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<document id="39AB376D7C4F2E45461998F9A8920CB4" ID="10.11646/zootaxa.4238.3.1SLASH10400" ID-CLB-Dataset="36345" ID-DOI="10.11646/zootaxa.4238.3.1" ID-GBIF-Dataset="030203e2-e6a1-4b59-bfa0-0689bf817b08" ID-ISSN="1175-5326" ID-Zenodo-Dep="345756" ID-ZooBank="D123950C-DF2B-45EE-9966-813339A83491" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1488787785636" checkinUser="plazi" docAuthor="Krikken, J. &amp; Huijbregts, J." docDate="2017" docId="03E5BB4DE833FFC5AAE4FF0FFBB1F86D" docLanguage="en" docName="zootaxa.4238.3.1SLASH10400.pdf" docOrigin="Zootaxa 4238 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus manguliensis Boucomont 1914" docType="treatment" docVersion="7" lastPageNumber="324" masterDocId="FFDCC335E824FFD2AA73FF98FFDAFFDA" masterDocTitle="Sulawesi Onthophagus: seven new species in select groups (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="334" masterPageNumber="301" pageNumber="324" updateTime="1698437609924" updateUser="ExternalLinkService">
<mods:mods id="D9DDE1C9184FB0F168BB971503291B94" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4B17557C65BBBA7EA68EBCE13D034A0B">Sulawesi Onthophagus: seven new species in select groups (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="2B0F978946B3BD8F124D44AB762B25A7" type="personal">
<mods:role id="C353B3AFAB54BF8798A6F79F1A9AA0D6">
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</mods:role>
<mods:namePart id="6D640F60A734A6601127C5D1D2CB09F3">Krikken, J.</mods:namePart>
</mods:name>
<mods:name id="147196CA9337756676E53ABD66915CC8" type="personal">
<mods:role id="1FF9B57ED5C3355286E59737F36D1C0E">
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<mods:namePart id="7D54F2E361340D68D498961E19F4C52D">Huijbregts, J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="2A1D8C85F2BB9FEAD2CFE20AB49695E6">text</mods:typeOfResource>
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<mods:title id="4EE825C6EE17BB01E4F708881944692C">Zootaxa</mods:title>
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<mods:part id="83132266A1252D3B74EC1E3BE4A54491">
<mods:date id="ADE4F53201DB0C17545AA997CE5CA8C4">2017</mods:date>
<mods:detail id="45C5EF6588AB7F5B1C4EAF8BBD8ECF9E" type="volume">
<mods:number id="2A289F21F309325B8EF0B41083FE2629">4238</mods:number>
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<mods:number id="F53308C81F6D80FB662B178FC116EA22">3</mods:number>
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<mods:start id="2B106D6B2E49CC8BA4D748A3EA4214DF">301</mods:start>
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<mods:classification id="BBA723BFC8BF611C15AE1444D7336CAE">journal article</mods:classification>
<mods:identifier id="61B6ADE01FFF43DDAA3199ED1547E0A5" type="CLB-Dataset">36345</mods:identifier>
<mods:identifier id="8288C9868E6BEBA49CB788C3C2704142" type="DOI">10.11646/zootaxa.4238.3.1</mods:identifier>
<mods:identifier id="E2A7A9CED2F175667DFED4A94E8ADA7A" type="GBIF-Dataset">030203e2-e6a1-4b59-bfa0-0689bf817b08</mods:identifier>
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<treatment id="03E5BB4DE833FFC5AAE4FF0FFBB1F86D" ID-DOI="http://doi.org/10.5281/zenodo.6030195" ID-GBIF-Taxon="127719082" ID-Zenodo-Dep="6030195" LSID="urn:lsid:plazi:treatment:03E5BB4DE833FFC5AAE4FF0FFBB1F86D" httpUri="http://treatment.plazi.org/id/03E5BB4DE833FFC5AAE4FF0FFBB1F86D" lastPageNumber="324" pageId="23" pageNumber="324">
<subSubSection id="C35659D0E833FFC5AAE4FF0FFEAEFF09" pageId="23" pageNumber="324" type="nomenclature">
<paragraph id="8BF30A5BE833FFC5AAE4FF0FFD1EFF68" blockId="23.[151,708,151,211]" box="[151,708,151,178]" pageId="23" pageNumber="324">
<heading id="D0BBBD37E833FFC5AAE4FF0FFD1EFF68" bold="true" box="[151,708,151,178]" fontSize="11" level="1" pageId="23" pageNumber="324" reason="1">
<taxonomicName id="4C4C71D8E833FFC5AAE4FF0FFD1EFF68" ID-CoL="7XNGY" authority="Boucomont, 1914" authorityName="Boucomont" authorityYear="1914" box="[151,708,151,178]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="manguliensis">
<emphasis id="B938D649E833FFC5AAE4FF0FFD1EFF68" bold="true" box="[151,708,151,178]" pageId="23" pageNumber="324">
<emphasis id="B938D649E833FFC5AAE4FF0FFE3EFF6B" bold="true" box="[151,484,151,177]" italics="true" pageId="23" pageNumber="324">Onthophagus manguliensis</emphasis>
<bibRefCitation id="EFDD77AAE833FFC5AB98FF00FD1EFF68" author="Boucomont" box="[491,708,152,178]" pageId="23" pageNumber="324" refString="Boucomont, A. (1914) Les Coprophages de l'Archipel malais (Coleopt.). Annales de la Societe Entomologique de France, 73, 238 - 350." type="journal article" year="1914">Boucomont, 1914</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAE4FF22FEAEFF09" blockId="23.[151,708,151,211]" box="[151,372,186,211]" pageId="23" pageNumber="324">
(
<figureCitation id="137716DEE833FFC5AAECFF22FF37FF09" box="[159,237,186,211]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,264,1919,1942]" captionTargetBox="[207,1396,233,1891]" captionTargetId="figure@5.[177,1413,195,1894]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Sulawesi Onthophagus species, oblique habitus. 1, O. bisscrutator, holotype male from Toraut. 2, O. scrutator, male from Moramo. 3, O. bongkudai, holotype male from Moajat. 4, O. sangirensis, male from Toraut. 5, O. holosericus, male from Marena, 6, female from Marena. 7, O. sopu, holotype male from Sopu. 8, O. manguliensis, male from Mangoli. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345757/files/figure.png" pageId="23" pageNumber="324">Figs. 8</figureCitation>
,
<figureCitation id="137716DEE833FFC5AA89FF23FEC3FF09" box="[250,281,187,211]" captionStart="FIGURES 25 – 30" captionStartId="9.[151,264,1925,1948]" captionTargetBox="[170,1436,198,1875]" captionTargetId="figure@9.[151,1436,193,1904]" captionTargetPageId="9" captionText="FIGURES 25 – 30. Sulawesi Onthophagus species, dorsal forebody. 25, O. sopu, holotype male from Sopu. 26, O. sopu, paratype male from Sopu. 27, O. manguliensis, male from Mangoli. 28, O. annulopunctatus, paratype male from Mogogonipa. 29, O. hollowayi, holotype male from Dumoga. 30, O. seseba, holotype male from Batui. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345761/files/figure.png" pageId="23" pageNumber="324">27</figureCitation>
,
<figureCitation id="137716DEE833FFC5AB56FF22FEB6FF09" box="[293,364,186,211]" captionStart="FIGURES 47 – 55" captionStartId="22.[151,264,1855,1878]" captionTargetBox="[154,1435,291,1792]" captionTargetId="figure@22.[151,1436,267,1834]" captionTargetPageId="22" captionText="FIGURES 47 – 55. Sulawesi Onthophagu s body parts, male, female. 47 – 52, O. sopu, all Sopu, paratype male (47, 51), holotype male (48 – 50), paratype female (52). 53 – 55, O. manguliensis, male Mangoli (53), male Matanyo (55), female Matanyo (54). — 47, 52, 53 - 54, head full-face, 48, left elytron dorsal, 49, protibial upper side, 50, metatibial upperside, 51, 55, aedeagus lateral. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345764/files/figure.png" pageId="23" pageNumber="324">53–55</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35659D0E833FFC5AAE4FE9AFD15FC31" pageId="23" pageNumber="324" type="materials_examined">
<paragraph id="8BF30A5BE833FFC5AAE4FE9AFC42FE5D" blockId="23.[151,1437,258,1975]" pageId="23" pageNumber="324">
<materialsCitation id="3B240006E833FFC5AAE4FE9AFEF9FEE5" ID-GBIF-Occurrence="1438340100" collectionCode="MNHN, MUSEO" country="Indonesia" location="Sula Mangoli" pageId="23" pageNumber="324" specimenCount="1" typeStatus="syntype">
<emphasis id="B938D649E833FFC5AAE4FE9AFE5AFEC1" bold="true" box="[151,384,258,283]" pageId="23" pageNumber="324">Material examined.</emphasis>
<typeStatus id="54F7B4F9E833FFC5ABF5FE9BFE38FEC1" box="[390,482,259,283]" pageId="23" pageNumber="324" type="syntype">Syntype</typeStatus>
male (
<collectionCode id="ED5D929EE833FFC5A85EFE9BFD5CFEC1" box="[557,646,259,283]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="324">MNHN</collectionCode>
) from
<collectingCountry id="F35B4ACBE833FFC5A8A2FE9AFC9BFEC1" box="[721,833,258,283]" name="Indonesia" pageId="23" pageNumber="324">Indonesia</collectingCountry>
, labelled: &quot;
<location id="8E935C80E833FFC5A9CFFE9AFB8CFEC1" LSID="urn:lsid:plazi:treatment:03E5BB4DE833FFC5AAE4FF0FFBB1F86D:8E935C80E833FFC5A9CFFE9AFB8CFEC1" box="[956,1110,258,283]" country="Indonesia" name="Sula Mangoli" pageId="23" pageNumber="324">Sula Mangoli</location>
\ Oct.-Novbr. Doherty&quot;, &quot;EX
<collectionCode id="ED5D929EE833FFC5AAE4FEBFFF23FEE5" box="[151,249,295,319]" pageId="23" pageNumber="324">MUSEO</collectionCode>
\ N
</materialsCitation>
.VAN DE POLL&quot;, &quot;
<taxonomicName id="4C4C71D8E833FFC5AB8CFEBEFD4DFEE5" box="[511,663,294,319]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="genus">Onthophagus</taxonomicName>
\
<taxonomicName id="4C4C71D8E833FFC5A8DEFEBEFC9BFEE5" box="[685,833,294,319]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="manguliensis">manguliensis</taxonomicName>
\ B. \ COTYPE&quot;, &quot;MUSEUM PARIS \ 1938 \ COLL.
<materialsCitation id="3B240006E833FFC5AAE4FED3FC42FE5D" ID-GBIF-Occurrence="1438340096" collectionCode="RMNH" collectorName="Doherty &amp; W. Doherty" country="Indonesia" location="Sula Islands" pageId="23" pageNumber="324" specimenCount="1" typeStatus="holotype">
A. BOUCOMONT&quot;. Additional material (
<collectionCode id="ED5D929EE833FFC5A804FED3FD15FEB9" box="[631,719,331,355]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="324">RMNH</collectionCode>
) with a similar
<collectorName id="26B96F8DE833FFC5A9F5FED2FC38FEB9" box="[902,994,330,355]" pageId="23" pageNumber="324">Doherty</collectorName>
collection label (
<collectorName id="26B96F8DE833FFC5AEDAFED3FAF5FEB9" box="[1193,1327,330,355]" pageId="23" pageNumber="324">W. Doherty</collectorName>
collected on the
<location id="8E935C80E833FFC5AA90FEF6FEB1FE5D" LSID="urn:lsid:plazi:treatment:03E5BB4DE833FFC5AAE4FF0FFBB1F86D:8E935C80E833FFC5AA90FEF6FEB1FE5D" box="[227,363,366,391]" country="Indonesia" name="Sula Islands" pageId="23" pageNumber="324">Sula Islands</location>
around 1890); see also note on
<typeStatus id="54F7B4F9E833FFC5A8A3FEF7FCDAFE5D" box="[720,768,367,391]" pageId="23" pageNumber="324">type</typeStatus>
status below.
</materialsCitation>
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAB4FE0AFD6AFE71" blockId="23.[151,1437,258,1975]" box="[199,688,402,427]" pageId="23" pageNumber="324">
The
<specimenCount id="9D4AC1D2E833FFC5AA8AFE0AFE4BFE71" box="[249,401,402,427]" pageId="23" pageNumber="324" type="generic">35 specimens</specimenCount>
examined are as follows:
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAE4FE2FFD65FE15" blockId="23.[151,1437,258,1975]" box="[151,703,439,463]" pageId="23" pageNumber="324">
<materialsCitation id="3B240006E833FFC5AAE4FE2FFD65FE15" ID-GBIF-Occurrence="1438340109" box="[151,703,439,463]" location="CENTRAL SULAWESI" pageId="23" pageNumber="324" specimenCount="1" stateProvince="CENTRAL SULAWESI">
<collectingRegion id="4988C4B9E833FFC5AAE4FE2FFE75FE15" box="[151,431,439,463]" country="Indonesia" name="Sulawesi Tengah" pageId="23" pageNumber="324">CENTRAL SULAWESI</collectingRegion>
: BANGGAI DISTRICT
</materialsCitation>
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAE4FE43FEF1FDCC" blockId="23.[151,1437,258,1975]" pageId="23" pageNumber="324">
- Batui:
<taxonomicName id="4C4C71D8E833FFC5AB60FE42FEBBFE29" box="[275,353,474,499]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="331" phylum="Arthropoda" rank="species" species="seseba">Seseba</taxonomicName>
Estate,
<quantity id="4CB4A7BEE833FFC5ABCFFE43FE22FE28" box="[444,504,475,499]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="324" unit="m" value="80.0">80 m</quantity>
,
<date id="FFF22C9BE833FFC5A876FE42FDABFE29" box="[517,625,474,499]" pageId="23" pageNumber="324" value="1989-11-09">9.xi.1989</date>
, Krikken &amp; Van der Blom, sw31, grassland, cattle excrement extracted,
<quantity id="4CB4A7BEE833FFC5AAE4FE66FF0AFDCD" box="[151,208,510,535]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="23" pageNumber="324" unit="in" value="16.0">16 in</quantity>
<collectionCode id="ED5D929EE833FFC5AAA4FE67FEF1FDCC" box="[215,299,511,534]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="324">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAE4FDBBFD38FD84" blockId="23.[151,1437,258,1975]" pageId="23" pageNumber="324">
- Matanyo Forest, N of Kayutanyo,
<quantity id="4CB4A7BEE833FFC5A835FDBBFD57FDE0" box="[582,653,547,571]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="23" pageNumber="324" unit="m" value="110.0">110 m</quantity>
,
<date id="FFF22C9BE833FFC5A8EAFDBAFCFAFDE1" box="[665,800,546,571]" pageId="23" pageNumber="324" value="1985-11-02" valueMax="1985-11-04" valueMin="1985-11-02">2–4.xi.1985</date>
, Krikken &amp; Van der Blom, sw19a, multistratal evergreen forest, river, 2 human excrement traps,
<quantity id="4CB4A7BEE833FFC5A83DFDDFFD52FD85" box="[590,648,582,607]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="23" pageNumber="324" unit="in" value="12.0">12 in</quantity>
<collectionCode id="ED5D929EE833FFC5A8FDFDDFFD38FD84" box="[654,738,583,606]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="324">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAE4FDF3FDEDFD7C" blockId="23.[151,1437,258,1975]" pageId="23" pageNumber="324">
- Matanyo Forest, N of Kayutanyo,
<quantity id="4CB4A7BEE833FFC5A836FDF3FD57FD58" box="[581,653,619,643]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="23" pageNumber="324" unit="m" value="110.0">110 m</quantity>
,
<date id="FFF22C9BE833FFC5A8EBFDF2FCFAFD59" box="[664,800,618,643]" pageId="23" pageNumber="324" value="1989-11-02" valueMax="1989-11-04" valueMin="1989-11-02">2–4.xi.1989</date>
, Krikken &amp; Van der Blom, sw19b, multistratal evergreen forest, river, 2 fish traps,
<quantity id="4CB4A7BEE833FFC5ABC2FD17FE06FD7D" box="[433,476,654,679]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="23" pageNumber="324" unit="in" value="2.0">2 in</quantity>
<collectionCode id="ED5D929EE833FFC5AB90FD17FDEDFD7C" box="[483,567,655,678]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="324">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAE4FD2BFDAAFD11" blockId="23.[151,1437,258,1975]" box="[151,624,691,715]" pageId="23" pageNumber="324">
<collectingRegion id="4988C4B9E833FFC5AAE4FD2BFE75FD11" box="[151,431,691,715]" country="Indonesia" name="Sulawesi Tengah" pageId="23" pageNumber="324">
<collectionCode id="ED5D929EE833FFC5AAE4FD2BFE75FD11" box="[151,431,691,715]" pageId="23" pageNumber="324">CENTRAL SULAWESI</collectionCode>
</collectingRegion>
:
<collectionCode id="ED5D929EE833FFC5ABCAFD2BFDAAFD11" box="[441,624,691,715]" pageId="23" pageNumber="324">PALU REGION</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAE4FD4FFD46FCC8" blockId="23.[151,1437,258,1975]" pageId="23" pageNumber="324">
- Tawaeli-Toboli road (km 29),
<quantity id="4CB4A7BEE833FFC5A857FD4EFDAAFD34" box="[548,624,726,751]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="23" pageNumber="324" unit="m" value="250.0">250 m</quantity>
,
<date id="FFF22C9BE833FFC5A80CFD4EFCF1FD35" box="[639,811,726,751]" pageId="23" pageNumber="324" value="1985-12-20" valueMax="1985-12-22" valueMin="1985-12-20">20–22.xii.1985</date>
, J. Krikken, pw68a, second-growth forest/coffee, near rivulet, 2 human excrement traps,
<quantity id="4CB4A7BEE833FFC5A865FD63FD9BFCC9" box="[534,577,762,787]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="23" pageNumber="324" unit="in" value="2.0">2 in</quantity>
<collectionCode id="ED5D929EE833FFC5A83BFD63FD46FCC8" box="[584,668,763,786]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="324">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAE4FC87FD0CFCED" blockId="23.[151,1437,258,1975]" box="[151,726,799,823]" pageId="23" pageNumber="324">
<materialsCitation id="3B240006E833FFC5AAE4FC87FD0CFCED" ID-GBIF-Occurrence="1438340116" box="[151,726,799,823]" location="SOUTHEAST SULAWESI" pageId="23" pageNumber="324" specimenCount="1" stateProvince="SOUTHEAST SULAWESI">
<collectingRegion id="4988C4B9E833FFC5AAE4FC87FE0BFCED" box="[151,465,799,823]" country="Indonesia" name="Sulawesi Tenggara" pageId="23" pageNumber="324">SOUTHEAST SULAWESI</collectingRegion>
: KOLAKA DISTRICT
</materialsCitation>
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAE4FCDBFD48FCA4" blockId="23.[151,1437,258,1975]" pageId="23" pageNumber="324">
- Watuwila Mosquito Camp,
<quantity id="4CB4A7BEE833FFC5A877FCDAFD87FC80" box="[516,605,834,859]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="23" pageNumber="324" unit="m" value="1150.0">1150 m</quantity>
,
<date id="FFF22C9BE833FFC5A81FFCDAFCD2FC81" box="[620,776,834,859]" pageId="23" pageNumber="324" value="1989-10-13" valueMax="1989-10-15" valueMin="1989-10-13">13–15.x.1989</date>
, Krikken &amp; Van der Blom, sw11a, multistratal evergreen forest, 2 human excrement traps,
<quantity id="4CB4A7BEE833FFC5A87EFCFFFDE2FCA5" box="[525,568,870,895]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="23" pageNumber="324" unit="in" value="1.0">1 in</quantity>
<collectionCode id="ED5D929EE833FFC5A84DFCFFFD48FCA4" box="[574,658,871,894]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="324">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAE4FC13FE87FC79" blockId="23.[151,1437,258,1975]" box="[151,349,907,931]" pageId="23" pageNumber="324">SULA ISLANDS</paragraph>
<paragraph id="8BF30A5BE833FFC5AAE4FC37FD86FC1C" blockId="23.[151,1437,258,1975]" box="[151,604,942,967]" pageId="23" pageNumber="324">
- Mangoli, ix-x, Doherty,
<quantity id="4CB4A7BEE833FFC5ABA4FC37FDD8FC1D" box="[471,514,942,967]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="23" pageNumber="324" unit="in" value="1.0">1 in</quantity>
<collectionCode id="ED5D929EE833FFC5A87BFC37FD86FC1C" box="[520,604,943,966]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="324">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAE4FC4BFD15FC31" blockId="23.[151,1437,258,1975]" box="[151,719,978,1003]" pageId="23" pageNumber="324">
- Mangoli, x-xi, Doherty,
<quantity id="4CB4A7BEE833FFC5ABA4FC4BFDD8FC31" box="[471,514,978,1003]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="23" pageNumber="324" unit="in" value="1.0">1 in</quantity>
MNHN (syntype)
</paragraph>
</subSubSection>
<subSubSection id="C35659D0E833FFC5AAB4FC6EFBB1F86D" pageId="23" pageNumber="324" type="description">
<paragraph id="8BF30A5BE833FFC5AAB4FC6EFB25FBE8" blockId="23.[151,1437,258,1975]" pageId="23" pageNumber="324">
<emphasis id="B938D649E833FFC5AAB4FC6EFE83FBD5" bold="true" box="[199,345,1014,1039]" pageId="23" pageNumber="324">Description.</emphasis>
Body length
<quantity id="4CB4A7BEE833FFC5A874FC6EFDB5FBD4" box="[519,623,1014,1039]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="23" pageNumber="324" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
. Colour symmetrically patterned brown and orange-yellow, slightly variable, largely matt due to microreticulation. Dorsal side with numerous distinct pale-yellow setae.
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAB4FBA6FCA2FB19" blockId="23.[151,1437,258,1975]" pageId="23" pageNumber="324">
Clypeus anteromedially with pair of apically round denticles, emargination moderately deep; clypeogenal transition continuous at border; low clypeofrontal ridge present, slightly arcuate; paraocular protrusions between anterior border of eyes slight, more distinct in female; clypeus rugulate-punctate, shiny; remainder of head with sparse, coarse seta-bearing punctures and microreticulation (
<date id="FFF22C9BE833FFC5A94EFB32FCB1FB19" box="[829,875,1194,1219]" pageId="23" pageNumber="324" value="1960-10">x60</date>
).
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAB4FB57FBEBFA89" blockId="23.[151,1437,258,1975]" pageId="23" pageNumber="324">Pronotum slightly, evenly convex; anterior section of lateral border slightly round, posterior section of lateral border scarcely sinuate; punctation double; entire pronotal derm with abundant, seta-bearing, ocellate primary punctation; setae approximately 3 times puncture diameters, punctures mostly separated by 23 diameters; punctures on shallowly depressed, matt basomedian area slightly less pronounced.</paragraph>
<paragraph id="8BF30A5BE833FFC5AAB4FAC6FDACFA39" blockId="23.[151,1437,258,1975]" pageId="23" pageNumber="324">
Elytral striae discally well defined, with distinct punctures, mostly separated by 2–4 times their diameter, slightly crenulated on interstriae; interstriae almost flat, microreticulate (
<date id="FFF22C9BE833FFC5A9CCFA1BFC37FA41" box="[959,1005,1411,1435]" pageId="23" pageNumber="324" value="1940-10">x40</date>
), with numerous seta-bearing granules (asperate,
<date id="FFF22C9BE833FFC5AB79FA3FFEEFFA65" box="[266,309,1447,1471]" pageId="23" pageNumber="324" value="1940-10">x40</date>
) arranged in indistinct rows; peripheral surfaces of elytra with distinct yellow patches (like humerus, posterior declivity,
<emphasis id="B938D649E833FFC5AB1CFA53FE4AFA38" box="[367,400,1483,1506]" italics="true" pageId="23" pageNumber="324">etc</emphasis>
.), remainder brown.
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAB4FA76FD14F9A9" blockId="23.[151,1437,258,1975]" pageId="23" pageNumber="324">Antennal club gray yellowish brown. Metaventrite disc with numerous seta-bearing punctures on anterior lobe; lateral wings with sparse, seta-bearing, subocellate punctures, microreticulate. Abdominal ventrites each with transverse row of seta-bearing punctures. Pygidium yellow, matt, virtually flat, subtriangular; surface with sparse, seta-bearing punctures; base with transverse ridge.</paragraph>
<paragraph id="8BF30A5BE833FFC5AAB4F9E6FBB7F905" blockId="23.[151,1437,258,1975]" pageId="23" pageNumber="324">Protibia with 3+(1) larger denticles, slightly curved, proximal serration with approximately 10 small denticles; spur elongate-acuminate. Femora largely yellow-brown. Mesotibiae and metatibiae and their spurs unmodified. Metatarsomere 1 and spur almost equal in lenth, shorter than tarsomeres 2–5 combined.</paragraph>
<paragraph id="8BF30A5BE833FFC5AAB4F972FE40F8D9" blockId="23.[151,1437,258,1975]" box="[199,410,1770,1795]" pageId="23" pageNumber="324">
Aedeagus,
<figureCitation id="137716DEE833FFC5AB30F972FE4FF8D9" box="[323,405,1770,1795]" captionStart="FIGURES 47 – 55" captionStartId="22.[151,264,1855,1878]" captionTargetBox="[154,1435,291,1792]" captionTargetId="figure@22.[151,1436,267,1834]" captionTargetPageId="22" captionText="FIGURES 47 – 55. Sulawesi Onthophagu s body parts, male, female. 47 – 52, O. sopu, all Sopu, paratype male (47, 51), holotype male (48 – 50), paratype female (52). 53 – 55, O. manguliensis, male Mangoli (53), male Matanyo (55), female Matanyo (54). — 47, 52, 53 - 54, head full-face, 48, left elytron dorsal, 49, protibial upper side, 50, metatibial upperside, 51, 55, aedeagus lateral. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345764/files/figure.png" pageId="23" pageNumber="324">Fig. 55</figureCitation>
.
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAB4F896FAD5F891" blockId="23.[151,1437,258,1975]" pageId="23" pageNumber="324">
<emphasis id="B938D649E833FFC5AAB4F896FD1BF8FD" bold="true" box="[199,705,1806,1831]" pageId="23" pageNumber="324">
Variation, sexual dimorphism,
<typeStatus id="54F7B4F9E833FFC5A84AF896FDB7F8FD" box="[569,621,1806,1831]" pageId="23" pageNumber="324">type</typeStatus>
status.
</emphasis>
Size slightly variable—Boucomont (1914: 23) mentioned “3 ½– 4 ½ mm”. Colour pattern also only slightly variable.
<emphasis id="B938D649E833FFC5A894F8ABFCE1F891" box="[743,827,1843,1867]" italics="true" pageId="23" pageNumber="324">Female</emphasis>
similar to male, as shown in
<figureCitation id="137716DEE833FFC5AEF7F8AAFAD6F891" box="[1156,1292,1842,1867]" captionStart="FIGURES 47 – 55" captionStartId="22.[151,264,1855,1878]" captionTargetBox="[154,1435,291,1792]" captionTargetId="figure@22.[151,1436,267,1834]" captionTargetPageId="22" captionText="FIGURES 47 – 55. Sulawesi Onthophagu s body parts, male, female. 47 – 52, O. sopu, all Sopu, paratype male (47, 51), holotype male (48 – 50), paratype female (52). 53 – 55, O. manguliensis, male Mangoli (53), male Matanyo (55), female Matanyo (54). — 47, 52, 53 - 54, head full-face, 48, left elytron dorsal, 49, protibial upper side, 50, metatibial upperside, 51, 55, aedeagus lateral. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345764/files/figure.png" pageId="23" pageNumber="324">Figs. 53–54</figureCitation>
.
</paragraph>
<paragraph id="8BF30A5BE833FFC5AAB4F8CEFBB1F86D" blockId="23.[151,1437,258,1975]" pageId="23" pageNumber="324">
Clearly Boucomont had more specimens before him when he wrote the original description, but lacking details we prefer to maintain their
<typeStatus id="54F7B4F9E833FFC5ABAEF8E3FDEEF849" box="[477,564,1915,1939]" pageId="23" pageNumber="324" type="syntype">syntype</typeStatus>
(“cotype”) status. Especially since
<taxonomicName id="4C4C71D8E833FFC5A9A5F8E3FB4AF849" box="[982,1168,1915,1939]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="manguliensis">
<emphasis id="B938D649E833FFC5A9A5F8E3FC30F848" box="[982,1002,1915,1938]" italics="true" pageId="23" pageNumber="324">O</emphasis>
.
<emphasis id="B938D649E833FFC5A98FF8E3FB4AF849" box="[1020,1168,1915,1939]" italics="true" pageId="23" pageNumber="324">manguliensis</emphasis>
</taxonomicName>
-like material requiring study was recently reported from other Moluccan islands (Huijbregts &amp; Krikken 2012).
</paragraph>
</subSubSection>
</treatment>
</document>
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