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<document id="E99F622AF29C537F96EC7FB87489C561" ID-CLB-Dataset="7182" ID-DOI="10.11646/zootaxa.4963.1.5" ID-GBIF-Dataset="ee9bfd7b-3b05-4403-9af2-e54beaae5e69" ID-ISSN="1175-5326" ID-Zenodo-Dep="4719924" ID-ZooBank="6B4B8F71-7B08-4027-A76B-7EE8C987C9BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618562743172" checkinUser="plazi" docAuthor="Zhu, Wenhui &amp; Li, Shuqiang" docDate="2021" docId="4E3C87F3FFC0FF95C6ECFBFFFC7BFD5C" docLanguage="en" docName="zootaxa.4963.1.5.pdf" docOrigin="Zootaxa 4963 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belisana bachma Zhu &amp; Li 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="119" masterDocId="B205FF8BFFC1FF91C67BFFC1FFB6FF80" masterDocTitle="Six new species of the spider genus Belisana (Araneae: Pholcidae) from Southeast Asia" masterLastPageNumber="137" masterPageNumber="115" pageNumber="116" updateTime="1698937141523" updateUser="ExternalLinkService">
<mods:mods id="7448A5B365ED6933F1FDD401A5B651C4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="928DC22B78F917E481BE4864F0D93C31">
<mods:title id="9A1FC76E698E05E84579600C20DD8F8C">Six new species of the spider genus Belisana (Araneae: Pholcidae) from Southeast Asia</mods:title>
</mods:titleInfo>
<mods:name id="A74CC9857B639067A7FCEE723EA7134B" type="personal">
<mods:role id="DAA18D79D403202D221C7B20F25A9AC5">
<mods:roleTerm id="2B25B8FEC4937543BBB2F71C8D384F30">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E3E5B4EFBE54AE24E498B1EFACAEF43C">Zhu, Wenhui</mods:namePart>
<mods:nameIdentifier id="32E4B0E72FFED7B422405ECA3C337386" type="ORCID">0000-0001-5650-1670</mods:nameIdentifier>
<mods:affiliation id="26B0FC707347A32A39CFE455D8FB8F53">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar &amp; zhuwh 1016 @ outlook. com; https: // orcid. org / 0000 - 0001 - 5650 - 1670</mods:affiliation>
<mods:nameIdentifier id="17323894C33077A1D1EEA41D3FBD793B" type="email">zhuwh1016@outlook.com</mods:nameIdentifier>
</mods:name>
<mods:name id="BD7EBAC7ED02750902B9E0B079FE2DCA" type="personal">
<mods:role id="2DA1D8770EF62695ED69EBDBF59FE234">
<mods:roleTerm id="7CD1B8BB410592F540C0D0A08EA78596">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1D6BB7E7756010F482393EC7D8835380">Li, Shuqiang</mods:namePart>
<mods:nameIdentifier id="E1A65F063773C73F0F909833A13BB9AA" type="ORCID">0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation id="24E96F80319506780F2A2254BE495F0E">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; lisq @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 3290 - 5416</mods:affiliation>
<mods:nameIdentifier id="D32689688AAFBCD7F15A8866B018EED8" type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="E4B881353E7F1AF5B7AA0D448E6865E2">text</mods:typeOfResource>
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<mods:title id="442457362C701E2369F606568FF5C232">Zootaxa</mods:title>
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<mods:part id="4DFB69059C997AF9771A91A71A8B75E9">
<mods:date id="E4C37923A0331CFE5A7781187AC6EB13">2021</mods:date>
<mods:detail id="FC4BACF1F1E47119F3AFC3073C2E8BDF" type="pubDate">
<mods:number id="6E19D65624CB18CDBB023B4F2A062841">2021-04-16</mods:number>
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<mods:detail id="7ADE2E28D53A96C04888A81218C436B4" type="volume">
<mods:number id="917822139BAEEBD12A2E896B50B4AB02">4963</mods:number>
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<mods:number id="5FB8392BFD3158F6D22DEB393B70E7D6">1</mods:number>
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<mods:classification id="91A46C1187A0EB85428FA2F0FFF213A9">journal article</mods:classification>
<mods:identifier id="CDBEBE3091FC0E91B4AB018C6E5701AE" type="CLB-Dataset">7182</mods:identifier>
<mods:identifier id="86894FAC5774D09CE79483A5CDFD357C" type="DOI">10.11646/zootaxa.4963.1.5</mods:identifier>
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<mods:identifier id="6CF95A1A713705B469D547211418790A" type="ISSN">1175-5326</mods:identifier>
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<treatment id="4E3C87F3FFC0FF95C6ECFBFFFC7BFD5C" ID-DOI="http://doi.org/10.5281/zenodo.4696996" ID-GBIF-Taxon="181600554" ID-Zenodo-Dep="4696996" LSID="urn:lsid:plazi:treatment:4E3C87F3FFC0FF95C6ECFBFFFC7BFD5C" httpUri="http://treatment.plazi.org/id/4E3C87F3FFC0FF95C6ECFBFFFC7BFD5C" lastPageId="4" lastPageNumber="119" pageId="1" pageNumber="116">
<subSubSection id="8E8F656EFFC0FF90C6ECFBFFFE78FBD9" box="[151,462,1086,1113]" pageId="1" pageNumber="116" type="nomenclature">
<paragraph id="C62A36E5FFC0FF90C6ECFBFFFE78FBD9" blockId="1.[151,500,1086,1149]" box="[151,462,1086,1113]" pageId="1" pageNumber="116">
<heading id="9D628189FFC0FF90C6ECFBFFFE78FBD9" bold="true" box="[151,462,1086,1113]" fontSize="11" level="1" pageId="1" pageNumber="116" reason="1">
<emphasis id="F4E1EAF7FFC0FF90C6ECFBFFFE78FBD9" bold="true" box="[151,462,1086,1113]" pageId="1" pageNumber="116">
<taxonomicName id="01954D66FFC0FF90C6ECFBFFFEDEFBD9" ID-CoL="98YXW" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[151,360,1086,1113]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="116" phylum="Arthropoda" rank="species" species="bachma" status="sp. nov.">
<emphasis id="F4E1EAF7FFC0FF90C6ECFBFFFEDEFBD9" bold="true" box="[151,360,1086,1113]" italics="true" pageId="1" pageNumber="116">Belisana bachma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EFD2578CFFC0FF90C70BFBFEFE78FBD9" box="[368,462,1087,1113]" pageId="1" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC0FF90C6ECFBA2FE42FBFD" box="[151,500,1122,1149]" pageId="1" pageNumber="116" type="description">
<paragraph id="C62A36E5FFC0FF90C6ECFBA2FE42FBFD" blockId="1.[151,500,1086,1149]" box="[151,500,1122,1149]" pageId="1" pageNumber="116">
<figureCitation id="5EAE2A60FFC0FF90C6ECFBA2FF4CFBFD" box="[151,250,1123,1149]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1860,1884]" captionStartId-1="3.[151,250,1826,1850]" captionTargetBox-0="[231,1357,179,1834]" captionTargetBox-1="[243,1345,181,1802]" captionTargetId-0="figure-19@2.[231,1357,179,1834]" captionTargetId-1="figure-17@3.[243,1345,181,1802]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Belisana bachma sp. nov., holotype male.A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." captionText-1="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi-0="http://doi.org/10.5281/zenodo.4719926" figureDoi-1="http://doi.org/10.5281/zenodo.4719928" httpUri-0="https://zenodo.org/record/4719926/files/figure.png" httpUri-1="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Figs 1–2</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C77DFBA2FEECFBFD" box="[262,346,1123,1149]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="1" pageNumber="116">13A–B</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C71EFBA2FE0EFBFD" box="[357,440,1123,1149]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="1" pageNumber="116">14A–B</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C7BFFBA3FE42FBFD" box="[452,500,1122,1149]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="1" pageNumber="116">15A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC0FF90C6ECFB6AFCBEFA8D" pageId="1" pageNumber="116" type="materials_examined">
<paragraph id="C62A36E5FFC0FF90C6ECFB6AFCBEFA8D" blockId="1.[151,1437,1194,1977]" pageId="1" pageNumber="116">
<emphasis id="F4E1EAF7FFC0FF90C6ECFB6AFEFFFB44" bold="true" box="[151,329,1194,1221]" pageId="1" pageNumber="116">Type material.</emphasis>
<materialsCitation id="76FD3CB8FFC0FF90C728FB6AFC45FB69" ID-GBIF-Occurrence="3091104307" collectingDate="2015-08-27" collectionCode="IZCAS" collectorName="D. S. Pham" country="Vietnam" elevation="371" latitude="16.229383" location="Male" longLatPrecision="1" longitude="107.84818" municipality="Bach Ma National Park" pageId="1" pageNumber="116" specimenCount="1" typeStatus="holotype">
<emphasis id="F4E1EAF7FFC0FF90C728FB6AFE7FFB45" bold="true" box="[339,457,1195,1221]" pageId="1" pageNumber="116">
<typeStatus id="192E8847FFC0FF90C728FB6AFE72FB45" box="[339,452,1195,1221]" pageId="1" pageNumber="116" type="holotype">Holotype</typeStatus>
:
</emphasis>
<location id="C34A603EFFC0FF90C7AFFB6AFDB8FB45" LSID="urn:lsid:plazi:treatment:4E3C87F3FFC0FF95C6ECFBFFFC7BFD5C:C34A603EFFC0FF90C7AFFB6AFDB8FB45" box="[468,526,1195,1221]" country="Vietnam" latitude="16.229383" longLatPrecision="1" longitude="107.84818" municipality="Bach Ma National Park" name="Male" pageId="1" pageNumber="116">Male</location>
(
<collectionCode id="A084AE20FFC0FF90C45BFB6AFDC5FB45" box="[544,627,1195,1221]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="116">IZCAS</collectionCode>
Ar41901),
<collectingMunicipality id="264EAC9FFFC0FF90C481FB6AFBBAFB45" box="[762,1036,1195,1221]" pageId="1" pageNumber="116">Bach Ma National Park</collectingMunicipality>
(
<geoCoordinate id="A3A15022FFC0FF90C25BFB6BFB79FB45" box="[1056,1231,1194,1221]" degrees="16" direction="north" minutes="13" orientation="latitude" pageId="1" pageNumber="116" precision="1" seconds="45.78" value="16.229383">16°13’45.78”N</geoCoordinate>
,
<geoCoordinate id="A3A15022FFC0FF90C2A5FB6BFA2EFB45" box="[1246,1432,1194,1221]" degrees="107" direction="east" minutes="50" orientation="longitude" pageId="1" pageNumber="116" precision="1" seconds="53.46" value="107.84818">107°50’53.46”E</geoCoordinate>
,
<elevation id="4DB8D1D6FFC0FF90C6ECFB0EFEFBFB69" box="[151,333,1231,1257]" metricMagnitude="2" metricUnit="m" metricValue="3.71" pageId="1" pageNumber="116" unit="m" value="371.0">
elevation
<quantity id="016D9B00FFC0FF90C77FFB0EFEFBFB69" box="[260,333,1231,1257]" metricMagnitude="2" metricUnit="m" metricValue="3.71" pageId="1" pageNumber="116" unit="m" value="371.0">371 m</quantity>
</elevation>
), Thua Thien Hue,
<collectingCountry id="BE827675FFC0FF90C459FB0EFD32FB69" box="[546,644,1231,1257]" name="Vietnam" pageId="1" pageNumber="116">Vietnam</collectingCountry>
,
<date id="B22B1025FFC0FF90C4F5FB0EFCF7FB69" box="[654,833,1230,1257]" pageId="1" pageNumber="116" value="2015-08-27">
<collectingDate id="A26FE9CDFFC0FF90C4F5FB0EFCF7FB69" box="[654,833,1230,1257]" pageId="1" pageNumber="116" value="2015-08-27">27 August 2015</collectingDate>
</date>
,
<collectorName id="6B605333FFC0FF90C530FB0EFC75FB69" box="[843,963,1231,1257]" pageId="1" pageNumber="116">D.S. Pham</collectorName>
leg.
</materialsCitation>
<materialsCitation id="76FD3CB8FFC0FF90C583FB0EFF09FA8D" ID-GBIF-Occurrence="3091104313" collectionCode="IZCAS" country="Vietnam" pageId="1" pageNumber="116" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="F4E1EAF7FFC0FF90C583FB0EFBCFFB69" bold="true" box="[1016,1145,1231,1257]" pageId="1" pageNumber="116">
<typeStatus id="192E8847FFC0FF90C583FB0EFBC2FB69" box="[1016,1140,1231,1257]" pageId="1" pageNumber="116" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="D093FD6CFFC0FF90C2FBFB0EFB7CFB69" box="[1152,1226,1231,1257]" pageId="1" pageNumber="116" type="male">1 male</specimenCount>
(
<collectionCode id="A084AE20FFC0FF90C2A3FB0EFA9DFB69" box="[1240,1323,1231,1257]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="116">IZCAS</collectionCode>
Ar41902) and
</materialsCitation>
<materialsCitation id="76FD3CB8FFC0FF90C6BCFB32FCB2FA8D" ID-GBIF-Occurrence="3091104303" box="[199,772,1267,1293]" collectingDate="2015-08-27" collectionCode="IZCAS" collectorName="D. S. Pham" country="Vietnam" elevation="371" latitude="16.229383" location="Male" longLatPrecision="1" longitude="107.84818" municipality="Bach Ma National Park" pageId="1" pageNumber="116" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="D093FD6CFFC0FF90C6BCFB32FE9EFA8D" box="[199,296,1267,1293]" pageId="1" pageNumber="116" type="female">1 female</specimenCount>
(
<collectionCode id="A084AE20FFC0FF90C74DFB32FE3FFA8D" box="[310,393,1267,1293]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="116">IZCAS</collectionCode>
Ar41903), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC0FF90C6BCFAD6FBCAFAB1" box="[199,1148,1303,1329]" pageId="1" pageNumber="116" type="etymology">
<paragraph id="C62A36E5FFC0FF90C6BCFAD6FBCAFAB1" blockId="1.[151,1437,1194,1977]" box="[199,1148,1303,1329]" pageId="1" pageNumber="116">
<emphasis id="F4E1EAF7FFC0FF90C6BCFAD6FEF8FAB1" bold="true" box="[199,334,1303,1329]" pageId="1" pageNumber="116">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="192E8847FFC0FF90C4CEFAD6FD53FAB1" box="[693,741,1303,1329]" pageId="1" pageNumber="116">type</typeStatus>
locality and is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC0FF90C6BCFAFBFA9DF9D1" pageId="1" pageNumber="116" type="diagnosis">
<paragraph id="C62A36E5FFC0FF90C6BCFAFBFA9DF9D1" blockId="1.[151,1437,1194,1977]" pageId="1" pageNumber="116">
<emphasis id="F4E1EAF7FFC0FF90C6BCFAFBFEF4FAD4" bold="true" box="[199,322,1338,1364]" pageId="1" pageNumber="116">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="01954D66FFC0FF90C436FAFAFC7BFAD5" authority="Yao, Pham &amp; Li, 2015" authorityName="Yao, Pham &amp; Li" authorityYear="2015" box="[589,973,1338,1365]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="116" phylum="Arthropoda" rank="species" species="babensis">
<emphasis id="F4E1EAF7FFC0FF90C436FAFAFD78FAD5" box="[589,718,1339,1365]" italics="true" pageId="1" pageNumber="116">B. babensis</emphasis>
<bibRefCitation id="A2044B14FFC0FF90C4A8FAFAFC7BFAD5" author="Yao, Z. &amp; Pham, D. S. &amp; Li, S." box="[723,973,1338,1365]" journalOrPublisher="Zootaxa" pageId="1" pageNumber="116" pagination="1 - 82" part="3909" refId="ref8999" refString="Yao, Z., Pham, D. S. &amp; Li, S. (2015) Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species. Zootaxa, 3909 (1), 1 - 82. https: // doi. org / 10.11646 / zootaxa. 3909.1.1" title="Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species" type="journal article" year="2015">Yao, Pham &amp; Li, 2015</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="A2044B14FFC0FF90C27DFAFAFB7AFAD5" author="Yao, Z. &amp; Pham, D. S. &amp; Li, S." box="[1030,1228,1338,1365]" journalOrPublisher="Zootaxa" pageId="1" pageNumber="116" pagination="1 - 82" part="3909" refId="ref8999" refString="Yao, Z., Pham, D. S. &amp; Li, S. (2015) Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species. Zootaxa, 3909 (1), 1 - 82. https: // doi. org / 10.11646 / zootaxa. 3909.1.1" title="Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species" type="journal article" year="2015">
Yao
<emphasis id="F4E1EAF7FFC0FF90C243FAFAFBDFFAD5" box="[1080,1129,1339,1365]" italics="true" pageId="1" pageNumber="116">et al</emphasis>
. 2015: 5
</bibRefCitation>
, figs 1–3) with the elongated bulbal apophyses (
<figureCitation id="5EAE2A60FFC0FF90C7A5FA9EFD80FAF9" box="[478,566,1375,1401]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Fig. 2D</figureCitation>
) and vulval anterior arch of a similar dark internal shade (
<figureCitation id="5EAE2A60FFC0FF90C2B2FA9EFA90FAF9" box="[1225,1318,1375,1401]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Figs 2B</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C349FA9EFAD5FAF9" box="[1330,1379,1375,1401]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="1" pageNumber="116">14B</figureCitation>
), but males can be distinguished by the presence of a strongly conical subdisto-dorsally protruding (arrow in
<figureCitation id="5EAE2A60FFC0FF90C347FA42FA25FA1D" box="[1340,1427,1411,1437]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="1" pageNumber="116">Fig. 1A</figureCitation>
) and bifid disto-prolateral spine (arrow
<quantity id="016D9B00FFC0FF90C430FA66FDC1FA41" box="[587,631,1447,1473]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="116" unit="in" value="1.0">1 in</quantity>
<figureCitation id="5EAE2A60FFC0FF90C404FA66FD6EFA41" box="[639,728,1447,1473]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="1" pageNumber="116">Figs 1C</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C49DFA66FCA0FA41" box="[742,790,1447,1473]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="1" pageNumber="116">13A</figureCitation>
) on the procursus, by the presence of a frontal apophysis (arrow in
<figureCitation id="5EAE2A60FFC0FF90C77DFA0AFEEFFA65" box="[262,345,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Fig. 2E</figureCitation>
) and a pair of small ventral apophyses (arrow in
<figureCitation id="5EAE2A60FFC0FF90C504FA0AFC62FA65" box="[895,980,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Fig. 2C</figureCitation>
) on the clypeus, and the cheliceral distal apophyses are directed laterally (arrow da in
<figureCitation id="5EAE2A60FFC0FF90C4E6FA2EFCB6F989" box="[669,768,1519,1545]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Figs. 2C</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C575FA2FFCF6F989" box="[782,832,1518,1545]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="1" pageNumber="116">15A</figureCitation>
; distal apophyses directed anteriorly in
<taxonomicName id="01954D66FFC0FF90C372FA2EFA3AF989" authorityName="Yao, Pham &amp; Li" authorityYear="2015" box="[1289,1420,1519,1545]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="116" phylum="Arthropoda" rank="species" species="babensis">
<emphasis id="F4E1EAF7FFC0FF90C372FA2EFA3AF989" box="[1289,1420,1519,1545]" italics="true" pageId="1" pageNumber="116">B. babensis</emphasis>
</taxonomicName>
). Females can be distinguished by having frontal “epigynal” pockets laterally on a sclerotized plate (arrow ep in
<figureCitation id="5EAE2A60FFC0FF90C317F9D2FF6BF9D1" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Figs 2A–B</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C693F9F6FE8CF9D1" box="[232,314,1591,1617]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="1" pageNumber="116">14A–B</figureCitation>
) and the vulval pore plates are approximately as wide as long (arrow pp in
<figureCitation id="5EAE2A60FFC0FF90C2FDF9F6FB57F9D1" box="[1158,1249,1591,1617]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Figs 2B</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C297F9F6FAA8F9D1" box="[1260,1310,1591,1617]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="1" pageNumber="116">14B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC0FF95C6BCF99BFABFFD39" lastPageId="4" lastPageNumber="119" pageId="1" pageNumber="116" type="description">
<paragraph id="C62A36E5FFC0FF95C6BCF99BFE75FE01" blockId="1.[151,1437,1194,1977]" lastBlockId="4.[151,1437,151,732]" lastPageId="4" lastPageNumber="119" pageId="1" pageNumber="116">
<emphasis id="F4E1EAF7FFC0FF90C6BCF99BFE29F9F5" bold="true" box="[199,415,1626,1653]" pageId="1" pageNumber="116">Description. Male</emphasis>
(
<emphasis id="F4E1EAF7FFC0FF90C7C9F99AFDAFF9F5" bold="true" box="[434,537,1627,1653]" pageId="1" pageNumber="116">
<typeStatus id="192E8847FFC0FF90C7C9F99AFDAFF9F5" box="[434,537,1627,1653]" pageId="1" pageNumber="116" type="holotype">holotype</typeStatus>
</emphasis>
, IZCAS Ar41901): Total length 2.01 (2.16 with clypeus), carapace 0.98 long, 0.65 wide, opisthosoma 1.18 long, 0.80 wide. Leg I: 16.89 (4.55 + 0.19 + 4.42 + 6.67 + 1.15), leg II: 11.08 (3.21 + 0.18 + 2.75 + 4.17 + 0.77), leg III: 7.40 (2.24 + 0.16 + 1.86 + 2.56 + 0.58), leg IV: 10.61 (3.14 + 0.17 + 2.69 + 3.97 + 0.64); tibia I L/d: 55. Distance PME-PME 0.17, diameter PME 0.08, distance PME-ALE 0.11, AME absent. Habitus as in
<figureCitation id="5EAE2A60FFC0FF90C74CF92AFE19F885" box="[311,431,1771,1797]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Figs 2E–F</figureCitation>
. Carapace pale, with brown lateral margins. Ocular area with indistinct dark marks and not elevated. Clypeus brown, with frontal apophysis (arrow in
<figureCitation id="5EAE2A60FFC0FF90C510F8CEFC76F8A9" box="[875,960,1807,1833]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Fig. 2E</figureCitation>
) and pair of small ventral apophyses (arrow in
<figureCitation id="5EAE2A60FFC0FF90C693F8F2FEF6F8CD" box="[232,320,1843,1869]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Fig. 2C</figureCitation>
). Chelicerae (
<figureCitation id="5EAE2A60FFC0FF90C7A7F8F2FD88F8CD" box="[476,574,1843,1869]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Figs. 2C</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C430F8F3FDCAF8CD" box="[587,636,1842,1869]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="1" pageNumber="116">15A</figureCitation>
) with pair of small proximo-lateral apophyses and pair of long, curved distal apophyses, each with sclerotized and pointed tip (distance between tips of distal apophyses: 0.38). Sternum pale, approximately as wide as long (0.50). Thoracic furrow absent. Legs pale. Opisthosoma pale, with indistinct large spots. Palp as in
<figureCitation id="5EAE2A60FFC0FF90C7E9F85EFDA6F839" box="[402,528,1951,1977]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="1" pageNumber="116">Figs 1A–B</figureCitation>
; coxa unmodified; trochanter with short ventral apophysis with small retrolateral apophysis (arrow
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<figureCitation id="5EAE2A60FFC5FF95C7EDFF56FE5BFF31" box="[406,493,151,177]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="4" pageNumber="119">Fig. 1B</figureCitation>
); femur with small proximo-dorsal apophysis (arrow
<quantity id="016D9B00FFC5FF95C237FF56FBCEFF31" box="[1100,1144,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="119" unit="in" value="2.0">2 in</quantity>
<figureCitation id="5EAE2A60FFC5FF95C2FBFF56FB60FF31" box="[1152,1238,151,177]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="4" pageNumber="119">Fig. 1B</figureCitation>
); procursus (
<figureCitation id="5EAE2A60FFC5FF95C310FF56FF68FF53" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="4" pageNumber="119">Figs 1A–D</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C690FF78FE8BFF53" box="[235,317,185,211]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13A–B</figureCitation>
) simple proximally but complex distally, strongly protruding subdisto-dorsally (arrow in
<figureCitation id="5EAE2A60FFC5FF95C34EFF78FA38FF53" box="[1333,1422,185,211]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="4" pageNumber="119">Fig. 1A</figureCitation>
), with bifid disto-prolateral spine (arrow
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<figureCitation id="5EAE2A60FFC5FF95C4EFFF1DFD59FF76" box="[660,751,220,246]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="4" pageNumber="119">Figs 1C</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C485FF1DFC87FF76" box="[766,817,220,246]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13A</figureCitation>
), large, curved subdisto-prolateral sclerite (arrow
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<figureCitation id="5EAE2A60FFC5FF95C6ECFF3EFF47FE99" box="[151,241,255,281]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="4" pageNumber="119">Figs 1C</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C685FF3EFE87FE99" box="[254,305,255,281]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13A</figureCitation>
), disto-retrolateral membranous lamella (arrow in
<figureCitation id="5EAE2A60FFC5FF95C510FF3EFC70FE99" box="[875,966,255,281]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="4" pageNumber="119">Figs 1D</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C5AFFF3EFBB5FE99" box="[980,1027,255,281]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13B</figureCitation>
) and tongue-shaped retrolateral flap (arrow f in
<figureCitation id="5EAE2A60FFC5FF95C76DFEE0FEC7FEBB" box="[278,369,289,315]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="4" pageNumber="119">Figs 1D</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C706FEE0FE18FEBB" box="[381,430,289,315]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13B</figureCitation>
); bulb (
<figureCitation id="5EAE2A60FFC5FF95C47EFEE0FDEAFEBB" box="[517,604,289,315]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="4" pageNumber="119">Fig. 2D</figureCitation>
) with hooked apophysis and simple embolus. Retrolateral trichobothria of tibia Iat 6% proximally; legs with short, vertical setae on metatarsi, without spines or curved setae; tarsus Iwith 11 distinct pseudopodomeres.
</paragraph>
<caption id="92EA666DFFC3FF93C6ECF885FB23F86C" ID-DOI="http://doi.org/10.5281/zenodo.4719926" ID-Zenodo-Dep="4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="2" pageNumber="117" startId="2.[151,250,1860,1884]" targetBox="[231,1357,179,1834]" targetPageId="2">
<paragraph id="C62A36E5FFC3FF93C6ECF885FB23F86C" blockId="2.[151,1437,1860,2028]" pageId="2" pageNumber="117">
<emphasis id="F4E1EAF7FFC3FF93C6ECF885FEA5F8DC" bold="true" box="[151,275,1860,1884]" pageId="2" pageNumber="117">FIGURE 1.</emphasis>
<taxonomicName id="01954D66FFC3FF93C763F885FE71F8DC" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[280,455,1860,1884]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="117" phylum="Arthropoda" rank="species" species="bachma" status="sp. nov.">
<emphasis id="F4E1EAF7FFC3FF93C763F885FE71F8DC" box="[280,455,1860,1884]" italics="true" pageId="2" pageNumber="117">Belisana bachma</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFC3FF93C7B7F885FDAAF8DC" bold="true" box="[460,540,1860,1884]" pageId="2" pageNumber="117">
<taxonomicNameLabel id="EFD2578CFFC3FF93C7B7F885FDAAF8DC" box="[460,540,1860,1884]" pageId="2" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D).
</paragraph>
</caption>
<caption id="92EA666DFFC2FF92C6ECF8E3FEA6F86E" ID-DOI="http://doi.org/10.5281/zenodo.4719928" ID-Zenodo-Dep="4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="3" pageNumber="118" startId="3.[151,250,1826,1850]" targetBox="[243,1345,181,1802]" targetPageId="3">
<paragraph id="C62A36E5FFC2FF92C6ECF8E3FEA6F86E" blockId="3.[151,1436,1826,2030]" pageId="3" pageNumber="118">
<emphasis id="F4E1EAF7FFC2FF92C6ECF8E3FEA2F8BA" bold="true" box="[151,276,1826,1850]" pageId="3" pageNumber="118">FIGURE 2.</emphasis>
<taxonomicName id="01954D66FFC2FF92C767F8E3FE7AF8BA" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[284,460,1826,1850]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="bachma" status="sp. nov.">
<emphasis id="F4E1EAF7FFC2FF92C767F8E3FE7AF8BA" box="[284,460,1826,1850]" italics="true" pageId="3" pageNumber="118">Belisana bachma</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFC2FF92C7AFF8E3FD90F8BA" bold="true" box="[468,550,1826,1850]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="EFD2578CFFC2FF92C7AFF8E3FD90F8BA" box="[468,550,1826,1850]" pageId="3" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyseson clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H).
</paragraph>
</caption>
<paragraph id="C62A36E5FFC5FF95C6BCFE48FD9DFDF3" blockId="4.[151,1437,151,732]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6BCFE48FEA8FE23" bold="true" box="[199,286,393,419]" pageId="4" pageNumber="119">Female</emphasis>
(IZCAS Ar41903): Similar to male, habitus as in
<figureCitation id="5EAE2A60FFC5FF95C51BFE48FC57FE23" box="[864,993,393,419]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="4" pageNumber="119">Figs 2G–H</figureCitation>
. Total length 2.19 (2.32 with clypeus), carapace 0.75 long, 0.63 wide, opisthosoma 1.16 long, 0.81 wide. Tibia I: 3.70; tibia I L/d: 46. Distance PME-PME 0.13, diameter PME 0.08, distance PME-ALE 0.09, AME absent. Sternum approximately as wide as long (0.56). Ocular area without marks. External female genitalia (
<figureCitation id="5EAE2A60FFC5FF95C489FE30FCF8FD8B" box="[754,846,497,523]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="4" pageNumber="119">Figs 2A</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C522FE30FC3CFD8B" box="[857,906,497,523]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14A</figureCitation>
) simple and flat, with pair of frontal “epigynal” pockets (arrow ep in
<figureCitation id="5EAE2A60FFC5FF95C7F3FDD5FDB0FDAE" box="[392,518,532,558]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="4" pageNumber="119">Figs 2A–B</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C469FDD5FDD1FDAE" box="[530,615,532,558]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14A–B</figureCitation>
), 0.48 apart. Vulva (
<figureCitation id="5EAE2A60FFC5FF95C535FDD5FC1DFDAE" box="[846,939,532,558]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="4" pageNumber="119">Figs 2B</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C5CCFDD5FC51FDAE" box="[951,999,532,558]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14B</figureCitation>
) with W-shaped anterior arch (median ridge strongly protruding, arrow in
<figureCitation id="5EAE2A60FFC5FF95C459FDF6FDCBFDD1" box="[546,637,567,593]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="4" pageNumber="119">Figs 2B</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C4F3FDF6FD01FDD1" box="[648,695,567,593]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14B</figureCitation>
) and pair of large, nearly trapezoidal pore plates (arrow pp in
<figureCitation id="5EAE2A60FFC5FF95C317FDF6FF0CFDF3" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; E–H. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="4" pageNumber="119">Figs 2B</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C6BDFD98FF41FDF4" box="[198,247,601,628]" captionStart="FIGURE 16" captionStartId="21.[151,250,1791,1815]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-17@21.[151,1435,181,1768]" captionTargetPageId="21" captionText="FIGURE 16. Distribution records of six new species of the genus Belisana from Southern Asia. 1. B. bachma sp. nov.; 2. B. cucphuong sp. nov.; 3. B. jaegeri sp. nov.; 4. B. kachin sp. nov.; 5. B. putao sp. nov.; 6. B. tarang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4719963" httpUri="https://zenodo.org/record/4719963/files/figure.png" pageId="4" pageNumber="119">16B</figureCitation>
), without serrated sclerites.
</paragraph>
<paragraph id="C62A36E5FFC5FF95C6BCFDBDFCD4FD16" blockId="4.[151,1437,151,732]" box="[199,866,636,663]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6BCFDBDFEF7FD16" bold="true" box="[199,321,636,662]" pageId="4" pageNumber="119">Variation:</emphasis>
Tibia Iin another male (IZCAS Ar41902): 4.47.
</paragraph>
<paragraph id="C62A36E5FFC5FF95C6BCFD5EFABFFD39" blockId="4.[151,1437,151,732]" box="[199,1289,670,697]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6BCFD5EFE33FD38" bold="true" box="[199,389,670,697]" pageId="4" pageNumber="119">Natural history.</emphasis>
The species was found in leaf litter and collected by small plot direct searching.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC5FF95C6BCFD00FC7BFD5C" box="[199,973,705,732]" pageId="4" pageNumber="119" type="distribution">
<paragraph id="C62A36E5FFC5FF95C6BCFD00FC7BFD5C" blockId="4.[151,1437,151,732]" box="[199,973,705,732]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6BCFD00FED7FD5B" bold="true" box="[199,353,705,731]" pageId="4" pageNumber="119">Distribution.</emphasis>
<collectingCountry id="BE827675FFC5FF95C71CFD00FE70FD5B" box="[359,454,705,731]" name="Vietnam" pageId="4" pageNumber="119">Vietnam</collectingCountry>
(Thua Thien Hue,
<typeStatus id="192E8847FFC5FF95C4E0FD03FD7DFD5C" box="[667,715,706,732]" pageId="4" pageNumber="119">type</typeStatus>
locality only;
<figureCitation id="5EAE2A60FFC5FF95C515FD00FC76FD5C" box="[878,960,705,732]" captionStart="FIGURE 16" captionStartId="21.[151,250,1791,1815]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-17@21.[151,1435,181,1768]" captionTargetPageId="21" captionText="FIGURE 16. Distribution records of six new species of the genus Belisana from Southern Asia. 1. B. bachma sp. nov.; 2. B. cucphuong sp. nov.; 3. B. jaegeri sp. nov.; 4. B. kachin sp. nov.; 5. B. putao sp. nov.; 6. B. tarang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4719963" httpUri="https://zenodo.org/record/4719963/files/figure.png" pageId="4" pageNumber="119">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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