268 lines
33 KiB
XML
268 lines
33 KiB
XML
<document id="12BC6071B2E97AB57B13532926F1C9EC" ID-DOI="10.11646/zootaxa.4034.1.11" ID-GBIF-Dataset="6a397dbc-ec50-4694-b08a-a1590b8812e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="243749" ID-ZooBank="61D0F8E1-12BB-42B3-9768-18FE47901A3E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461299537489" checkinUser="plazi" docAuthor="Ćurčić, Srećko, Ilić, Nastas, Vesović, Nikola & Antić, Dragan" docDate="2015" docId="03EA3245FF86FFDBF09D7DFAFCADFE57" docLanguage="en" docName="zt04034p200.pdf" docOrigin="Zootaxa 4034 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tapinopterus (Tapinopterus) cognatus subsp. pancici Curcic and Ilic, ssp. n." docType="treatment" docVersion="8" lastPageNumber="200" masterDocId="FFD34A3DFF84FFD8F00A7E22FFC4FFB0" masterDocTitle="Two new high-altitude subspecies of Tapinopterus (Tapinopterus) cognatus (Dejean, 1831) (Coleoptera: Carabidae: Pterostichinae) from Serbia" masterLastPageNumber="200" masterPageNumber="197" pageNumber="199" updateTime="1698617118831" updateUser="plazi">
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<mods:titleInfo id="75D2C8BC36207EE947BF1C1756240300">
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<mods:title id="D870AA1B6DC20006D7EC435CF99B2173">Two new high-altitude subspecies of Tapinopterus (Tapinopterus) cognatus (Dejean, 1831) (Coleoptera: Carabidae: Pterostichinae) from Serbia</mods:title>
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<mods:name id="89FB9DE7305B9EE2FC9D9BFC10747E05" type="personal">
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<mods:role id="AD62F8C0012B60E0D28C21559B528982">
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<mods:namePart id="68BDF9E42D54BBC7B07087EE159BC8F3">Ćurčić, Srećko</mods:namePart>
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</mods:name>
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<mods:namePart id="454B194F096F555C389419DDF95B6035">Ilić, Nastas</mods:namePart>
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</mods:name>
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<mods:name id="C4539BF7AD163114810AD05BF620EA0B" type="personal">
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</mods:role>
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<mods:namePart id="28761616CDD79F025DF5C801E90186A3">Vesović, Nikola</mods:namePart>
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</mods:name>
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<mods:name id="F630894B36D8794534DA121C985BE142" type="personal">
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<mods:roleTerm id="9E35F7EC9E9124D34D32AC23459035F4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="13A4341F55F0C683D4C6A626F24651EC">Antić, Dragan</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="F32900BF4178AAC960D08CA3B2551C39">text</mods:typeOfResource>
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<mods:title id="63D8427C9745BE2D9004F668A5566BC3">Zootaxa</mods:title>
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<mods:part id="8DFDD42E337834A6127F9B0ACB3D1202">
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<mods:date id="E2F2A099DAB7853604147FDF52246765">2015</mods:date>
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<mods:detail id="6E670A8150FF997AE83C512F0BF0F0C9" type="volume">
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<mods:number id="F00413CBFF088BF1F1DCC8FBCAA6C35C">4034</mods:number>
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</mods:detail>
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<mods:detail id="A2D47EEF958D82D3EE9E7A99676B58D8" type="issue">
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<mods:number id="B2F823F47083AA018541BA96E6EDB0AE">1</mods:number>
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<mods:classification id="E929538597EF39DF0BCFAFB9608C7CB7">journal article</mods:classification>
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<mods:identifier id="3F2B5432E27226F6BDBBEE6067842ABE" type="DOI">10.11646/zootaxa.4034.1.11</mods:identifier>
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<mods:identifier id="770DE77B23A73216AF49BA363C2747C5" type="Zenodo-Dep">243749</mods:identifier>
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<treatment id="03EA3245FF86FFDBF09D7DFAFCADFE57" ID-DOI="http://doi.org/10.5281/zenodo.6096544" ID-GBIF-Taxon="119643204" ID-Zenodo-Dep="6096544" LSID="urn:lsid:plazi:treatment:03EA3245FF86FFDBF09D7DFAFCADFE57" httpUri="http://treatment.plazi.org/id/03EA3245FF86FFDBF09D7DFAFCADFE57" lastPageId="3" lastPageNumber="200" pageId="2" pageNumber="199">
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<subSubSection id="C359D0D8FF86FFDAF09D7DFAFBBCFC42" box="[151,1144,984,1010]" pageId="2" pageNumber="199" type="nomenclature">
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<paragraph id="8BFC8353FF86FFDAF09D7DFAFBBCFC42" blockId="2.[151,1144,984,1010]" box="[151,1144,984,1010]" pageId="2" pageNumber="199">
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<heading id="D0B4343FFF86FFDAF09D7DFAFBBCFC42" bold="true" box="[151,1144,984,1010]" fontSize="11" level="1" pageId="2" pageNumber="199" reason="1">
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<emphasis id="B9375F41FF86FFDAF09D7DFAFBBCFC42" bold="true" box="[151,1144,984,1010]" pageId="2" pageNumber="199">
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<taxonomicName id="4C43F8D0FF86FFDAF09D7DFAFC42FC42" ID-CoL="8MF84" authority="Curcic and Ilic" authorityName="Curcic and Ilic" box="[151,902,984,1010]" class="Insecta" family="Carabidae" genus="Tapinopterus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="subSpecies" species="cognatus" status="subsp. nov." subGenus="Tapinopterus" subSpecies="pancici">
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||
<emphasis id="B9375F41FF86FFDAF09D7DFAFEF1FC42" bold="true" box="[151,309,984,1010]" italics="true" pageId="2" pageNumber="199">Tapinopterus</emphasis>
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||
(
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||
<emphasis id="B9375F41FF86FFDAF14C7DFAFE20FC42" bold="true" box="[326,484,984,1010]" italics="true" pageId="2" pageNumber="199">Tapinopterus</emphasis>
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)
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<emphasis id="B9375F41FF86FFDAF1FC7DFAFD06FC42" bold="true" box="[502,706,984,1010]" italics="true" pageId="2" pageNumber="199">cognatus pancici</emphasis>
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Ćurčić and Ilić
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||
</taxonomicName>
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||
,
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<taxonomicNameLabel id="A204E23AFF86FFDAF3997DFAFC25FC42" box="[915,993,984,1010]" pageId="2" pageNumber="199" rank="subSpecies">ssp. n.</taxonomicNameLabel>
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(
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||
<figureCitation id="13789FD6FF86FFDAF3FA7DFAFBB4FC42" box="[1008,1136,984,1010]" captionStart="FIGURES 1 – 12" captionStartId="1.[151,258,1840,1862]" captionTargetBox="[172,1432,703,1813]" captionTargetId="figure@1.[151,1436,684,1818]" captionTargetPageId="1" captionText="FIGURES 1 – 12. Tapinopterus (Tapinopterus) cognatus vlasinensis ssp. n. from the village of Vlasina Rid, Vlasina Lake Plateau, Mt. Čemernik, 1,200 m a. s. l., Surdulica, SE Serbia (1 – 6) and T. (T.) cognatus pancici ssp. n. from by the road Jelak Reserve - village of Brzeće, Mt. Kopaonik, 1,100 m a. s. l., Jošanička Banja, C Serbia (7 – 12). 1, 7 — holotype male, habitus (dorsal view); 2, 8 — holotype male, aedeagus (dorsal view); 3, 9 — holotype male, aedeagus (lateral view); 4, 10 — holotype male, right paramere (external lateral view); 5, 11 — holotype male, abdominal sternite IX (urite); 6, 12 — paratype female, genitalia (dorsal view). Scales = 5 mm (Figs. 1 and 7), 1 mm (Figs. 2, 3, 5, 6, 8, 9, 11, and 12), and 0.5 mm (Figs. 4 and 10)." httpUri="https://zenodo.org/record/243750/files/figure.png" pageId="2" pageNumber="199">Figs. 7–12</figureCitation>
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)
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C359D0D8FF86FFDAF09D7A3FFD21FA3F" pageId="2" pageNumber="199" type="materials_examined">
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<paragraph id="8BFC8353FF86FFDAF09D7A3FFBFCFBC9" blockId="2.[151,1436,1053,2012]" pageId="2" pageNumber="199">
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||
<emphasis id="B9375F41FF86FFDAF09D7A3FFE84FB84" bold="true" box="[151,320,1053,1076]" pageId="2" pageNumber="199">
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||
<typeStatus id="54F83DF1FF86FFDAF09D7A3FFF0BFB84" box="[151,207,1053,1076]" pageId="2" pageNumber="199">Type</typeStatus>
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||
localities.
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||
</emphasis>
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||
By the road Jelak Reserve - village of Brzeće, Mt. Kopaonik,
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||
<quantity id="4CBB2EB6FF86FFDAF3D47A3FFBF2FB84" box="[990,1078,1053,1076]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="199" unit="m" value="1100.0">1,100 m</quantity>
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||
a.s.l., Jošanička Banja, C
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||
<collectingCountry id="F354C3C3FF86FFDAF55A7A3FFA5CFB84" box="[1360,1432,1053,1076]" name="Serbia" pageId="2" pageNumber="199">Serbia</collectingCountry>
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||
; village of Brzeće, Mt. Kopaonik,
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||
<quantity id="4CBB2EB6FF86FFDAF20C7A62FD9BFBE7" box="[518,607,1088,1111]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="199" unit="m" value="1100.0">1,100 m</quantity>
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||
a.s.l., Jošanička Banja, C
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||
<collectingCountry id="F354C3C3FF86FFDAF38A7A62FC03FBE7" box="[896,967,1088,1111]" name="Serbia" pageId="2" pageNumber="199">Serbia</collectingCountry>
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||
; village of Đerekari, Mt. Kopaonik,
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||
<quantity id="4CBB2EB6FF86FFDAF55C7A62FA58FBE7" box="[1366,1436,1088,1111]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="199" unit="m" value="800.0">800 m</quantity>
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||
a.s.l., Brus, C
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||
<collectingCountry id="F354C3C3FF86FFDAF1247A40FEB1FBC9" box="[302,373,1122,1145]" name="Serbia" pageId="2" pageNumber="199">Serbia</collectingCountry>
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||
; village of Razbojna, Mt. Kopaonik,
|
||
<quantity id="4CBB2EB6FF86FFDAF30B7A40FC80FBC9" box="[769,836,1122,1145]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="199" unit="m" value="600.0">600 m</quantity>
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||
a.s.l., Blace, C
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||
<collectingCountry id="F354C3C3FF86FFDAF3E67A40FBF7FBC9" box="[1004,1075,1122,1145]" name="Serbia" pageId="2" pageNumber="199">Serbia</collectingCountry>
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||
.
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||
</paragraph>
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||
<paragraph id="8BFC8353FF86FFDAF0CD7AA7FCA9FB51" blockId="2.[151,1436,1053,2012]" pageId="2" pageNumber="199">
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<emphasis id="B9375F41FF86FFDAF0CD7AA7FEA9FB2C" bold="true" box="[199,365,1157,1180]" pageId="2" pageNumber="199">
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||
<typeStatus id="54F83DF1FF86FFDAF0CD7AA7FF3BFB2C" box="[199,255,1157,1180]" pageId="2" pageNumber="199">Type</typeStatus>
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||
material.
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||
</emphasis>
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||
<typeStatus id="54F83DF1FF86FFDAF17E7AA7FE13FB2C" box="[372,471,1157,1180]" pageId="2" pageNumber="199" type="holotype">Holotype</typeStatus>
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||
♂, by the road Jelak Reserve—village of Brzeće, Mt. Kopaonik,
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||
<quantity id="4CBB2EB6FF86FFDAF49C7AA7FB2AFB2C" box="[1174,1262,1157,1180]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="199" unit="m" value="1100.0">1,100 m</quantity>
|
||
a.s.l., Jošanička Banja, C
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||
<collectingCountry id="F354C3C3FF86FFDAF10C7A8AFE89FB0F" box="[262,333,1192,1215]" name="Serbia" pageId="2" pageNumber="199">Serbia</collectingCountry>
|
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,
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||
<date id="FFFDA593FF86FFDAF1577A8AFDF2FB0F" box="[349,566,1192,1215]" pageId="2" pageNumber="199" value="2014-06-28" valueMax="2014-08-09" valueMin="2014-06-28">28.VI–09.VIII.2014</date>
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||
, from pitfall traps, leg. N. Ilić (p)
|
||
<typeStatus id="54F83DF1FF86FFDAF3C77A8AFB9EFB0F" box="[973,1114,1192,1215]" pageId="2" pageNumber="199" type="holotype">HOLOTYPE</typeStatus>
|
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<taxonomicName id="4C43F8D0FF86FFDAF46C7A8AFE88FB51" class="Insecta" family="Carabidae" genus="Tapinopterus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="subSpecies" species="cognatus" status="subsp. nov." subGenus="Tapinopterus" subSpecies="pancici">
|
||
<emphasis id="B9375F41FF86FFDAF46C7A8AFB35FB0F" box="[1126,1265,1192,1215]" italics="true" pageId="2" pageNumber="199">Tapinopterus</emphasis>
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(
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<emphasis id="B9375F41FF86FFDAF50D7A8AFA56FB0F" box="[1287,1426,1192,1215]" italics="true" pageId="2" pageNumber="199">Tapinopterus</emphasis>
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||
)
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||
<emphasis id="B9375F41FF86FFDAF09D7AE8FE88FB51" box="[151,332,1226,1249]" italics="true" pageId="2" pageNumber="199">cognatus pancici</emphasis>
|
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</taxonomicName>
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<taxonomicNameLabel id="A204E23AFF86FFDAF1597AE8FE5DFB51" box="[339,409,1226,1249]" pageId="2" pageNumber="199" rank="subSpecies">
|
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<emphasis id="B9375F41FF86FFDAF1597AE8FE5DFB51" bold="true" box="[339,409,1226,1249]" pageId="2" pageNumber="199">ssp. n.</emphasis>
|
||
</taxonomicNameLabel>
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||
Ćurčić & Ilić, 2015 (red p) (
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||
<collectionCode id="ED521B96FF86FFDAF2C17AE8FCCCFB51" box="[715,776,1226,1249]" pageId="2" pageNumber="199">IZFB</collectionCode>
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) (
|
||
