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<document id="AC23F9BC5B5D26AE79D39FD3CA237C0B" ID-CLB-Dataset="7182" ID-DOI="10.11646/zootaxa.4963.1.5" ID-GBIF-Dataset="ee9bfd7b-3b05-4403-9af2-e54beaae5e69" ID-ISSN="1175-5326" ID-Zenodo-Dep="4719924" ID-ZooBank="6B4B8F71-7B08-4027-A76B-7EE8C987C9BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618562743172" checkinUser="plazi" docAuthor="Zhu, Wenhui &amp; Li, Shuqiang" docDate="2021" docId="4E3C87F3FFCBFF9BC6ECFF56FCC0FA42" docLanguage="en" docName="zootaxa.4963.1.5.pdf" docOrigin="Zootaxa 4963 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belisana kachin Zhu &amp; Li 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="125" masterDocId="B205FF8BFFC1FF91C67BFFC1FFB6FF80" masterDocTitle="Six new species of the spider genus Belisana (Araneae: Pholcidae) from Southeast Asia" masterLastPageNumber="137" masterPageNumber="115" pageNumber="125" updateTime="1698937141523" updateUser="ExternalLinkService">
<mods:mods id="EF261C7B36C8F58004753CCAE86F1644" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D6FE3084FE7D7BA8D675E3D56E081ADE">
<mods:title id="AB39FDD08F6516AC8BDD3EF4136E449D">Six new species of the spider genus Belisana (Araneae: Pholcidae) from Southeast Asia</mods:title>
</mods:titleInfo>
<mods:name id="4AF045DE5F6D3654B2A2419611D52A1B" type="personal">
<mods:role id="42CA176A3A0BF58CC1BC67AA582AE2E7">
<mods:roleTerm id="68D2776ED7BBAD4F9DA1C8378F5DA60D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E80C66711854756C26873094196680FB">Zhu, Wenhui</mods:namePart>
<mods:nameIdentifier id="54BA676909476F30B0223FB06BB571EA" type="ORCID">0000-0001-5650-1670</mods:nameIdentifier>
<mods:affiliation id="B9A453AF2A226651CDBE4302C312A54E">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar &amp; zhuwh 1016 @ outlook. com; https: // orcid. org / 0000 - 0001 - 5650 - 1670</mods:affiliation>
<mods:nameIdentifier id="8DF175C692FCCCAC90E3690C342751F1" type="email">zhuwh1016@outlook.com</mods:nameIdentifier>
</mods:name>
<mods:name id="2A529AF8B996559B33EEB7848920B3B1" type="personal">
<mods:role id="A3920A1293527E062D4915973231F112">
<mods:roleTerm id="E7CE13B1BB5E1E76D325A4DCCBEB4C19">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9AE43EF015A3BCBF0AEF95E1639DDB19">Li, Shuqiang</mods:namePart>
<mods:nameIdentifier id="75DFEB79205BB310FAD69F64AF402A6E" type="ORCID">0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation id="DC6AA030C75F6A88572341B5E3A89456">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; lisq @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 3290 - 5416</mods:affiliation>
<mods:nameIdentifier id="F29D24011595D78AB03709C8B0EAD055" type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
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<mods:typeOfResource id="BC9DEC9792071E7E196319E8324237D6">text</mods:typeOfResource>
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<mods:title id="D62D7322BDC300974AF7A6BDF4F65F2B">Zootaxa</mods:title>
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<mods:part id="A53CACAC4A5123A7EC32681D985F862D">
<mods:date id="881DDDA0C3D5A69C451C448FFE3EA0AA">2021</mods:date>
<mods:detail id="A915336928F5F0DCA2091BE0918237AE" type="pubDate">
<mods:number id="37EDB244549D41B2C25A13FE2DDD4497">2021-04-16</mods:number>
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<mods:detail id="65E0F868C854328F420E00650D7CDAC7" type="volume">
<mods:number id="A78B4A18CCDEC803FFE8E1571588B10D">4963</mods:number>
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<mods:number id="46247F0AAA30308F7C6710F977911719">1</mods:number>
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<mods:classification id="2480964AA677B6159166E12A0C64A462">journal article</mods:classification>
<mods:identifier id="E44FF92F25439BDEB0EBBD2B86F6BB51" type="CLB-Dataset">7182</mods:identifier>
<mods:identifier id="A61167B8C3C38CDA493F7D19C65E92CC" type="DOI">10.11646/zootaxa.4963.1.5</mods:identifier>
<mods:identifier id="9259A752309C783F589CC92512D317F1" type="GBIF-Dataset">ee9bfd7b-3b05-4403-9af2-e54beaae5e69</mods:identifier>
<mods:identifier id="659E8FD186F9A4EA140BC1FE823C1F0E" type="ISSN">1175-5326</mods:identifier>
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<treatment id="4E3C87F3FFCBFF9BC6ECFF56FCC0FA42" ID-DOI="http://doi.org/10.5281/zenodo.4697002" ID-GBIF-Taxon="181600551" ID-Zenodo-Dep="4697002" LSID="urn:lsid:plazi:treatment:4E3C87F3FFCBFF9BC6ECFF56FCC0FA42" httpUri="http://treatment.plazi.org/id/4E3C87F3FFCBFF9BC6ECFF56FCC0FA42" lastPageNumber="125" pageId="10" pageNumber="125">
<subSubSection id="8E8F656EFFCBFF9BC6ECFF56FE76FF31" box="[151,448,151,178]" pageId="10" pageNumber="125" type="nomenclature">
<paragraph id="C62A36E5FFCBFF9BC6ECFF56FE76FF31" blockId="10.[151,504,151,214]" box="[151,448,151,178]" pageId="10" pageNumber="125">
<heading id="9D628189FFCBFF9BC6ECFF56FE76FF31" bold="true" box="[151,448,151,178]" fontSize="11" level="1" pageId="10" pageNumber="125" reason="1">
<emphasis id="F4E1EAF7FFCBFF9BC6ECFF56FE76FF31" bold="true" box="[151,448,151,178]" pageId="10" pageNumber="125">
<taxonomicName id="01954D66FFCBFF9BC6ECFF56FEEDFF32" ID-CoL="98YXZ" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[151,347,151,178]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="125" phylum="Arthropoda" rank="species" species="kachin" status="sp. nov.">
<emphasis id="F4E1EAF7FFCBFF9BC6ECFF56FEEDFF32" bold="true" box="[151,347,151,178]" italics="true" pageId="10" pageNumber="125">Belisana kachin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EFD2578CFFCBFF9BC718FF56FE76FF31" box="[355,448,151,177]" pageId="10" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6ECFF7AFE4EFF56" box="[151,504,187,214]" pageId="10" pageNumber="125" type="description">
<paragraph id="C62A36E5FFCBFF9BC6ECFF7AFE4EFF56" blockId="10.[151,504,151,214]" box="[151,504,187,214]" pageId="10" pageNumber="125">
<figureCitation id="5EAE2A60FFCBFF9BC6ECFF7AFF4DFF55" box="[151,251,187,213]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="11.[151,250,1875,1899]" captionStartId-1="12.[151,250,1875,1899]" captionTargetBox-0="[226,1361,181,1851]" captionTargetBox-1="[226,1361,181,1851]" captionTargetId-0="figure-17@11.[226,1361,181,1851]" captionTargetId-1="figure-19@12.[226,1361,181,1851]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURE 7. Belisana kachin sp. nov., holotype male. A–B. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." captionText-1="FIGURE 8. Belisana kachin sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi-0="http://doi.org/10.5281/zenodo.4719944" figureDoi-1="http://doi.org/10.5281/zenodo.4719947" httpUri-0="https://zenodo.org/record/4719944/files/figure.png" httpUri-1="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 7–8</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC77DFF7AFEEAFF55" box="[262,348,187,213]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13G–H</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC71CFF7AFE0AFF55" box="[359,444,187,213]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="10" pageNumber="125">14G–H</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC7BCFF7AFE4EFF56" box="[455,504,187,214]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="10" pageNumber="125">15D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6ECFEC2FB23FEE5" pageId="10" pageNumber="125" type="materials_examined">
<paragraph id="C62A36E5FFCBFF9BC6ECFEC2FB23FEE5" blockId="10.[151,1437,259,1474]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6ECFEC2FEFDFE9D" bold="true" box="[151,331,259,285]" pageId="10" pageNumber="125">Type material.</emphasis>
<materialsCitation id="76FD3CB8FFCBFF9BC72CFEC2FED0FEE5" ID-GBIF-Occurrence="3091104304" collectingDate="2017-05-12" collectionCode="IZCAS" collectorName="Z. Chen &amp; J. Wu" country="Myanmar" county="Putao" elevation="1171" latitude="27.577435" location="Roadside" longLatPrecision="1" longitude="97.07573" municipality="Hponkanrazi Wildlife Sanctuary" pageId="10" pageNumber="125" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="F4E1EAF7FFCBFF9BC72CFEC2FE78FE9D" bold="true" box="[343,462,259,285]" pageId="10" pageNumber="125">
<typeStatus id="192E8847FFCBFF9BC72CFEC2FE7EFE9D" box="[343,456,259,285]" pageId="10" pageNumber="125" type="holotype">Holotype</typeStatus>
:
</emphasis>
Male (
<collectionCode id="A084AE20FFCBFF9BC453FEC2FDCDFE9D" box="[552,635,259,285]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="10" pageNumber="125">IZCAS</collectionCode>
Ar41907),
<location id="C34A603EFFCBFF9BC57CFEC2FCD9FE9D" LSID="urn:lsid:plazi:treatment:4E3C87F3FFCBFF9BC6ECFF56FCC0FA42:C34A603EFFCBFF9BC57CFEC2FCD9FE9D" box="[775,879,259,285]" country="Myanmar" county="Putao" latitude="27.577435" longLatPrecision="1" longitude="97.07573" municipality="Hponkanrazi Wildlife Sanctuary" name="Roadside" pageId="10" pageNumber="125" stateProvince="Kachin">Roadside</location>
between
<location id="C34A603EFFCBFF9BC59DFEC2FB9FFE9D" LSID="urn:lsid:plazi:treatment:4E3C87F3FFCBFF9BC6ECFF56FCC0FA42:C34A603EFFCBFF9BC59DFEC2FB9FFE9D" box="[998,1065,259,285]" country="Myanmar" county="Putao" latitude="27.577435" longLatPrecision="1" longitude="97.07573" municipality="Hponkanrazi Wildlife Sanctuary" name="Camp" pageId="10" pageNumber="125" stateProvince="Kachin">Camp</location>
2 to
<location id="C34A603EFFCBFF9BC208FEC2FB65FE9D" LSID="urn:lsid:plazi:treatment:4E3C87F3FFCBFF9BC6ECFF56FCC0FA42:C34A603EFFCBFF9BC208FEC2FB65FE9D" box="[1139,1235,259,285]" country="Myanmar" county="Putao" latitude="27.577435" longLatPrecision="1" longitude="97.07573" municipality="Hponkanrazi Wildlife Sanctuary" name="Ziradum" pageId="10" pageNumber="125" stateProvince="Kachin">Ziradum</location>
(
<geoCoordinate id="A3A15022FFCBFF9BC293FEC2FA21FE9D" box="[1256,1431,259,285]" degrees="27" direction="north" minutes="34" orientation="latitude" pageId="10" pageNumber="125" precision="1" seconds="38.76" value="27.577435">27°34’38.76”N</geoCoordinate>
,
<geoCoordinate id="A3A15022FFCBFF9BC6ECFEE6FEF5FEC1" box="[151,323,295,321]" degrees="97" direction="east" minutes="04" orientation="longitude" pageId="10" pageNumber="125" precision="1" seconds="32.64" value="97.07573">97°04’32.64”E</geoCoordinate>
,
<elevation id="4DB8D1D6FFCBFF9BC72BFEE6FDACFEC2" box="[336,538,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.1709999999999998" pageId="10" pageNumber="125" unit="m" value="1171.0">
elevation
<quantity id="016D9B00FFCBFF9BC7BBFEE6FDACFEC2" box="[448,538,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.1709999999999998" pageId="10" pageNumber="125" unit="m" value="1171.0">1171 m</quantity>
</elevation>
),
<collectingMunicipality id="264EAC9FFFCBFF9BC44BFEE6FC16FEC1" box="[560,928,295,321]" pageId="10" pageNumber="125">Hponkanrazi Wildlife Sanctuary</collectingMunicipality>
,
<collectingCounty id="2F4B4E69FFCBFF9BC5D6FEE9FC46FEC2" box="[941,1008,296,322]" pageId="10" pageNumber="125">Putao</collectingCounty>
,
<collectingRegion id="0451F807FFCBFF9BC586FEE6FBE7FEC1" box="[1021,1105,295,321]" country="Myanmar" name="Kachin" pageId="10" pageNumber="125">Kachin</collectingRegion>
,
<collectingCountry id="BE827675FFCBFF9BC225FEE9FB7BFEC2" box="[1118,1229,296,322]" name="Myanmar" pageId="10" pageNumber="125">Myanmar</collectingCountry>
,
<date id="B22B1025FFCBFF9BC2A1FEE6FAC1FEC1" box="[1242,1399,295,322]" pageId="10" pageNumber="125" value="2017-05-12">
<collectingDate id="A26FE9CDFFCBFF9BC2A1FEE6FAC1FEC1" box="[1242,1399,295,322]" pageId="10" pageNumber="125" value="2017-05-12">12 May 2017</collectingDate>
</date>
,
<collectorName id="6B605333FFCBFF9BC3FFFEE9FF64FEE5" pageId="10" pageNumber="125">Z. Chen</collectorName>
&amp;
<collectorName id="6B605333FFCBFF9BC68DFE8DFE83FEE6" box="[246,309,332,358]" pageId="10" pageNumber="125">J. Wu</collectorName>
leg.
</materialsCitation>
<materialsCitation id="76FD3CB8FFCBFF9BC716FE8AFB27FEE5" ID-GBIF-Occurrence="3091104305" box="[365,1169,331,358]" collectingDate="2017-05-12" collectionCode="IZCAS" collectorName="Z. Chen &amp; J. Wu" country="Myanmar" county="Putao" elevation="1171" latitude="27.577435" location="Roadside" longLatPrecision="1" longitude="97.07573" municipality="Hponkanrazi Wildlife Sanctuary" pageId="10" pageNumber="125" specimenCount="2" specimenCount-female="2" stateProvince="Kachin" typeStatus="paratype">
<emphasis id="F4E1EAF7FFCBFF9BC716FE8AFE58FEE5" bold="true" box="[365,494,331,357]" pageId="10" pageNumber="125">
<typeStatus id="192E8847FFCBFF9BC716FE8AFE5FFEE5" box="[365,489,331,357]" pageId="10" pageNumber="125" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="D093FD6CFFCBFF9BC78EFE8AFDD7FEE5" box="[501,609,331,357]" pageId="10" pageNumber="125" type="female">2 females</specimenCount>
(
<collectionCode id="A084AE20FFCBFF9BC40BFE8AFD75FEE5" box="[624,707,331,357]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="10" pageNumber="125">IZCAS</collectionCode>
Ar41908–41909), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6BCFEAEFBCAFE09" box="[199,1148,367,394]" pageId="10" pageNumber="125" type="etymology">
<paragraph id="C62A36E5FFCBFF9BC6BCFEAEFBCAFE09" blockId="10.[151,1437,259,1474]" box="[199,1148,367,394]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCFEAEFEF8FE09" bold="true" box="[199,334,367,393]" pageId="10" pageNumber="125">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="192E8847FFCBFF9BC4CEFEB1FD53FE0A" box="[693,741,368,394]" pageId="10" pageNumber="125">type</typeStatus>
locality and is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6BCFE52FD2EFDE1" pageId="10" pageNumber="125" type="diagnosis">
<paragraph id="C62A36E5FFCBFF9BC6BCFE52FD2EFDE1" blockId="10.[151,1437,259,1474]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCFE52FEF4FE2D" bold="true" box="[199,322,403,429]" pageId="10" pageNumber="125">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="01954D66FFCBFF9BC429FE55FCEDFE2E" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" box="[594,859,403,430]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="125" phylum="Arthropoda" rank="species" species="nujiang">
<emphasis id="F4E1EAF7FFCBFF9BC429FE55FD7EFE2D" box="[594,712,404,429]" italics="true" pageId="10" pageNumber="125">B. nujiang</emphasis>
<bibRefCitation id="A2044B14FFCBFF9BC4B4FE52FCEDFE2E" author="Huber, B. A." box="[719,859,403,430]" journalOrPublisher="Zoologica" pageId="10" pageNumber="125" pagination="1 - 126" part="155" refId="ref8453" refString="Huber, B. A. (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." title="High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae)" type="journal article" year="2005">Huber, 2005</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="A2044B14FFCBFF9BC5ECFE52FBFCFE2D" author="Huber, B. A." box="[919,1098,403,430]" journalOrPublisher="Zoologica" pageId="10" pageNumber="125" pagination="1 - 126" part="155" refId="ref8453" refString="Huber, B. A. (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." title="High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae)" type="journal article" year="2005">Huber 2005: 14</bibRefCitation>
, figs 7, 8, 57, 83–87, 93–111) with the elongated distal apophyses on the male chelicerae (
<figureCitation id="5EAE2A60FFCBFF9BC531FE76FC18FE51" box="[842,942,439,465]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs. 8C</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC5C6FE76FC58FE52" box="[957,1006,439,466]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="10" pageNumber="125">15D</figureCitation>
) and bulbal apophyses (
<figureCitation id="5EAE2A60FFCBFF9BC37DFE76FAD7FE51" box="[1286,1377,439,465]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Fig. 8D</figureCitation>
), but males can be distinguished by the absence of a distal spine and retrolateral flap on the procursus (
<figureCitation id="5EAE2A60FFCBFF9BC28FFE1AFAF8FE76" box="[1268,1358,475,502]" captionStart="FIGURE 7" captionStartId="11.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@11.[226,1361,181,1851]" captionTargetPageId="11" captionText="FIGURE 7. Belisana kachin sp. nov., holotype male. A–B. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="10" pageNumber="125">Fig. 7D</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC327FE1AFA26FE75" box="[1372,1424,475,501]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13H</figureCitation>
). Females can be distinguished from other species by the external genitalia with an anterior arch extending to the posterior part (arrow in
<figureCitation id="5EAE2A60FFCBFF9BC700FDE2FE63FDBD" box="[379,469,547,573]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8B</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC79AFDE2FDA3FDBD" box="[481,533,547,573]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="10" pageNumber="125">14H</figureCitation>
; anterior arch extending to the median part in
<taxonomicName id="01954D66FFCBFF9BC263FDE5FB3BFDBD" authorityName="Huber" authorityYear="2005" box="[1048,1165,548,573]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="125" phylum="Arthropoda" rank="species" species="nujiang">
<emphasis id="F4E1EAF7FFCBFF9BC263FDE5FB3BFDBD" box="[1048,1165,548,573]" italics="true" pageId="10" pageNumber="125">B. nujiang</emphasis>
</taxonomicName>
) and the different shape of the pore plates (arrow pp in
<figureCitation id="5EAE2A60FFCBFF9BC78AFD86FDFDFDE1" box="[497,587,583,609]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8B</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC42CFD86FD3DFDE1" box="[599,651,583,609]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="10" pageNumber="125">14H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6BCFDAAFB67FA1D" pageId="10" pageNumber="125" type="description">
<paragraph id="C62A36E5FFCBFF9BC6BCFDAAFC7DFBD9" blockId="10.[151,1437,259,1474]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCFDAAFE29FD05" bold="true" box="[199,415,619,645]" pageId="10" pageNumber="125">Description. Male</emphasis>
(
<emphasis id="F4E1EAF7FFCBFF9BC7C9FDAAFDAFFD05" bold="true" box="[434,537,619,645]" pageId="10" pageNumber="125">
<typeStatus id="192E8847FFCBFF9BC7C9FDAAFDAFFD05" box="[434,537,619,645]" pageId="10" pageNumber="125" type="holotype">holotype</typeStatus>
</emphasis>
, IZCAS Ar41907): Total length 1.70 (1.76 with clypeus), carapace 0.64 long, 0.62 wide, opisthosoma 1.12 long, 0.60 wide. Leg I: 16.34 (4.49 + 0.25 + 4.10 + 6.35 + 1.15), leg II: 11.13 (3.40 + 0.23 + 2.82 + 3.85 + 0.83), leg III: 7.44 (2.18 + 0.20 + 1.92 + 2.56 + 0.58), leg IV: 9.70 (2.88 + 0.21 + 2.69 + 3.21 + 0.71); tibia I L/d: 43. Distance PME-PME 0.19, diameter PME 0.06, distance PME-ALE 0.08, AME absent. Habitus as in
<figureCitation id="5EAE2A60FFCBFF9BC6AAFD3AFEF3FC95" box="[209,325,763,789]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8E–F</figureCitation>
. Carapace pale. Ocular area not elevated. Clypeus unmodified. Chelicerae (
<figureCitation id="5EAE2A60FFCBFF9BC2F7FD3AFB5BFC95" box="[1164,1261,763,789]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs. 8C</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC282FD3AFA9CFC96" box="[1273,1322,763,790]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="10" pageNumber="125">15D</figureCitation>
) with pair of small proximo-lateral apophyses and pair of long, curved distal apophyses, each with sclerotized and pointed tip (distance between tips of distal apophyses: 0.44). Sternum pale, slightly wider than long (0.53/0.52). Thoracic furrow absent. Legs pale. Opisthosoma pale, without spots. Palps as in
<figureCitation id="5EAE2A60FFCBFF9BC5E1FCA6FBAEFC01" box="[922,1048,871,897]" captionStart="FIGURE 7" captionStartId="11.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@11.[226,1361,181,1851]" captionTargetPageId="11" captionText="FIGURE 7. Belisana kachin sp. nov., holotype male. A–B. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="10" pageNumber="125">Figs 7A–B</figureCitation>
; coxa unmodified; trochanter with ventral apophysis (arrow in
<figureCitation id="5EAE2A60FFCBFF9BC7A7FC4AFD80FC26" box="[476,566,907,934]" captionStart="FIGURE 7" captionStartId="11.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@11.[226,1361,181,1851]" captionTargetPageId="11" captionText="FIGURE 7. Belisana kachin sp. nov., holotype male. A–B. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="10" pageNumber="125">Fig. 7B</figureCitation>
); femur without proximo-dorsal apophysis; procursus (
<figureCitation id="5EAE2A60FFCBFF9BC2CBFC4AFA85FC25" box="[1200,1331,907,933]" captionStart="FIGURE 7" captionStartId="11.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@11.[226,1361,181,1851]" captionTargetPageId="11" captionText="FIGURE 7. Belisana kachin sp. nov., holotype male. A–B. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="10" pageNumber="125">Figs 7A–D</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC33BFC4AFA22FC25" box="[1344,1428,907,933]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13G–H</figureCitation>
) simple proximally but complex distally, with curved subdisto-dorsal sclerite (arrow
<quantity id="016D9B00FFCBFF9BC23BFC6EFBDDFC49" box="[1088,1131,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="125" unit="in" value="1.0">1 in</quantity>
<figureCitation id="5EAE2A60FFCBFF9BC209FC6EFB7DFC49" box="[1138,1227,943,969]" captionStart="FIGURE 7" captionStartId="11.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@11.[226,1361,181,1851]" captionTargetPageId="11" captionText="FIGURE 7. Belisana kachin sp. nov., holotype male. A–B. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="10" pageNumber="125">Figs 7C</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC2ACFC6EFABCFC49" box="[1239,1290,943,969]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13G</figureCitation>
), large distoprolateral sclerite (arrow
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<figureCitation id="5EAE2A60FFCBFF9BC78BFC12FDFAFC6D" box="[496,588,979,1005]" captionStart="FIGURE 7" captionStartId="11.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@11.[226,1361,181,1851]" captionTargetPageId="11" captionText="FIGURE 7. Belisana kachin sp. nov., holotype male. A–B. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="10" pageNumber="125">Figs 7C</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC420FC12FD38FC6D" box="[603,654,979,1005]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13G</figureCitation>
), inverted L-shaped subdisto-prolateral sclerite (arrow
<quantity id="016D9B00FFCBFF9BC37EFC12FA84FC6D" box="[1285,1330,979,1005]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="125" unit="in" value="3.0">3 in</quantity>
<figureCitation id="5EAE2A60FFCBFF9BC340FC12FA21FC6D" box="[1339,1431,979,1005]" captionStart="FIGURE 7" captionStartId="11.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@11.[226,1361,181,1851]" captionTargetPageId="11" captionText="FIGURE 7. Belisana kachin sp. nov., holotype male. A–B. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="10" pageNumber="125">Figs 7C</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC6ECFC36FF7EFB91" box="[151,200,1015,1041]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13G</figureCitation>
) and disto-retrolateral membranous lamella (arrow in
<figureCitation id="5EAE2A60FFCBFF9BC564FC36FCCFFB92" box="[799,889,1015,1042]" captionStart="FIGURE 7" captionStartId="11.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@11.[226,1361,181,1851]" captionTargetPageId="11" captionText="FIGURE 7. Belisana kachin sp. nov., holotype male. A–B. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="10" pageNumber="125">Figs 7D</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC5FFFC36FC01FB91" box="[900,951,1015,1041]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13H</figureCitation>
); bulb (
<figureCitation id="5EAE2A60FFCBFF9BC271FC36FBD6FB91" box="[1034,1120,1015,1041]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Fig. 8D</figureCitation>
) with hooked apophysis and simple embolus. Retrolateral trichobothria of tibia Iat 5% proximally; legs with short vertical setae on metatarsi, without spines or curved setae; tarsus Iwith 9 distinct pseudopodomeres.
</paragraph>
<paragraph id="C62A36E5FFCBFF9BC6BCFBA2FE01FAD5" blockId="10.[151,1437,259,1474]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCFBA2FEA8FBFD" bold="true" box="[199,286,1123,1149]" pageId="10" pageNumber="125">Female</emphasis>
(IZCAS Ar41909): Similar to male, habitus as in
<figureCitation id="5EAE2A60FFCBFF9BC51BFBA2FC57FBFD" box="[864,993,1123,1149]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8G–H</figureCitation>
. Total length 1.60 (1.66 with clypeus), carapace 0.60 long, 0.52 wide, opisthosoma 1.06 long, 0.60 wide. Tibia I: 1.75; tibia I L/d: 21. Distance PME-PME 0.13, diameter PME 0.05, distance PME-ALE 0.04, AME absent. Sternum slightly wider than long (0.44/0.42). External female genitalia (
<figureCitation id="5EAE2A60FFCBFF9BC7C6FB0EFDACFB69" box="[445,538,1231,1257]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8A</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC45EFB0EFDE1FB69" box="[549,599,1231,1257]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="10" pageNumber="125">14G</figureCitation>
) simple and flat, with pair of frontal “epigynal” pockets laterally on sclerotized plate (arrow ep in
<figureCitation id="5EAE2A60FFCBFF9BC7DBFB32FDABFA8D" box="[416,541,1267,1293]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8A–B</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC453FB32FDC8FA8D" box="[552,638,1267,1293]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="10" pageNumber="125">14G–H</figureCitation>
), 0.42 apart. Vulva (
<figureCitation id="5EAE2A60FFCBFF9BC524FB32FC0DFA8D" box="[863,955,1267,1293]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8B</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC5BDFB32FC41FA8D" box="[966,1015,1267,1293]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="10" pageNumber="125">14H</figureCitation>
) with pointed anterior arch extending to posterior part, with scaly edge (arrow in
<figureCitation id="5EAE2A60FFCBFF9BC406FAD6FD61FAB1" box="[637,727,1303,1329]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8B</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC498FAD6FCA2FAB1" box="[739,788,1303,1329]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="10" pageNumber="125">14H</figureCitation>
) and pair of large pore plates (arrow pp in
<figureCitation id="5EAE2A60FFCBFF9BC28DFAD6FAE6FAB1" box="[1270,1360,1303,1329]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8B</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC327FAD6FA39FAB1" box="[1372,1423,1303,1329]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="10" pageNumber="125">14H</figureCitation>
), without serrated sclerites.
</paragraph>
<paragraph id="C62A36E5FFCBFF9BC6BCFA9EFCD5FAF9" blockId="10.[151,1437,259,1474]" box="[199,867,1375,1402]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCFA9EFEF7FAF9" bold="true" box="[199,321,1375,1401]" pageId="10" pageNumber="125">Variation:</emphasis>
Tibia Iin another male (IZCAS Ar41908): 4.04.
</paragraph>
<paragraph id="C62A36E5FFCBFF9BC6BCFA42FB67FA1D" blockId="10.[151,1437,259,1474]" box="[199,1233,1411,1438]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCFA42FE33FA1D" bold="true" box="[199,389,1411,1437]" pageId="10" pageNumber="125">Natural history.</emphasis>
The species was found on domed webs attached to the underside of leaves.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6BCFA66FCC0FA42" box="[199,886,1447,1474]" pageId="10" pageNumber="125" type="distribution">
<paragraph id="C62A36E5FFCBFF9BC6BCFA66FCC0FA42" blockId="10.[151,1437,259,1474]" box="[199,886,1447,1474]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCFA66FED7FA41" bold="true" box="[199,353,1447,1473]" pageId="10" pageNumber="125">Distribution.</emphasis>
<collectingCountry id="BE827675FFCBFF9BC71CFA69FE62FA42" box="[359,468,1448,1474]" name="Myanmar" pageId="10" pageNumber="125">Myanmar</collectingCountry>
(
<collectingRegion id="0451F807FFCBFF9BC798FA66FD8EFA41" box="[483,568,1447,1473]" country="Myanmar" name="Kachin" pageId="10" pageNumber="125">Kachin</collectingRegion>
,
<typeStatus id="192E8847FFCBFF9BC438FA69FDC5FA42" box="[579,627,1448,1474]" pageId="10" pageNumber="125">type</typeStatus>
locality only;
<figureCitation id="5EAE2A60FFCBFF9BC56DFA66FCDFFA42" box="[790,873,1447,1474]" captionStart="FIGURE 16" captionStartId="21.[151,250,1791,1815]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-17@21.[151,1435,181,1768]" captionTargetPageId="21" captionText="FIGURE 16. Distribution records of six new species of the genus Belisana from Southern Asia. 1. B. bachma sp. nov.; 2. B. cucphuong sp. nov.; 3. B. jaegeri sp. nov.; 4. B. kachin sp. nov.; 5. B. putao sp. nov.; 6. B. tarang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4719963" httpUri="https://zenodo.org/record/4719963/files/figure.png" pageId="10" pageNumber="125">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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