193 lines
17 KiB
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193 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.315.5293" ID-GBIF-Dataset="eed3d9a5-c6b2-45ac-af9f-79770c35df88" ID-PMC="PMC3713351" ID-Pensoft-Pub="1313-2970-315-17" ID-PubMed="23878508" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-315-17" ModsDocOrigin="ZooKeys 315" ModsDocTitle="Western Hemisphere Zuphiini: descriptions of Coarazuphium whiteheadi, new species, and Zuphioides, new genus, and classification of the genera (Coleoptera, Carabidae)" checkinTime="1451247157087" checkinUser="pensoft" docAuthor="Ball, George E. & Shpeley, Danny" docDate="2013" docId="E35ABDD156ECFDEDB8B2F46C66D6921F" docLanguage="en" docName="ZooKeys 315: 17-54" docOrigin="ZooKeys 315" docSource="http://dx.doi.org/10.3897/zookeys.315.5293" docTitle="Zuphioides mexicanum" docType="treatment" docVersion="3" lastPageNumber="38" masterDocId="0F6AFFB1B44A505EFFB3FF89734EE542" masterDocTitle="Western Hemisphere Zuphiini: descriptions of Coarazuphium whiteheadi, new species, and Zuphioides, new genus, and classification of the genera (Coleoptera, Carabidae)" masterLastPageNumber="54" masterPageNumber="17" pageNumber="36" updateTime="1668156099637" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Western Hemisphere Zuphiini: descriptions of Coarazuphium whiteheadi, new species, and Zuphioides, new genus, and classification of the genera (Coleoptera, Carabidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ball, George E.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shpeley, Danny</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>315</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>17</mods:start>
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<mods:end>54</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.315.5293</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.315.5293</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-315-17</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152045956" LSID="urn:lsid:plazi:treatment:E35ABDD156ECFDEDB8B2F46C66D6921F" httpUri="http://treatment.plazi.org/id/E35ABDD156ECFDEDB8B2F46C66D6921F" lastPageId="21" lastPageNumber="38" pageId="19" pageNumber="36">
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<subSubSection pageId="19" pageNumber="36" type="nomenclature">
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<paragraph pageId="19" pageNumber="36">
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<taxonomicName LSID="http://species-id.net/wiki/Zuphioides_mexicanum" authority="Chaudoir" class="Insecta" family="Carabidae" genus="Zuphioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zuphioides mexicanum" order="Coleoptera" pageId="19" pageNumber="36" phylum="Arthropoda" rank="species" species="mexicanum">Zuphioides mexicanum (Chaudoir)</taxonomicName>
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Figs 1B, 2B, 3B, 4C, 4D, 5C, 5D, 7
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<normalizedToken originalValue="A–">A-</normalizedToken>
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7F, 8
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<normalizedToken originalValue="D–">D-</normalizedToken>
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8F, 9A, 10
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<normalizedToken originalValue="D–">D-</normalizedToken>
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10F, 11
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<normalizedToken originalValue="D–">D-</normalizedToken>
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11F, 12B
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="36" type="reference_group">
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<paragraph pageId="19" pageNumber="36">
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<taxonomicName class="Insecta" family="Carabidae" genus="Zuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zuphium mexicanum" order="Coleoptera" pageId="19" pageNumber="36" phylum="Arthropoda" rank="species" species="mexicanum">Zuphium mexicanum</taxonomicName>
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<bibRefCitation pageId="19" pageNumber="36">Chaudoir, 1862: 314. Bates 1883</bibRefCitation>
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: 166.
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<bibRefCitation author="Mateu, J" journalOrPublisher="Folia Entomologica Mexicana" pageId="25" pageNumber="42" pagination="111 - 128" title="Revision de los Zuphium Latreille del Continente Americano (Coleoptera: Carabidae) 1 a. Nota." volume="47" year="1981">Mateu 1981</bibRefCitation>
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: 125.
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</paragraph>
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<paragraph pageId="19" pageNumber="36">
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<taxonomicName class="Insecta" family="Carabidae" genus="Zuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zuphium vicinum" order="Coleoptera" pageId="19" pageNumber="36" phylum="Arthropoda" rank="species" species="vicinum">Zuphium vicinum</taxonomicName>
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<bibRefCitation author="Mroczkowski, M" journalOrPublisher="Annales Zoologici" pageId="26" pageNumber="43" pagination="365 - 409" title="List of type specimens in the collection of the Institute of Zoology of the Polish Academy of Sciences in Warzawa III. Carabidae (Coleoptera)." volume="18" year="1960">Liebke, 1933: 471. Mroczkowski 1960</bibRefCitation>
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: 400.