<figureCitation id="13789FD6FF86FFDAF3147AE8FC9BFB51" box="[798,863,1226,1249]" captionStart="FIGURES 1 – 12" captionStartId="1.[151,258,1840,1862]" captionTargetBox="[172,1432,703,1813]" captionTargetId="figure@1.[151,1436,684,1818]" captionTargetPageId="1" captionText="FIGURES 1 – 12. Tapinopterus (Tapinopterus) cognatus vlasinensis ssp. n. from the village of Vlasina Rid, Vlasina Lake Plateau, Mt. Čemernik, 1,200 m a. s. l., Surdulica, SE Serbia (1 – 6) and T. (T.) cognatus pancici ssp. n. from by the road Jelak Reserve - village of Brzeće, Mt. Kopaonik, 1,100 m a. s. l., Jošanička Banja, C Serbia (7 – 12). 1, 7 — holotype male, habitus (dorsal view); 2, 8 — holotype male, aedeagus (dorsal view); 3, 9 — holotype male, aedeagus (lateral view); 4, 10 — holotype male, right paramere (external lateral view); 5, 11 — holotype male, abdominal sternite IX (urite); 6, 12 — paratype female, genitalia (dorsal view). Scales = 5 mm (Figs. 1 and 7), 1 mm (Figs. 2, 3, 5, 6, 8, 9, 11, and 12), and 0.5 mm (Figs. 4 and 10)." httpUri="https://zenodo.org/record/243750/files/figure.png" pageId="2" pageNumber="199">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFC8353FF86FFDAF0CD7ACFFD21FA3F" blockId="2.[151,1436,1053,2012]" pageId="2" pageNumber="199">
|
||
<typeStatus id="54F83DF1FF86FFDAF0CD7ACFFEF7FAB5" box="[199,307,1261,1285]" pageId="2" pageNumber="199" type="paratype">Paratypes</typeStatus>
|
||
: 5 ♂♂ 3 ♀♀, same data as for
|
||
<typeStatus id="54F83DF1FF86FFDAF28F7ACEFD22FAB4" box="[645,742,1260,1284]" pageId="2" pageNumber="199" type="holotype">holotype</typeStatus>
|
||
; 1 ♀, village of Brzeće, Mt. Kopaonik,
|
||
<quantity id="4CBB2EB6FF86FFDAF49E7ACEFB28FAB4" box="[1172,1260,1260,1284]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="199" unit="m" value="1100.0">1,100 m</quantity>
|
||
a.s.l., Jošanička Banja, C
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||
<collectingCountry id="F354C3C3FF86FFDAF10A7B2DFE83FA97" box="[256,327,1295,1319]" name="Serbia" pageId="2" pageNumber="199">Serbia</collectingCountry>
|
||
,
|
||
<date id="FFFDA593FF86FFDAF15E7B2DFE25FA97" box="[340,481,1295,1319]" pageId="2" pageNumber="199" value="1996-08-03">03.VIII.1996</date>
|
||
, from pitfall traps, leg. N. Ilić; 4 ♀♀, village of Đerekari, Mt. Kopaonik,
|
||
<quantity id="4CBB2EB6FF86FFDAF51E7B2DFA9EFA97" box="[1300,1370,1295,1319]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="199" unit="m" value="800.0">800 m</quantity>
|
||
a.s.l., Brus, C
|
||
<collectingCountry id="F354C3C3FF86FFDAF0FB7B13FEFCFAF9" box="[241,312,1329,1353]" name="Serbia" pageId="2" pageNumber="199">Serbia</collectingCountry>
|
||
,
|
||
<date id="FFFDA593FF86FFDAF14E7B13FE15FAF9" box="[324,465,1329,1353]" pageId="2" pageNumber="199" value="1996-08-03">03.VIII.1996</date>
|
||
, from pitfall traps, leg. N. Ilić;
|
||
<specimenCount id="9D4548DAFF86FFDAF32D7B13FC94FAF9" box="[807,848,1329,1353]" pageId="2" pageNumber="199" type="generic">1 ♂</specimenCount>
|
||
1 ♀, village of Razbojna, Mt. Kopaonik,
|
||
<quantity id="4CBB2EB6FF86FFDAF51D7B13FA98FAF9" box="[1303,1372,1329,1353]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="199" unit="m" value="600.0">600 m</quantity>
|
||
a.s.l., Blace, C
|
||
<collectingCountry id="F354C3C3FF86FFDAF10B7B77FE8CFADC" box="[257,328,1365,1388]" name="Serbia" pageId="2" pageNumber="199">Serbia</collectingCountry>
|
||
,
|
||
<date id="FFFDA593FF86FFDAF15C7B77FE20FADC" box="[342,484,1365,1388]" pageId="2" pageNumber="199" value="1996-08-03">03.VIII.1996</date>
|
||
, from pitfall traps, leg. N. Ilić (p)
|
||
<typeStatus id="54F83DF1FF86FFDAF3647B77FC36FADC" box="[878,1010,1365,1388]" pageId="2" pageNumber="199" type="paratype">PARATYPE</typeStatus>
|
||
<taxonomicName id="4C43F8D0FF86FFDAF3F77B77FF21FA3F" class="Insecta" family="Carabidae" genus="Tapinopterus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="subSpecies" species="cognatus" status="subsp. nov." subGenus="Tapinopterus" subSpecies="pancici">
|
||
<emphasis id="B9375F41FF86FFDAF3F77B77FB4CFADC" box="[1021,1160,1365,1388]" italics="true" pageId="2" pageNumber="199">Tapinopterus</emphasis>
|
||
(
|
||
<emphasis id="B9375F41FF86FFDAF4967B77FAE3FADC" box="[1180,1319,1365,1388]" italics="true" pageId="2" pageNumber="199">Tapinopterus</emphasis>
|
||
)
|
||
<emphasis id="B9375F41FF86FFDAF5367B77FF21FA3F" italics="true" pageId="2" pageNumber="199">cognatus pancici</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A204E23AFF86FFDAF0E67B5AFEF6FA3F" box="[236,306,1400,1423]" pageId="2" pageNumber="199" rank="subSpecies">
|
||
<emphasis id="B9375F41FF86FFDAF0E67B5AFEF6FA3F" bold="true" box="[236,306,1400,1423]" pageId="2" pageNumber="199">ssp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Ćurčić & Ilić, 2015 (red p) (
|
||
<collectionCode id="ED521B96FF86FFDAF26F7B5AFD67FA3F" box="[613,675,1400,1423]" pageId="2" pageNumber="199">IZFB</collectionCode>
|
||
, cNI).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C359D0D8FF86FFDBF0CD7BB8FEDAFF44" lastPageId="3" lastPageNumber="200" pageId="2" pageNumber="199" type="description">
|
||
<paragraph id="8BFC8353FF86FFDAF0CD7BB8FBBFF9A9" blockId="2.[151,1436,1053,2012]" pageId="2" pageNumber="199">
|
||
<emphasis id="B9375F41FF86FFDAF0CD7BB8FE96FA01" bold="true" box="[199,338,1434,1457]" pageId="2" pageNumber="199">Description.</emphasis>
|
||
BL M
|
||
<quantity id="4CBB2EB6FF86FFDAF1A27BB8FDDCFA01" box="[424,536,1434,1457]" metricMagnitude="-2" metricUnit="m" metricValue="1.053" pageId="2" pageNumber="199" unit="mm" value="10.53">10.53 mm</quantity>
|
||
(R
|
||
<quantity id="4CBB2EB6FF86FFDAF24C7BB8FCC7FA01" box="[582,771,1434,1457]" metricMagnitude="-2" metricUnit="m" metricValue="1.0539999999999998" metricValueMax="1.098" metricValueMin="1.01" pageId="2" pageNumber="199" unit="mm" value="10.54" valueMax="10.98" valueMin="10.1">10.10–10.98 mm</quantity>
|
||
), BW M
|
||
<quantity id="4CBB2EB6FF86FFDAF3647BB8FC17FA01" box="[878,979,1434,1457]" metricMagnitude="-3" metricUnit="m" metricValue="3.46" pageId="2" pageNumber="199" unit="mm" value="3.46">3.46 mm</quantity>
|
||
. Head, pronotum, and elytra dark brown, shiny, antennae of similar color, legs and abdomen piceous brown, palpi somewhat paler. Head large, robust, somewhat wider than long (ratio HL/HW M 0.92), smooth, with a fine isodiametric microsculpture. Antennae moderately long, somewhat exceeding elytral humeri, with dense, decumbent pubescence from antennomere 4.
|
||
</paragraph>
|
||
<paragraph id="8BFC8353FF86FFDAF0CD7807FE6EF8BC" blockId="2.[151,1436,1053,2012]" pageId="2" pageNumber="199">
|
||
Pronotum weakly transverse, somewhat wider than long, ratio PL/PW M 0.90, PL M
|
||
<quantity id="4CBB2EB6FF86FFDAF46C7807FB02F98C" box="[1126,1222,1573,1596]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="199" unit="mm" value="2.6">2.60 mm</quantity>
|
||
(R
|
||
<quantity id="4CBB2EB6FF86FFDAF4FA7807FA4BF98C" box="[1264,1423,1573,1596]" metricMagnitude="-3" metricUnit="m" metricValue="2.5949999999999998" metricValueMax="2.75" metricValueMin="2.44" pageId="2" pageNumber="199" unit="mm" value="2.5949999999999998" valueMax="2.75" valueMin="2.44">2.44–2.75 mm</quantity>
|
||
), PW M
|
||
<quantity id="4CBB2EB6FF86FFDAF0E9786AFE85F9EF" box="[227,321,1608,1631]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="199" unit="mm" value="2.9">2.90 mm</quantity>
|
||
(R
|
||
<quantity id="4CBB2EB6FF86FFDAF162786AFDC1F9EF" box="[360,517,1608,1631]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.05" metricValueMin="2.75" pageId="2" pageNumber="199" unit="mm" value="2.9" valueMax="3.05" valueMin="2.75">2.75–3.05 mm</quantity>
|
||
), PB M
|
||
<quantity id="4CBB2EB6FF86FFDAF256786AFD7FF9EF" box="[604,699,1608,1631]" metricMagnitude="-3" metricUnit="m" metricValue="1.93" pageId="2" pageNumber="199" unit="mm" value="1.93">1.93 mm</quantity>
|
||
(R
|
||
<quantity id="4CBB2EB6FF86FFDAF2EB786AFCBBF9EF" box="[737,895,1608,1631]" metricMagnitude="-3" metricUnit="m" metricValue="1.9449999999999998" metricValueMax="2.07" metricValueMin="1.82" pageId="2" pageNumber="199" unit="mm" value="1.9449999999999998" valueMax="2.07" valueMin="1.82">1.82–2.07 mm</quantity>
|
||
), with rounded lateral margins, smooth, with a fine isodiametric microsculpture, widest slightly before first third. Anterior margin straight or somewhat sinuated, with prominent, at apex rounded angles, basal margin straight or slightly sinuated. Posterior angles prominent, acute, almost rectangular. Pronotal base with single deep longitudinal posterolateral impression on each side of midline, reaching basal third of pronotum. Midline distinct, not reaching basal or anterior margins. Both midline and posterolateral impressions with transversal wrinkles.
|
||
</paragraph>
|
||
<paragraph id="8BFC8353FF86FFDAF0CD793AFB04F8C4" blockId="2.[151,1436,1053,2012]" pageId="2" pageNumber="199">Elytra oval, with an isodiametric microsculpture, ratio EW/EL M 0.57, with maximum width around at middle, with distinct, weakly punctate striae. Disc relatively flattened basally, while somewhat convex apically. Shoulders prominent, rounded. Second elytral striae with one setigerous pore each situated at the level of ¾ elytral length.</paragraph>
|
||
<paragraph id="8BFC8353FF86FFDAF0CD79A2FA85F86C" blockId="2.[151,1436,1053,2012]" pageId="2" pageNumber="199">
|
||
Median lobe of aedeagus in dorsal view relatively long, straight, with apical blade wide, curved to left (
|
||
<figureCitation id="13789FD6FF86FFDAF4E579A2FAE9F827" box="[1263,1325,1920,1943]" captionStart="FIGURES 1 – 12" captionStartId="1.[151,258,1840,1862]" captionTargetBox="[172,1432,703,1813]" captionTargetId="figure@1.[151,1436,684,1818]" captionTargetPageId="1" captionText="FIGURES 1 – 12. Tapinopterus (Tapinopterus) cognatus vlasinensis ssp. n. from the village of Vlasina Rid, Vlasina Lake Plateau, Mt. Čemernik, 1,200 m a. s. l., Surdulica, SE Serbia (1 – 6) and T. (T.) cognatus pancici ssp. n. from by the road Jelak Reserve - village of Brzeće, Mt. Kopaonik, 1,100 m a. s. l., Jošanička Banja, C Serbia (7 – 12). 1, 7 — holotype male, habitus (dorsal view); 2, 8 — holotype male, aedeagus (dorsal view); 3, 9 — holotype male, aedeagus (lateral view); 4, 10 — holotype male, right paramere (external lateral view); 5, 11 — holotype male, abdominal sternite IX (urite); 6, 12 — paratype female, genitalia (dorsal view). Scales = 5 mm (Figs. 1 and 7), 1 mm (Figs. 2, 3, 5, 6, 8, 9, 11, and 12), and 0.5 mm (Figs. 4 and 10)." httpUri="https://zenodo.org/record/243750/files/figure.png" pageId="2" pageNumber="199">Fig. 8</figureCitation>
|
||
), in lateral view arcuate, sub-apically abruptly broadened, with an apex abruptly bent upwards (
|
||
<figureCitation id="13789FD6FF86FFDAF4047980FB8AF809" box="[1038,1102,1954,1977]" captionStart="FIGURES 1 – 12" captionStartId="1.[151,258,1840,1862]" captionTargetBox="[172,1432,703,1813]" captionTargetId="figure@1.[151,1436,684,1818]" captionTargetPageId="1" captionText="FIGURES 1 – 12. Tapinopterus (Tapinopterus) cognatus vlasinensis ssp. n. from the village of Vlasina Rid, Vlasina Lake Plateau, Mt. Čemernik, 1,200 m a. s. l., Surdulica, SE Serbia (1 – 6) and T. (T.) cognatus pancici ssp. n. from by the road Jelak Reserve - village of Brzeće, Mt. Kopaonik, 1,100 m a. s. l., Jošanička Banja, C Serbia (7 – 12). 1, 7 — holotype male, habitus (dorsal view); 2, 8 — holotype male, aedeagus (dorsal view); 3, 9 — holotype male, aedeagus (lateral view); 4, 10 — holotype male, right paramere (external lateral view); 5, 11 — holotype male, abdominal sternite IX (urite); 6, 12 — paratype female, genitalia (dorsal view). Scales = 5 mm (Figs. 1 and 7), 1 mm (Figs. 2, 3, 5, 6, 8, 9, 11, and 12), and 0.5 mm (Figs. 4 and 10)." httpUri="https://zenodo.org/record/243750/files/figure.png" pageId="2" pageNumber="199">Fig. 9</figureCitation>
|
||
). Right non-conchoid paramere elongate, of almost equal width over its whole length, curved (
|
||
<figureCitation id="13789FD6FF86FFDAF32B79E7FCA9F86C" box="[801,877,1989,2012]" captionStart="FIGURES 1 – 12" captionStartId="1.[151,258,1840,1862]" captionTargetBox="[172,1432,703,1813]" captionTargetId="figure@1.[151,1436,684,1818]" captionTargetPageId="1" captionText="FIGURES 1 – 12. Tapinopterus (Tapinopterus) cognatus vlasinensis ssp. n. from the village of Vlasina Rid, Vlasina Lake Plateau, Mt. Čemernik, 1,200 m a. s. l., Surdulica, SE Serbia (1 – 6) and T. (T.) cognatus pancici ssp. n. from by the road Jelak Reserve - village of Brzeće, Mt. Kopaonik, 1,100 m a. s. l., Jošanička Banja, C Serbia (7 – 12). 1, 7 — holotype male, habitus (dorsal view); 2, 8 — holotype male, aedeagus (dorsal view); 3, 9 — holotype male, aedeagus (lateral view); 4, 10 — holotype male, right paramere (external lateral view); 5, 11 — holotype male, abdominal sternite IX (urite); 6, 12 — paratype female, genitalia (dorsal view). Scales = 5 mm (Figs. 1 and 7), 1 mm (Figs. 2, 3, 5, 6, 8, 9, 11, and 12), and 0.5 mm (Figs. 4 and 10)." httpUri="https://zenodo.org/record/243750/files/figure.png" pageId="2" pageNumber="199">Fig. 10</figureCitation>
|
||
), while left conchoid broad, ear-like (
|
||
<figureCitation id="13789FD6FF86FFDAF4FF79E7FAF7F86C" box="[1269,1331,1989,2012]" captionStart="FIGURES 1 – 12" captionStartId="1.[151,258,1840,1862]" captionTargetBox="[172,1432,703,1813]" captionTargetId="figure@1.[151,1436,684,1818]" captionTargetPageId="1" captionText="FIGURES 1 – 12. Tapinopterus (Tapinopterus) cognatus vlasinensis ssp. n. from the village of Vlasina Rid, Vlasina Lake Plateau, Mt. Čemernik, 1,200 m a. s. l., Surdulica, SE Serbia (1 – 6) and T. (T.) cognatus pancici ssp. n. from by the road Jelak Reserve - village of Brzeće, Mt. Kopaonik, 1,100 m a. s. l., Jošanička Banja, C Serbia (7 – 12). 1, 7 — holotype male, habitus (dorsal view); 2, 8 — holotype male, aedeagus (dorsal view); 3, 9 — holotype male, aedeagus (lateral view); 4, 10 — holotype male, right paramere (external lateral view); 5, 11 — holotype male, abdominal sternite IX (urite); 6, 12 — paratype female, genitalia (dorsal view). Scales = 5 mm (Figs. 1 and 7), 1 mm (Figs. 2, 3, 5, 6, 8, 9, 11, and 12), and 0.5 mm (Figs. 4 and 10)." httpUri="https://zenodo.org/record/243750/files/figure.png" pageId="2" pageNumber="199">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFC8353FF87FFDBF0CD7EBAFC50FF1F" blockId="3.[151,1436,152,487]" box="[199,916,152,175]" pageId="3" pageNumber="200">
|
||
Male abdominal sternite IX (urite) well-developed, ovoid (
|
||
<figureCitation id="13789FD6FF87FFDBF3337EBAFC43FF1F" box="[825,903,152,175]" captionStart="FIGURES 1 – 12" captionStartId="1.[151,258,1840,1862]" captionTargetBox="[172,1432,703,1813]" captionTargetId="figure@1.[151,1436,684,1818]" captionTargetPageId="1" captionText="FIGURES 1 – 12. Tapinopterus (Tapinopterus) cognatus vlasinensis ssp. n. from the village of Vlasina Rid, Vlasina Lake Plateau, Mt. Čemernik, 1,200 m a. s. l., Surdulica, SE Serbia (1 – 6) and T. (T.) cognatus pancici ssp. n. from by the road Jelak Reserve - village of Brzeće, Mt. Kopaonik, 1,100 m a. s. l., Jošanička Banja, C Serbia (7 – 12). 1, 7 — holotype male, habitus (dorsal view); 2, 8 — holotype male, aedeagus (dorsal view); 3, 9 — holotype male, aedeagus (lateral view); 4, 10 — holotype male, right paramere (external lateral view); 5, 11 — holotype male, abdominal sternite IX (urite); 6, 12 — paratype female, genitalia (dorsal view). Scales = 5 mm (Figs. 1 and 7), 1 mm (Figs. 2, 3, 5, 6, 8, 9, 11, and 12), and 0.5 mm (Figs. 4 and 10)." httpUri="https://zenodo.org/record/243750/files/figure.png" pageId="3" pageNumber="200">Fig. 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFC8353FF87FFDBF0CD7E98FEDAFF44" blockId="3.[151,1436,152,487]" pageId="3" pageNumber="200">
|
||
Gonocoxites IX long, gradually narrowing distally, somewhat curved, basally joined with massive gonosubcoxites IX (
|
||
<figureCitation id="13789FD6FF87FFDBF0C87EFFFED5FF44" box="[194,273,221,244]" captionStart="FIGURES 1 – 12" captionStartId="1.[151,258,1840,1862]" captionTargetBox="[172,1432,703,1813]" captionTargetId="figure@1.[151,1436,684,1818]" captionTargetPageId="1" captionText="FIGURES 1 – 12. Tapinopterus (Tapinopterus) cognatus vlasinensis ssp. n. from the village of Vlasina Rid, Vlasina Lake Plateau, Mt. Čemernik, 1,200 m a. s. l., Surdulica, SE Serbia (1 – 6) and T. (T.) cognatus pancici ssp. n. from by the road Jelak Reserve - village of Brzeće, Mt. Kopaonik, 1,100 m a. s. l., Jošanička Banja, C Serbia (7 – 12). 1, 7 — holotype male, habitus (dorsal view); 2, 8 — holotype male, aedeagus (dorsal view); 3, 9 — holotype male, aedeagus (lateral view); 4, 10 — holotype male, right paramere (external lateral view); 5, 11 — holotype male, abdominal sternite IX (urite); 6, 12 — paratype female, genitalia (dorsal view). Scales = 5 mm (Figs. 1 and 7), 1 mm (Figs. 2, 3, 5, 6, 8, 9, 11, and 12), and 0.5 mm (Figs. 4 and 10)." httpUri="https://zenodo.org/record/243750/files/figure.png" pageId="3" pageNumber="200">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C359D0D8FF87FFDBF0CD7F22FEDFFEEC" pageId="3" pageNumber="200" type="etymology">
|
||
<paragraph id="8BFC8353FF87FFDBF0CD7F22FEDFFEEC" blockId="3.[151,1436,152,487]" pageId="3" pageNumber="200">
|
||
<emphasis id="B9375F41FF87FFDBF0CD7F22FE83FEA7" bold="true" box="[199,327,256,279]" pageId="3" pageNumber="200">Etymology.</emphasis>
|
||
The subspecies is named after Academician Prof. Dr. Josif Pančić, a famous Serbian naturalist and the first president of the Serbian Royal Academy, whose mausoleum is located in the highest peak of Mt. Kopaonik, the
|
||
<emphasis id="B9375F41FF87FFDBF09D7F67FED0FEEC" box="[151,276,325,348]" italics="true" pageId="3" pageNumber="200">terra typica</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C359D0D8FF87FFDBF0CD7F4AFA48FE11" pageId="3" pageNumber="200" type="diagnosis">
|
||
<paragraph id="8BFC8353FF87FFDBF0CD7F4AFA48FE11" blockId="3.[151,1436,152,487]" pageId="3" pageNumber="200">
|
||
<emphasis id="B9375F41FF87FFDBF0CD7F4AFEF8FECF" bold="true" box="[199,316,360,383]" pageId="3" pageNumber="200">Diagnosis.</emphasis>
|
||
The diagnostic differences between
|
||
<taxonomicName id="4C43F8D0FF87FFDBF2CD7F4AFBF9FECF" box="[711,1085,360,383]" class="Insecta" family="Carabidae" genus="Tapinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="200" phylum="Arthropoda" rank="subSpecies" species="cognatus" status="subsp. nov." subGenus="Tapinopterus" subSpecies="pancici">
|
||
<emphasis id="B9375F41FF87FFDBF2CD7F4AFC96FECF" box="[711,850,360,383]" italics="true" pageId="3" pageNumber="200">Tapinopterus</emphasis>
|
||
(
|
||
<emphasis id="B9375F41FF87FFDBF36E7F4AFCB5FECF" box="[868,881,360,383]" italics="true" pageId="3" pageNumber="200">T</emphasis>
|
||
.)
|
||
<emphasis id="B9375F41FF87FFDBF3827F4AFBF9FECF" box="[904,1085,360,383]" italics="true" pageId="3" pageNumber="200">cognatus pancici</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A204E23AFF87FFDBF44C7F4AFB49FECF" box="[1094,1165,360,383]" pageId="3" pageNumber="200" rank="subSpecies">
|
||
<emphasis id="B9375F41FF87FFDBF44C7F4AFB49FECF" bold="true" box="[1094,1165,360,383]" pageId="3" pageNumber="200">ssp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and the morphologically and geographically closest taxa are previously presented in the differential diagnosis of
|
||
<taxonomicName id="4C43F8D0FF87FFDBF42C7FA8FA84FE11" box="[1062,1344,394,417]" class="Insecta" family="Carabidae" genus="Tapinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="200" phylum="Arthropoda" rank="subSpecies" species="cognatus" status="subsp. nov." subGenus="Tapinopterus" subSpecies="vlasinensis">
|
||
<emphasis id="B9375F41FF87FFDBF42C7FA8FBFCFE11" box="[1062,1080,394,417]" italics="true" pageId="3" pageNumber="200">T.</emphasis>
|
||
(
|
||
<emphasis id="B9375F41FF87FFDBF4427FA8FB90FE11" box="[1096,1108,394,417]" italics="true" pageId="3" pageNumber="200">T</emphasis>
|
||
.)
|
||
<emphasis id="B9375F41FF87FFDBF4607FA8FA84FE11" box="[1130,1344,394,417]" italics="true" pageId="3" pageNumber="200">cognatus vlasinensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9375F41FF87FFDBF54D7FA8FA48FE11" bold="true" box="[1351,1420,394,417]" pageId="3" pageNumber="200">
|
||
<taxonomicNameLabel id="A204E23AFF87FFDBF54D7FA8FA48FE11" box="[1351,1420,394,417]" pageId="3" pageNumber="200" rank="subSpecies">ssp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C359D0D8FF87FFDBF0CD7F8FFCADFE57" pageId="3" pageNumber="200" type="distribution">
|
||
<paragraph id="8BFC8353FF87FFDBF0CD7F8FFCADFE57" blockId="3.[151,1436,152,487]" pageId="3" pageNumber="200">
|
||
<emphasis id="B9375F41FF87FFDBF0CD7F8FFE36FE74" bold="true" box="[199,498,429,452]" pageId="3" pageNumber="200">Distribution and habitats.</emphasis>
|
||
The
|
||
<typeStatus id="54F83DF1FF87FFDBF2247F8FFD98FE74" box="[558,604,429,452]" pageId="3" pageNumber="200">type</typeStatus>
|
||
specimens have been collected at different altitudes (
|
||
<quantity id="4CBB2EB6FF87FFDBF4A37F8FFAFEFE74" box="[1193,1338,429,452]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" pageId="3" pageNumber="200" unit="m" value="850.0" valueMax="1100.0" valueMin="600.0">600–1,100 m</quantity>
|
||
a.s.l.) in mixed deciduous-coniferous forests at a few sites on Mt. Kopaonik.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |