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<document id="D3B706EC989643C4194CCB67943349CD" ID-CLB-Dataset="7182" ID-DOI="10.11646/zootaxa.4963.1.5" ID-GBIF-Dataset="ee9bfd7b-3b05-4403-9af2-e54beaae5e69" ID-ISSN="1175-5326" ID-Zenodo-Dep="4719924" ID-ZooBank="6B4B8F71-7B08-4027-A76B-7EE8C987C9BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618562743172" checkinUser="plazi" docAuthor="Zhu, Wenhui &amp; Li, Shuqiang" docDate="2021" docId="4E3C87F3FFCBFF9EC6ECF9D2FCC0FD3A" docLanguage="en" docName="zootaxa.4963.1.5.pdf" docOrigin="Zootaxa 4963 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belisana putao Zhu &amp; Li 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="130" masterDocId="B205FF8BFFC1FF91C67BFFC1FFB6FF80" masterDocTitle="Six new species of the spider genus Belisana (Araneae: Pholcidae) from Southeast Asia" masterLastPageNumber="137" masterPageNumber="115" pageNumber="125" updateTime="1698937141523" updateUser="ExternalLinkService">
<mods:mods id="76F0BEA142F1DDEAAAAA1BCC7FD9D676" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8E2AADFF4065388F39C334AD6A19BC1F">
<mods:title id="F60D4E035839374D42A6AFBACE401682">Six new species of the spider genus Belisana (Araneae: Pholcidae) from Southeast Asia</mods:title>
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<mods:name id="BA828C20A4D4E41FC4DAB812716AA9F1" type="personal">
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<mods:namePart id="31184255F84BBAD201AFF3E21B8C1356">Zhu, Wenhui</mods:namePart>
<mods:nameIdentifier id="5804A0BBC8A73318C85C326B2001719B" type="ORCID">0000-0001-5650-1670</mods:nameIdentifier>
<mods:affiliation id="F70409DCD6031848FE8607E1CF2B55DB">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar &amp; zhuwh 1016 @ outlook. com; https: // orcid. org / 0000 - 0001 - 5650 - 1670</mods:affiliation>
<mods:nameIdentifier id="CA2F093384EB11922BA8AEA0B40CC4C9" type="email">zhuwh1016@outlook.com</mods:nameIdentifier>
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<mods:name id="0B658DEC9AC1B92D2205014E20FF4999" type="personal">
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<mods:namePart id="80FE8C14DF51BC53547BF933477999D2">Li, Shuqiang</mods:namePart>
<mods:nameIdentifier id="DB8E3DD2AA3CEA6C254887A51C176818" type="ORCID">0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation id="C10994AD7E88EB62A1955759B8E042C8">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; lisq @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 3290 - 5416</mods:affiliation>
<mods:nameIdentifier id="632A2AFFF174C7AE90AA7E1CF5431029" type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
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<mods:title id="507FC4D5810391977C5B38A7C23A0DCE">Zootaxa</mods:title>
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<mods:date id="E9981A8AF140F74356C62E4AD5B3025A">2021</mods:date>
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<mods:number id="83AD9AE8A770D0075B8B77ABC6C22510">2021-04-16</mods:number>
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<treatment id="4E3C87F3FFCBFF9EC6ECF9D2FCC0FD3A" ID-DOI="http://doi.org/10.5281/zenodo.4697006" ID-GBIF-Taxon="181600550" ID-Zenodo-Dep="4697006" LSID="urn:lsid:plazi:treatment:4E3C87F3FFCBFF9EC6ECF9D2FCC0FD3A" httpUri="http://treatment.plazi.org/id/4E3C87F3FFCBFF9EC6ECF9D2FCC0FD3A" lastPageId="15" lastPageNumber="130" pageId="10" pageNumber="125">
<subSubSection id="8E8F656EFFCBFF9BC6ECF9D2FE04F9AD" box="[151,434,1555,1582]" pageId="10" pageNumber="125" type="nomenclature">
<paragraph id="C62A36E5FFCBFF9BC6ECF9D2FE04F9AD" blockId="10.[151,434,1555,1618]" box="[151,434,1555,1582]" pageId="10" pageNumber="125">
<heading id="9D628189FFCBFF9BC6ECF9D2FE04F9AD" bold="true" box="[151,434,1555,1582]" fontSize="11" level="1" pageId="10" pageNumber="125" reason="1">
<emphasis id="F4E1EAF7FFCBFF9BC6ECF9D2FE04F9AD" bold="true" box="[151,434,1555,1582]" pageId="10" pageNumber="125">
<taxonomicName id="01954D66FFCBFF9BC6ECF9D2FEFAF9AD" ID-CoL="98YXX" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[151,332,1555,1582]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="125" phylum="Arthropoda" rank="species" species="putao" status="sp. nov.">
<emphasis id="F4E1EAF7FFCBFF9BC6ECF9D2FEFAF9AD" bold="true" box="[151,332,1555,1582]" italics="true" pageId="10" pageNumber="125">Belisana putao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EFD2578CFFCBFF9BC72FF9D2FE04F9AD" box="[340,434,1555,1581]" pageId="10" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6ECF9F6FE38F9D2" box="[151,398,1591,1618]" pageId="10" pageNumber="125" type="description">
<paragraph id="C62A36E5FFCBFF9BC6ECF9F6FE38F9D2" blockId="10.[151,434,1555,1618]" box="[151,398,1591,1618]" pageId="10" pageNumber="125">
<figureCitation id="5EAE2A60FFCBFF9BC6ECF9F6FEBFF9D1" box="[151,265,1591,1617]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="13.[151,250,1875,1899]" captionStartId-1="14.[151,250,1910,1934]" captionTargetBox-0="[226,1361,181,1851]" captionTargetBox-1="[214,1374,181,1886]" captionTargetId-0="figure-17@13.[226,1361,181,1851]" captionTargetId-1="figure-19@14.[214,1374,181,1886]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURE 9. Belisana putao sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." captionText-1="FIGURE 10. Belisana putao sp. nov., holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; C–E. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (A–B), 0.20 (C–E)." figureDoi-0="http://doi.org/10.5281/zenodo.4719949" figureDoi-1="http://doi.org/10.5281/zenodo.4719951" httpUri-0="https://zenodo.org/record/4719949/files/figure.png" httpUri-1="https://zenodo.org/record/4719951/files/figure.png" pageId="10" pageNumber="125">Figs 9–10</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC76FF9F6FEE3F9D1" box="[276,341,1591,1617]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13I–J</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC71AF9F6FE38F9D2" box="[353,398,1591,1618]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="10" pageNumber="125">15E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6ECF9BEFB64F93D" pageId="10" pageNumber="125" type="materials_examined">
<paragraph id="C62A36E5FFCBFF9BC6ECF9BEFB64F93D" blockId="10.[151,1437,1663,2014]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6ECF9BEFEFEF919" bold="true" box="[151,328,1663,1689]" pageId="10" pageNumber="125">Type material.</emphasis>
<materialsCitation id="76FD3CB8FFCBFF9BC728F9BEFB64F93D" ID-GBIF-Occurrence="3091104312" collectingDate="2017-05-07" collectionCode="IZCAS" collectorName="Z. Chen &amp; J. Wu" country="Myanmar" elevation="874" latitude="27.496883" location="Male" longLatPrecision="1" longitude="97.17108" pageId="10" pageNumber="125" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="F4E1EAF7FFCBFF9BC728F9BEFE7FF919" bold="true" box="[339,457,1663,1689]" pageId="10" pageNumber="125">
<typeStatus id="192E8847FFCBFF9BC728F9BEFE72F919" box="[339,452,1663,1689]" pageId="10" pageNumber="125" type="holotype">Holotype</typeStatus>
:
</emphasis>
<location id="C34A603EFFCBFF9BC7AFF9BEFDB8F919" LSID="urn:lsid:plazi:treatment:4E3C87F3FFCBFF9EC6ECF9D2FCC0FD3A:C34A603EFFCBFF9BC7AFF9BEFDB8F919" box="[468,526,1663,1689]" country="Myanmar" latitude="27.496883" longLatPrecision="1" longitude="97.17108" name="Male" pageId="10" pageNumber="125" stateProvince="Kachin">Male</location>
(
<collectionCode id="A084AE20FFCBFF9BC464F9BEFDC4F919" box="[543,626,1663,1689]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="10" pageNumber="125">IZCAS</collectionCode>
Ar41910), roadside
<locationDeviation id="D7982FC7FFCBFF9BC519F9BEFB63F919" box="[866,1237,1663,1689]" firstLocation="Wasadum" pageId="10" pageNumber="125" secondLocation="Ziradum">between Wasadum and Ziradum</locationDeviation>
(
<geoCoordinate id="A3A15022FFCBFF9BC293F9BEFA21F919" box="[1256,1431,1663,1689]" degrees="27" direction="north" minutes="29" orientation="latitude" pageId="10" pageNumber="125" precision="1" seconds="48.78" value="27.496883">27°29’48.78”N</geoCoordinate>
,
<geoCoordinate id="A3A15022FFCBFF9BC6ECF962FEF5F93E" box="[151,323,1699,1726]" degrees="97" direction="east" minutes="10" orientation="longitude" pageId="10" pageNumber="125" precision="1" seconds="15.90" value="97.17108">97°10’15.90”E</geoCoordinate>
,
<elevation id="4DB8D1D6FFCBFF9BC735F962FDB0F93E" box="[334,518,1699,1726]" metricMagnitude="2" metricUnit="m" metricValue="8.74" pageId="10" pageNumber="125" unit="m" value="874.0">
elevation
<quantity id="016D9B00FFCBFF9BC7C7F962FDB0F93E" box="[444,518,1699,1726]" metricMagnitude="2" metricUnit="m" metricValue="8.74" pageId="10" pageNumber="125" unit="m" value="874.0">874 m</quantity>
</elevation>
), Putao,
<collectingRegion id="0451F807FFCBFF9BC413F962FD0AF93D" box="[616,700,1699,1725]" country="Myanmar" name="Kachin" pageId="10" pageNumber="125">Kachin</collectingRegion>
,
<collectingCountry id="BE827675FFCBFF9BC4BCF965FC80F93E" box="[711,822,1700,1726]" name="Myanmar" pageId="10" pageNumber="125">Myanmar</collectingCountry>
,
<date id="B22B1025FFCBFF9BC53BF962FC6FF93D" box="[832,985,1699,1726]" pageId="10" pageNumber="125" value="2017-05-07">
<collectingDate id="A26FE9CDFFCBFF9BC53BF962FC6FF93D" box="[832,985,1699,1726]" pageId="10" pageNumber="125" value="2017-05-07">07 May 2017</collectingDate>
</date>
,
<collectorName id="6B605333FFCBFF9BC59FF965FB88F93D" box="[996,1086,1699,1726]" pageId="10" pageNumber="125">Z. Chen</collectorName>
&amp;
<collectorName id="6B605333FFCBFF9BC219F965FB17F93E" box="[1122,1185,1700,1726]" pageId="10" pageNumber="125">J. Wu</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6BCF906FBCBF961" box="[199,1149,1735,1762]" pageId="10" pageNumber="125" type="etymology">
<paragraph id="C62A36E5FFCBFF9BC6BCF906FBCBF961" blockId="10.[151,1437,1663,2014]" box="[199,1149,1735,1762]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCF906FEF8F961" bold="true" box="[199,334,1735,1761]" pageId="10" pageNumber="125">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="192E8847FFCBFF9BC4CDF909FD50F962" box="[694,742,1736,1762]" pageId="10" pageNumber="125">type</typeStatus>
locality and is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6BCF92AFEF7F815" pageId="10" pageNumber="125" type="diagnosis">
<paragraph id="C62A36E5FFCBFF9BC6BCF92AFEF7F815" blockId="10.[151,1437,1663,2014]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCF92AFEF4F885" bold="true" box="[199,322,1771,1797]" pageId="10" pageNumber="125">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="01954D66FFCBFF9BC434F92DFCE5F886" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" box="[591,851,1771,1798]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="125" phylum="Arthropoda" rank="species" species="pianma">
<emphasis id="F4E1EAF7FFCBFF9BC434F92DFD74F885" box="[591,706,1772,1797]" italics="true" pageId="10" pageNumber="125">B. pianma</emphasis>
<bibRefCitation id="A2044B14FFCBFF9BC4B2F92AFCE5F886" author="Huber, B. A." box="[713,851,1771,1798]" journalOrPublisher="Zoologica" pageId="10" pageNumber="125" pagination="1 - 126" part="155" refId="ref8453" refString="Huber, B. A. (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." title="High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae)" type="journal article" year="2005">Huber, 2005</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="A2044B14FFCBFF9BC5F5F92AFBF6F885" author="Huber, B. A." box="[910,1088,1771,1798]" journalOrPublisher="Zoologica" pageId="10" pageNumber="125" pagination="1 - 126" part="155" refId="ref8453" refString="Huber, B. A. (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." title="High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae)" type="journal article" year="2005">Huber 2005: 14</bibRefCitation>
, figs 5, 6, 56, 88–92, 112–130) with the elongated distal apophyses on the male chelicerae (
<figureCitation id="5EAE2A60FFCBFF9BC551F8CEFC2CF8A9" box="[810,922,1807,1833]" captionStart="FIGURE 10" captionStartId="14.[151,250,1910,1934]" captionTargetBox="[214,1374,181,1886]" captionTargetId="figure-19@14.[214,1374,181,1886]" captionTargetPageId="14" captionText="FIGURE 10. Belisana putao sp. nov., holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; C–E. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (A–B), 0.20 (C–E)." figureDoi="http://doi.org/10.5281/zenodo.4719951" httpUri="https://zenodo.org/record/4719951/files/figure.png" pageId="10" pageNumber="125">Figs. 10B</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC5DFF8CEFC64F8AA" box="[932,978,1807,1834]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="10" pageNumber="125">15E</figureCitation>
) but can be distinguished by the presence of a subdisto-ventral membranous lamella (arrow
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<figureCitation id="5EAE2A60FFCBFF9BC494F8F2FCFEF8CD" box="[751,840,1843,1869]" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="10" pageNumber="125">Figs 9C</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC528F8F2FCCFF8CD" box="[851,889,1843,1869]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13I</figureCitation>
) on the procursus, a pair of angular apophyses on the bulbal apophyses (arrow in
<figureCitation id="5EAE2A60FFCBFF9BC78AF896FDEDF8F1" box="[497,603,1879,1905]" captionStart="FIGURE 10" captionStartId="14.[151,250,1910,1934]" captionTargetBox="[214,1374,181,1886]" captionTargetId="figure-19@14.[214,1374,181,1886]" captionTargetPageId="14" captionText="FIGURE 10. Belisana putao sp. nov., holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; C–E. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (A–B), 0.20 (C–E)." figureDoi="http://doi.org/10.5281/zenodo.4719951" httpUri="https://zenodo.org/record/4719951/files/figure.png" pageId="10" pageNumber="125">Figs 10A</figureCitation>
), the absence of a ventral hump on the palpal femur and a pair of apophyses on the clypeus.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9EC6BCF85EFB67FD16" lastPageId="15" lastPageNumber="130" pageId="10" pageNumber="125" type="description">
<paragraph id="C62A36E5FFCBFF9EC6BCF85EFAEDFDD1" blockId="10.[151,1437,1663,2014]" lastBlockId="15.[151,1437,150,698]" lastPageId="15" lastPageNumber="130" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCF85EFE29F839" bold="true" box="[199,415,1951,1977]" pageId="10" pageNumber="125">Description. Male</emphasis>
(
<emphasis id="F4E1EAF7FFCBFF9BC7C9F85EFDAFF839" bold="true" box="[434,537,1951,1977]" pageId="10" pageNumber="125">
<typeStatus id="192E8847FFCBFF9BC7C9F85EFDAFF839" box="[434,537,1951,1977]" pageId="10" pageNumber="125" type="holotype">holotype</typeStatus>
</emphasis>
, IZCAS Ar41910): Total length 1.60 (1.66 with clypeus), carapace 0.66 long, 0.66 wide, opisthosoma 1.00 long, 0.53 wide. Leg I: 11.39 (3.00 + 0.17 + 2.97 + 4.09 + 1.16), leg II: 7.65 (2.03 + 0.15 + 1.91 + 2.72 + 0.84), leg III: 4.85 (1.41 + 0.09 + 1.13 + 1.59 + 0.63), leg IV: 8.04 (2.28 + 0.16 + 1.94 + 2.75 + 0.91); tibia I L/d: 50. Distance PME-PME 0.14, diameter PME 0.06, distance PME-ALE 0.09, AME absent. Habitus as in
<figureCitation id="5EAE2A60FFCEFF9EC6A8FF1DFEEBFF76" box="[211,349,220,246]" captionStart="FIGURE 10" captionStartId="14.[151,250,1910,1934]" captionTargetBox="[214,1374,181,1886]" captionTargetId="figure-19@14.[214,1374,181,1886]" captionTargetPageId="14" captionText="FIGURE 10. Belisana putao sp. nov., holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; C–E. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (A–B), 0.20 (C–E)." figureDoi="http://doi.org/10.5281/zenodo.4719951" httpUri="https://zenodo.org/record/4719951/files/figure.png" pageId="15" pageNumber="130">Figs 10C–E</figureCitation>
. Carapace pale. Ocular area not elevated. Clypeus unmodified. Chelicerae (
<figureCitation id="5EAE2A60FFCEFF9EC2D4FF1DFA97FF76" box="[1199,1313,220,246]" captionStart="FIGURE 10" captionStartId="14.[151,250,1910,1934]" captionTargetBox="[214,1374,181,1886]" captionTargetId="figure-19@14.[214,1374,181,1886]" captionTargetPageId="14" captionText="FIGURE 10. Belisana putao sp. nov., holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; C–E. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (A–B), 0.20 (C–E)." figureDoi="http://doi.org/10.5281/zenodo.4719951" httpUri="https://zenodo.org/record/4719951/files/figure.png" pageId="15" pageNumber="130">Figs. 10B</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC357FF1DFAECFF77" box="[1324,1370,220,247]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="15" pageNumber="130">15E</figureCitation>
) with pair of small proximo-lateral apophyses and pair of long, curved distal apophyses, each with sclerotized and pointed tip (distance between tips of distal apophyses: 0.39). Sternum pale, approximately as wide as long (0.41). Thoracic furrow absent. Legs pale. Opisthosoma pale, without spots. Palps as in
<figureCitation id="5EAE2A60FFCEFF9EC5B6FE85FBFAFEDE" box="[973,1100,324,350]" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Figs 9A–B</figureCitation>
; coxa unmodified; trochanter with short retrolatero-ventral apophysis; femur with retrolatero-dorsal apophysis (arrow in
<figureCitation id="5EAE2A60FFCEFF9EC2F0FEA6FB6CFE01" box="[1163,1242,359,385]" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Fig 9B</figureCitation>
); procursus (
<figureCitation id="5EAE2A60FFCEFF9EC311FEA6FF6BFE23" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Figs 9A–D</figureCitation>
) curved (arrow in
<figureCitation id="5EAE2A60FFCEFF9EC7CFFE48FDB8FE23" box="[436,526,393,419]" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Fig. 9A</figureCitation>
), complex distally, with droplet-shaped subdisto-dorsal sclerite (arrow
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<figureCitation id="5EAE2A60FFCEFF9EC317FE48FF0CFE46" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Figs 9C</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC6BDFE6DFF58FE46" box="[198,238,428,454]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13I</figureCitation>
), small subdisto-prolateral sclerite (arrow
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<figureCitation id="5EAE2A60FFCEFF9EC486FE6DFCE1FE46" box="[765,855,428,454]" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Figs 9C</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC51FFE6DFC3AFE46" box="[868,908,428,454]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13I</figureCitation>
), semicircular subdisto-prolateral membranous lamella (arrow
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<figureCitation id="5EAE2A60FFCEFF9EC70DFE0EFE67FE69" box="[374,465,463,489]" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Figs 9C</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC7A6FE0EFDB3FE69" box="[477,517,463,489]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13I</figureCitation>
), subdisto-ventral membranous lamella (arrow in
<figureCitation id="5EAE2A60FFCEFF9EC243FE0EFB22FE69" box="[1080,1172,463,489]" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Figs 9D</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC2DAFE0EFB7EFE69" box="[1185,1224,463,489]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13J</figureCitation>
) and with retrolateral membranous flap (arrow f in
<figureCitation id="5EAE2A60FFCEFF9EC46BFE30FDDDFD8B" box="[528,619,497,523]" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Figs 9D</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC40CFE30FD17FD8B" box="[631,673,497,523]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13J</figureCitation>
); bulb (
<figureCitation id="5EAE2A60FFCEFF9EC48DFE30FCEAFD8B" box="[758,860,497,523]" captionStart="FIGURE 10" captionStartId="14.[151,250,1910,1934]" captionTargetBox="[214,1374,181,1886]" captionTargetId="figure-19@14.[214,1374,181,1886]" captionTargetPageId="14" captionText="FIGURE 10. Belisana putao sp. nov., holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; C–E. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (A–B), 0.20 (C–E)." figureDoi="http://doi.org/10.5281/zenodo.4719951" httpUri="https://zenodo.org/record/4719951/files/figure.png" pageId="15" pageNumber="130">Fig. 10A</figureCitation>
) with hooked apophysis (pair of angular apophyses subdistally, arrow in
<figureCitation id="5EAE2A60FFCEFF9EC7FBFDD5FE5EFDAE" box="[384,488,532,558]" captionStart="FIGURE 10" captionStartId="14.[151,250,1910,1934]" captionTargetBox="[214,1374,181,1886]" captionTargetId="figure-19@14.[214,1374,181,1886]" captionTargetPageId="14" captionText="FIGURE 10. Belisana putao sp. nov., holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; C–E. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (A–B), 0.20 (C–E)." figureDoi="http://doi.org/10.5281/zenodo.4719951" httpUri="https://zenodo.org/record/4719951/files/figure.png" pageId="15" pageNumber="130">Figs 10A</figureCitation>
) and simple embolus. Retrolateral trichobothria of tibia Iat 6% proximally; legs with short vertical setae on metatarsi, without spines and curved setae; tarsus Iwith 15 distinct pseudopodomeres.
</paragraph>
<caption id="92EA666DFFCAFF9AC6ECF892FDC0F87B" ID-DOI="http://doi.org/10.5281/zenodo.4719944" ID-Zenodo-Dep="4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="11" pageNumber="126" startId="11.[151,250,1875,1899]" targetBox="[226,1361,181,1851]" targetPageId="11">
<paragraph id="C62A36E5FFCAFF9AC6ECF892FDC0F87B" blockId="11.[151,1436,1875,2043]" pageId="11" pageNumber="126">
<emphasis id="F4E1EAF7FFCAFF9AC6ECF892FEA6F8EB" bold="true" box="[151,272,1875,1899]" pageId="11" pageNumber="126">FIGURE 7</emphasis>
.
<taxonomicName id="01954D66FFCAFF9AC764F892FE73F8EB" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[287,453,1875,1899]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="126" phylum="Arthropoda" rank="species" species="kachin" status="sp. nov.">
<emphasis id="F4E1EAF7FFCAFF9AC764F892FE73F8EB" box="[287,453,1875,1899]" italics="true" pageId="11" pageNumber="126">Belisana kachin</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFCAFF9AC7B6F892FD97F8EB" bold="true" box="[461,545,1875,1899]" pageId="11" pageNumber="126">
<taxonomicNameLabel id="EFD2578CFFCAFF9AC7B6F892FD97F8EB" box="[461,545,1875,1899]" pageId="11" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. A–B. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D).
</paragraph>
</caption>
<caption id="92EA666DFFCDFF9DC6ECF892FEA6F87B" ID-DOI="http://doi.org/10.5281/zenodo.4719947" ID-Zenodo-Dep="4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="12" pageNumber="127" startId="12.[151,250,1875,1899]" targetBox="[226,1361,181,1851]" targetPageId="12">
<paragraph id="C62A36E5FFCDFF9DC6ECF892FEA6F87B" blockId="12.[151,1437,1875,2043]" pageId="12" pageNumber="127">
<emphasis id="F4E1EAF7FFCDFF9DC6ECF892FEA3F8EB" bold="true" box="[151,277,1875,1899]" pageId="12" pageNumber="127">FIGURE 8.</emphasis>
<taxonomicName id="01954D66FFCDFF9DC766F892FE74F8EB" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[285,450,1875,1899]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="127" phylum="Arthropoda" rank="species" species="kachin" status="sp. nov.">
<emphasis id="F4E1EAF7FFCDFF9DC766F892FE74F8EB" box="[285,450,1875,1899]" italics="true" pageId="12" pageNumber="127">Belisana kachin</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFCDFF9DC7B1F892FDABF8EB" bold="true" box="[458,541,1875,1899]" pageId="12" pageNumber="127">
<taxonomicNameLabel id="EFD2578CFFCDFF9DC7B1F892FDABF8EB" box="[458,541,1875,1899]" pageId="12" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H).
</paragraph>
</caption>
<caption id="92EA666DFFCCFF9CC6ECF892FC50F87B" ID-DOI="http://doi.org/10.5281/zenodo.4719949" ID-Zenodo-Dep="4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="13" pageNumber="128" startId="13.[151,250,1875,1899]" targetBox="[226,1361,181,1851]" targetPageId="13">
<paragraph id="C62A36E5FFCCFF9CC6ECF892FC50F87B" blockId="13.[151,1436,1875,2043]" pageId="13" pageNumber="128">
<emphasis id="F4E1EAF7FFCCFF9CC6ECF892FEB9F8EB" bold="true" box="[151,271,1875,1899]" pageId="13" pageNumber="128">FIGURE 9</emphasis>
.
<taxonomicName id="01954D66FFCCFF9CC766F892FE0EF8EB" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[285,440,1875,1899]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="128" phylum="Arthropoda" rank="species" species="putao" status="sp. nov.">
<emphasis id="F4E1EAF7FFCCFF9CC766F892FE0EF8EB" box="[285,440,1875,1899]" italics="true" pageId="13" pageNumber="128">Belisana putao</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFCCFF9CC7BBF892FDA5F8EB" bold="true" box="[448,531,1875,1899]" pageId="13" pageNumber="128">
<taxonomicNameLabel id="EFD2578CFFCCFF9CC7BBF892FDA5F8EB" box="[448,531,1875,1899]" pageId="13" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D).
</paragraph>
</caption>
<caption id="92EA666DFFCFFF9FC6ECF8B7FAD9F856" ID-DOI="http://doi.org/10.5281/zenodo.4719951" ID-Zenodo-Dep="4719951" httpUri="https://zenodo.org/record/4719951/files/figure.png" pageId="14" pageNumber="129" startId="14.[151,250,1910,1934]" targetBox="[214,1374,181,1886]" targetPageId="14">
<paragraph id="C62A36E5FFCFFF9FC6ECF8B7FAD9F856" blockId="14.[151,1436,1910,2006]" pageId="14" pageNumber="129">
<emphasis id="F4E1EAF7FFCFFF9FC6ECF8B7FE97F80E" bold="true" box="[151,289,1910,1934]" pageId="14" pageNumber="129">FIGURE 10.</emphasis>
<taxonomicName id="01954D66FFCFFF9FC752F8B7FE75F80E" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[297,451,1910,1934]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="129" phylum="Arthropoda" rank="species" species="putao" status="sp. nov.">
<emphasis id="F4E1EAF7FFCFFF9FC752F8B7FE75F80E" box="[297,451,1910,1934]" italics="true" pageId="14" pageNumber="129">Belisana putao</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFCFFF9FC7B0F8B7FDABF80E" bold="true" box="[459,541,1910,1934]" pageId="14" pageNumber="129">
<taxonomicNameLabel id="EFD2578CFFCFFF9FC7B0F8B7FDABF80E" box="[459,541,1910,1934]" pageId="14" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; C–E. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (A–B), 0.20 (C–E).
</paragraph>
</caption>
<paragraph id="C62A36E5FFCEFF9EC6BCFD98FE13FDF3" blockId="15.[151,1437,150,698]" box="[199,421,601,627]" pageId="15" pageNumber="130">
<emphasis id="F4E1EAF7FFCEFF9EC6BCFD98FE9EFDF3" bold="true" box="[199,296,601,627]" pageId="15" pageNumber="130">Female:</emphasis>
Unknown.
</paragraph>
<paragraph id="C62A36E5FFCEFF9EC6BCFDBDFB67FD16" blockId="15.[151,1437,150,698]" box="[199,1233,636,663]" pageId="15" pageNumber="130">
<emphasis id="F4E1EAF7FFCEFF9EC6BCFDBDFE33FD16" bold="true" box="[199,389,636,662]" pageId="15" pageNumber="130">Natural history.</emphasis>
The species was found on domed webs attached to the underside of leaves.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCEFF9EC6BCFD5FFCC0FD3A" box="[199,886,670,698]" pageId="15" pageNumber="130" type="distribution">
<paragraph id="C62A36E5FFCEFF9EC6BCFD5FFCC0FD3A" blockId="15.[151,1437,150,698]" box="[199,886,670,698]" pageId="15" pageNumber="130">
<emphasis id="F4E1EAF7FFCEFF9EC6BCFD5FFED7FD38" bold="true" box="[199,353,670,696]" pageId="15" pageNumber="130">Distribution.</emphasis>
<collectingCountry id="BE827675FFCEFF9EC71CFD5EFE62FD39" box="[359,468,671,697]" name="Myanmar" pageId="15" pageNumber="130">Myanmar</collectingCountry>
(
<collectingRegion id="0451F807FFCEFF9EC798FD5EFD8EFD39" box="[483,568,671,697]" country="Myanmar" name="Kachin" pageId="15" pageNumber="130">Kachin</collectingRegion>
,
<typeStatus id="192E8847FFCEFF9EC438FD5EFDC5FD39" box="[579,627,671,697]" pageId="15" pageNumber="130">type</typeStatus>
locality only;
<figureCitation id="5EAE2A60FFCEFF9EC56DFD5EFCDFFD3A" box="[790,873,671,698]" captionStart="FIGURE 16" captionStartId="21.[151,250,1791,1815]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-17@21.[151,1435,181,1768]" captionTargetPageId="21" captionText="FIGURE 16. Distribution records of six new species of the genus Belisana from Southern Asia. 1. B. bachma sp. nov.; 2. B. cucphuong sp. nov.; 3. B. jaegeri sp. nov.; 4. B. kachin sp. nov.; 5. B. putao sp. nov.; 6. B. tarang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4719963" httpUri="https://zenodo.org/record/4719963/files/figure.png" pageId="15" pageNumber="130">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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