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<bibRefCitation author="Mateu, J" journalOrPublisher=")" pageId="25" pageNumber="42" pagination="329 - 330" title="A propos de trois especes du genre Z uphium d'Amerique du Sud decrites par M. Liebke (Col. Carabidae). Nouvelles Revue d'Entomologie (N. S." volume="2" year="1985">Mateu 1985</bibRefCitation>
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: 329.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="36" type="type material">
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<paragraph pageId="19" pageNumber="36">Type material.</paragraph>
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<paragraph pageId="19" pageNumber="36">
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For
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<taxonomicName class="Insecta" family="Carabidae" genus="Zuphioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zuphioides mexicanum" order="Coleoptera" pageId="19" pageNumber="36" phylum="Arthropoda" rank="species" species="mexicanum">Zuphioides mexicanum</taxonomicName>
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, two females,
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<normalizedToken originalValue="Oberthür-Chaudoir">Oberthuer-Chaudoir</normalizedToken>
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Collection, in front of the following box label: "mexicanum/ Chaud/ Mexique/ 57.
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<normalizedToken originalValue="Sallé”">Salle"</normalizedToken>
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. LECTOTYPE female, labeled as follows:
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<normalizedToken originalValue="“33”">"33"</normalizedToken>
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/ "Ex Musaeo/Chaudoir" [red print];
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<normalizedToken originalValue="“Lecto/type”">"Lecto/type"</normalizedToken>
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[circular, ringed with red]; in Museum National
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<normalizedToken originalValue="d’Histoire">d'Histoire</normalizedToken>
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Naturelle, Paris; PARALECTOTYPE labeled " Ex Musaeo/Chaudoir" [red print]. Designated by
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<bibRefCitation author="Mateu, J" journalOrPublisher=")" pageId="25" pageNumber="42" pagination="329 - 330" title="A propos de trois especes du genre Z uphium d'Amerique du Sud decrites par M. Liebke (Col. Carabidae). Nouvelles Revue d'Entomologie (N. S." volume="2" year="1985">Mateu (1985</bibRefCitation>
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: 330). For
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<taxonomicName class="Insecta" family="Carabidae" genus="Zuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zuphium vicinum" order="Coleoptera" pageId="19" pageNumber="36" phylum="Arthropoda" rank="species" species="vicinum">Zuphium vicinum</taxonomicName>
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, holotype, in Institute of Zoology, Polish Academy of Sciences, Warsawa, #1711.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="36" type="type area">
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<paragraph pageId="19" pageNumber="36">Type area.</paragraph>
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<paragraph pageId="19" pageNumber="36">
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Indicated by
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<bibRefCitation author="Mateu, J" journalOrPublisher=")" pageId="25" pageNumber="42" pagination="329 - 330" title="A propos de trois especes du genre Z uphium d'Amerique du Sud decrites par M. Liebke (Col. Carabidae). Nouvelles Revue d'Entomologie (N. S." volume="2" year="1985">Mateu (1985</bibRefCitation>
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: 330) as Veracruz, for
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<taxonomicName class="Insecta" family="Carabidae" genus="Zuphioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zuphioides mexicanum" order="Coleoptera" pageId="19" pageNumber="36" phylum="Arthropoda" rank="species" species="mexicanum">Zuphioides mexicanum</taxonomicName>
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; for
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<taxonomicName class="Insecta" family="Carabidae" genus="Zuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zuphium vicinum" order="Coleoptera" pageId="19" pageNumber="36" phylum="Arthropoda" rank="species" species="vicinum">Zuphium vicinum</taxonomicName>
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,
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<normalizedToken originalValue="“Mexico”">"Mexico"</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="36" type="specific epithet">
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<paragraph pageId="19" pageNumber="36">Specific epithet.</paragraph>
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<paragraph pageId="19" pageNumber="36">A Latinized eponym, nominative case, based on the name of the country in which the type locality is located.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="21" lastPageNumber="38" pageId="19" pageNumber="36" type="description">
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<paragraph pageId="19" pageNumber="36">Description.</paragraph>
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<paragraph pageId="19" pageNumber="36">
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Size and proportions. Small, OBL 4.88-4.96 mm; EW 1.60-1.62 mm A1L/A2-4L 0.90-1.00; HW/PW 0.92-0.97. (For seven species of
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<taxonomicName class="Insecta" family="Carabidae" genus="Zuphioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zuphioides" order="Coleoptera" pageId="19" pageNumber="36" phylum="Arthropoda" rank="genus">Zuphioides</taxonomicName>
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: OBL 4.64-6.586 mm; EW 1.60-1.84 mm A1L/A2-4L 0.88-1.12; HW/PW 0.91-0.97).
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</paragraph>
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<paragraph pageId="19" pageNumber="36">Color. Body rufotestaceous; appendages slightly paler; vestiture testaceous.</paragraph>
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<paragraph pageId="20" pageNumber="37">
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<pageBreakToken pageId="20" pageNumber="37" start="start">Habitus</pageBreakToken>
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(Fig. 1B). Flat, overall. Head capsule (dorsal aspect, Fig. 2B) trapezoidal, obtusely angulate posteriolaterally.
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</paragraph>
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<paragraph pageId="20" pageNumber="37">
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Eyes (Figs 3B). Convex, easily seen, ommatidia clearly evident at 50
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<normalizedToken originalValue="×">x</normalizedToken>
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(about 16 ommatidia are crossed on a horizontal diameter).
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</paragraph>
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<paragraph pageId="20" pageNumber="37">Pronotum. Narrow.</paragraph>
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<paragraph pageId="20" pageNumber="37">Elytra (Fig. 1B). Lateral margins straight, not bowed, parallel to one another.</paragraph>
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<paragraph pageId="20" pageNumber="37">
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Microsculpture. Dorsal surface of head capsule (including clypeus) and pronotum smooth, microlines not evident; labrum and elytra with microlines fine (not easily seen at 50
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<normalizedToken originalValue="×">x</normalizedToken>
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or lower magnification), mesh pattern isodiametric. Ventral surface with microlines fine, generally transverse.
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</paragraph>
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<paragraph pageId="20" pageNumber="37">Luster. Dorsal surface of head and pronotum shiny, elytra somewhat duller.</paragraph>
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<paragraph pageId="20" pageNumber="37">Body vestiture and punctation. Dorsal surface of head and pronotum sparsely punctate, vestiture sparse. Elytra and abdominal sterna III-VII with punctation and vestiture dense, vestitural setae decumbent.</paragraph>
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<paragraph pageId="20" pageNumber="37">Fixed setae. Head (Fig. 4C): clypeus one pair; head capsule with anterior pair of supraorbitals (asos) above eyes; posterior pair of supraorbitals (psos) posteriad eyes; one pair of postocular setae (pos) immediately behind eyes, laterally; one pair of occipital setae (ocs) posteriorly and mediad eyes; posterior supernumerary setae lacking. Antennomere 1 (Figs 4B, 4D, as1) with single long seta distally, in addition to decumbent vestiture and row of small erect setae (as2, as3). Pronotum with two pairs of lateral marginal setae, anterior pair in anterior 1/8, posterior pair at posteriolateral angles. Prosternum with one pair setae anterioventrally (Fig. 9B). Elytra: each elytron anteriorly with parascutellar seta; lateral setae about 21 (one group anteriorly, one group posteriorly, and single seta medially. Abdominal sterna IV-VI without evident ambulatory setae, sternum VII with pair of long setae near posterior margin.</paragraph>
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<paragraph pageId="20" pageNumber="37">
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Head, dorsal aspect (Fig. 4C). Occiput posteriorly and postocciput markedly constricted, in form of narrow neck, postoccipital suture evident. Frontoclypeal suture present. Genal sulcus (gs) prominent, but less so than in
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<taxonomicName class="Insecta" family="Carabidae" genus="Coarazuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarazuphium" order="Coleoptera" pageId="20" pageNumber="37" phylum="Arthropoda" rank="genus">Coarazuphium</taxonomicName>
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(cf. Fig. 4A).
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</paragraph>
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<paragraph pageId="20" pageNumber="37">
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Eyes (Fig. 4B). Macrophthalmous, markedly convex, readily seen, ommatidia evident at 50
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<normalizedToken originalValue="×">x</normalizedToken>
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, or lower magnification.
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</paragraph>
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<paragraph pageId="20" pageNumber="37">Antennae (Fig. 4D). Antennae filiform, extended about body length. Antennomere 1 rather slender, slightly longer than antennomeres a2-a4; antennomere 2 short, about half length of a3; antennomeres 3-11 narrow, cylindrical, distinctly longer than wide.</paragraph>
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<paragraph pageId="20" pageNumber="37">
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Mouthparts. As described for
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<taxonomicName class="Insecta" family="Carabidae" genus="Zuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zuphium" order="Coleoptera" pageId="20" pageNumber="37" phylum="Arthropoda" rank="genus">Zuphium</taxonomicName>
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genus-group, above. Labrum (Fig. 5C). Anterior margin concave. Epipharynx (Fig. 5D). Mandibles (Figs 7
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<normalizedToken originalValue="A–">A-</normalizedToken>
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7F). Maxillae (Figs 8D, 8E). Labium, ventral aspect (Fig. 8F).
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</paragraph>
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<paragraph pageId="20" pageNumber="37">Pronotum (Fig. 1B). Anterior margin truncate, lateral margins markedly sinuate posteriorly, posteriolateral angles prominent, dentiform, slightly anteriad posterior margin; surface impressions (anterior and posterior transverse and median longitudinal) shallow; lateral grooves and posteriolateral impressions moderately deep.</paragraph>
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<paragraph pageId="21" pageNumber="38">
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<pageBreakToken pageId="21" pageNumber="38" start="start">Pterothorax</pageBreakToken>
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. Metasternum of average length; metepisternum elongate, lateral margin longer than anterior margin.
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</paragraph>
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<paragraph pageId="21" pageNumber="38">Elytra (Fig. 1B). Separate from one another, not fused along suture. Each elytron more or less rectangular, lateral margin straight; humerus projected anteriorly, basal ridge straight; apical margin truncate, with narrow band of membrane. Surface with striae very shallow, intervals almost flat.</paragraph>
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<paragraph pageId="21" pageNumber="38">Hind wings. Macropterous; wings folded beneath elytra at rest.</paragraph>
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<paragraph pageId="21" pageNumber="38">Legs. Male fore-tarsus (Fig. 9A).</paragraph>
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<paragraph pageId="21" pageNumber="38">
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Abdomen. As described for Western Hemisphere
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<taxonomicName class="Insecta" family="Carabidae" genus="Zuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zuphium" order="Coleoptera" pageId="21" pageNumber="38" phylum="Arthropoda" rank="genus">Zuphium</taxonomicName>
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genus-group.
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</paragraph>
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<paragraph pageId="21" pageNumber="38">
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Male genitalia (Figs 10
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<normalizedToken originalValue="D–">D-</normalizedToken>
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10F). Phallus narrow (PW/PL 0.254), slightly curved ventrally, narrowed apically, apical margin rounded, apical portion very short (AL/PL 0.036), ostial membrane extensive (OM/PL 0.317). Left paramere (lp) conchoid. Right paramere (rp) short, ovoid in form (RP/LP 0.317).
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</paragraph>
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<paragraph pageId="21" pageNumber="38">
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Female genitalia: ovipositor (Figs 11
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<normalizedToken originalValue="D–">D-</normalizedToken>
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11F). Gonocoxite 1 (gc1) with patch of long trichoid setae distally on ventral surface. Gonocoxite 2 (gc2) short, thick; in lateral aspect falciform, apex pointed, row of four long trichoid setae dorso- and ventro-laterally; in dorso-ventral aspect, broad, paddle-like, apex broadly rounded, not notched; otherwise, as described for Western Hemisphere
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<taxonomicName class="Insecta" family="Carabidae" genus="Zuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zuphium" order="Coleoptera" pageId="21" pageNumber="38" phylum="Arthropoda" rank="genus">Zuphium</taxonomicName>
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genus-group.
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</paragraph>
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<paragraph pageId="21" pageNumber="38">Female genital tract (Fig. 12B). Bursa copulatrix (bc) bulbous (collapsed in Fig. 12B), inserted at base of common oviduct (co). Spermatheca (sp) inserted near junction of common oviduct and bursa copulatrix, markedly elongate and slender, with helminthoid sclerite (hs) at base. Spermathecal gland duct inserted near base of spermatheca, with distinct swelling proximad spermathecal gland (spg). Without secondary spermathecal gland.</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="38" type="geographical distribution">
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<paragraph pageId="21" pageNumber="38">Geographical distribution.</paragraph>
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<paragraph pageId="21" pageNumber="38">
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This species ranges from northern Mexico (states of Nuevo Leon, Nayarit, and Tamaulipas) northward to southwestern U.S.A (states of Arizona, New Mexico, and Texas;
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<bibRefCitation author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="24" pageNumber="41" pagination="1 - 1722" title="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico." url="10.3897/zookeys.245.3416" volume="245" year="2012">Bousquet (2012</bibRefCitation>
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: 1357).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |