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<document id="18FC904CABE886A15B429B4047A9E5EC" ID-CLB-Dataset="29552" ID-DOI="10.11646/zootaxa.4483.1.3" ID-GBIF-Dataset="48be31ab-c94e-4044-bc82-751bd71fe938" ID-ISSN="1175-5326" ID-Zenodo-Dep="1436632" ID-ZooBank="85962DD1-E874-4273-8B1A-FB45A7B965A3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538071052148" checkinUser="plazi" docAuthor="Knutson, Lloyd V., Deeming, John C. &amp; Ebejer, Martin J." docDate="2018" docId="03D187E9FFB1406DBC873CD816AFFB92" docLanguage="en" docName="zootaxa.4483.1.3.pdf.imf" docOrigin="Zootaxa 4483 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sepedon (Mesosepedon) knutsoni Vala, Gbedjissi and Dossou 1994" docType="treatment" docVersion="10" lastPageNumber="90" masterDocId="FFE8FF91FFA7407ABC1038591210FF82" masterDocTitle="The Snail-killing Flies (Diptera: Sciomyzidae) of West Africa" masterLastPageNumber="100" masterPageNumber="67" pageNumber="89" updateTime="1698554682055" updateUser="ExternalLinkService">
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<mods:title id="C6EC26684DC894384BD9B902ED25B52D">The Snail-killing Flies (Diptera: Sciomyzidae) of West Africa</mods:title>
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<mods:namePart id="1A82F9EA30089863E54692C5C078C938">Knutson, Lloyd V.</mods:namePart>
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<mods:namePart id="18B7CAAC9328CCFD4E7C8E80FB3F249D">Deeming, John C.</mods:namePart>
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<mods:namePart id="A1ACDB1708CCD81730BAE71FE4D3795C">Ebejer, Martin J.</mods:namePart>
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<treatment id="03D187E9FFB1406DBC873CD816AFFB92" ID-DOI="http://doi.org/10.5281/zenodo.5981724" ID-GBIF-Taxon="148402985" ID-Zenodo-Dep="5981724" LSID="urn:lsid:plazi:treatment:03D187E9FFB1406DBC873CD816AFFB92" httpUri="http://treatment.plazi.org/id/03D187E9FFB1406DBC873CD816AFFB92" lastPageId="23" lastPageNumber="90" pageId="22" pageNumber="89">
<subSubSection id="C3626574FFB1406CBC873CD813B8FB3E" pageId="22" pageNumber="89" type="nomenclature">
<paragraph id="8BC736FFFFB1406CBC873CD811C6FB19" blockId="22.[151,982,1153,1213]" box="[151,982,1153,1179]" pageId="22" pageNumber="89">
<heading id="D08F8193FFB1406CBC873CD811C6FB19" bold="true" box="[151,982,1153,1179]" fontSize="11" level="1" pageId="22" pageNumber="89" reason="1">
<emphasis id="B90CEAEDFFB1406CBC873CD811C6FB19" bold="true" box="[151,982,1153,1179]" pageId="22" pageNumber="89">
<taxonomicName id="4C784D7CFFB1406CBC873CD811C2FB19" ID-CoL="8DVN3" authority="Vala, Gbedjissi and Dossou, 1994" authorityName="Vala, Gbedjissi and Dossou" authorityYear="1994" box="[151,978,1153,1179]" class="Insecta" family="Sciomyzidae" genus="Sepedon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="89" phylum="Arthropoda" rank="species" species="knutsoni" subGenus="Mesosepedon">
<emphasis id="B90CEAEDFFB1406CBC873CD812EEFB19" bold="true" box="[151,254,1153,1179]" italics="true" pageId="22" pageNumber="89">Sepedon</emphasis>
(
<emphasis id="B90CEAEDFFB1406CBD1E3CD813A1FB19" bold="true" box="[270,433,1153,1179]" italics="true" pageId="22" pageNumber="89">Mesosepedon</emphasis>
)
<emphasis id="B90CEAEDFFB1406CBDD23CD8103BFB19" bold="true" box="[450,555,1153,1179]" italics="true" pageId="22" pageNumber="89">knutsoni</emphasis>
<bibRefCitation id="EFE94B0EFFB1406CBE233CD811C2FB19" box="[563,978,1153,1179]" pageId="22" pageNumber="89" refId="ref26981" refString="Vala, J-C., Gasc, C., Gbedjissi, G. &amp; Dossou, C. (1994) Les Sciomyzidae du Benin, description de Sepedon (Mesosepedon) knutsoni n. sp. (Diptera). Bulletin de la Societe entomologique de France, 99 (5), 497 - 504." type="journal article">Vala, Gbedjissi and Dossou, 1994</bibRefCitation>
</taxonomicName>
.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC736FFFFB1406CBC873CFD13B8FB3E" blockId="22.[151,982,1153,1213]" box="[151,424,1188,1213]" pageId="22" pageNumber="89">
(Figs 2225,
<figureCitation id="13432A7AFFB1406CBD3B3CFD138EFB3E" box="[299,414,1188,1213]" captionStart="PLATE 8" captionStartId="27.[151,234,1915,1937]" captionTargetBox="[164,1422,826,1873]" captionTargetId="figure@27.[150,1436,826,1893]" captionTargetPageId="27" captionText="PLATE 8. Figs 3538, wings of Sciomyzidae: 35—Pherbellia (Graphomyzina) kivuana Verbeke, 36—Pteromicra zariae sp. n., 37—Sepedon (Mesosepedon) knutsoni Vala, Gbedjissi &amp; Dossou, 38—Sepedonella nana (Verbeke); 39—Sepedon (Mesosepedon) knutsoni, habitus, 40—Sepedon (Sepedomyia) nasuta (Verbeke) head, lateral." httpUri="https://zenodo.org/record/1436650/files/figure.png" pageId="22" pageNumber="89">plate 8c, e</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3626574FFB1406CBC873CB51170FA80" box="[151,864,1260,1282]" pageId="22" pageNumber="89" type="reference_group">
<paragraph id="8BC736FFFFB1406CBC873CB51170FA80" blockId="22.[151,864,1260,1282]" box="[151,864,1260,1282]" pageId="22" pageNumber="89">
References to figures:
<bibRefCitation id="EFE94B0EFFB1406CBD6B3CB5100AFA80" box="[379,538,1260,1282]" pageId="22" pageNumber="89" refId="ref26981" refString="Vala, J-C., Gasc, C., Gbedjissi, G. &amp; Dossou, C. (1994) Les Sciomyzidae du Benin, description de Sepedon (Mesosepedon) knutsoni n. sp. (Diptera). Bulletin de la Societe entomologique de France, 99 (5), 497 - 504." type="journal article">
Vala
<emphasis id="B90CEAEDFFB1406CBDBE3CB413CBFA80" box="[430,475,1260,1282]" italics="true" pageId="22" pageNumber="89">et al</emphasis>
. 1994
</bibRefCitation>
(head, postabdomen, genitalia).
</paragraph>
</subSubSection>
<subSubSection id="C3626574FFB1406DBC873D771322FC4A" lastPageId="23" lastPageNumber="90" pageId="22" pageNumber="89" type="materials_examined">
<paragraph id="8BC736FFFFB1406CBC873D7712F9FA55" blockId="22.[151,1437,1326,2035]" pageId="22" pageNumber="89">
<typeStatus id="54C3885DFFB1406CBC873D771314FAC5" box="[151,260,1326,1351]" pageId="22" pageNumber="89" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D7EFD76FFB1406CBD1D3D761334FAC4" box="[269,292,1327,1350]" pageId="22" pageNumber="89" type="male"></specimenCount>
,
<date id="FFC6103FFFB1406CBD203D7713BAFAC5" box="[304,426,1326,1351]" pageId="22" pageNumber="89" value="1991-06-10">10.vi.1991</date>
,
<collectingCountry id="F36F766FFFB1406CBDA63D77101EFAC5" box="[438,526,1326,1351]" name="Benin" pageId="22" pageNumber="89">
<collectionCode id="ED69AE3AFFB1406CBDA63D77101EFAC5" box="[438,526,1326,1351]" country="Benin" httpUri="http://grbio.org/cool/rx6h-yh4d" name="Universit<69> National du B<>nin" pageId="22" pageNumber="89">BÉNIN</collectionCode>
</collectingCountry>
, Agnavo, L.G. Gbedjissi (
<collectionCode id="ED69AE3AFFB1406CBF3E3D76118AFAC5" box="[814,922,1327,1351]" country="0" httpUri="http://grbio.org/cool/bmqw-2x87" name="Museo Nacional de Historia Natural del Paraguay" pageId="22" pageNumber="89">MNHNP</collectionCode>
). This species, first discovered by Deeming in
<collectingCountry id="F36F766FFFB1406CBC873D0B12FBFAE9" box="[151,235,1362,1387]" name="Nigeria" pageId="22" pageNumber="89">Nigeria</collectingCountry>
in 1979, was described by
<bibRefCitation id="EFE94B0EFFB1406CBE2C3D0B10EBFAE9" box="[572,763,1362,1387]" pageId="22" pageNumber="89" refId="ref26981" refString="Vala, J-C., Gasc, C., Gbedjissi, G. &amp; Dossou, C. (1994) Les Sciomyzidae du Benin, description de Sepedon (Mesosepedon) knutsoni n. sp. (Diptera). Bulletin de la Societe entomologique de France, 99 (5), 497 - 504." type="journal article">
Vala
<emphasis id="B90CEAEDFFB1406CBE6A3D0D10BFFAE9" box="[634,687,1362,1387]" italics="true" pageId="22" pageNumber="89">et al</emphasis>
. 1994
</bibRefCitation>
on the material they collected in the
<collectingCountry id="F36F766FFFB1406CB8D43D0B178CFAE9" box="[1220,1436,1362,1387]" name="Benin" pageId="22" pageNumber="89">Republic of Benin</collectingCountry>
(formerly
<collectingCountry id="F36F766FFFB1406CBD1E3D2F136DFA0D" box="[270,381,1398,1423]" name="Benin" pageId="22" pageNumber="89">Dahomey</collectingCountry>
) in
<date id="FFC6103FFFB1406CBDA33D2E1074FA0D" box="[435,612,1398,1423]" pageId="22" pageNumber="89" value="1991" valueMax="1993">1991 and 1993</date>
and some specimens collected by Deeming. Our original (previously unpublished) description presented below is based on
<typeStatus id="54C3885DFFB1406CBEEC3DC21177FA31" box="[764,871,1435,1459]" pageId="22" pageNumber="89" type="paratype">paratypes</typeStatus>
from
<collectingCountry id="F36F766FFFB1406CBFBE3DC31612FA31" box="[942,1026,1434,1459]" name="Nigeria" pageId="22" pageNumber="89">Nigeria</collectingCountry>
and complements that of
<bibRefCitation id="EFE94B0EFFB1406CB9393DC312F4FA55" pageId="22" pageNumber="89" refId="ref26981" refString="Vala, J-C., Gasc, C., Gbedjissi, G. &amp; Dossou, C. (1994) Les Sciomyzidae du Benin, description de Sepedon (Mesosepedon) knutsoni n. sp. (Diptera). Bulletin de la Societe entomologique de France, 99 (5), 497 - 504." type="journal article">
Vala
<emphasis id="B90CEAEDFFB1406CB9733DC51785FA31" box="[1379,1429,1434,1459]" italics="true" pageId="22" pageNumber="89">et al</emphasis>
. (1994)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BC736FFFFB1406DBCD73DBB13C8FCBA" blockId="22.[151,1437,1326,2035]" lastBlockId="23.[151,1437,151,1040]" lastPageId="23" lastPageNumber="90" pageId="22" pageNumber="89">
Body length 5.47.0mm, wing length
<quantity id="4C809B1AFFB1406CBE943DBB10C6FA79" box="[644,726,1506,1531]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="22" pageNumber="89" unit="mm" value="5.5">5.5mm</quantity>
. Head yellow; frons shiny except for very slightly yellowish pruinose frontal plates and velvety, yellow spot in front of orbital plate on either side, these spots touching eye margin and with 7 or 8 short black setulae; frons strongly concave; eye margin slightly indented at level of base of antenna; no fronto-orbital spot present, but frons narrowly but densely white pruinose from frontal spot to this indentation; ocellar triangle yellowish-white pruinose; vertex slightly brownish, rest of occiput yellowish; posterior portion of gena with dense silvery-white velvety area that extends along eye margin almost to base of antenna; anterior part of gena and face yellowish; gena and lower portion of face shiny, upper portion of face lightly whitish pruinose; 1 strong fronto-orbital; internal and external verticals strong, of equal size, no ocellar setae or setulae; postocellars slightly stronger than verticals; group of 4 or 5 moderately strong setae and 2 or 3 setulae on either side of cervix; gena with a few fine black setulae, these extending onto face along inner margin of silvery pruinosity beside eye to antennal base, rest of face bare. Scape yellow, in length slightly less than distance between bases of antennae; pedicel fuscous, 4 times length of first, postpedicel 2/3 times length of second, dark brown, but yellowish around base of arista; segments 1 and 2 and base of segment 3 of arista yellowish, rest of arista white, including the very short setulae. Palpus yellow. Thorax yellowish brown, a little whiter ventrally, pleura with dense white pruinosity on lower half, dorsum with faint brownish pattern consisting of a pair of narrow longitudinal vittae connected by broad brownish area beginning at posterior 2/3 and extending onto scutellum; transverse suture well developed, moderately strong ntpl, sa and pa setae, 1 pair weak prsc dc setae, row of short fine setulae extending from these last setae anteriorly to anterior declivity; sides of scutum with scattered fine black setulae; groups of several setulae including 1 somewhat longer in presutural position; pleura with scattered short black setulae, distinct row on ridge behind posterior spiracle; 3 minute setulae at apex of anterior portion of prosternum; no setae on katatergite callus; disc of scutellum flat, with a few fine black setulae. Haltere densely pruinose yellowish-white including stem, knob slightly infuscate, group of 7 or 8 short black setulae dorsally at base of knob. Wing membrane greyish, somewhat more yellowish anteriorly and slightly darker towards apex, with brownish veins and even distribution of microtrichiae; anal vein becoming evanescent beyond middle but distinctly reaching wing margin; alula narrow, about equal in width to second basal cell; posterior crossvein upright, not meeting fourth vein at right angles. Coxae yellow, each with 1 strong marginal seta and several smaller setae; middle femur yellow; hind femur yellowish brown; strong seta on anterior surface of mid femur at mid length with double row of very short, sparsely distributed spinules from near base to apex; all femora with double row of strong ventral spinules from basal 1/3 to apex; front tibia yellowish basally becoming brownish apically; fore tarsus unicolorous brown; mid and hind tarsi unicolorous brown. Abdominal tergites yellowish, posterior margins somewhat darker, uniformly covered with short black setulae, shiny and with very sparse pruinosity, base of abdomen dorsally lacking setulae except for patch of 1012 setulae in anterolateral corners; setulae along margin of second tergite twice as long as other setulae; sterna yellowish, reduced, each about 1/5 as wide as long. Female with cercus yellowish white.
</paragraph>
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<collectingRegion id="49BCF81DFFB0406DBCD73B1D1303FCDE" box="[199,275,836,860]" country="Tanzania" name="Mjini Magharibi" pageId="23" pageNumber="90">WEST</collectingRegion>
AFRICA
</collectionCode>
:
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: Zaria, Samaru,
<specimenCount id="9D7EFD76FFB0406DBED63B1D10FAFCDE" box="[710,746,836,860]" pageId="23" pageNumber="90" type="male">1♂</specimenCount>
,
<date id="FFC6103FFFB0406DBEE83B1D117CFCDF" box="[760,876,836,861]" pageId="23" pageNumber="90" value="1969-10-14">
<collectingDate id="EF82E9D7FFB0406DBEE83B1D117CFCDF" box="[760,876,836,861]" pageId="23" pageNumber="90" value="1969-10-14">14.x.1969</collectingDate>
</date>
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; 1♂,
<date id="FFC6103FFFB0406DBFBB3B1D1636FCDF" box="[939,1062,836,861]" pageId="23" pageNumber="90" value="1970-11-12">12.xi.1970</date>
; 1♀,
<date id="FFC6103FFFB0406DB8763B1D16CDFCDF" box="[1126,1245,836,861]" pageId="23" pageNumber="90" value="1966-06-18">18.vi.1966</date>
(USNM); Zaria, Dumbi Wood, pond, 1♂,
<date id="FFC6103FFFB0406DBDD53B3E102DFC02" box="[453,573,871,896]" pageId="23" pageNumber="90" value="1969-11-17">17.xi.1969</date>
(USNM); Zaria, Dumbi Wood, 2♀,
<date id="FFC6103FFFB0406DBFE23B3E1671FC02" box="[1010,1121,871,896]" pageId="23" pageNumber="90" value="1979-10-25">25.x.1979</date>
(NMWC);
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<collectorName id="268D5329FFB0406DB8FD3B311788FC02" box="[1261,1432,871,896]" pageId="23" pageNumber="90">Nr. Kafanchan</collectorName>
,
<location id="8EA76024FFB0406DBC873BD4132CFC26" LSID="urn:lsid:plazi:treatment:03D187E9FFB1406DBC873CD816AFFB92:8EA76024FFB0406DBC873BD4132CFC26" box="[151,316,909,933]" country="Nigeria" name="Kagoro Forest" pageId="23" pageNumber="90" stateProvince="WEST">Kagoro Forest</location>
,
<specimenCount id="9D7EFD76FFB0406DBD583BD5137CFC26" box="[328,364,908,932]" pageId="23" pageNumber="90" type="male">1♂</specimenCount>
,
<date id="FFC6103FFFB0406DBD6A3BD513FBFC27" box="[378,491,908,933]" pageId="23" pageNumber="90" value="1977-02-26">
<collectingDate id="EF82E9D7FFB0406DBD6A3BD513FBFC27" box="[378,491,908,933]" pageId="23" pageNumber="90" value="1977-02-26">26.ii.1977</collectingDate>
</date>
(
<collectionCode id="ED69AE3AFFB0406DBDEB3BD51050FC26" box="[507,576,908,932]" pageId="23" pageNumber="90">IARS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B103CA2FFB0406DBE453BD5133EFC4A" ID-GBIF-Occurrence="1913813778" collectingDate="1971-12-26" collectionCode="USNM" collectorName="Deeming" country="Niger" location="Niger" pageId="23" pageNumber="90" specimenCount="1" specimenCount-male="1" stateProvince="Niger">
<collectingCountry id="F36F766FFFB0406DBE453BD51085FC27" box="[597,661,908,933]" name="Niger" pageId="23" pageNumber="90">
<collectingRegion id="49BCF81DFFB0406DBE453BD51085FC27" box="[597,661,908,933]" country="Nigeria" name="Niger" pageId="23" pageNumber="90">Niger</collectingRegion>
</collectingCountry>
Prov., nr. Mokwa, Zugurma,
<specimenCount id="9D7EFD76FFB0406DBFF53BD51619FC26" box="[997,1033,908,932]" pageId="23" pageNumber="90" type="male">1♂</specimenCount>
,
<date id="FFC6103FFFB0406DB8073BD51687FC27" box="[1047,1175,908,933]" pageId="23" pageNumber="90" value="1971-12-26">
<collectingDate id="EF82E9D7FFB0406DB8073BD51687FC27" box="[1047,1175,908,933]" pageId="23" pageNumber="90" value="1971-12-26">26.xii.1971</collectingDate>
</date>
(
<collectionCode id="ED69AE3AFFB0406DB8B63BD516EAFC26" box="[1190,1274,908,932]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="90">USNM</collectionCode>
) (all collected by
<collectorName id="268D5329FFB0406DBCAA3BF61336FC4A" box="[186,294,943,968]" pageId="23" pageNumber="90">Deeming</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3626574FFB0406DBCD73B8D168FFC6E" box="[199,1183,980,1005]" pageId="23" pageNumber="90" type="description">
<paragraph id="8BC736FFFFB0406DBCD73B8D168FFC6E" blockId="23.[151,1437,151,1040]" box="[199,1183,980,1005]" pageId="23" pageNumber="90">
<emphasis id="B90CEAEDFFB0406DBCD73B8D13B5FC6F" bold="true" box="[199,421,980,1005]" pageId="23" pageNumber="90">Published records.</emphasis>
<collectingCountry id="F36F766FFFB0406DBDBC3B8D1016FC6F" box="[428,518,980,1005]" name="Benin" pageId="23" pageNumber="90">BÉNIN</collectingCountry>
: Agnavo and
<collectingCountry id="F36F766FFFB0406DBE8F3B8D1107FC6E" box="[671,791,980,1004]" name="Nigeria" pageId="23" pageNumber="90">NIGERIA</collectingCountry>
: Zaria (
<bibRefCitation id="EFE94B0EFFB0406DBF7A3B8D160CFC6E" box="[874,1052,980,1005]" pageId="23" pageNumber="90" refId="ref26981" refString="Vala, J-C., Gasc, C., Gbedjissi, G. &amp; Dossou, C. (1994) Les Sciomyzidae du Benin, description de Sepedon (Mesosepedon) knutsoni n. sp. (Diptera). Bulletin de la Societe entomologique de France, 99 (5), 497 - 504." type="journal article">
Vala
<emphasis id="B90CEAEDFFB0406DBFB53B8C11C6FC6F" box="[933,982,980,1005]" italics="true" pageId="23" pageNumber="90">et al</emphasis>
. 1994
</bibRefCitation>
<typeStatus id="54C3885DFFB0406DB8333B8D1683FC6E" box="[1059,1171,980,1004]" pageId="23" pageNumber="90" type="paratype">paratypes</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="C3626574FFB0406DBCD73BAE16AFFB92" box="[199,1215,1015,1040]" pageId="23" pageNumber="90" type="biology_ecology">
<paragraph id="8BC736FFFFB0406DBCD73BAE16AFFB92" blockId="23.[151,1437,151,1040]" box="[199,1215,1015,1040]" pageId="23" pageNumber="90">
<emphasis id="B90CEAEDFFB0406DBCD73BAE1337FB92" bold="true" box="[199,295,1015,1040]" pageId="23" pageNumber="90">Biology.</emphasis>
Larvae feed on fresh-water oligochaete worms (Vala
<emphasis id="B90CEAEDFFB0406DBF903BA011A0FB92" box="[896,944,1015,1040]" italics="true" pageId="23" pageNumber="90">et al</emphasis>
. 2002;
<bibRefCitation id="EFE94B0EFFB0406DB8163BAE16A3FB92" box="[1030,1203,1015,1040]" pageId="23" pageNumber="90" refId="ref25510" refString="Gbedjissi, L. G. (2003) Relations mollusciques-Dipteres Sciomyzidae. Implication dans la luttre contre les diastomatoses au Benin. Ph. D. thesis, UniVersite d'AVignon et des pays du Vaucluse, AVignon, 153 pp." type="book">Gbedjissi 2003</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>