionTargetBox="[115,1451,190,1191]" captionTargetPageId="3" captionText="Fig. 2. Geosesarma tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves. A, overall habitus; B, ventral view of cephalothorax; C, left chela and carpus, dorsal view; D, left chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.5355490" httpUri="https://zenodo.org/record/5355490/files/figure.png" pageId="5" pageNumber="340">Fig. 2B</figureCitation>
|
||
) with telson ca. 1.15–1.25 wider than long, lateral margins gently convex on distal part, almost straight proximally, tip rounded; somite 6 slightly shorter than telson, distal part of lateral margin convex, proximal part almost straight; somites 3–5 increasingly trapezoidal; somites 1 and 2 transversely narrow.
|
||
</paragraph>
|
||
<paragraph id="D4079D6D902DFFD99879F8B1EEA99D58" blockId="5.[813,1408,1821,1845]" box="[813,1408,1821,1845]" pageId="5" pageNumber="340">
|
||
Male chelipeds (
|
||
<figureCitation id="4C8381E8902DFFD998B5F8B1EF539D58" box="[993,1146,1821,1845]" captionStart="Fig" captionStartId="3.[113,148,1206,1228]" captionTargetBox="[115,1451,190,1191]" captionTargetPageId="3" captionText="Fig. 2. Geosesarma tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves. A, overall habitus; B, ventral view of cephalothorax; C, left chela and carpus, dorsal view; D, left chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.5355490" httpUri="https://zenodo.org/record/5355490/files/figure.png" pageId="5" pageNumber="340">Fig. 2A, C, D</figureCitation>
|
||
) as in previous species.
|
||
</paragraph>
|
||
<paragraph id="D4079D6D902DFFD99879F8F1EE339DB8" blockId="5.[811,1453,1885,2005]" pageId="5" pageNumber="340">
|
||
Ambulatory legs (
|
||
<figureCitation id="4C8381E8902DFFD99F52F8F1EF4D9D18" box="[1030,1124,1885,1909]" captionStart="Fig" captionStartId="3.[113,148,1206,1228]" captionTargetBox="[115,1451,190,1191]" captionTargetPageId="3" captionText="Fig. 2. Geosesarma tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves. A, overall habitus; B, ventral view of cephalothorax; C, left chela and carpus, dorsal view; D, left chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.5355490" httpUri="https://zenodo.org/record/5355490/files/figure.png" pageId="5" pageNumber="340">Fig. 2A</figureCitation>
|
||
) slender, very long; fourth ambulatory merus about 0.9 times as long as carapace width, lateral margins of meri subparallel at least from basal 0.3 to 0.8. Otherwise similar to previous species.
|
||
</paragraph>
|
||
<paragraph id="D4079D6D902DFFDA9879F851E93D9BD8" blockId="5.[813,1451,2045,2101]" lastBlockId="6.[136,776,189,437]" lastPageId="6" lastPageNumber="341" pageId="5" pageNumber="340">
|
||
G1 (
|
||
<figureCitation id="4C8381E8902DFFD9980FF851E8EA9278" box="[859,963,2045,2069]" captionStart="Fig" captionStartId="4.[135,170,1184,1206]" captionTargetBox="[136,775,189,1168]" captionTargetId="figure-262@4.[135,775,189,1168]" captionTargetPageId="4" captionText="Fig. 4. A–D, Geosesarma batak, new species, holotype male (22.8 × 22.4 mm) (NMCR-40105), Concepcion, near Cleopatra Needle; E, F, G. batak, new species, paratype male (18.9 × 18.5 mm) (ZRC 2015.291), from Laptay/Napsan; G–J, G. tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves; K, L, G. tagbanua, new species, paratype male (21.2 × 21.7 mm) (NMCR-25052), Narra, Tuas, Estrella Creek. A–C, F, G–I, L, left G1 (from different angles); D, E, J, K, left G2." figureDoi="http://doi.org/10.5281/zenodo.5355494" httpUri="https://zenodo.org/record/5355494/files/figure.png" pageId="5" pageNumber="340">Fig. 4G–I</figureCitation>
|
||
) long, slender, straight, reaching distal end of abdominal somite 6; subdistal inner margin with long setae that do not overreach tip; basal part rounded, prominent (
|
||
<figureCitation id="4C8381E8902EFFDA9988FF11EB849A98" captionStart="Fig" captionStartId="4.[135,170,1184,1206]" captionTargetBox="[136,775,189,1168]" captionTargetId="figure-262@4.[135,775,189,1168]" captionTargetPageId="4" captionText="Fig. 4. A–D, Geosesarma batak, new species, holotype male (22.8 × 22.4 mm) (NMCR-40105), Concepcion, near Cleopatra Needle; E, F, G. batak, new species, paratype male (18.9 × 18.5 mm) (ZRC 2015.291), from Laptay/Napsan; G–J, G. tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves; K, L, G. tagbanua, new species, paratype male (21.2 × 21.7 mm) (NMCR-25052), Narra, Tuas, Estrella Creek. A–C, F, G–I, L, left G1 (from different angles); D, E, J, K, left G2." figureDoi="http://doi.org/10.5281/zenodo.5355494" httpUri="https://zenodo.org/record/5355494/files/figure.png" pageId="6" pageNumber="341">Fig. 4H</figureCitation>
|
||
); shaft rather abruptly tapering subapically near distal chitinous part, apically rather abruptly angled lateral at 45° in ventral view (
|
||
<figureCitation id="4C8381E8902EFFDA9A10FEB1EAB69B58" box="[324,415,285,309]" captionStart="Fig" captionStartId="4.[135,170,1184,1206]" captionTargetBox="[136,775,189,1168]" captionTargetId="figure-262@4.[135,775,189,1168]" captionTargetPageId="4" captionText="Fig. 4. A–D, Geosesarma batak, new species, holotype male (22.8 × 22.4 mm) (NMCR-40105), Concepcion, near Cleopatra Needle; E, F, G. batak, new species, paratype male (18.9 × 18.5 mm) (ZRC 2015.291), from Laptay/Napsan; G–J, G. tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves; K, L, G. tagbanua, new species, paratype male (21.2 × 21.7 mm) (NMCR-25052), Narra, Tuas, Estrella Creek. A–C, F, G–I, L, left G1 (from different angles); D, E, J, K, left G2." figureDoi="http://doi.org/10.5281/zenodo.5355494" httpUri="https://zenodo.org/record/5355494/files/figure.png" pageId="6" pageNumber="341">Fig. 4G</figureCitation>
|
||
); distal part moderately slender, moderately long, slightly bent dorsad in lateral view (
|
||
<figureCitation id="4C8381E8902EFFDA9988FE91EB829B18" captionStart="Fig" captionStartId="4.[135,170,1184,1206]" captionTargetBox="[136,775,189,1168]" captionTargetId="figure-262@4.[135,775,189,1168]" captionTargetPageId="4" captionText="Fig. 4. A–D, Geosesarma batak, new species, holotype male (22.8 × 22.4 mm) (NMCR-40105), Concepcion, near Cleopatra Needle; E, F, G. batak, new species, paratype male (18.9 × 18.5 mm) (ZRC 2015.291), from Laptay/Napsan; G–J, G. tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves; K, L, G. tagbanua, new species, paratype male (21.2 × 21.7 mm) (NMCR-25052), Narra, Tuas, Estrella Creek. A–C, F, G–I, L, left G1 (from different angles); D, E, J, K, left G2." figureDoi="http://doi.org/10.5281/zenodo.5355494" httpUri="https://zenodo.org/record/5355494/files/figure.png" pageId="6" pageNumber="341">Fig. 4H</figureCitation>
|
||
), overall rather straight, but apex distinctly curved. G2 (
|
||
<figureCitation id="4C8381E8902EFFDA9BC4FED1EBF69BF8" box="[144,223,381,405]" captionStart="Fig" captionStartId="4.[135,170,1184,1206]" captionTargetBox="[136,775,189,1168]" captionTargetId="figure-262@4.[135,775,189,1168]" captionTargetPageId="4" captionText="Fig. 4. A–D, Geosesarma batak, new species, holotype male (22.8 × 22.4 mm) (NMCR-40105), Concepcion, near Cleopatra Needle; E, F, G. batak, new species, paratype male (18.9 × 18.5 mm) (ZRC 2015.291), from Laptay/Napsan; G–J, G. tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves; K, L, G. tagbanua, new species, paratype male (21.2 × 21.7 mm) (NMCR-25052), Narra, Tuas, Estrella Creek. A–C, F, G–I, L, left G1 (from different angles); D, E, J, K, left G2." figureDoi="http://doi.org/10.5281/zenodo.5355494" httpUri="https://zenodo.org/record/5355494/files/figure.png" pageId="6" pageNumber="341">Fig. 4J</figureCitation>
|
||
) moderately short; base subtriangular; distal part slender, outer margin gently curved.
|
||
</paragraph>
|
||
<paragraph id="D4079D6D902EFFDA9BDCFE71EA9A9B98" blockId="6.[136,435,477,501]" box="[136,435,477,501]" pageId="6" pageNumber="341">
|
||
<emphasis id="E6CC417F902EFFDA9BDCFE71EA039B98" bold="true" box="[136,298,477,501]" pageId="6" pageNumber="341">Colour in life.</emphasis>
|
||
Not known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CA2CEE6902EFFDA9BDCFDB1E9FE9818" pageId="6" pageNumber="341" type="etymology">
|
||
<paragraph id="D4079D6D902EFFDA9BDCFDB1E9FE9818" blockId="6.[136,775,541,629]" pageId="6" pageNumber="341">
|
||
<emphasis id="E6CC417F902EFFDA9BDCFDB1EA239858" bold="true" box="[136,266,541,565]" pageId="6" pageNumber="341">Etymology.</emphasis>
|
||
Named after the Tagbanua, an indigenous tribe in
|
||
<collectingRegion id="167C538F902EFFDA9BF1FD91EA2B9838" box="[165,258,573,597]" country="Philippines" name="Palawan" pageId="6" pageNumber="341">Palawan</collectingRegion>
|
||
, who like the Bataks, get their sustenance from the forests. The name is used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CA2CEE6902EFFDA9BDCFD31E9219E78" pageId="6" pageNumber="341" type="distribution">
|
||
<paragraph id="D4079D6D902EFFDA9BDCFD31EA219898" blockId="6.[134,775,669,757]" pageId="6" pageNumber="341">
|
||
<emphasis id="E6CC417F902EFFDA9BDCFD31EA3C98D8" bold="true" box="[136,277,669,693]" pageId="6" pageNumber="341">Distribution.</emphasis>
|
||
Only known from two sites in the Municipalities of Narra and
|
||
<collectingRegion id="167C538F902EFFDA9A7DFD11EAA998B8" box="[297,384,701,725]" country="Philippines" name="Quezon" pageId="6" pageNumber="341">Quezon</collectingRegion>
|
||
of the Province of
|
||
<collectingRegion id="167C538F902EFFDA9924FD11E9FA98B8" box="[624,723,701,725]" country="Philippines" name="Palawan" pageId="6" pageNumber="341">Palawan</collectingRegion>
|
||
, the
|
||
<collectingCountry id="ACAFDDFD902EFFDA9BDCFD71EA2D9898" box="[136,260,733,757]" name="Philippines" pageId="6" pageNumber="341">Philippines</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D4079D6D902EFFDA9BDCFCB1E9219E78" blockId="6.[135,776,797,1045]" pageId="6" pageNumber="341">
|
||
<emphasis id="E6CC417F902EFFDA9BDCFCB1EA6E9958" bold="true" box="[136,327,797,821]" pageId="6" pageNumber="341">Ecological notes.</emphasis>
|
||
Specimens were collected in a limestone cave and at a montane forest. One specimen (NMCR-25052) was collected across a creek at ca.
|
||
<quantity id="13403088902EFFDA995FFCF1E9789918" box="[523,593,861,885]" metricMagnitude="2" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="341" unit="m" value="370.0">370 m</quantity>
|
||
elevation at Mt. Victoria. The site was surrounded by huge boulders of algae-covered rocks where dipterocarps, bamboos and thorny rattans dominated. The specimen was found there by the first author among the fallen barks of “almaciga” (
|
||
<taxonomicName id="13B8E6EE902EFFDA99E1FC71EA269E78" class="Pinopsida" family="Araucariaceae" genus="Agathis" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="341" phylum="Tracheophyta" rank="species" species="loranthifolia">
|
||
<emphasis id="E6CC417F902EFFDA99E1FC71EA269E78" italics="true" pageId="6" pageNumber="341">Agathis loranthifolia</emphasis>
|
||
</taxonomicName>
|
||
) near the creek banks.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CA2CEE6902EFFDA9BDCFB91EA1C9CD8" pageId="6" pageNumber="341" type="discussion">
|
||
<paragraph id="D4079D6D902EFFDA9BDCFB91E91A9C38" blockId="6.[135,777,1085,1621]" pageId="6" pageNumber="341">
|
||
<emphasis id="E6CC417F902EFFDA9BDCFB91EBD09E38" bold="true" box="[136,249,1085,1109]" pageId="6" pageNumber="341">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="0B0323CF902EFFDA9A6CFB92EABF9E3B" box="[312,406,1086,1110]" pageId="6" pageNumber="341" type="paratype">paratype</typeStatus>
|
||
male specimen of
|
||
<taxonomicName id="13B8E6EE902EFFDA9920FB92E8299E38" authority="Manuel-Santos & Ng & Freitag, 2016" authorityName="Manuel-Santos & Ng & Freitag" authorityYear="2016" box="[628,768,1085,1109]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="tagbanua" status="sp. nov.">
|
||
<emphasis id="E6CC417F902EFFDA9920FB92E8299E38" box="[628,768,1085,1109]" italics="true" pageId="6" pageNumber="341">G. tagbanua</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="FDFFFC04902EFFDA9BDCFBF2EA259E18" box="[136,268,1117,1141]" pageId="6" pageNumber="341" rank="species">new species</taxonomicNameLabel>
|
||
, from Narra (NMCR-25052) (
|
||
<figureCitation id="4C8381E8902EFFDA9900FBF1E9FA9E18" box="[596,723,1117,1141]" captionStart="Fig" captionStartId="3.[812,847,1206,1228]" captionText="Fig. 3. Geosesarma tagbanua, new species, paratype male (21.2 × 21.7 mm) (NMCR-25052), Narra, Tuas, Estrella Creek. A, overall habitus; B, ventral view of cephalothorax; C, right chela, outer view." pageId="6" pageNumber="341">Figs. 3A–C</figureCitation>
|
||
,
|
||
<figureCitation id="4C8381E8902EFFDA99B4FBF1EBB19EF8" captionStart="Fig" captionStartId="4.[135,170,1184,1206]" captionTargetBox="[136,775,189,1168]" captionTargetId="figure-262@4.[135,775,189,1168]" captionTargetPageId="4" captionText="Fig. 4. A–D, Geosesarma batak, new species, holotype male (22.8 × 22.4 mm) (NMCR-40105), Concepcion, near Cleopatra Needle; E, F, G. batak, new species, paratype male (18.9 × 18.5 mm) (ZRC 2015.291), from Laptay/Napsan; G–J, G. tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves; K, L, G. tagbanua, new species, paratype male (21.2 × 21.7 mm) (NMCR-25052), Narra, Tuas, Estrella Creek. A–C, F, G–I, L, left G1 (from different angles); D, E, J, K, left G2." figureDoi="http://doi.org/10.5281/zenodo.5355494" httpUri="https://zenodo.org/record/5355494/files/figure.png" pageId="6" pageNumber="341">4K, L</figureCitation>
|
||
) differs slightly from the remaining type material in its carapace shape, which appears to be somewhat rounder and is 1.02 times longer than broad. The carapaces of the remaining specimens are distinctly more squarish and almost exactly as wide as long. Furthermore, its dorsal surface is slightly more concave, the external orbital teeth are more acute and pronounced (
|
||
<figureCitation id="4C8381E8902EFFDA9AD0FA91EAF19F38" box="[388,472,1341,1365]" captionStart="Fig" captionStartId="3.[812,847,1206,1228]" captionText="Fig. 3. Geosesarma tagbanua, new species, paratype male (21.2 × 21.7 mm) (NMCR-25052), Narra, Tuas, Estrella Creek. A, overall habitus; B, ventral view of cephalothorax; C, right chela, outer view." pageId="6" pageNumber="341">Fig. 3A</figureCitation>
|
||
right side; cf.
|
||
<figureCitation id="4C8381E8902EFFDA992CFA91E9E59F38" box="[632,716,1341,1365]" captionStart="Fig" captionStartId="3.[113,148,1206,1228]" captionTargetBox="[115,1451,190,1191]" captionTargetPageId="3" captionText="Fig. 2. Geosesarma tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves. A, overall habitus; B, ventral view of cephalothorax; C, left chela and carpus, dorsal view; D, left chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.5355490" httpUri="https://zenodo.org/record/5355490/files/figure.png" pageId="6" pageNumber="341">Fig. 2C</figureCitation>
|
||
), and the dactylar finger of the chela is slightly stouter and lacks the dorsal row of granules (
|
||
<figureCitation id="4C8381E8902EFFDA9AE8FAD1E93A9FF8" box="[444,531,1405,1429]" captionStart="Fig" captionStartId="3.[812,847,1206,1228]" captionText="Fig. 3. Geosesarma tagbanua, new species, paratype male (21.2 × 21.7 mm) (NMCR-25052), Narra, Tuas, Estrella Creek. A, overall habitus; B, ventral view of cephalothorax; C, right chela, outer view." pageId="6" pageNumber="341">Fig. 3C</figureCitation>
|
||
; cf.
|
||
<figureCitation id="4C8381E8902EFFDA9912FAD1E9B79FF8" box="[582,670,1405,1429]" captionStart="Fig" captionStartId="3.[113,148,1206,1228]" captionTargetBox="[115,1451,190,1191]" captionTargetPageId="3" captionText="Fig. 2. Geosesarma tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves. A, overall habitus; B, ventral view of cephalothorax; C, left chela and carpus, dorsal view; D, left chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.5355490" httpUri="https://zenodo.org/record/5355490/files/figure.png" pageId="6" pageNumber="341">Fig. 2D</figureCitation>
|
||
). The G1 of this specimen is also not as strongly chitinised as the other specimens. These observations suggest this specimen had recently moulted. However, all the other key diagnostic characters, notably that of the G1 and G2 (
|
||
<figureCitation id="4C8381E8902EFFDA9931FA51E9F49C78" box="[613,733,1533,1557]" captionStart="Fig" captionStartId="4.[135,170,1184,1206]" captionTargetBox="[136,775,189,1168]" captionTargetId="figure-262@4.[135,775,189,1168]" captionTargetPageId="4" captionText="Fig. 4. A–D, Geosesarma batak, new species, holotype male (22.8 × 22.4 mm) (NMCR-40105), Concepcion, near Cleopatra Needle; E, F, G. batak, new species, paratype male (18.9 × 18.5 mm) (ZRC 2015.291), from Laptay/Napsan; G–J, G. tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves; K, L, G. tagbanua, new species, paratype male (21.2 × 21.7 mm) (NMCR-25052), Narra, Tuas, Estrella Creek. A–C, F, G–I, L, left G1 (from different angles); D, E, J, K, left G2." figureDoi="http://doi.org/10.5281/zenodo.5355494" httpUri="https://zenodo.org/record/5355494/files/figure.png" pageId="6" pageNumber="341">Fig. 4L, K</figureCitation>
|
||
; cf.
|
||
<figureCitation id="4C8381E8902EFFDA9BDCF9B1EBD19C58" box="[136,248,1565,1589]" captionStart="Fig" captionStartId="4.[135,170,1184,1206]" captionTargetBox="[136,775,189,1168]" captionTargetId="figure-262@4.[135,775,189,1168]" captionTargetPageId="4" captionText="Fig. 4. A–D, Geosesarma batak, new species, holotype male (22.8 × 22.4 mm) (NMCR-40105), Concepcion, near Cleopatra Needle; E, F, G. batak, new species, paratype male (18.9 × 18.5 mm) (ZRC 2015.291), from Laptay/Napsan; G–J, G. tagbanua, new species, holotype male (21.8 × 21.8 mm) (NMCR-40107), Lipuun Point, Tabon Caves; K, L, G. tagbanua, new species, paratype male (21.2 × 21.7 mm) (NMCR-25052), Narra, Tuas, Estrella Creek. A–C, F, G–I, L, left G1 (from different angles); D, E, J, K, left G2." figureDoi="http://doi.org/10.5281/zenodo.5355494" httpUri="https://zenodo.org/record/5355494/files/figure.png" pageId="6" pageNumber="341">Fig. 4G, J</figureCitation>
|
||
) is consistent with the other specimens and we regard them all as conspecific for now.
|
||
</paragraph>
|
||
<paragraph id="D4079D6D902EFFDA9BDCF9D2EA1C9CD8" blockId="6.[136,775,1661,1717]" pageId="6" pageNumber="341">
|
||
For comparisons with
|
||
<taxonomicName id="13B8E6EE902EFFDA9AD5F9D2EAC99CF8" authority="Manuel-Santos & Ng & Freitag, 2016" authorityName="Manuel-Santos & Ng & Freitag" authorityYear="2016" box="[385,480,1661,1685]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="batak" status="sp. nov.">
|
||
<emphasis id="E6CC417F902EFFDA9AD5F9D2EAC99CF8" box="[385,480,1661,1685]" italics="true" pageId="6" pageNumber="341">G. batak</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="FDFFFC04902EFFDA9AA5F9D2E95E9CF8" box="[497,631,1661,1685]" pageId="6" pageNumber="341" rank="species">new species</taxonomicNameLabel>
|
||
, see remarks for that species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |