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<document id="39BCEFA6A20A23B4AB3BA882657B35B5" ID-CLB-Dataset="32043" ID-DOI="10.11646/zootaxa.4323.1.4" ID-GBIF-Dataset="ce6f1bf6-5cf5-4cad-9f75-759930ba4a09" ID-ISSN="1175-5326" ID-Zenodo-Dep="897327" ID-ZooBank="C21944C7-F089-4185-Bd36-7A81870D039F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505881933849" checkinUser="plazi" docAuthor="Chatzimanolis, Stylianos" docDate="2017" docId="03ED6514630FF974BB81FC52FBAEF82A" docLanguage="en" docName="zootaxa.4323.1.4.pdf" docOrigin="Zootaxa 4323 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phanolinopsis goniakos Chatzimanolis 2017, new species" docType="treatment" docVersion="7" lastPageNumber="61" masterDocId="FFD41D6C6304F978BB16FFA5FFEEFF80" masterDocTitle="And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="67" masterPageNumber="49" pageNumber="60" updateTime="1698468730623" updateUser="ExternalLinkService">
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<mods:title id="C087B3483B52698BC27A9F8A3C9F39EB">And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="C74393F970176C3DA923A80038E41A48">Chatzimanolis, Stylianos</mods:namePart>
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<mods:title id="DB326CEDFFDB3AD041D62BD771F3A357">Zootaxa</mods:title>
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<mods:part id="F6A25933FD8C9002D054213DBECAB6D7">
<mods:date id="500165636C82949C1273CFDD941C8AFA">2017</mods:date>
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<mods:number id="9D60C357FB7B7045C02B558B47010864">2017-09-20</mods:number>
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<mods:number id="74BE1B2024452257A5DB72914C9A5D6A">4323</mods:number>
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<treatment id="03ED6514630FF974BB81FC52FBAEF82A" ID-DOI="http://doi.org/10.5281/zenodo.6041895" ID-GBIF-Taxon="134313852" ID-Zenodo-Dep="6041895" LSID="urn:lsid:plazi:treatment:03ED6514630FF974BB81FC52FBAEF82A" httpUri="http://treatment.plazi.org/id/03ED6514630FF974BB81FC52FBAEF82A" lastPageId="12" lastPageNumber="61" pageId="11" pageNumber="60">
<subSubSection id="C35E8789630FF973BB81FC52FE26FBB3" pageId="11" pageNumber="60" type="nomenclature">
<paragraph id="8BFBD402630FF973BB81FC52FCFCFB92" blockId="11.[151,786,1015,1075]" box="[151,786,1015,1042]" pageId="11" pageNumber="60">
<heading id="D0B3636E630FF973BB81FC52FCFCFB92" bold="true" box="[151,786,1015,1042]" fontSize="11" level="1" pageId="11" pageNumber="60" reason="1">
<emphasis id="B9300810630FF973BB81FC52FCFCFB92" bold="true" box="[151,786,1015,1042]" pageId="11" pageNumber="60">
<taxonomicName id="4C44AF81630FF973BB81FC52FD98FB92" ID-CoL="4FTY6" authority="Chatzimanolis" authorityName="Chatzimanolis" authorityYear="2017" box="[151,630,1015,1042]" class="Insecta" family="Staphylinidae" genus="Phanolinopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="goniakos" status="sp. nov.">
<emphasis id="B9300810630FF973BB81FC52FE56FB91" bold="true" box="[151,440,1015,1041]" italics="true" pageId="11" pageNumber="60">Phanolinopsis goniakos</emphasis>
Chatzimanolis
</taxonomicName>
,
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</emphasis>
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<paragraph id="8BFBD402630FF973BB81FBBFFE26FBB3" blockId="11.[151,786,1015,1075]" box="[151,456,1050,1075]" pageId="11" pageNumber="60">
(
<figureCitation id="137FC887630FF973BB89FBBFFF00FBB3" box="[159,238,1050,1075]" captionStart="FIGURES 3 4" captionStartId="3.[151,264,1372,1395]" captionTargetBox="[340,1247,157,1354]" captionTargetId="figure@3.[340,1247,157,1354]" captionTargetPageId="3" captionText="FIGURES 3 4. Habitus photographs of species of Phanolinopsis Scheerpeltz. 3. The holotype of P. fassli (Bernhauer), total length = 14.0 mm. 4. The holotype of P. goniakos Chatzimanolis, total length = 12.1 mm." httpUri="https://zenodo.org/record/897333/files/figure.png" pageId="11" pageNumber="60">Figs. 4</figureCitation>
,
<figureCitation id="137FC887630FF973BBECFBBEFEF6FBB3" box="[250,280,1051,1075]" captionStart="FIGURES 7 12" captionStartId="5.[151,264,1243,1266]" captionTargetBox="[340,1247,193,1222]" captionTargetId="figure@5.[340,1247,193,1222]" captionTargetPageId="5" captionText="FIGURES 7 12. Heads and pronota of Phanolinopsis Scheerpeltz. 7. P. discedens (Sharp). 8. P. erythros Chatzimanolis. 9. P. fassli (Bernhauer). 10. P. goniakos Chatzimanolis. 11. P. metaksenios Chatzimanolis. 12. P. norahae Chatzimanolis. Not to scale." httpUri="https://zenodo.org/record/897337/files/figure.png" pageId="11" pageNumber="60">10</figureCitation>
,
<figureCitation id="137FC887630FF973BA33FBBFFEADFBB3" box="[293,323,1050,1075]" captionStart="FIGURES 13 18" captionStartId="6.[151,264,1356,1379]" captionTargetBox="[330,1256,375,1335]" captionTargetId="figure@6.[330,1256,375,1335]" captionTargetPageId="6" captionText="FIGURES 13 18. Lateral view of heads of Phanolinopsis Scheerpeltz indicating the differences in the temple area and specifically the distance between the eye and postmandibular ridge. White arrows point to the extent of the postmandibular ridge. 13. P. discedens (Sharp). 14. P. erythros Chatzimanolis. 15. P. fassli (Bernhauer). 16. P. goniakos Chatzimanolis. 17. P. metaksenios Chatzimanolis. 18. P. norahae Chatzimanolis. Not to scale." httpUri="https://zenodo.org/record/897339/files/figure.png" pageId="11" pageNumber="60">16</figureCitation>
,
<figureCitation id="137FC887630FF973BA58FBBFFE79FBB3" box="[334,407,1050,1075]" captionStart="FIGURES 34 36" captionStartId="12.[151,264,940,963]" captionTargetBox="[325,1260,193,904]" captionTargetId="figure@12.[323,1263,193,919]" captionTargetPageId="12" captionText="FIGURES 34 36. Aedeagus of P. goniakos Chatzimanolis. 34. Dorsal view. 35. Lateral view. 36. Detail of paramere, ventral view." httpUri="https://zenodo.org/record/897349/files/figure.png" pageId="11" pageNumber="60">3436</figureCitation>
,
<figureCitation id="137FC887630FF973BAB4FBBEFE51FBB3" box="[418,447,1051,1075]" captionStart="FIGURE 43" captionStartId="17.[151,250,940,962]" captionTargetBox="[340,1247,193,919]" captionTargetId="figure@17.[340,1247,193,919]" captionTargetPageId="17" captionText="FIGURE 43. Distribution map of P. discedens (Sharp), P. erythros Chatzimanolis and P. goniakos Chatzimanolis." httpUri="https://zenodo.org/record/897355/files/figure.png" pageId="11" pageNumber="60">43</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35E8789630FF973BB81FBC7FE65F8EF" pageId="11" pageNumber="60" type="materials_examined">
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material
</emphasis>
.
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<emphasis id="B9300810630FF973BA5CFBC7FE59FBFB" bold="true" box="[330,439,1122,1147]" pageId="11" pageNumber="60">Holotype</emphasis>
</typeStatus>
, here designated, male, with labels: “
<collectingCountry id="F3539492630FF973B841FBC6FC39FBFB" box="[855,983,1122,1147]" name="Costa Rica" pageId="11" pageNumber="60">Costa Rica</collectingCountry>
:
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, Monteverde, Estacion Biol. Monteverde,
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,
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,
<date id="FFFAF2C2630FF973B9D0FB23FCAEFB1F" box="[710,832,1158,1183]" pageId="11" pageNumber="60" value="2001-06-12">12.vi.2001</date>
, montane forest litter, R. Anderson, CR1A01 107” / “[barcode label] SM0516461”
<emphasis id="B9300810630FF973BAE1FB0FFE11FB43" bold="true" box="[503,511,1194,1219]" pageId="11" pageNumber="60">/</emphasis>
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<taxonomicName id="4C44AF81630FF973B990FB0FFBADFB43" authority="Chatzimanolis" authorityName="Chatzimanolis" box="[646,1091,1194,1219]" class="Insecta" family="Staphylinidae" genus="Phanolinopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="goniakos">
<emphasis id="B9300810630FF973B990FB0FFC7AFB43" box="[646,916,1194,1219]" italics="true" pageId="11" pageNumber="60">Phanolinopsis goniakos</emphasis>
Chatzimanolis
</taxonomicName>
, des. Chatzimanolis 2017”. In the collection of
<collectionCode id="ED554CC7630FF973BA42FB6AFE4AFB67" box="[340,420,1231,1255]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="11" pageNumber="60">SEMC</collectionCode>
.
</paragraph>
<paragraph id="8BFBD402630FF973BBD1FB57FE65F8EF" blockId="11.[151,1437,1122,2011]" pageId="11" pageNumber="60">
<emphasis id="B9300810630FF973BBD1FB57FEA8FA8B" bold="true" box="[199,326,1266,1291]" pageId="11" pageNumber="60">
<typeStatus id="54FF6AA0630FF973BBD1FB57FEAFFA8B" box="[199,321,1266,1291]" pageId="11" pageNumber="60" type="paratype">Paratypes</typeStatus>
.
</emphasis>
45: same label as
<typeStatus id="54FF6AA0630FF973B904FB57FD99FA8B" box="[530,631,1266,1291]" pageId="11" pageNumber="60" type="holotype">holotype</typeStatus>
, SM0516460 (
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); “
<collectingCountry id="F3539492630FF973B8AAFB56FBD3FA8B" box="[956,1085,1266,1291]" name="Costa Rica" pageId="11" pageNumber="60">Costa Rica</collectingCountry>
:
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, Monteverde, Biol. Station,
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,
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,
<date id="FFFAF2C2630FF973B987FAB3FCDBFAAF" box="[657,821,1302,1327]" pageId="11" pageNumber="60" value="2001-06-10" valueMax="2001-06-17" valueMin="2001-06-10">1017.vi.2001</date>
, S &amp; J. Peck, 01-09, FIT, cloud forest, CR1P01 001”, barcode label SM0459582 (
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); “
<collectingCountry id="F3539492630FF973B96CFA9EFD13FAD3" box="[634,765,1338,1363]" name="Costa Rica" pageId="11" pageNumber="60">Costa Rica</collectingCountry>
:
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, Monteverde,
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,
<date id="FFFAF2C2630FF973BF9FFA9FFB14FAD3" box="[1161,1274,1338,1363]" pageId="11" pageNumber="60" value="1989-05-15">15.v.1989</date>
, J.S. Ashe, R. Brooks, R Leschen, pitfall traps” / “Snow Entomol. Mus.
<collectingCountry id="F3539492630FF973B839FAFAFC44FAF7" box="[815,938,1374,1399]" name="Costa Rica" pageId="11" pageNumber="60">Costa Rica</collectingCountry>
Exped. #205”, barcode labels SM0079890, SM0079896, SM0079886, SM0079905, SM0079897, SM0079906, SM0079885, SM0079887, SM0079889, SM0079899, SM0079900, SM0079901, SM0079895, SM0079888, SM0079893, SM0079894, SM0079891, SM0079892, SM0079903, SM0079904, SM0079902, SM0079898 (
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,
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<collectionCode id="ED554CC7630FF973BF09FA6EFB8AFA63" box="[1055,1124,1483,1507]" pageId="11" pageNumber="60">UTCI</collectionCode>
;
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,
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); same locality and collectors,
<date id="FFFAF2C2630FF973BAB8FA4AFDF0F987" box="[430,542,1519,1543]" pageId="11" pageNumber="60" value="1989-05-10">10.v.1989</date>
, #101, barcode labels SM0079919, SM0079924, SM0079922, SM0079921, SM0079925, SM0079923, SM0079920 (
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<collectionCode id="ED554CC7630FF973B9B2F9B6FD07F9AB" box="[676,745,1555,1579]" pageId="11" pageNumber="60">UTCI</collectionCode>
;
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,
<specimenCount id="9D421F8B630FF973B823F9B6FC8DF9AA" box="[821,867,1555,1579]" pageId="11" pageNumber="60" type="female">1 ♀</specimenCount>
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); same locality and collectors,
<date id="FFFAF2C2630FF973BE31F9B6FA79F9AB" box="[1319,1431,1555,1579]" pageId="11" pageNumber="60" value="1989-05-12">12.v.1989</date>
, #129, barcode labels SM0079910, SM0079911, SM0079909 (
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); same locality and collectors,
<date id="FFFAF2C2630FF973BE23F992FA76F9CF" box="[1333,1432,1591,1615]" pageId="11" pageNumber="60" value="1989-05-09">9.v.1989</date>
, #074, barcode labels SM0079908, SM0079907 (
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,
<specimenCount id="9D421F8B630FF973B9ECF9FEFCCBF9F2" box="[762,805,1627,1651]" pageId="11" pageNumber="60" type="female">1 ♀</specimenCount>
<collectionCode id="ED554CC7630FF973B838F9FEFC91F9F3" box="[814,895,1627,1651]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="11" pageNumber="60">SEMC</collectionCode>
); same locality and collectors,
<date id="FFFAF2C2630FF973BFF6F9FEFABFF9F3" box="[1248,1361,1627,1651]" pageId="11" pageNumber="60" value="1989-05-23">23.v.1989</date>
, #412, barcode labels SM0079918, SM0079917 (
<specimenCount id="9D421F8B630FF973B993F9DAFD5BF916" box="[645,693,1663,1687]" pageId="11" pageNumber="60" type="female">2 ♀</specimenCount>
<collectionCode id="ED554CC7630FF973B9D7F9DAFCFCF917" box="[705,786,1663,1687]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="11" pageNumber="60">SEMC</collectionCode>
); same locality and collectors, Chomogo Trail,
<quantity id="4CBC79E7630FF973BE51F9DBFA76F917" box="[1351,1432,1662,1687]" metricMagnitude="3" metricUnit="m" metricValue="1.69" pageId="11" pageNumber="60" unit="m" value="1690.0">1690m</quantity>
,
<date id="FFFAF2C2630FF973BB81F906FEE6F93B" box="[151,264,1699,1723]" pageId="11" pageNumber="60" value="1989-05-23">23.v.1989</date>
, #410, barcode labels SM0079915, SM0079912, SM0079914, SM0079913 (
<specimenCount id="9D421F8B630FF973BF94F906FB5CF93A" box="[1154,1202,1699,1723]" pageId="11" pageNumber="60" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D421F8B630FF973BFD4F906FB1EF93A" box="[1218,1264,1699,1723]" pageId="11" pageNumber="60" type="female">2 ♀</specimenCount>
<collectionCode id="ED554CC7630FF973BFECF906FAA5F93B" box="[1274,1355,1699,1723]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="11" pageNumber="60">SEMC</collectionCode>
); same locality,
<quantity id="4CBC79E7630FF973BBEFF963FEA7F95F" box="[249,329,1734,1759]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="11" pageNumber="60" unit="m" value="1520.0">1520m</quantity>
,
<date id="FFFAF2C2630FF973BA41F963FE21F95F" box="[343,463,1734,1759]" pageId="11" pageNumber="60" value="1986-04-30">30.iv.1986</date>
, J.S. Ashe, leaf litter, barcode label SM0079926 (
<specimenCount id="9D421F8B630FF973BF18F962FBD6F95E" box="[1038,1080,1735,1759]" pageId="11" pageNumber="60" type="male">1 ♂</specimenCount>
<collectionCode id="ED554CC7630FF973BF57F962FB7CF95F" box="[1089,1170,1735,1759]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="11" pageNumber="60">SEMC</collectionCode>
); same locality,
<quantity id="4CBC79E7630FF973BE5AF963FA76F95F" box="[1356,1432,1734,1759]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="11" pageNumber="60" unit="ft" value="5000.0">5000ft</quantity>
,
<date id="FFFAF2C2630FF973BB81F94EFEDCF883" box="[151,306,1771,1795]" pageId="11" pageNumber="60" value="1979-05-23" valueMax="1979-05-24" valueMin="1979-05-23">2324.v.1979</date>
, J.M. &amp; B.A. Campbell (
<specimenCount id="9D421F8B630FF973B945F94EFD93F882" box="[595,637,1771,1795]" pageId="11" pageNumber="60" type="male">1 ♂</specimenCount>
<collectionCode id="ED554CC7630FF973B993F94EFD2DF883" box="[645,707,1771,1795]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="11" pageNumber="60">CNC</collectionCode>
); “
<collectingCountry id="F3539492630FF973B9F4F94EFC8AF883" box="[738,868,1770,1795]" name="Costa Rica" pageId="11" pageNumber="60">Costa Rica</collectingCountry>
:
<collectingRegion id="49801AE0630FF973B879F94EFC1DF883" box="[879,1011,1771,1795]" country="Costa Rica" name="Guanacaste" pageId="11" pageNumber="60">Guanacaste</collectingRegion>
,
<collectingRegion id="49801AE0630FF973BF16F94EFB67F883" box="[1024,1161,1770,1795]" country="Saint Helena, Ascension and Tristan da Cunha" name="Saint Helena" pageId="11" pageNumber="60">
<collectingCountry id="F3539492630FF973BF16F94EFB67F883" box="[1024,1161,1770,1795]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="11" pageNumber="60">Santa Elena</collectingCountry>
</collectingRegion>
,
<collectingRegion id="49801AE0630FF973BF83F94EFAF5F883" box="[1173,1307,1770,1795]" country="Saint Helena, Ascension and Tristan da Cunha" name="Saint Helena" pageId="11" pageNumber="60">
<collectingCountry id="F3539492630FF973BF83F94EFAF5F883" box="[1173,1307,1770,1795]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="11" pageNumber="60">Santa Elena</collectingCountry>
</collectingRegion>
Cloud For. Reserve,
<quantity id="4CBC79E7630FF973BA15F8ABFEBDF8A7" box="[259,339,1806,1831]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="11" pageNumber="60" unit="m" value="1650.0">1650m</quantity>
,
<geoCoordinate id="EE70B2C5630FF973BA75F8AAFE1EF8A7" box="[355,496,1807,1831]" direction="north" orientation="latitude" pageId="11" pageNumber="60" precision="15" value="10.344999">10°2042N</geoCoordinate>
<geoCoordinate id="EE70B2C5630FF973BAEAF8ABFD61F8A7" box="[508,655,1806,1831]" direction="west" orientation="longitude" pageId="11" pageNumber="60" precision="15" value="-84.79806">84°4753W</geoCoordinate>
,
<date id="FFFAF2C2630FF973B989F8ABFCF6F8A7" box="[671,792,1806,1831]" pageId="11" pageNumber="60" value="2001-06-11">11.vi.2001</date>
, cloud for. litter, R. Anderson, CR1A01 104”, barcode label SM0516591 (
<specimenCount id="9D421F8B630FF973BA60F896FE4FF8CA" box="[374,417,1843,1867]" pageId="11" pageNumber="60" type="male">1 ♂</specimenCount>
<collectionCode id="ED554CC7630FF973BABCF896FE15F8CB" box="[426,507,1843,1867]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="11" pageNumber="60">SEMC</collectionCode>
). All
<typeStatus id="54FF6AA0630FF973B92BF896FD47F8CB" box="[573,681,1843,1867]" pageId="11" pageNumber="60" type="paratype">paratypes</typeStatus>
with label “
<typeStatus id="54FF6AA0630FF973B821F896FC72F8CB" box="[823,924,1843,1867]" pageId="11" pageNumber="60" type="paratype">Paratype</typeStatus>
<taxonomicName id="4C44AF81630FF973B8B3F897FA8CF8CB" authority="Chatzimanolis" authorityName="Chatzimanolis" box="[933,1378,1842,1867]" class="Insecta" family="Staphylinidae" genus="Phanolinopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="goniakos">
<emphasis id="B9300810630FF973B8B3F897FB5AF8CB" box="[933,1204,1842,1867]" italics="true" pageId="11" pageNumber="60">Phanolinopsis goniakos</emphasis>
Chatzimanolis
</taxonomicName>
, des. Chatzimanolis 2017”.
</paragraph>
</subSubSection>
<subSubSection id="C35E8789630FF973BBD1F8DFFE04F85B" pageId="11" pageNumber="60" type="diagnosis">
<paragraph id="8BFBD402630FF973BBD1F8DFFE04F85B" blockId="11.[151,1437,1122,2011]" pageId="11" pageNumber="60">
<emphasis id="B9300810630FF973BBD1F8DFFEAFF813" bold="true" box="[199,321,1914,1939]" pageId="11" pageNumber="60">Diagnosis.</emphasis>
This species can be easily distinguished from all other taxa of
<taxonomicName id="4C44AF81630FF973BF42F8DFFB1AF813" box="[1108,1268,1914,1939]" class="Insecta" family="Staphylinidae" genus="Phanolinopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="B9300810630FF973BF42F8DFFB1AF813" box="[1108,1268,1914,1939]" italics="true" pageId="11" pageNumber="60">Phanolinopsis</emphasis>
</taxonomicName>
based on the morphology of the head. In
<taxonomicName id="4C44AF81630FF973BAC7F83AFDBCF837" box="[465,594,1950,1975]" class="Insecta" family="Staphylinidae" genus="Phanolinopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="goniakos">
<emphasis id="B9300810630FF973BAC7F83AFDBCF837" box="[465,594,1950,1975]" italics="true" pageId="11" pageNumber="60">P. goniakos</emphasis>
</taxonomicName>
the posterolateral corners of the head are pointed (
<figureCitation id="137FC887630FF973BF99F83BFB30F837" box="[1167,1246,1950,1975]" captionStart="FIGURES 3 4" captionStartId="3.[151,264,1372,1395]" captionTargetBox="[340,1247,157,1354]" captionTargetId="figure@3.[340,1247,157,1354]" captionTargetPageId="3" captionText="FIGURES 3 4. Habitus photographs of species of Phanolinopsis Scheerpeltz. 3. The holotype of P. fassli (Bernhauer), total length = 14.0 mm. 4. The holotype of P. goniakos Chatzimanolis, total length = 12.1 mm." httpUri="https://zenodo.org/record/897333/files/figure.png" pageId="11" pageNumber="60">Figs. 4</figureCitation>
,
<figureCitation id="137FC887630FF973BFFDF83AFAE7F837" box="[1259,1289,1951,1975]" captionStart="FIGURES 7 12" captionStartId="5.[151,264,1243,1266]" captionTargetBox="[340,1247,193,1222]" captionTargetId="figure@5.[340,1247,193,1222]" captionTargetPageId="5" captionText="FIGURES 7 12. Heads and pronota of Phanolinopsis Scheerpeltz. 7. P. discedens (Sharp). 8. P. erythros Chatzimanolis. 9. P. fassli (Bernhauer). 10. P. goniakos Chatzimanolis. 11. P. metaksenios Chatzimanolis. 12. P. norahae Chatzimanolis. Not to scale." httpUri="https://zenodo.org/record/897337/files/figure.png" pageId="11" pageNumber="60">10</figureCitation>
,
<figureCitation id="137FC887630FF973BE00F83BFADDF837" box="[1302,1331,1950,1975]" captionStart="FIGURES 13 18" captionStartId="6.[151,264,1356,1379]" captionTargetBox="[330,1256,375,1335]" captionTargetId="figure@6.[330,1256,375,1335]" captionTargetPageId="6" captionText="FIGURES 13 18. Lateral view of heads of Phanolinopsis Scheerpeltz indicating the differences in the temple area and specifically the distance between the eye and postmandibular ridge. White arrows point to the extent of the postmandibular ridge. 13. P. discedens (Sharp). 14. P. erythros Chatzimanolis. 15. P. fassli (Bernhauer). 16. P. goniakos Chatzimanolis. 17. P. metaksenios Chatzimanolis. 18. P. norahae Chatzimanolis. Not to scale." httpUri="https://zenodo.org/record/897339/files/figure.png" pageId="11" pageNumber="60">16</figureCitation>
) more so than any other known species.
</paragraph>
</subSubSection>
<caption id="DF3B848A6308F974BB81FC09FF20FC61" httpUri="https://zenodo.org/record/897349/files/figure.png" pageId="12" pageNumber="61" targetBox="[325,1260,193,904]" targetPageId="12">
<paragraph id="8BFBD4026308F974BB81FC09FF20FC61" blockId="12.[151,1436,940,993]" pageId="12" pageNumber="61">
<emphasis id="B93008106308F974BB81FC09FEBBFC41" bold="true" box="[151,341,940,963]" pageId="12" pageNumber="61">FIGURES 3436.</emphasis>
Aedeagus of
<taxonomicName id="4C44AF816308F974BAF5FC08FD17FC42" authority="Chatzimanolis." authorityName="Chatzimanolis." box="[483,761,940,962]" class="Insecta" family="Staphylinidae" genus="Phanolinopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="goniakos">
<emphasis id="B93008106308F974BAF5FC08FDB7FC42" box="[483,601,940,962]" italics="true" pageId="12" pageNumber="61">P. goniakos</emphasis>
Chatzimanolis.
</taxonomicName>
34. Dorsal view. 35. Lateral view. 36. Detail of paramere, ventral view.
</paragraph>
</caption>
<subSubSection id="C35E87896308F974BBD1FBA8FC91F975" pageId="12" pageNumber="61" type="description">
<paragraph id="8BFBD4026308F974BBD1FBA8FC91F975" blockId="12.[151,1437,1037,1962]" pageId="12" pageNumber="61">
<emphasis id="B93008106308F974BBD1FBA8FEB7FBA6" bold="true" box="[199,345,1037,1062]" pageId="12" pageNumber="61">Description.</emphasis>
Body length 10.513.0mm. Head and pronotum metallic brown, sometimes with green overtones; elytra dark metallic brown-green. Mouthparts, mesoscutellum, ventral surface of thorax and legs brown. Antennomeres 16 brown; antennomeres 711 dark yellow. Abdominal terga and sterna brown; posterior 1/5 of segment VII and segment VIII orange. Head transverse, width: length ratio = 1.45. Epicranium with transverse and polygon-shaped microsculpture and sparse micropunctures; with large to medium-sized punctures around margin of head (becoming more numerous near posterior margin); with 34 other large puncture near center but with no other punctures on epicranium. Eyes medium-sized, length of eyes / length of head ratio = 0.48, distance between eyes as wide as 2.07 times length of eye. Area between postmandibular ridge and eye (lateral side of head) wide, with polygon-shaped microsculpture; posterolateral corner of head pointed. Antennomeres 19, 11 longer than wide; antennomere 10 subquadrate. Neck with micropunctures, microsculpture, and with sparse small punctures. Pronotum subquadrate, width: length ratio = 0.98; surface of pronotum uniformly covered with sparse micropunctures and dense polygon-shaped microsculpture; appearing matte due to microsculpture. Pronotum with few large punctures around margin; disc of pronotum with four large punctures at center, each one delimiting corner of square. Elytra with medium-sized punctures (about 89 punctures / elytron width); distance between punctures 0.51 width of puncture. Elytra appearing shiny; with sparse longitudinal microsculpture. Abdominal terga sparsely punctate with small to medium-sized punctures. Male secondary sexual structures with sternum VIII having shallow emargination medially; sternum IX with deep U-shaped emargination medially. Female without obvious sexual structures. Aedeagus as in
<figureCitation id="137FC8876308F974B983F9D4FCCCF90A" box="[661,802,1649,1674]" captionStart="FIGURES 34 36" captionStartId="12.[151,264,940,963]" captionTargetBox="[325,1260,193,904]" captionTargetId="figure@12.[323,1263,193,919]" captionTargetPageId="12" captionText="FIGURES 34 36. Aedeagus of P. goniakos Chatzimanolis. 34. Dorsal view. 35. Lateral view. 36. Detail of paramere, ventral view." httpUri="https://zenodo.org/record/897349/files/figure.png" pageId="12" pageNumber="61">Figs. 3436</figureCitation>
; in dorsal view paramere converging to rounded tip; paramere about as long as and narrower than median lobe; in lateral view paramere slightly concave apically; paramere with peg setae as in
<figureCitation id="137FC8876308F974BAE1F91CFDA2F952" box="[503,588,1721,1746]" captionStart="FIGURES 34 36" captionStartId="12.[151,264,940,963]" captionTargetBox="[325,1260,193,904]" captionTargetId="figure@12.[323,1263,193,919]" captionTargetPageId="12" captionText="FIGURES 34 36. Aedeagus of P. goniakos Chatzimanolis. 34. Dorsal view. 35. Lateral view. 36. Detail of paramere, ventral view." httpUri="https://zenodo.org/record/897349/files/figure.png" pageId="12" pageNumber="61">Fig. 36</figureCitation>
. Median lobe in dorsal view wide, converging to rounded apex, with two broad teeth apically; in lateral view becoming narrower near apex.
</paragraph>
</subSubSection>
<subSubSection id="C35E87896308F974BBD1F8A4FE79F8E2" pageId="12" pageNumber="61" type="distribution">
<paragraph id="8BFBD4026308F974BBD1F8A4FB7FF89A" blockId="12.[151,1437,1037,1962]" box="[199,1169,1793,1818]" pageId="12" pageNumber="61">
<emphasis id="B93008106308F974BBD1F8A4FE8EF89A" bold="true" box="[199,352,1793,1818]" pageId="12" pageNumber="61">Distribution.</emphasis>
Known from the provinces of
<collectingRegion id="49801AE06308F974B9AFF8A7FCD4F89A" box="[697,826,1794,1818]" country="Costa Rica" name="Guanacaste" pageId="12" pageNumber="61">Guanacaste</collectingRegion>
and
<collectingRegion id="49801AE06308F974B867F8A7FC02F899" box="[881,1004,1794,1817]" country="Costa Rica" name="Puntarenas" pageId="12" pageNumber="61">Puntarenas</collectingRegion>
in
<collectingCountry id="F35394926308F974BF06F8A7FB63F89A" box="[1040,1165,1793,1818]" name="Costa Rica" pageId="12" pageNumber="61">Costa Rica</collectingCountry>
.
</paragraph>
<paragraph id="8BFBD4026308F974BBD1F880FE79F8E2" blockId="12.[151,1437,1037,1962]" pageId="12" pageNumber="61">
<emphasis id="B93008106308F974BBD1F880FEC4F8BE" bold="true" box="[199,298,1829,1854]" pageId="12" pageNumber="61">Habitat.</emphasis>
Collected with pitfall traps, flight intercept traps and in leaf litter in cloud forests at elevations between
<quantity id="4CBC79E76308F974BBEDF8ECFE7CF8E2" box="[251,402,1865,1890]" metricMagnitude="3" metricUnit="m" metricValue="1.605" metricValueMax="1.69" metricValueMin="1.52" pageId="12" pageNumber="61" unit="m" value="1605.0" valueMax="1690.0" valueMin="1520.0">15201690m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C35E87896308F974BBD1F8C8FBAEF82A" pageId="12" pageNumber="61" type="etymology">
<paragraph id="8BFBD4026308F974BBD1F8C8FBAEF82A" blockId="12.[151,1437,1037,1962]" pageId="12" pageNumber="61">
<emphasis id="B93008106308F974BBD1F8C8FEA0F806" bold="true" box="[199,334,1901,1926]" pageId="12" pageNumber="61">Etymology.</emphasis>
The specific epithet is derived from the Greek word γωνία (angle) and refers to the sharp angle of the posterolateral corners of the head. The epithet is treated as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>