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<mods:title id="BFA305B19F1D0DE68A9B92A88912602D">A new species of Onuphis (Polychaeta: Onuphidae) from Southern Portugal, with comments on the validity of O. pancerii Claparède, 1868</mods:title>
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<taxonomicName id="4C5288BB6C33A666FF037746553D1921" ID-CoL="49QKV" authority="Claparede, 1868" authorityName="Claparede" authorityYear="1868" box="[151,573,1321,1348]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="356" phylum="Annelida" rank="species" species="pancerii">
<emphasis id="B9262F2A6C33A666FF037746553D1921" bold="true" box="[151,573,1321,1348]" pageId="13" pageNumber="356">
<emphasis id="B9262F2A6C33A666FF03774656691926" bold="true" box="[151,361,1321,1347]" italics="true" pageId="13" pageNumber="356">Onuphis pancerii</emphasis>
<bibRefCitation id="EFC38EC96C33A666FEE57745553D1921" author="Claparede" box="[369,573,1322,1348]" pageId="13" pageNumber="356" refString="Claparede, E. (1868) Les annelides chetopodes du Golfe de Naples. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 19 (2), 313 - 584. http: // dx. doi. org / 10.5962 / bhl. title. 2142" type="journal article" year="1868">Claparède, 1868</bibRefCitation>
</emphasis>
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<taxonomicName id="4C5288BB6C33A666FF03771F564A19E3" ID-CoL="84R8T" authority="Pancerii" authorityName="Pancerii" box="[151,330,1392,1414]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="356" phylum="Annelida" rank="genus">
<emphasis id="B9262F2A6C33A666FF03771F564A19E3" box="[151,330,1392,1414]" italics="true" pageId="13" pageNumber="356">Onuphis Pancerii</emphasis>
</taxonomicName>
<treatmentCitation id="0AF3D5296C33A666FEF0771F553719E3" author="Claparede" box="[356,567,1392,1414]" page="438" pageId="13" pageNumber="356" year="1868">
<bibRefCitation id="EFC38EC96C33A666FEF0771F550619E3" author="Claparede" box="[356,518,1392,1414]" pageId="13" pageNumber="356" refString="Claparede, E. (1868) Les annelides chetopodes du Golfe de Naples. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 19 (2), 313 - 584. http: // dx. doi. org / 10.5962 / bhl. title. 2142" type="journal article" year="1868">Claparède 1868</bibRefCitation>
: 438
</treatmentCitation>
440, pl. 8 figs. 1, 2
<emphasis id="B9262F2A6C33A666FC95771D541319E2" box="[769,787,1394,1415]" italics="true" pageId="13" pageNumber="356">H</emphasis>
;
<treatmentCitation id="0AF3D5296C33A666FCB5771F54F719E3" author="Claparede" box="[801,1015,1392,1414]" page="202" pageId="13" pageNumber="356" year="1869">
<bibRefCitation id="EFC38EC96C33A666FCB5771F54C419E3" author="Claparede" box="[801,964,1392,1414]" pageId="13" pageNumber="356" refString="Claparede, E. (1869) Les annelides chetopodes du Golfe de Naples. Ordre II me. Annelides Sedentaires (Aud. et Edw.). Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 20 (1), 1 - 225. http: // dx. doi. org / 10.5962 / bhl. title. 2142" type="journal article" year="1869">Claparède 1869</bibRefCitation>
: 202
</treatmentCitation>
;
<treatmentCitation id="0AF3D5296C33A666FB95771F53D619E3" author="Claparede" box="[1025,1238,1392,1414]" page="387" pageId="13" pageNumber="356" year="1870">
<bibRefCitation id="EFC38EC96C33A666FB95771F53A419E3" author="Claparede" box="[1025,1188,1392,1414]" pageId="13" pageNumber="356" refString="Claparede, E. (1870) Les annelides chetopodes du Golfe de Naples. Supplement. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 20 (2), 365 - 542. http: // dx. doi. org / 10.5962 / bhl. title. 2142" type="journal article" year="1870">Claparède 1870</bibRefCitation>
: 387
</treatmentCitation>
, pl. 5 fig. 5. Not
<taxonomicName id="4C5288BB6C33A666FF5777E0567619C0" ID-CoL="84R8T" authority="Pancerii" authorityName="Pancerii" box="[195,374,1423,1445]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="356" phylum="Annelida" rank="genus">
<emphasis id="B9262F2A6C33A666FF5777E0567619C0" box="[195,374,1423,1445]" italics="true" pageId="13" pageNumber="356">Onuphis Pancerii</emphasis>
</taxonomicName>
<treatmentCitation id="0AF3D5296C33A666FE1B77E0552719C0" author="Rioja" box="[399,551,1423,1445]" page="39" pageId="13" pageNumber="356" year="1918">
<bibRefCitation id="EFC38EC96C33A666FE1B77E0550319C0" author="Rioja" box="[399,515,1423,1445]" pageId="13" pageNumber="356" refString="Rioja, E. (1918) Datos para el conocimiento de la fauna de anelidos poliquetos del Cantabrico (2 a Parte). Trabajos del Museo Nacional de Ciencias Naturales, Madrid, Serie Zoologica, 37, 1 - 99." type="journal article" year="1918">Rioja 1918</bibRefCitation>
: 39
</treatmentCitation>
43, figs. 10
<emphasis id="B9262F2A6C33A666FD3077E055CA19C0" box="[676,714,1423,1445]" italics="true" pageId="13" pageNumber="356">ah</emphasis>
(=
<taxonomicName id="4C5288BB6C33A666FD7A77E053F619C0" ID-CoL="49QJ7" authority="Audouin &amp; Milne-Edwards, 1833" authorityName="Audouin &amp; Milne-Edwards" authorityYear="1833" box="[750,1270,1423,1445]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="356" phylum="Annelida" rank="species" species="eremita">
<emphasis id="B9262F2A6C33A666FD7A77E0549619C0" box="[750,918,1423,1445]" italics="true" pageId="13" pageNumber="356">Onuphis eremita</emphasis>
<bibRefCitation id="EFC38EC96C33A666FC0977E053F619C0" author="Audouin" box="[925,1270,1423,1445]" pageId="13" pageNumber="356" refString="Audouin, J. V. &amp; Milne Edwards, H. (1833) Classification des Annelides, et description des celles qui habitent les cotes de la France (Suite). Annales des Sciences Naturelles, Paris, 28, 187 - 247." type="journal article" year="1833">Audouin &amp; Milne-Edwards, 1833</bibRefCitation>
</taxonomicName>
).
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<paragraph id="8BEDF3386C33A666FF0377BD55E41B6E" blockId="13.[151,1436,1490,2019]" pageId="13" pageNumber="356">
<emphasis id="B9262F2A6C33A666FF0377BD560B198E" bold="true" box="[151,267,1490,1515]" pageId="13" pageNumber="356">Remarks.</emphasis>
<taxonomicName id="4C5288BB6C33A666FE8577BD5591198E" authority="Claparede, 1868" authorityName="Claparede" authorityYear="1868" box="[273,657,1490,1515]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="356" phylum="Annelida" rank="species" species="pancerii">
<emphasis id="B9262F2A6C33A666FE8577BD56D3198F" box="[273,467,1490,1514]" italics="true" pageId="13" pageNumber="356">Onuphis pancerii</emphasis>
<bibRefCitation id="EFC38EC96C33A666FE4E77BD5591198E" author="Claparede" box="[474,657,1490,1515]" pageId="13" pageNumber="356" refString="Claparede, E. (1868) Les annelides chetopodes du Golfe de Naples. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 19 (2), 313 - 584. http: // dx. doi. org / 10.5962 / bhl. title. 2142" type="journal article" year="1868">Claparède, 1868</bibRefCitation>
</taxonomicName>
is normally considered as a junior synonym of
<taxonomicName id="4C5288BB6C33A666FB3577BD56921A6B" authority="Audouin &amp; Milne-Edwards, 1833" authorityName="Audouin &amp; Milne-Edwards" authorityYear="1833" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="356" phylum="Annelida" rank="species" species="eremita">
<emphasis id="B9262F2A6C33A666FB3577BD5218198F" box="[1185,1304,1490,1514]" italics="true" pageId="13" pageNumber="356">
<collectingCountry id="F345B3A86C33A666FB3577BD53B8198C" box="[1185,1208,1490,1513]" name="Iceland" pageId="13" pageNumber="356">O</collectingCountry>
. eremita
</emphasis>
<bibRefCitation id="EFC38EC96C33A666FA8A77BD56921A6B" author="Audouin" pageId="13" pageNumber="356" refString="Audouin, J. V. &amp; Milne Edwards, H. (1833) Classification des Annelides, et description des celles qui habitent les cotes de la France (Suite). Annales des Sciences Naturelles, Paris, 28, 187 - 247." type="journal article" year="1833">Audouin &amp; Milne-Edwards, 1833</bibRefCitation>
</taxonomicName>
, but taking into account the more restrictive definition of this species accepted today, it should be considered as a valid Mediterranean species.
<taxonomicName id="4C5288BB6C33A666FC80747554951A57" box="[788,917,1562,1586]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="356" phylum="Annelida" rank="species" species="pancerii">
<emphasis id="B9262F2A6C33A666FC80747554951A57" box="[788,917,1562,1586]" italics="true" pageId="13" pageNumber="356">
<collectingCountry id="F345B3A86C33A666FC807475542B1A54" box="[788,811,1562,1585]" name="Iceland" pageId="13" pageNumber="356">O</collectingCountry>
. pancerii
</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFC38EC96C33A666FBF8747552311A56" author="Claparede" box="[1132,1329,1562,1587]" pageId="13" pageNumber="356" refString="Claparede, E. (1868) Les annelides chetopodes du Golfe de Naples. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 19 (2), 313 - 584. http: // dx. doi. org / 10.5962 / bhl. title. 2142" type="journal article" year="1868">Claparède (1868)</bibRefCitation>
from the Gulf of Naples, who refers to it as being very common but normally misidentified as
<taxonomicName id="4C5288BB6C33A666FBC27450524A1A33" box="[1110,1354,1598,1622]" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="356" phylum="Annelida" rank="species" species="neapolitana">
<emphasis id="B9262F2A6C33A666FBC27450524A1A33" box="[1110,1354,1598,1622]" italics="true" pageId="13" pageNumber="356">Diopatra neapolitana</emphasis>
</taxonomicName>
. It was commonly used as bait by the Napolitan fishermen, who named it as “
<emphasis id="B9262F2A6C33A666FC77740C53C61A1F" box="[995,1222,1634,1658]" italics="true" pageId="13" pageNumber="356">esca di palo canita</emphasis>
”, whilst
<taxonomicName id="4C5288BB6C33A666FAA1740C561D1AFB" class="Polychaeta" family="Onuphidae" genus="Diopatra" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="356" phylum="Annelida" rank="species" species="neapolitana">
<emphasis id="B9262F2A6C33A666FAA1740C561D1AFB" italics="true" pageId="13" pageNumber="356">Diopatra neapolitana</emphasis>
</taxonomicName>
was known by the less elaborate name “
<emphasis id="B9262F2A6C33A666FD6374E854841AFB" box="[759,900,1670,1694]" italics="true" pageId="13" pageNumber="356">esca di palo</emphasis>
” (
<bibRefCitation id="EFC38EC96C33A666FC3574EA53961AFB" author="Claparede" box="[929,1174,1669,1694]" pageId="13" pageNumber="356" refString="Claparede, E. (1869) Les annelides chetopodes du Golfe de Naples. Ordre II me. Annelides Sedentaires (Aud. et Edw.). Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 20 (1), 1 - 225. http: // dx. doi. org / 10.5962 / bhl. title. 2142" type="journal article" year="1869">Claparède 1869: 202</bibRefCitation>
). This denotes that not only the species was very common and easily accessible for bait digging, occurring probably at shallow water, but also that fishermen could easily recognize both species with the naked eye. This could be due to differences in morphology, ethology or ecology of the two species.
</paragraph>
<paragraph id="8BEDF3386C33A666FF53757A56E91BFF" blockId="13.[151,1436,1490,2019]" pageId="13" pageNumber="356">
The original description of
<taxonomicName id="4C5288BB6C33A666FE687579557D1B4B" box="[508,637,1814,1838]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="356" phylum="Annelida" rank="species" species="pancerii">
<emphasis id="B9262F2A6C33A666FE687579557D1B4B" box="[508,637,1814,1838]" italics="true" pageId="13" pageNumber="356">
<collectingCountry id="F345B3A86C33A666FE68757955131B48" box="[508,531,1814,1837]" name="Iceland" pageId="13" pageNumber="356">O</collectingCountry>
. pancerii
</emphasis>
</taxonomicName>
does not detail some important taxonomic characters useful today, but the species can still be separated from
<taxonomicName id="4C5288BB6C33A666FDC2755555CF1B37" box="[598,719,1850,1874]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="356" phylum="Annelida" rank="species" species="eremita">
<emphasis id="B9262F2A6C33A666FDC2755555CF1B37" box="[598,719,1850,1874]" italics="true" pageId="13" pageNumber="356">
<collectingCountry id="F345B3A86C33A666FDC27555556D1B34" box="[598,621,1850,1873]" name="Iceland" pageId="13" pageNumber="356">O</collectingCountry>
. eremita
</emphasis>
</taxonomicName>
based on the presence of two anterior chaetigers with bi and tridentate pseudocompound hooks, as opposed to 3 chaetigers possessing tridentate pseudocompound hooks, respectively (see also table 1).
</paragraph>
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In the original description
<bibRefCitation id="EFC38EC96C33A666FE6E75CA54611BDB" author="Claparede" box="[506,865,1957,1982]" pageId="13" pageNumber="356" refString="Claparede, E. (1868) Les annelides chetopodes du Golfe de Naples. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 19 (2), 313 - 584. http: // dx. doi. org / 10.5962 / bhl. title. 2142" type="journal article" year="1868">
Claparède (1868: pl. 8 fig. 2
<emphasis id="B9262F2A6C33A666FCD075C854581BDB" box="[836,856,1959,1982]" italics="true" pageId="13" pageNumber="356">H</emphasis>
)
</bibRefCitation>
pictured the species as having a right maxilla III present, something that would be unique in the family. However, Claparède himself rectified that lapsus in a proceeding work, providing a corrected drawing of the maxillae (
<bibRefCitation id="EFC38EC96C30A665FC1372F853CA1CD5" author="Claparede" box="[903,1226,151,176]" pageId="14" pageNumber="357" refString="Claparede, E. (1870) Les annelides chetopodes du Golfe de Naples. Supplement. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 20 (2), 365 - 542. http: // dx. doi. org / 10.5962 / bhl. title. 2142" type="journal article" year="1870">Claparède 1870: pl. 5 fig. 5</bibRefCitation>
).
<collectingCountry id="F345B3A86C30A665FB7572F752001CD5" box="[1249,1280,152,176]" name="American Samoa" pageId="14" pageNumber="357">As</collectingCountry>
stated by the author (
<bibRefCitation id="EFC38EC96C30A665FF6572D356E91CB1" author="Claparede" box="[241,489,188,213]" pageId="14" pageNumber="357" refString="Claparede, E. (1870) Les annelides chetopodes du Golfe de Naples. Supplement. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 20 (2), 365 - 542. http: // dx. doi. org / 10.5962 / bhl. title. 2142" type="journal article" year="1870">Claparède 1870: 387</bibRefCitation>
), “
<emphasis id="B9262F2A6C30A665FD9A72D2557B1CB1" box="[526,635,188,212]" italics="true" pageId="14" pageNumber="357">Le dessin</emphasis>
[from the publication of 1868]
<emphasis id="B9262F2A6C30A665FC6D72D256721D25" italics="true" pageId="14" pageNumber="357">avait été fait à Genève daprès une préparation de Naples. Je suis frappé de labsence du paragnathe droit, absence que je navais pas remarquée à Naples. Persuadé que le paragnathe avait disparu par un accident de la préparation, je lai rétabli à tort dans le dessin du côté droit</emphasis>
.”
</paragraph>
<paragraph id="8BEDF3386C30A665FF53732356EA1E59" blockId="14.[151,1437,151,1293]" pageId="14" pageNumber="357">
After its description,
<taxonomicName id="4C5288BB6C30A665FE277323552A1D01" box="[435,554,332,356]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="357" phylum="Annelida" rank="species" species="panceri">
<emphasis id="B9262F2A6C30A665FE277323552A1D01" box="[435,554,332,356]" italics="true" pageId="14" pageNumber="357">
<collectingCountry id="F345B3A86C30A665FE27732356CA1D06" box="[435,458,332,355]" name="Iceland" pageId="14" pageNumber="357">O</collectingCountry>
. panceri
</emphasis>
</taxonomicName>
has only been referenced once, by
<bibRefCitation id="EFC38EC96C30A665FC26732353421D01" author="Rioja" box="[946,1090,332,357]" pageId="14" pageNumber="357" refString="Rioja, E. (1918) Datos para el conocimiento de la fauna de anelidos poliquetos del Cantabrico (2 a Parte). Trabajos del Museo Nacional de Ciencias Naturales, Madrid, Serie Zoologica, 37, 1 - 99." type="journal article" year="1918">Rioja (1918)</bibRefCitation>
, who identified it, within some specimens collected at Santander (
<collectingCountry id="F345B3A86C30A665FDBB730055741DED" box="[559,628,367,392]" name="Spain" pageId="14" pageNumber="357">Spain</collectingCountry>
). He compared his specimens with one specimen from the
<emphasis id="B9262F2A6C30A665FAAA731F56A41DC9" italics="true" pageId="14" pageNumber="357">Stazione Zoologica Anton Dohrn</emphasis>
, in Naples, and found both populations to be similar, but differing slightly to Claparèdes description. The description given by
<bibRefCitation id="EFC38EC96C30A665FDDA73D855E01DB5" author="Rioja" box="[590,736,439,464]" pageId="14" pageNumber="357" refString="Rioja, E. (1918) Datos para el conocimiento de la fauna de anelidos poliquetos del Cantabrico (2 a Parte). Trabajos del Museo Nacional de Ciencias Naturales, Madrid, Serie Zoologica, 37, 1 - 99." type="journal article" year="1918">Rioja (1918)</bibRefCitation>
fits quite well the description of
<taxonomicName id="4C5288BB6C30A665FBFC73D753E11DAA" box="[1128,1249,440,463]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="357" phylum="Annelida" rank="species" species="eremita">
<emphasis id="B9262F2A6C30A665FBFC73D753E11DAA" box="[1128,1249,440,463]" italics="true" pageId="14" pageNumber="357">
<collectingCountry id="F345B3A86C30A665FBFC73D7537F1DAA" box="[1128,1151,440,463]" name="Iceland" pageId="14" pageNumber="357">O</collectingCountry>
. eremita
</emphasis>
</taxonomicName>
, except that the pseudocompound hooks of chaetigers 13 are stated to be bi and tridentate, instead of only tridentate.
<bibRefCitation id="EFC38EC96C30A665FAC473B357E11E7D" author="Fauvel" pageId="14" pageNumber="357" refString="Fauvel, P. (1923) Polychetes Errantes. Faune de France, 5, 1 - 488." type="journal article" year="1923">Fauvel (1923)</bibRefCitation>
referred the population from Santander to the species
<taxonomicName id="4C5288BB6C30A665FCCF706F54D41E72" box="[859,980,512,535]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="357" phylum="Annelida" rank="species" species="eremita">
<emphasis id="B9262F2A6C30A665FCCF706F54D41E72" box="[859,980,512,535]" italics="true" pageId="14" pageNumber="357">
<collectingCountry id="F345B3A86C30A665FCCF706F54721E72" box="[859,882,512,535]" name="Iceland" pageId="14" pageNumber="357">O</collectingCountry>
. eremita
</emphasis>
</taxonomicName>
, and considered
<taxonomicName id="4C5288BB6C30A665FB0E706F521C1E72" box="[1178,1308,512,535]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="357" phylum="Annelida" rank="species" species="pancerii">
<emphasis id="B9262F2A6C30A665FB0E706F521C1E72" box="[1178,1308,512,535]" italics="true" pageId="14" pageNumber="357">
<collectingCountry id="F345B3A86C30A665FB0E706F53B11E72" box="[1178,1201,512,535]" name="Iceland" pageId="14" pageNumber="357">O</collectingCountry>
. pancerii
</emphasis>
</taxonomicName>
as being a junior synonym of
<taxonomicName id="4C5288BB6C30A665FEF8704B56E31E59" box="[364,483,548,572]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="357" phylum="Annelida" rank="species" species="eremita">
<emphasis id="B9262F2A6C30A665FEF8704B56E31E59" box="[364,483,548,572]" italics="true" pageId="14" pageNumber="357">
<collectingCountry id="F345B3A86C30A665FEF8704B56831E5E" box="[364,387,548,571]" name="Iceland" pageId="14" pageNumber="357">O</collectingCountry>
. eremita
</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BEDF3386C30A665FF53702756AD1F70" blockId="14.[151,1437,151,1293]" pageId="14" pageNumber="357">
A search for specimens that could fit the original description of
<taxonomicName id="4C5288BB6C30A665FC45702752641E05" authority="Claparede, 1868" authorityName="Claparede" authorityYear="1868" box="[977,1380,583,608]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="357" phylum="Annelida" rank="species" species="pancerii">
<emphasis id="B9262F2A6C30A665FC457027539A1E3A" box="[977,1178,584,608]" italics="true" pageId="14" pageNumber="357">Onuphis pancerii</emphasis>
<bibRefCitation id="EFC38EC96C30A665FB32702852641E05" author="Claparede" box="[1190,1380,583,608]" pageId="14" pageNumber="357" refString="Claparede, E. (1868) Les annelides chetopodes du Golfe de Naples. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 19 (2), 313 - 584. http: // dx. doi. org / 10.5962 / bhl. title. 2142" type="journal article" year="1868">Claparède, 1868</bibRefCitation>
</taxonomicName>
was performed through loan of material from the collections of
<emphasis id="B9262F2A6C30A665FCA0700353A71EE1" box="[820,1191,620,644]" italics="true" pageId="14" pageNumber="357">Stazione Zoologica Anton Dohrn</emphasis>
, in Naples. However, all studied specimens (
<specimenCount id="9D5438B16C30A665FE3D70FF55481ECD" box="[425,584,655,680]" pageId="14" pageNumber="357" type="generic">12 specimens</specimenCount>
from Salerno, Campania,
<specimenCount id="9D5438B16C30A665FC1C70FF53181ECD" box="[904,1048,655,680]" pageId="14" pageNumber="357" type="generic">3 specimens</specimenCount>
from Uccellina, Toscana, and
<specimenCount id="9D5438B16C30A665FA1A70FF560C1EA8" pageId="14" pageNumber="357" type="generic">3 specimens</specimenCount>
from the mouth of the Tiber, Lazio; no further information available) were identified as
<taxonomicName id="4C5288BB6C30A665FB7270DB525C1EA9" box="[1254,1372,692,716]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="357" phylum="Annelida" rank="species" species="eremita">
<emphasis id="B9262F2A6C30A665FB7270DB525C1EA9" box="[1254,1372,692,716]" italics="true" pageId="14" pageNumber="357">
<collectingCountry id="F345B3A86C30A665FB7270DB53FD1EAE" box="[1254,1277,692,715]" name="Iceland" pageId="14" pageNumber="357">O</collectingCountry>
. eremita
</emphasis>
</taxonomicName>
, with the exception of an
<taxonomicName id="4C5288BB6C30A665FEE970B756F51E95" box="[381,501,728,752]" class="Polychaeta" family="Onuphidae" genus="Aponuphis" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="357" phylum="Annelida" rank="genus">
<emphasis id="B9262F2A6C30A665FEE970B756F51E95" box="[381,501,728,752]" italics="true" pageId="14" pageNumber="357">Aponuphis</emphasis>
</taxonomicName>
sp. specimen from Ucellina. Nevertheless, only one of the studied lots was from Southern
<collectingCountry id="F345B3A86C30A665FE97709356361F70" box="[259,310,764,789]" name="Italy" pageId="14" pageNumber="357">Italy</collectingCountry>
(Salerno).
</paragraph>
<paragraph id="8BEDF3386C30A665FF53714F53CF1875" blockId="14.[151,1437,151,1293]" pageId="14" pageNumber="357">
A definitive statement on the validity of
<taxonomicName id="4C5288BB6C30A665FD1A714F540D1F52" box="[654,781,800,823]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="357" phylum="Annelida" rank="species" species="pancerii">
<emphasis id="B9262F2A6C30A665FD1A714F540D1F52" box="[654,781,800,823]" italics="true" pageId="14" pageNumber="357">
<collectingCountry id="F345B3A86C30A665FD1A714F55A51F52" box="[654,677,800,823]" name="Iceland" pageId="14" pageNumber="357">O</collectingCountry>
. pancerii
</emphasis>
</taxonomicName>
will depend on the discovery of new material, either fresh or within existing collections, that could fit the original description, as no
<typeStatus id="54E94D9A6C30A665FB8A712A534E1F38" box="[1054,1102,837,861]" pageId="14" pageNumber="357">type</typeStatus>
material is known to exist. Particularly important would be the study of topotype material from the Gulf of Naples, but it is possible that the species is restricted to a specific habitat at the
<typeStatus id="54E94D9A6C30A665FD3071E255D41FC0" box="[676,724,909,933]" pageId="14" pageNumber="357">type</typeStatus>
locality itself, or that it has a patchy distribution that renders a new finding difficult, as in the case of
<taxonomicName id="4C5288BB6C30A665FDC571DF54101FAD" box="[593,784,944,968]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="357" phylum="Annelida" rank="species" species="farensis" status="sp. nov.">
<emphasis id="B9262F2A6C30A665FDC571DF54101FAD" box="[593,784,944,968]" italics="true" pageId="14" pageNumber="357">Onuphis farensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21592516C30A665FC8C71C054731FAD" box="[792,883,943,968]" pageId="14" pageNumber="357" rank="species">
<emphasis id="B9262F2A6C30A665FC8C71C054731FAD" bold="true" box="[792,883,943,968]" pageId="14" pageNumber="357">sp. nov.</emphasis>
</taxonomicNameLabel>
at the Ria
<collectingCountry id="F345B3A86C30A665FC6571DF53571FA2" box="[1009,1111,944,967]" name="Taiwan" pageId="14" pageNumber="357">Formosa</collectingCountry>
. However, it is also possible that the original description by
<bibRefCitation id="EFC38EC96C30A665FE6771BB55B51F88" author="Claparede" box="[499,693,980,1005]" pageId="14" pageNumber="357" refString="Claparede, E. (1868) Les annelides chetopodes du Golfe de Naples. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 19 (2), 313 - 584. http: // dx. doi. org / 10.5962 / bhl. title. 2142" type="journal article" year="1868">Claparède (1868)</bibRefCitation>
included several errors or misinterpretations of the morphology of the worms, and that it is a junior synonym of
<taxonomicName id="4C5288BB6C30A665FD007197540A186A" box="[660,778,1016,1039]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="357" phylum="Annelida" rank="species" species="eremita">
<emphasis id="B9262F2A6C30A665FD007197540A186A" box="[660,778,1016,1039]" italics="true" pageId="14" pageNumber="357">
<collectingCountry id="F345B3A86C30A665FD00719755AB186A" box="[660,683,1016,1039]" name="Iceland" pageId="14" pageNumber="357">O</collectingCountry>
. eremita
</emphasis>
</taxonomicName>
, as it has been considered and accepted.
</paragraph>
<paragraph id="8BEDF3386C30A665FF53767356501818" blockId="14.[151,1437,151,1293]" pageId="14" pageNumber="357">
The possibility that the species was described based on specimens not fully developed can be disregarded, as Claparède described the worms as being
<quantity id="4CAA5EDD6C30A665FDFD762F55AF1832" box="[617,687,1088,1112]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="357" unit="cm" value="12.0">12 cm</quantity>
long, for
<quantity id="4CAA5EDD6C30A665FCB6765054801832" box="[802,896,1087,1112]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="14" pageNumber="357" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
wide and 130 chaetigers, very likely indicating full sized adults.
</paragraph>
<paragraph id="8BEDF3386C30A665FF5376E856861968" blockId="14.[151,1437,151,1293]" pageId="14" pageNumber="357">
Finally, the fact that in other geographic regions many species have very restricted distribution patterns (
<bibRefCitation id="EFC38EC96C30A665FF0A76C356CC18A1" author="Maekawa" box="[158,460,1196,1221]" pageId="14" pageNumber="357" refString="Maekawa, N. &amp; Hayashi, I. (1999) Taxonomic study on the genus Onuphis (Polychaeta, Onuphidae) from Japan and adjacent seas, with descriptions of six new species. Bulletin of the National Science Museum, Series A (Zoology), 25 (3), 163 - 214." type="journal article" year="1999">Maekawa &amp; Hayashi 1999</bibRefCitation>
) could explain the difficulty in finding further specimens of
<taxonomicName id="4C5288BB6C30A665FBE676C353F118A1" box="[1138,1265,1196,1220]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="357" phylum="Annelida" rank="species" species="pancerii">
<emphasis id="B9262F2A6C30A665FBE676C353F118A1" box="[1138,1265,1196,1220]" italics="true" pageId="14" pageNumber="357">
<collectingCountry id="F345B3A86C30A665FBE676C3538918A6" box="[1138,1161,1196,1219]" name="Iceland" pageId="14" pageNumber="357">O</collectingCountry>
. pancerii
</emphasis>
</taxonomicName>
. The species is herein considered as being likely valid, and is therefore included in the synoptic table of
<taxonomicName id="4C5288BB6C30A665FBEB76BF53DE188D" box="[1151,1246,1232,1256]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="357" phylum="Annelida" rank="genus">
<emphasis id="B9262F2A6C30A665FBEB76BF53DE188D" box="[1151,1246,1232,1256]" italics="true" pageId="14" pageNumber="357">Onuphis</emphasis>
</taxonomicName>
species (table 1) and in the key below.
</paragraph>
</subSubSection>
</treatment>
</document>