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<document id="8F8B78F983614EBF25C9DDF2BE21BD67" ID-DOI="10.1206/3953.1" ID-ISSN="0003-0082" ID-Zenodo-Dep="5370347" approvalRequired="6" approvalRequired_for_textStreams="6" checkinTime="1630268857110" checkinUser="carolina" docAuthor="Long, Khuat Dang, Van Achterberg, Cornelis, Carpenter, James M. &amp; Oanh, Nguyen Thi" docDate="2020" docId="03D3F52DF604FFE3BDF8FE7FFE21F8FD" docLanguage="en" docName="N3953.pdf" docOrigin="American Museum Novitates 2020 (3953)" docSource="https://bioone.org/journals/american-museum-novitates/volume-2020/issue-3953/3953.1/Review-of-the-Genus-Wroughtonia-Cameron-1899-Hymenoptera-Braconidae-Helconinae/10.1206/3953.1.full" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.4:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="4" docTitle="Wroughtonia undulata Long &amp; Van Achterberg &amp; Carpenter &amp; Oanh 2020, sp. nov." docType="treatment" docVersion="4" lastPageNumber="41" masterDocId="FFEA8D55F622FFCBBC2CFFACFFC7FE19" masterDocTitle="Review of the genus Wroughtonia Cameron, 1899 (Hymenoptera, Braconidae, Helconinae), with the description of 12 new species from Vietnam" masterLastPageNumber="56" masterPageNumber="1" pageNumber="39" updateTime="1698468785609" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="48B1A81F9EB75B261968778B1CF44E30">Review of the genus Wroughtonia Cameron, 1899 (Hymenoptera, Braconidae, Helconinae), with the description of 12 new species from Vietnam</mods:title>
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<mods:namePart id="FA9A110F78A061D8727ED79CAB3208B5">Long, Khuat Dang</mods:namePart>
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<mods:namePart id="97278B61F40443F6C20B9A720292FC35">Van Achterberg, Cornelis</mods:namePart>
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<mods:namePart id="C6422BED4A282BD76E013DF24849C6A5">Carpenter, James M.</mods:namePart>
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<mods:namePart id="1239BE20F4E45E6C9AACA8C6CD713817">Oanh, Nguyen Thi</mods:namePart>
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<mods:url id="FED8D018ECC1CD9D89C7A77A6232A4FA">https://bioone.org/journals/american-museum-novitates/volume-2020/issue-3953/3953.1/Review-of-the-Genus-Wroughtonia-Cameron-1899-Hymenoptera-Braconidae-Helconinae/10.1206/3953.1.full</mods:url>
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<treatment id="03D3F52DF604FFE3BDF8FE7FFE21F8FD" ID-DOI="http://doi.org/10.5281/zenodo.5461937" ID-Zenodo-Dep="5461937" LSID="urn:lsid:plazi:treatment:03D3F52DF604FFE3BDF8FE7FFE21F8FD" httpUri="http://treatment.plazi.org/id/03D3F52DF604FFE3BDF8FE7FFE21F8FD" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
<subSubSection id="C36017B0F604FFEDBDF8FE7FFC96FFF6" box="[468,849,467,495]" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="8BC5443BF604FFEDBDF8FE7FFC96FFF6" blockId="38.[468,849,467,495]" box="[468,849,467,495]" pageId="38" pageNumber="39">
<heading id="D08DF357F604FFEDBDF8FE7FFC96FFF6" box="[468,849,467,495]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="39" reason="2">
<emphasis id="B90E9829F604FFEDBDF8FE7FFD37FFF7" bold="true" box="[468,752,467,494]" pageId="38" pageNumber="39">
<taxonomicName id="4C7A3FB8F604FFEDBDF8FE7FFD2DFFF7" authority="Long &amp; Van Achterberg &amp; Carpenter &amp; Oanh, 2020" authorityName="Long &amp; Van Achterberg &amp; Carpenter &amp; Oanh" authorityYear="2020" box="[468,746,467,494]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="undulata" status="sp. nov.">
<emphasis id="B90E9829F604FFEDBDF8FE7FFD2DFFF7" bold="true" box="[468,746,467,494]" italics="true" pageId="38" pageNumber="39">Wroughtonia undulata</emphasis>
</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A23D2552F604FFEDBED4FE79FC96FFF6" box="[760,849,469,495]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36017B0F604FFEDBE1EFDA0FD33FC3E" box="[562,756,524,551]" pageId="38" pageNumber="39" type="description">
<paragraph id="8BC5443BF604FFEDBE1EFDA0FD33FC3E" blockId="38.[562,756,524,551]" box="[562,756,524,551]" pageId="38" pageNumber="39">
<heading id="D08DF357F604FFEDBE1EFDA0FD33FC3E" box="[562,756,524,551]" centered="true" fontSize="36" level="3" pageId="38" pageNumber="39" reason="7">
<figureCitation id="134158BEF604FFEDBE1EFDA0FD33FC3E" box="[562,756,524,551]" captionStart-0="FIG" captionStart-1="FIGS" captionStartId-0="40.[112,154,1160,1181]" captionStartId-1="41.[108,162,1520,1541]" captionTargetBox-0="[207,1114,224,1133]" captionTargetBox-1="[185,1138,224,1492]" captionTargetId-0="figure-157@40.[207,1114,224,1133]" captionTargetId-1="figure-14@41.[502,1137,737,945]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionText-0="FIG. 116. Wroughtonia undulata, sp. nov., habitus (holotype, female, lateral)" captionText-1="FIGS. 117125. Wroughtonia undulata holotype, female. 117. Head, dorsal. 118. Head, frontal. 119. Head, lateral. 120. Mesoscutum. 121. Mesopleuron. 122. Hind femur, lateral. 123. Propodeum. 124. Metasoma. 125. Hind wing." figureDoi-0="http://doi.org/10.5281/zenodo.5370389" figureDoi-1="http://doi.org/10.5281/zenodo.5370391" httpUri-0="https://zenodo.org/record/5370389/files/figure.png" httpUri-1="https://zenodo.org/record/5370391/files/figure.png" pageId="38" pageNumber="39">Figures 116125</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36017B0F604FFEDBCB0FDE9FE86FCD6" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="8BC5443BF604FFEDBCB0FDE9FE86FCD6" blockId="38.[107,1211,579,1727]" pageId="38" pageNumber="39">
<materialsCitation id="3B124E66F604FFEDBCB0FDE9FEF9FCD6" collectingDate="1998-05-18" collectionCode="IEBR, AMNH, VNMN" collectorName="K. Long. &amp; K. Long" country="Vietnam" elevation="900" latitude="18.366667" location="Huong Son" longLatPrecision="1275" longitude="106.21667" pageId="38" pageNumber="39" specimenCount="3" specimenCount-female="3" stateProvince="Ha Tinh" typeStatus="holotype">
TYPE MATERIAL:
<typeStatus id="54C1FA99F604FFEDBD5EFDE8FE21FC47" box="[370,486,580,606]" pageId="38" pageNumber="39">Holotype</typeStatus>
,
<specimenCount id="9D7C8FB2F604FFEDBDD8FDEFFDCBFC47" box="[500,524,579,606]" pageId="38" pageNumber="39" type="female"></specimenCount>
, “Hel.065” (
<collectionCode id="ED6BDCFEF604FFEDBE89FDE8FD2BFC47" box="[677,748,580,606]" pageId="38" pageNumber="39">IEBR</collectionCode>
), NC
<collectingCountry id="F36D04ABF604FFEDBF1AFDE8FC65FC47" box="[822,930,580,606]" name="Vietnam" pageId="38" pageNumber="39">Vietnam</collectingCountry>
:
<collectingRegion id="49BE8AD9F604FFEDBF83FDE9FBDDFC47" box="[943,1050,580,607]" country="Vietnam" name="Ha Tinh" pageId="38" pageNumber="39">Ha Tinh</collectingRegion>
,
<location id="8EA512E0F604FFEDB804FDE9FB72FC47" LSID="urn:lsid:plazi:treatment:03D3F52DF604FFE3BDF8FE7FFE21F8FD:8EA512E0F604FFEDB804FDE9FB72FC47" box="[1064,1205,580,607]" country="Vietnam" latitude="18.366667" longLatPrecision="1275" longitude="106.21667" name="Huong Son" pageId="38" pageNumber="39" stateProvince="Ha Tinh">Huong Son</location>
,
<geoCoordinate id="EE4E22FCF604FFEDBC41FDC5FF17FC9D" box="[109,208,617,644]" degrees="18" direction="north" minutes="22" orientation="latitude" pageId="38" pageNumber="39" precision="925" value="18.366667">18°22N</geoCoordinate>
<geoCoordinate id="EE4E22FCF604FFEDBCF7FDC5FE8BFC9D" box="[219,332,617,644]" degrees="106" direction="east" minutes="13" orientation="longitude" pageId="38" pageNumber="39" precision="925" value="106.21667">106°13E</geoCoordinate>
,
<quantity id="4C82E9DEF604FFEDBD70FDC5FE6BFC9D" box="[348,428,617,644]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="38" pageNumber="39" unit="m" value="900.0">
<elevation id="0057A308F604FFEDBD70FDC5FE6BFC9D" box="[348,428,617,644]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="38" pageNumber="39" unit="m" value="900.0">900 m</elevation>
</quantity>
,
<date id="FFC462FBF604FFEDBD90FDC6FDA0FC9A" box="[444,615,617,644]" pageId="38" pageNumber="39" value="1998-05-18">
<collectingDate id="EF809B13F604FFEDBD90FDC6FDA0FC9A" box="[444,615,617,644]" pageId="38" pageNumber="39" value="1998-05-18">May 18, 1998</collectingDate>
</date>
,
<collectingMethod id="523B3C2CF604FFEDBE5AFDC5FCD6FC9D" box="[630,785,617,644]" pageId="38" pageNumber="39">Malaise trap</collectingMethod>
, AMNH,
<collectorName id="268F21EDF604FFEDBFA3FDC6FC3DFC9D" box="[911,1018,618,644]" pageId="38" pageNumber="39">K. Long.</collectorName>
<typeStatus id="54C1FA99F604FFEDB829FDC6FBB9FC9D" box="[1029,1150,618,644]" pageId="38" pageNumber="39">Paratypes</typeStatus>
:
<specimenCount id="9D7C8FB2F604FFEDB8A0FDC5FB74FC9D" box="[1164,1203,617,644]" pageId="38" pageNumber="39" type="female">2♀</specimenCount>
, “Hel.077” (
<collectionCode id="ED6BDCFEF604FFEDBCD5FD23FE99FCB0" box="[249,350,655,681]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="38" pageNumber="39">AMNH</collectionCode>
), same place; “Hel.078” (
<collectionCode id="ED6BDCFEF604FFEDBE87FD23FCD7FCB0" box="[683,784,655,681]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="38" pageNumber="39">VNMN</collectionCode>
), same place, but
<date id="FFC462FBF604FFEDB82FFD23FB71FCB0" box="[1027,1206,655,681]" pageId="38" pageNumber="39" value="1998-05-18">
<collectingDate id="EF809B13F604FFEDB82FFD23FB71FCB0" box="[1027,1206,655,681]" pageId="38" pageNumber="39" value="1998-05-18">May 18, 1998</collectingDate>
</date>
, AMNH,
<collectorName id="268F21EDF604FFEDBCF4FD19FEF9FCD6" box="[216,318,693,719]" pageId="38" pageNumber="39">K. Long</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36017B0F604FFEDBCB7FD75FCC0FCEA" box="[155,775,729,756]" pageId="38" pageNumber="39" type="distribution">
<paragraph id="8BC5443BF604FFEDBCB7FD75FCC0FCEA" blockId="38.[107,1211,579,1727]" box="[155,775,729,756]" pageId="38" pageNumber="39">
DISTRIBUTION: NC
<collectingCountry id="F36D04ABF604FFEDBDAAFD75FE28FCEA" box="[390,495,729,755]" name="Vietnam" pageId="38" pageNumber="39">Vietnam</collectingCountry>
:
<collectingRegion id="49BE8AD9F604FFEDBDD6FD76FD9AFCEA" box="[506,605,729,756]" country="Vietnam" name="Ha Tinh" pageId="38" pageNumber="39">Ha Tinh</collectingRegion>
(Huong Son).
</paragraph>
</subSubSection>
<subSubSection id="C36017B0F604FFEDBCB7FD53FE5EFD00" box="[155,409,767,793]" pageId="38" pageNumber="39" type="biology_ecology">
<paragraph id="8BC5443BF604FFEDBCB7FD53FE5EFD00" blockId="38.[107,1211,579,1727]" box="[155,409,767,793]" pageId="38" pageNumber="39">BIOLOGY: Unknown.</paragraph>
</subSubSection>
<subSubSection id="C36017B0F604FFECBCB7FC88FC55FF38" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="8BC5443BF604FFEDBCB7FC88FD29F830" blockId="38.[107,1211,579,1727]" pageId="38" pageNumber="39">
DIAGNOSIS: Antenna with 3435 segments, antennal segments 1023 cream white; maxillary palp 1.6
<emphasis id="B90E9829F604FFEDBD64FCE5FE9EFD7A" bold="true" box="[328,345,841,867]" pageId="38" pageNumber="39">×</emphasis>
as long as head; in frontal view height of eye 2.2
<emphasis id="B90E9829F604FFEDBF87FCE6FC7AFD7D" bold="true" box="[939,957,842,868]" italics="true" pageId="38" pageNumber="39">×</emphasis>
its transverse width; width of face 2.2
<emphasis id="B90E9829F604FFEDBD1AFCC3FE8FFD90" bold="true" box="[310,328,879,905]" italics="true" pageId="38" pageNumber="39">×</emphasis>
length of face and clypeus combined; frontal protuberance lamelliform, lower than level of lateral carina of frons; in dorsal view width of head 1.6
<emphasis id="B90E9829F604FFEDB830FC38FBEAFDB7" bold="true" box="[1052,1069,916,942]" pageId="38" pageNumber="39">×</emphasis>
its median length; height of eye 1.45
<emphasis id="B90E9829F604FFEDBD88FC15FE72FDCA" bold="true" box="[420,437,953,979]" pageId="38" pageNumber="39">×</emphasis>
temple; in lateral view height of eye 1.3
<emphasis id="B90E9829F604FFEDBF8BFC15FC7FFDCA" bold="true" box="[935,952,953,979]" pageId="38" pageNumber="39">×</emphasis>
its transverse width; transverse width of eye 1.4
<emphasis id="B90E9829F604FFEDBD92FC73FE08FDE0" bold="true" box="[446,463,991,1017]" pageId="38" pageNumber="39">×</emphasis>
temple; face coarsely rugose; lateral area between antennal sockets and eye margin with transverse rugosities; clypeus rugose-punctate, malar space coarsely rugose.
<taxonomicName id="4C7A3FB8F604FFEDBD14FB86FE7DFA5D" box="[312,442,1066,1092]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="38" pageNumber="47" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
1.65
<emphasis id="B90E9829F604FFEDBDDBFB85FDCFFA5A" bold="true" box="[503,520,1065,1091]" pageId="38" pageNumber="39">×</emphasis>
longer than high; notauli wide, sparsely crenulate anteriorly, fused posteriorly with two parallel rugosities in large longitudinally rugose area close to scutellar sulcus; scutellar sulcus wide, 0.7
<emphasis id="B90E9829F604FFEDBEE1FBD8FD19FA97" bold="true" box="[717,734,1140,1166]" pageId="38" pageNumber="39">×</emphasis>
as long as scutellum; propodeum with basal median carina and large areola; propodeal areola nearly smooth, coriaceous; lateral areas of propodeum with transverse rugosities. Length of forewing 2.4
<emphasis id="B90E9829F604FFEDBFDAFB13FBC0FAC0" bold="true" box="[1014,1031,1215,1241]" pageId="38" pageNumber="39">×</emphasis>
its maximum width; vein 3-SR short, 0.5
<emphasis id="B90E9829F604FFEDBD9EFB48FE04FAE7" bold="true" box="[434,451,1252,1278]" pageId="38" pageNumber="39">×</emphasis>
vein r; vein 2-M 3.4
<emphasis id="B90E9829F604FFEDBE93FB48FD17FAE7" bold="true" box="[703,720,1252,1278]" pageId="38" pageNumber="39">×</emphasis>
3-SR; hind wing broad, vein 1-M short, 0.3
<emphasis id="B90E9829F604FFEDBCA3FAA5FF67FB3A" bold="true" box="[143,160,1289,1315]" pageId="38" pageNumber="39">×</emphasis>
vein 1r-m; vein SR of hind wing sinuate; vein 2-SC+R transverse; hind wing with 4 hamuli. Foretarsus 1.3
<emphasis id="B90E9829F604FFEDBD56FA83FE4CFB50" bold="true" box="[378,395,1327,1353]" pageId="38" pageNumber="39">×</emphasis>
as long as foretibia; hind femur robust, with ventral serrations and stout tooth-shaped protuberance; length of hind femur (without protuberance and serrations) 3.2
<emphasis id="B90E9829F604FFEDBCCCFAD5FF36FB8A" bold="true" box="[224,241,1401,1427]" pageId="38" pageNumber="39">×</emphasis>
its maximum width; hind coxa sparsely punctate laterally; hind femur reticulate-rugulose laterally. First metasomal tergite 1.4
<emphasis id="B90E9829F604FFEDBEE1FA33FD19FBA0" bold="true" box="[717,734,1439,1465]" pageId="38" pageNumber="39">×</emphasis>
longer than its apical width, with long dorsal carinae almost up to apex of tergite; first tergite largely smooth, somewhat coriaceous; median length of second tergite 0.7
<emphasis id="B90E9829F604FFEDBE43FA45FD47F81A" bold="true" box="[623,640,1513,1539]" pageId="38" pageNumber="39">×</emphasis>
its apical width, and 1.1
<emphasis id="B90E9829F604FFEDBF86FA45FC7CF81A" bold="true" box="[938,955,1513,1539]" pageId="38" pageNumber="39">×</emphasis>
third tergite; second tergite smooth, with sparse large punctures apically.
</paragraph>
<paragraph id="8BC5443BF604FFECBCB7F998FC55FF38" blockId="38.[107,1211,579,1727]" lastBlockId="39.[107,1214,225,1745]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
NOTES:
<emphasis id="B90E9829F604FFEDBCD3F999FE67F857" box="[255,416,1588,1614]" italics="true" pageId="38" pageNumber="39">
<taxonomicName id="4C7A3FB8F604FFEDBCD3F999FE5BF857" authority="Long &amp; Van Achterberg &amp; Carpenter &amp; Oanh, 2020" authorityName="Long &amp; Van Achterberg &amp; Carpenter &amp; Oanh" authorityYear="2020" box="[255,412,1588,1614]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="undulata" status="sp. nov.">W. undulata</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A23D2552F604FFEDBD80F999FDC9F856" box="[428,526,1589,1615]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
, runs to
<taxonomicName id="4C7A3FB8F604FFEDBEA8F999FBA9F857" authority="(Chou and Hsu, 1998)" baseAuthorityName="Chou and Hsu" baseAuthorityYear="1998" box="[644,1134,1588,1615]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pterolophiae">
<emphasis id="B90E9829F604FFEDBEA8F999FC82F857" box="[644,837,1588,1614]" italics="true" pageId="38" pageNumber="39">W. pterolophiae</emphasis>
(
<bibRefCitation id="EFEB39CAF604FFEDBF74F998FBA2F857" author="Chou, L. Y. &amp; T. C. Hsu" box="[856,1125,1588,1615]" pageId="38" pageNumber="39" pagination="283 - 314" refId="ref27076" refString="Chou, L. Y., and T. C. Hsu. 1998. The Braconidae (Hymenoptera) of Taiwan 8. Brulleiini, Diospilini and Helconini. Journal of Agricultural Research of China 47 (3): 283 - 314." type="journal article" year="1998">Chou and Hsu, 1998</bibRefCitation>
)
</taxonomicName>
, from
<collectingCountry id="F36D04ABF604FFEDBC47F9F5FF72F86A" box="[107,181,1625,1651]" name="China" pageId="38" pageNumber="39">China</collectingCountry>
(couplet 20a) in the key by
<bibRefCitation id="EFEB39CAF604FFEDBE3AF9F6FD23F86A" author="Yan, C. - J. &amp; C. van Achterberg &amp; J. - H. He &amp; X. - X. Chen" box="[534,740,1625,1652]" pageId="38" pageNumber="39" pagination="401 - 457" refId="ref27775" refString="Yan, C. - J., C. van Achterberg, J. - H. He, and X. - X. Chen. 2017. Review of the tribe Helconini Foerster s. s. from China, with the description of 18 new species. Zootaxa 4291 (3): 401 - 457." type="journal article" year="2017">Yan et al. (2017)</bibRefCitation>
, but differs as follows: (1) first tergite largely smooth or coriaceous with sparse punctures (smooth to moderately foveolate-punctate in
<taxonomicName id="4C7A3FB8F604FFEDBCE8F909FE45F8A7" baseAuthorityName="Chou and Hsu" baseAuthorityYear="1998" box="[196,386,1700,1726]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pterolophiae">
<emphasis id="B90E9829F604FFEDBCE8F909FE45F8A7" box="[196,386,1700,1726]" italics="true" pageId="38" pageNumber="39">W. pterolophiae</emphasis>
</taxonomicName>
); (2) hind tarsus whitish yellow (fourth and fifth tarsal hind segments brown, darker than third segment in
<taxonomicName id="4C7A3FB8F605FFECBEA9FF4EFC87FEE2" baseAuthorityName="Chou and Hsu" baseAuthorityYear="1998" box="[645,832,225,251]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="pterolophiae">
<emphasis id="B90E9829F605FFECBEA9FF4EFC87FEE2" box="[645,832,225,251]" italics="true" pageId="39" pageNumber="40">W. pterolophiae</emphasis>
</taxonomicName>
); and (3) vein SR of hind wing sinuate (vein SR of hind wing slightly curved in
<taxonomicName id="4C7A3FB8F605FFECBEEAFEABFC46FF38" baseAuthorityName="Chou and Hsu" baseAuthorityYear="1998" box="[710,897,263,289]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="pterolophiae">
<emphasis id="B90E9829F605FFECBEEAFEABFC46FF38" box="[710,897,263,289]" italics="true" pageId="39" pageNumber="40">W. pterolophiae</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36017B0F605FFE3BCB7FE80FEA7F830" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="description">
<paragraph id="8BC5443BF605FFECBCB7FE80FE63FF75" blockId="39.[107,1214,225,1745]" pageId="39" pageNumber="40">
DESCRIPTION:
<typeStatus id="54C1FA99F605FFECBD7CFE80FE03FF5F" box="[336,452,300,326]" pageId="39" pageNumber="40">Holotype</typeStatus>
,
<specimenCount id="9D7C8FB2F605FFECBDFEFE87FE2DFF5F" box="[466,490,299,326]" pageId="39" pageNumber="40" type="female"></specimenCount>
, body length
<quantity id="4C82E9DEF605FFECBEBBFE80FD30FF5E" box="[663,759,300,327]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="39" pageNumber="40" unit="mm" value="9.7">9.7 mm</quantity>
, forewing length 8.0 mm, ovipositor sheath
<quantity id="4C82E9DEF605FFECBCE9FEFDFEE4FF75" box="[197,291,337,364]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="39" pageNumber="40" unit="mm" value="4.5">4.5 mm</quantity>
(fig. 116).
</paragraph>
<paragraph id="8BC5443BF605FFECBCB7FEDBFC03FD35" blockId="39.[107,1214,225,1745]" pageId="39" pageNumber="40">
<emphasis id="B90E9829F605FFECBCB7FEDBFF18FF88" box="[155,223,375,401]" italics="true" pageId="39" pageNumber="40">Head:</emphasis>
Antenna with 35 segments; third antennal segment 1.1
<emphasis id="B90E9829F605FFECBF5DFEDBFC45FF88" bold="true" box="[881,898,375,401]" pageId="39" pageNumber="40">×</emphasis>
as long as fourth segment (18:16); in frontal view height of eye 2.2
<emphasis id="B90E9829F605FFECBE15FE30FD8DFFAF" bold="true" box="[569,586,412,438]" pageId="39" pageNumber="40">×</emphasis>
its transverse width (40:18); width of face 2.2
<emphasis id="B90E9829F605FFECB875FE30FBADFFAF" bold="true" box="[1113,1130,412,438]" pageId="39" pageNumber="40">×</emphasis>
length of face and clypeus combined (53:24); malar space as long as mandible width (15:15) and 0.4
<emphasis id="B90E9829F605FFECB886FE6DFB7CFFC2" bold="true" box="[1194,1211,449,475]" pageId="39" pageNumber="40">×</emphasis>
height of eye (15:40) (fig. 118); maxillary palp 1.6
<emphasis id="B90E9829F605FFECBE9BFE4BFD0FFC18" bold="true" box="[695,712,487,513]" pageId="39" pageNumber="40">×</emphasis>
as long as head (51:31); tentorial pits distinct, distance between tentorial pits 0.9
<emphasis id="B90E9829F605FFECBE6EFDA0FD94FC3F" bold="true" box="[578,595,524,550]" pageId="39" pageNumber="40">×</emphasis>
distance between pit and eye (17:19); frontal protuberance lamelliform, lower than level of lateral carina of frons; in dorsal view width of head 1.6
<emphasis id="B90E9829F605FFECBCA1FDFBFF59FC68" bold="true" box="[141,158,599,625]" pageId="39" pageNumber="40">×</emphasis>
its median length (87:54); height of eye 1.45
<emphasis id="B90E9829F605FFECBE8FFDFBFD73FC68" bold="true" box="[675,692,599,625]" pageId="39" pageNumber="40">×</emphasis>
as long as temple (29:20); ocelli rather large, POL:OD:OOL = 10:7:20; distance between front and hind ocelli 0.25
<emphasis id="B90E9829F605FFECBFB1FDD0FC69FC8F" bold="true" box="[925,942,636,662]" pageId="39" pageNumber="40">×</emphasis>
OOL (5:20) (fig. 117); in lateral view height of eye 1.3
<emphasis id="B90E9829F605FFECBDC0FD0DFE3AFCA2" bold="true" box="[492,509,673,699]" pageId="39" pageNumber="40">×</emphasis>
its transverse width (39:29); transverse width of eye 1.2
<emphasis id="B90E9829F605FFECB886FD0DFB7CFCA2" bold="true" box="[1194,1211,673,699]" pageId="39" pageNumber="40">×</emphasis>
temple (29:24) (fig. 119); face coarsely rugose; lateral area between antennal sockets and eye margin with transverse rugosities; clypeus rugose-punctate, malar space coarsely rugose; frons smooth; vertex and temple almost smooth with scattered fine punctures.
</paragraph>
<paragraph id="8BC5443BF605FFECBCB7FC9BFEA4FA62" blockId="39.[107,1214,225,1745]" pageId="39" pageNumber="40">
<taxonomicName id="4C7A3FB8F605FFECBCB7FC9BFE62FD48" authority=": Mesosoma" authorityName="Mesosoma" box="[155,421,823,849]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B90E9829F605FFECBCB7FC9BFEDCFD49" box="[155,283,823,848]" italics="true" pageId="39" pageNumber="40">Mesosoma:</emphasis>
Mesosoma
</taxonomicName>
1.65
<emphasis id="B90E9829F605FFECBDCDFC9BFE35FD48" bold="true" box="[481,498,823,849]" pageId="39" pageNumber="40">×</emphasis>
longer than high (48:29) (fig. 121); pronotal side coarsely rugose, densely punctate dorsally; mesopleuron smooth medially, punctate-reticulate ventrally; metapleuron areolate-rugulose; median lobe of mesoscutum sparsely punctate, reticulate-rugulose posteriorly; lateral lobes of mesoscutum almost smooth; notauli wide, sparsely crenulate anteriorly, fused posteriorly with two parallel rugosities in large longitudinally rugose area close to scutellar sulcus (fig. 120); scutellar sulcus wide, with 3 carinae, two lateral carinae sinuate, 0.7
<emphasis id="B90E9829F605FFECBCA2FBBBFF58FA28" bold="true" box="[142,159,1047,1073]" pageId="39" pageNumber="40">×</emphasis>
as long as scutellum (15:21); scutellum sparsely punctate; propodeum with mediobasal carina and large areola (fig. 123); areola smooth or coriaceous; lateral areas of propodeum with transverse rugosities.
</paragraph>
<paragraph id="8BC5443BF605FFECBCB7FB2BFE1DFB2F" blockId="39.[107,1214,225,1745]" pageId="39" pageNumber="40">
<emphasis id="B90E9829F605FFECBCB7FB2BFF2EFAB9" box="[155,233,1159,1184]" italics="true" pageId="39" pageNumber="40">Wings:</emphasis>
Forewing: length of forewing 2.4
<emphasis id="B90E9829F605FFECBE59FB2BFD41FAB8" bold="true" box="[629,646,1159,1185]" pageId="39" pageNumber="40">×</emphasis>
its maximum width (120:49); pterostigma 3.4
<emphasis id="B90E9829F605FFECB886FB2BFB7CFAB8" bold="true" box="[1194,1211,1159,1185]" pageId="39" pageNumber="40">×</emphasis>
longer than wide (41:12); vein 3-SR short, 0.5
<emphasis id="B90E9829F605FFECBEBEFB00FD64FADF" bold="true" box="[658,675,1196,1222]" pageId="39" pageNumber="40">×</emphasis>
vein r; r:2-SR:3-SR:SR1:r-m = 15:18:8:60:12; 1-CU1:cu-a:2-CU1 = 4:13:21; second submarginal cell trapezium-shaped, vein 2-M 3.4
<emphasis id="B90E9829F605FFECB846FB7DFBBCFAF2" bold="true" box="[1130,1147,1233,1259]" pageId="39" pageNumber="40">×</emphasis>
3-SR (27:8); hind wing broad, vein 1-M short, 0.3
<emphasis id="B90E9829F605FFECBE51FB5BFD49FB08" bold="true" box="[637,654,1271,1297]" pageId="39" pageNumber="40">×</emphasis>
vein 1r-m (3:10); vein 2-SC+R transverse (fig. 125); hind wing with 4 hamuli.
</paragraph>
<paragraph id="8BC5443BF605FFECBCB0FAEEFF21F825" blockId="39.[107,1214,225,1745]" pageId="39" pageNumber="40">
<emphasis id="B90E9829F605FFECBCB0FAEEFF12FB42" box="[156,213,1346,1371]" italics="true" pageId="39" pageNumber="40">Legs:</emphasis>
Foretarsus 1.3
<emphasis id="B90E9829F605FFECBDAAFAEDFE50FB42" bold="true" box="[390,407,1345,1371]" pageId="39" pageNumber="40">×</emphasis>
longer than foretibia (45:35); hind femur robust, with ventral serrations and stout tooth-shaped protuberance (fig. 122); length of hind femur (without protuberance or serrations), tibia, and basitarsus 3.2 (57:18), 9.9 (89:9), and 7.25
<emphasis id="B90E9829F605FFECB82DFA20FBD5FBBF" bold="true" box="[1025,1042,1420,1446]" pageId="39" pageNumber="40">×</emphasis>
(29:4) as long as their maximum width, respectively; hind coxa sparsely punctate laterally; hind femur reticulate-rugulose laterally; hind basitarsus 0.3
<emphasis id="B90E9829F605FFECBE81FA7BFD79FBE8" bold="true" box="[685,702,1495,1521]" pageId="39" pageNumber="40">×</emphasis>
as long as hind tibia (29:89), and 0.7
<emphasis id="B90E9829F605FFECB8A4FA7BFB5EFBE8" bold="true" box="[1160,1177,1495,1521]" pageId="39" pageNumber="40">×</emphasis>
as long as hind tarsal segments 25 (29:41); fourth hind tarsal segment 0.4
<emphasis id="B90E9829F605FFECBFE1FA50FC19F80F" bold="true" box="[973,990,1532,1558]" pageId="39" pageNumber="40">×</emphasis>
as long as telotarsus (5:14).
</paragraph>
<paragraph id="8BC5443BF605FFE3BCB0F9EBFE2CFAE6" blockId="39.[107,1214,225,1745]" lastBlockId="40.[107,1212,1215,1764]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<emphasis id="B90E9829F605FFECBCB0F9EBFEDCF879" box="[156,283,1607,1632]" italics="true" pageId="39" pageNumber="40">Metasoma:</emphasis>
Metasoma 0.7
<emphasis id="B90E9829F605FFECBDE4F9EBFE1EF878" bold="true" box="[456,473,1607,1633]" pageId="39" pageNumber="40">×</emphasis>
as long as head and mesosoma combined (45:63); first tergite 1.4
<emphasis id="B90E9829F605FFECBCA2F9C0FF58F89F" bold="true" box="[142,159,1644,1670]" pageId="39" pageNumber="40">×</emphasis>
longer than its apical width (55:40), with long dorsal carinae almost up to apex of tergite, converging apically (fig. 124); first tergite almost smooth or coriaceous (fig. 124); median length of second tergite 0.7
<emphasis id="B90E9829F605FFECBD99F91BFE01F8C8" bold="true" box="[437,454,1719,1745]" pageId="39" pageNumber="40">×</emphasis>
its apical width (28:42), and 1.1
<emphasis id="B90E9829F605FFECBF63F91BFCA7F8C8" bold="true" box="[847,864,1719,1745]" pageId="39" pageNumber="40">×</emphasis>
third tergite (28:25); second tergite smooth, with sparse large punctures apically; third tergite smooth; ovipositor sheath 0.6
<emphasis id="B90E9829F60AFFE3BCA2FB49FF58FAE6" bold="true" box="[142,159,1253,1279]" pageId="40" pageNumber="41">×</emphasis>
as long as forewing (45:80).
</paragraph>
<caption id="DF0514B3F60AFFE3BC5CFB24FCB3FA87" ID-DOI="http://doi.org/10.5281/zenodo.5370389" ID-Zenodo-Dep="5370389" box="[112,884,1158,1182]" httpUri="https://zenodo.org/record/5370389/files/figure.png" pageId="40" pageNumber="41" startId="40.[112,154,1160,1181]" targetBox="[207,1114,224,1133]" targetPageId="40">
<paragraph id="8BC5443BF60AFFE3BC5CFB24FCB3FA87" blockId="40.[112,884,1158,1182]" box="[112,884,1158,1182]" pageId="40" pageNumber="41">
FIG. 116.
<taxonomicName id="4C7A3FB8F60AFFE3BCF9FB2AFE7FFA87" authority="Long &amp; Van Achterberg &amp; Carpenter &amp; Oanh, 2020" authorityName="Long &amp; Van Achterberg &amp; Carpenter &amp; Oanh" authorityYear="2020" box="[213,440,1158,1182]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="undulata" status="sp. nov.">
<emphasis id="B90E9829F60AFFE3BCF9FB2AFE7FFA87" box="[213,440,1158,1182]" italics="true" pageId="40" pageNumber="41">Wroughtonia undulata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A23D2552F60AFFE3BDE9FB2BFDD3FA87" box="[453,532,1159,1182]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
, habitus (holotype, female, lateral)
</paragraph>
</caption>
<paragraph id="8BC5443BF60AFFE3BCB0FAA6FE2BF81D" blockId="40.[107,1212,1215,1764]" pageId="40" pageNumber="41">
<emphasis id="B90E9829F60AFFE3BCB0FAA6FF23FB3D" box="[156,228,1290,1316]" italics="true" pageId="40" pageNumber="41">Color:</emphasis>
Body dark brown; scapus light brown; antenna brown, with 11th23rd antennal segments cream white; palpi yellow; fore- and middle legs yellow, but tibia pale yellow, tarsus whitish yellow; hind coxa, trochanter, and trochantellus yellow; hind femur yellow basally and ventrally, brown dorsoapically; hind tibia yellow basally, infuscate medially, brown apically; hind tarsus whitish yellow; pterostigma and wing veins brown; wing membrane yellow; first metasomal tergite yellow; second and third tergites dark brown; remainder of metasoma light brown, ovipositor sheath brown.
</paragraph>
<paragraph id="8BC5443BF60AFFE3BCB7F9A3FEA7F830" blockId="40.[107,1212,1215,1764]" box="[155,352,1551,1577]" pageId="40" pageNumber="41">
<emphasis id="B90E9829F60AFFE3BCB7F9A3FF1CF830" box="[155,219,1551,1577]" italics="true" pageId="40" pageNumber="41">Male:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36017B0F60AFFE3BCB7F999FF37F86D" pageId="40" pageNumber="41" type="etymology">
<paragraph id="8BC5443BF60AFFE3BCB7F999FF37F86D" blockId="40.[107,1212,1215,1764]" pageId="40" pageNumber="41">
ETYMOLOGY: From
<emphasis id="B90E9829F60AFFE3BDA7F998FDC3F857" box="[395,516,1588,1614]" italics="true" pageId="40" pageNumber="41">undulatus</emphasis>
(Latin for “wavy”), because of the sinuate vein SR of the hind wing.
</paragraph>
</subSubSection>
<subSubSection id="C36017B0F60AFFE3BCB1F9D3FE21F8FD" pageId="40" pageNumber="41" type="description">
<paragraph id="8BC5443BF60AFFE3BCB1F9D3FE21F8FD" blockId="40.[107,1212,1215,1764]" pageId="40" pageNumber="41">
VARIATION: Female
<typeStatus id="54C1FA99F60AFFE3BDA3F92CFDCFF883" box="[399,520,1664,1690]" pageId="40" pageNumber="41">paratypes</typeStatus>
: antenna with 3435 segments and 10th22nd segments cream white; body length 8.0
<quantity id="4C82E9DEF60AFFE3BDC2F908FD97F8A6" box="[494,592,1700,1727]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="40" pageNumber="41" unit="mm" value="8.5">8.5 mm</quantity>
; forewing
<quantity id="4C82E9DEF60AFFE3BEF5F908FCB7F8A6" box="[729,880,1700,1727]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" metricValueMax="6.8" metricValueMin="6.4" pageId="40" pageNumber="41" unit="mm" value="6.6" valueMax="6.8" valueMin="6.4">6.46.8 mm</quantity>
; ovipositor sheath
<quantity id="4C82E9DEF60AFFE3B84FF908FF59F8FD" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.6" metricValueMin="4.4" pageId="40" pageNumber="41" unit="mm" value="4.5" valueMax="4.6" valueMin="4.4">4.44.6 mm</quantity>
; hind wing with 3 hamuli.
</paragraph>
</subSubSection>
</treatment>
</document>