200 lines
40 KiB
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200 lines
40 KiB
XML
<document id="DD2414EB42FA66E8B0F57277B739771E" ID-DOI="10.1649/934.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5461668" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630266972764" checkinUser="valdenar" docAuthor="Liang, Hongbin & Kavanaugh, David H." docDate="2007" docId="03D187BCFFD0480728A7BCCF256BFB5F" docLanguage="en" docName="934.1.pdf" docOrigin="The Coleopterists Bulletin (Oxford, England) 61 (1)" docSource="http://dx.doi.org/10.1649/934.1" docStyle="DocumentStyle:76939A7658C5F80CF4BB6818E728552A.2:TheColeopBull.2003-2009.journal_article.1cover" docStyleId="76939A7658C5F80CF4BB6818E728552A" docStyleName="TheColeopBull.2003-2009.journal_article.1cover" docStyleVersion="2" docTitle="Desera Hope 1831" docType="treatment" docVersion="3" lastPageNumber="39" masterDocId="FFE8FFC4FFD348042808BC0D270CFFA4" masterDocTitle="Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species" masterLastPageNumber="39" masterPageNumber="1" pageNumber="1" updateTime="1699204912656" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="084BF9A8EF172A0A1C961AEF0CE076C8">Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species</mods:title>
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<mods:namePart id="BCBB3CD1F080B39D6156C617E9D68C00">Liang, Hongbin</mods:namePart>
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<mods:namePart id="D185DF2FC3660A5A029F2B1FB0474A97">Kavanaugh, David H.</mods:namePart>
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<mods:typeOfResource id="1A197D1FB17A80A1196556C7DBFED0EF">text</mods:typeOfResource>
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<mods:title id="9DD8522CBC98DB5856092BED7FED2CD5">The Coleopterists Bulletin</mods:title>
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<mods:part id="29A3C01421CDAAB039E6A17869C801C7">
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<mods:date id="07BF78ECB5910871C887E2158A0BED6F">2007</mods:date>
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<mods:title id="1414CFBD168D24AD11D17718196DF1C0">Oxford, England</mods:title>
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<mods:number id="036D9E581C3EF6BC67206CDC4F905693">2007-03-31</mods:number>
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<mods:number id="1CBCB2D905F6F0341D75B575717A01F9">61</mods:number>
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<mods:url id="89C3E806331C14BB419A8424D74B3E04">http://dx.doi.org/10.1649/934.1</mods:url>
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<mods:identifier id="A32741190F1BD1C206F48049136D061A" type="DOI">10.1649/934.1</mods:identifier>
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<mods:identifier id="16B94269B6FC483D9D5070BAEFAC995E" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="B7DDC649F835086A575A66AAA49DCD25" type="Zenodo-Dep">5461668</mods:identifier>
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<treatment id="03D187BCFFD0480728A7BCCF256BFB5F" LSID="urn:lsid:plazi:treatment:03D187BCFFD0480728A7BCCF256BFB5F" httpUri="http://treatment.plazi.org/id/03D187BCFFD0480728A7BCCF256BFB5F" pageId="3">
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<subSubSection id="C3626521FFD0480728A7BCCF24E0FF57" pageId="3" type="nomenclature">
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<paragraph id="8BC736AAFFD0480728A7BCCF24E0FF7C" blockId="3.[175,1005,194,1275]" box="[175,1004,194,216]" pageId="3">
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<heading id="D08F81C6FFD0480728A7BCCF24E0FF7C" box="[175,1004,194,216]" centered="true" fontSize="9" level="2" pageId="3" reason="2">
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<treatmentCitation id="0AD910BBFFD0480728A7BCCF2691FF7C" author="Hope" box="[175,413,194,216]" page="21" pageId="3" year="1831">
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<taxonomicName id="4C784D29FFD0480728A7BCCF2691FF7C" ID-CoL="8KVSP" authority="Hope 1831: 21" authorityName="Hope" authorityPageNumber="21" authorityYear="1831" box="[175,413,194,216]" class="Insecta" family="Carabidae" genus="Desera" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B90CEAB8FFD0480728A7BCCF27F9FF73" box="[175,245,194,215]" italics="true" pageId="3">Desera</emphasis>
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<bibRefCitation id="EFE94B5BFFD04807290DBCCF2691FF7C" author="Hope" box="[261,413,194,216]" pageId="3" pagination="21 - 32" refId="ref31055" refString="Hope, F. W. 1831. Synopsis of the new species of Nepal insects in the collection of Major General Hardwicke. Zoological Miscellany 1: 21 - 32." type="journal article" year="1831">Hope 1831:21</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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;
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<treatmentCitation id="0AD910BBFFD0480729B9BCCF2565FF7C" author="Dupuis" box="[433,617,194,216]" page="319" pageId="3" year="1912">
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<bibRefCitation id="EFE94B5BFFD0480729B9BCCF2565FF7C" author="Dupuis" box="[433,617,194,216]" pageId="3" pagination="308 - 338" refId="ref30901" refString="Dupuis, P. 1912. H. Sauter's Formosa-Ausbeute Carabidae (2 me Contribution). Annales de la Societe Entomologique de Belgique 56: 308 - 338." type="journal article" year="1912">Dupuis 1912:319</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AD910BBFFD048072A74BCCF2424FF7C" author="Heller" box="[636,808,194,216]" page="303" pageId="3" year="1923">
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<bibRefCitation id="EFE94B5BFFD048072A74BCCF2424FF7C" author="Heller" box="[636,808,194,216]" pageId="3" pagination="295 - 305" refId="ref31028" refString="Heller, K. M. 1923. Some new Malayan Carabidae, especially Philippine. The Philippine Journal of Science 23: 295 - 305." type="journal article" year="1923">Heller 1923:303</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AD910BBFFD048072B33BCCF24E4FF7C" author="Csiki" box="[827,1000,194,216]" page="1553" pageId="3" year="1932">
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<bibRefCitation id="EFE94B5BFFD048072B33BCCF24E4FF7C" author="Csiki" box="[827,1000,194,216]" pageId="3" pagination="1023 - 1933" refId="ref30809" refString="Csiki, E. 1932. Pars 124. Carabidae: Harpalinae VII [pp. 1279 - 1598]. In: Coleopterorum catalogus, Volumen III (W. Junk and S. Schenkling, editors). Carabidae III. W. Junk, Berlin. 1023 - 1933 pp." type="book chapter" year="1932">Csiki 1932:1553</bibRefCitation>
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</treatmentCitation>
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;
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</treatmentCitationGroup>
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</heading>
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</paragraph>
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<paragraph id="8BC736AAFFD0480728E7BCD024E0FF57" blockId="3.[175,1005,194,1275]" box="[239,1004,221,243]" pageId="3">
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<bibRefCitation id="EFE94B5BFFD0480728E7BCD026C7FF57" author="Basilewsky" box="[239,459,221,243]" pageId="3" pagination="133 - 182" refId="ref30614" refString="Basilewsky, P. 1960. Etude des Dryptinae d'Afrique (Coleoptera: Carabidae). Bulletin et Annales de la Societe Entomologique de Belgique 96: 133 - 182." type="journal article" year="1960">Basilewsky 1960:177</bibRefCitation>
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;
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||
<bibRefCitation id="EFE94B5BFFD0480729D6BCD02591FF57" author="Jedlicka" box="[478,669,221,243]" pageId="3" pagination="269 - 568" refId="ref31181" refString="Jedlicka, A. 1963. Monographie der Truncatipennen aus Ostasien, Lebiinae - Odacanthinae - Brachyninae (Coleoptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatliches Museum fur Tierkunde in Dresden 28: 269 - 568." type="journal article" year="1963">Jedlicka 1963:486</bibRefCitation>
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;
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||
<bibRefCitation id="EFE94B5BFFD048072AB8BCD02466FF57" author="Hansen" box="[688,874,221,243]" pageId="3" pagination="397 - 413" refId="ref30984" refString="Hansen, W. 1967. Contribution a la connaissance des coleopteres carabiques du sud-est asiatique, sur le genre Desera Hope (Coleoptera Carabidae Dryptinae). Bulletin et Annales de la Societe Royal Entomologique de Belgique 103: 397 - 413." type="journal article" year="1967">Hansen 1967:400</bibRefCitation>
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; and Habu
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3626521FFD0480728E7BCF5265DFEAA" box="[239,337,248,270]" pageId="3" type="description">
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<paragraph id="8BC736AAFFD0480728E7BCF5265DFEAA" blockId="3.[175,1005,194,1275]" box="[239,337,248,270]" pageId="3">1967:277.</paragraph>
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</subSubSection>
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<subSubSection id="C3626521FFD0480728CFBD19257CFEE1" pageId="3" type="type_taxon">
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<paragraph id="8BC736AAFFD0480728CFBD19257CFEE1" blockId="3.[175,1005,194,1275]" pageId="3">
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<emphasis id="B90CEAB8FFD0480728CFBD192641FE8E" bold="true" box="[199,333,276,298]" pageId="3">
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<typeStatus id="54C38808FFD0480728CFBD1927F5FE8E" box="[199,249,276,298]" pageId="3">Type</typeStatus>
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species.
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</emphasis>
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||
<taxonomicName id="4C784D29FFD04807295EBD192407FE8E" ID-CoL="34P57" authority="Schmidt-Gobel 1846" authorityName="Schmidt-Gobel" authorityYear="1846" box="[342,779,276,298]" class="Insecta" family="Carabidae" genus="Dendrocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="discolor">
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<emphasis id="B90CEAB8FFD04807295EBD192522FE8D" box="[342,558,276,297]" italics="true" pageId="3">Dendrocellus discolor</emphasis>
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Schmidt-Göbel 1846
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</taxonomicName>
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(5
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<taxonomicName id="4C784D29FFD048072B31BD19262FFEE1" ID-CoL="34ZFF" authority="Hope 1831" authorityName="Hope, Kryzhanovskij" authorityYear="1831" baseAuthorityName="Hope" class="Insecta" family="Carabidae" genus="Desera" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="nepalensis">
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||
<emphasis id="B90CEAB8FFD048072B31BD1924E0FE8D" box="[825,1004,276,297]" italics="true" pageId="3">Desera nepalensis</emphasis>
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||
<bibRefCitation id="EFE94B5BFFD0480728A7BD22262FFEE1" author="Hope" box="[175,291,303,325]" pageId="3" pagination="21 - 32" refId="ref31055" refString="Hope, F. W. 1831. Synopsis of the new species of Nepal insects in the collection of Major General Hardwicke. Zoological Miscellany 1: 21 - 32." type="journal article" year="1831">Hope 1831</bibRefCitation>
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||
</taxonomicName>
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), designated by Andrewes 1939.
|
||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3626521FFD0480728CFBD472617FB29" pageId="3" type="description">
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||
<paragraph id="8BC736AAFFD0480728CFBD472617FB29" blockId="3.[175,1005,194,1275]" pageId="3">
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||
<emphasis id="B90CEAB8FFD0480728CFBD47264CFEC4" bold="true" box="[199,320,330,352]" pageId="3">Description.</emphasis>
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Body narrow, elongate (
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<figureCitation id="13432A2FFFD048072A47BD4725B1FEC4" box="[591,701,330,352]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="4.[167,206,1376,1395]" captionStartId-1="5.[196,235,1430,1449]" captionTargetBox-0="[171,950,195,1353]" captionTargetBox-1="[184,993,197,1407]" captionTargetId-0="figure-1@4.[168,953,194,1354]" captionTargetId-1="figure-0@5.[183,996,194,1407]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Figs. 1–12. Habitus of Dendrocellus species. 1) D. australis (male, Congo); 2) D. coelestinus (syntype); 3) D. coelestinus (type of D. paralellus); 4) D. confusus (male, Thailand); 5) D. crassus (male, Borneo); 6) D. elegans (male, New Guinea); 7) D. rugicollis (lectotype); 8) D. geniculatus (holotype); 9) D. geniculatus (holotype of D. gilsoni); 10) D. gestroi (lectotype); 11) D. inexpectus (holotype); 12) D. javanus (holotype). Scale line 5 2.0 mm." captionText-1="Figs. 13–24. Habitus of Dendrocellus species. 13) D. kulti (male, Yunnan); 14) D. kulti (holotype of D. kulti reconditus); 15) D. micropectinatus (holotype); 16) D. nepalensis (male, Pedong, India); 17) D. nigripennis (holotype); 18) D. queenslandicus (holotype); 19) D. schultzei (lectotype); 20) D. sinicus (holotype); 21) D. smaragdinus (holotype); 22) D. ternatensis (holotype); 23) D. unidentatus (male, Java); 24) D. viridipennis (male, Congo). Scale line 5 2.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5461670" figureDoi-1="http://doi.org/10.5281/zenodo.5461672" httpUri-0="https://zenodo.org/record/5461670/files/figure.png" httpUri-1="https://zenodo.org/record/5461672/files/figure.png" pageId="3">Figs. 1–24</figureCitation>
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), length 8.0–14.0 mm, width
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<quantity id="4C809B4FFFD0480728A7BD6B2627FED8" box="[175,299,358,380]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="4.2" metricValueMin="2.5" pageId="3" unit="mm" value="3.35" valueMax="4.2" valueMin="2.5">2.5–4.2 mm</quantity>
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. Black with blue or green luster, in some species mixed with coppery or purple luster; in some species, the pronotum red or brown, sternum brown with blue luster. Microsculpture distinct, forming isodiametric meshes. Head punctate and pubescent; Ant 1 elongate, 2.5–3.8 times as long as Ant 3; mandibles about 3.0 times as long as labrum, hooked at apex; maxillary and labial apical palpomeres securiform; labrum with medial lobe slightly (
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<figureCitation id="13432A2FFFD048072B1CBDE02482FDA7" box="[788,910,493,515]" captionStart="Figs" captionStartId="6.[167,206,487,506]" captionTargetBox="[189,932,195,464]" captionTargetId="figure-1@6.[187,933,194,465]" captionTargetPageId="6" captionText="Figs. 25–30. Labrum of Dendrocellus species. 25) D. australis (male, Congo); 26) D. geniculatus (male, Tibet); 27) D. sinicus (holotype); 28) D. coelestinus (syntype); 29) D. rugicollis (lectotype, India); 30) D. confusus (male, Thailand). Scale line 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5461674" httpUri="https://zenodo.org/record/5461674/files/figure.png" pageId="3">Figs. 25–27</figureCitation>
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), moderately (
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||
<figureCitation id="13432A2FFFD0480728F9BE052662FDBA" box="[241,366,520,542]" captionStart="Figs" captionStartId="6.[167,206,487,506]" captionTargetBox="[189,932,195,464]" captionTargetId="figure-1@6.[187,933,194,465]" captionTargetPageId="6" captionText="Figs. 25–30. Labrum of Dendrocellus species. 25) D. australis (male, Congo); 26) D. geniculatus (male, Tibet); 27) D. sinicus (holotype); 28) D. coelestinus (syntype); 29) D. rugicollis (lectotype, India); 30) D. confusus (male, Thailand). Scale line 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5461674" httpUri="https://zenodo.org/record/5461674/files/figure.png" pageId="3">Figs. 28, 29</figureCitation>
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), or strongly (
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<figureCitation id="13432A2FFFD048072A0EBE05255CFDBA" box="[518,592,520,542]" captionStart="Figs" captionStartId="6.[167,206,487,506]" captionTargetBox="[189,932,195,464]" captionTargetId="figure-1@6.[187,933,194,465]" captionTargetPageId="6" captionText="Figs. 25–30. Labrum of Dendrocellus species. 25) D. australis (male, Congo); 26) D. geniculatus (male, Tibet); 27) D. sinicus (holotype); 28) D. coelestinus (syntype); 29) D. rugicollis (lectotype, India); 30) D. confusus (male, Thailand). Scale line 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5461674" httpUri="https://zenodo.org/record/5461674/files/figure.png" pageId="3">Fig. 30</figureCitation>
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) protruded anteriorly; genae short and oblique (
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||
<figureCitation id="13432A2FFFD048072907BE2E2683FD9D" box="[271,399,547,569]" captionStart="Figs" captionStartId="6.[167,206,1502,1521]" captionTargetBox="[191,931,732,1478]" captionTargetId="figure-42@6.[188,932,731,1479]" captionTargetPageId="6" captionText="Figs. 31–34. Head of Dendrocellus species. 31) D. queenslandicus (holotype); 32) D. ternatensis (male, Batchian, Indonesia); 33) D. rugicollis (lectotype, India); 34) D. javanus (holotype). Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461676" httpUri="https://zenodo.org/record/5461676/files/figure.png" pageId="3">Figs. 31, 32</figureCitation>
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||
), or long and distinctly tumid (
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<figureCitation id="13432A2FFFD048072AE5BE2E2461FD9D" box="[749,877,547,569]" captionStart="Figs" captionStartId="6.[167,206,1502,1521]" captionTargetBox="[191,931,732,1478]" captionTargetId="figure-42@6.[188,932,731,1479]" captionTargetPageId="6" captionText="Figs. 31–34. Head of Dendrocellus species. 31) D. queenslandicus (holotype); 32) D. ternatensis (male, Batchian, Indonesia); 33) D. rugicollis (lectotype, India); 34) D. javanus (holotype). Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461676" httpUri="https://zenodo.org/record/5461676/files/figure.png" pageId="3">Figs. 33, 34</figureCitation>
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). Pronotum cylindrical, about 1.3–1.8 times as long as wide, punctate and pubescent, punctures larger than those on head and elytra. Elytra truncate at apex, outer angles more or less round (
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<figureCitation id="13432A2FFFD0480729DBBE78255DFD2F" box="[467,593,629,651]" captionStart="Figs" captionStartId="7.[196,235,713,732]" captionTargetBox="[191,987,195,690]" captionTargetId="figure-169@7.[191,988,194,690]" captionTargetPageId="7" captionText="Figs. 35–43. Left elytron apex of Dendrocellus species. 35) D. kulti (male, Yunnan); 36) D. nepalensis (male, Pedong, India); 37) D. rugicollis (lectotype, India); 38) D. micropectinatus (holotype); 39) D. australis (male, Congo); 40) D. geniculatus (male, Tibet); 41) D. sinicus (holotype); 42) D. elegans (male, New Guinea); 43) D. unidentatus (male, Java). Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461678" httpUri="https://zenodo.org/record/5461678/files/figure.png" pageId="3">Figs. 35, 36</figureCitation>
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), obtuse (
|
||
<figureCitation id="13432A2FFFD048072AB3BE782434FD2F" box="[699,824,629,651]" captionStart="Figs" captionStartId="7.[196,235,713,732]" captionTargetBox="[191,987,195,690]" captionTargetId="figure-169@7.[191,988,194,690]" captionTargetPageId="7" captionText="Figs. 35–43. Left elytron apex of Dendrocellus species. 35) D. kulti (male, Yunnan); 36) D. nepalensis (male, Pedong, India); 37) D. rugicollis (lectotype, India); 38) D. micropectinatus (holotype); 39) D. australis (male, Congo); 40) D. geniculatus (male, Tibet); 41) D. sinicus (holotype); 42) D. elegans (male, New Guinea); 43) D. unidentatus (male, Java). Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461678" httpUri="https://zenodo.org/record/5461678/files/figure.png" pageId="3">Figs. 37, 38</figureCitation>
|
||
), slightly dentate (
|
||
<figureCitation id="13432A2FFFD0480728BFBE9D263AFD02" box="[183,310,656,678]" captionStart="Figs" captionStartId="7.[196,235,713,732]" captionTargetBox="[191,987,195,690]" captionTargetId="figure-169@7.[191,988,194,690]" captionTargetPageId="7" captionText="Figs. 35–43. Left elytron apex of Dendrocellus species. 35) D. kulti (male, Yunnan); 36) D. nepalensis (male, Pedong, India); 37) D. rugicollis (lectotype, India); 38) D. micropectinatus (holotype); 39) D. australis (male, Congo); 40) D. geniculatus (male, Tibet); 41) D. sinicus (holotype); 42) D. elegans (male, New Guinea); 43) D. unidentatus (male, Java). Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461678" httpUri="https://zenodo.org/record/5461678/files/figure.png" pageId="3">Figs. 39, 40</figureCitation>
|
||
), or strongly dentate and sharply pointed at apex (
|
||
<figureCitation id="13432A2FFFD048072B6FBE9D24EDFD02" box="[871,993,656,678]" captionStart="Figs" captionStartId="7.[196,235,713,732]" captionTargetBox="[191,987,195,690]" captionTargetId="figure-169@7.[191,988,194,690]" captionTargetPageId="7" captionText="Figs. 35–43. Left elytron apex of Dendrocellus species. 35) D. kulti (male, Yunnan); 36) D. nepalensis (male, Pedong, India); 37) D. rugicollis (lectotype, India); 38) D. micropectinatus (holotype); 39) D. australis (male, Congo); 40) D. geniculatus (male, Tibet); 41) D. sinicus (holotype); 42) D. elegans (male, New Guinea); 43) D. unidentatus (male, Java). Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461678" httpUri="https://zenodo.org/record/5461678/files/figure.png" pageId="3">Figs. 41–43</figureCitation>
|
||
); striae shallow, punctate; intervals flat or slightly convex, punctate and pubescent; in some species, punctures nearly equal in size, quite dense, interspace between two adjacent punctures narrower than diameter of punctures (
|
||
<figureCitation id="13432A2FFFD048072B32BEEC248AFD53" box="[826,902,737,759]" captionStart="Figs" captionStartId="7.[196,235,1525,1544]" captionTargetBox="[197,983,1132,1502]" captionTargetId="figure-228@7.[197,983,1132,1503]" captionTargetPageId="7" captionText="Figs. 44–46. Elytral punctation of Dendrocellus species. 44) D. geniculatus (male, Tibet);" figureDoi="http://doi.org/10.5281/zenodo.5461680" httpUri="https://zenodo.org/record/5461680/files/figure.png" pageId="3">Fig. 44</figureCitation>
|
||
), in other species, punctures unequal in size, sparse, interspace between two adjacent punctures often wider than diameter of punctures (
|
||
<figureCitation id="13432A2FFFD048072AB6BF1A2436FC89" box="[702,826,791,813]" captionStart="Figs" captionStartId="7.[196,235,1525,1544]" captionTargetBox="[197,983,1132,1502]" captionTargetId="figure-228@7.[197,983,1132,1503]" captionTargetPageId="7" captionText="Figs. 44–46. Elytral punctation of Dendrocellus species. 44) D. geniculatus (male, Tibet);" figureDoi="http://doi.org/10.5281/zenodo.5461680" httpUri="https://zenodo.org/record/5461680/files/figure.png" pageId="3">Figs. 45, 46</figureCitation>
|
||
). Hindwings fullsized, functional. Sterna densely punctate and pubescent; apical margin of terminal sternum slightly protruded posteriad medially in male (
|
||
<figureCitation id="13432A2FFFD048072B6ABF4024EDFCC7" box="[866,993,845,867]" captionStart="Figs" captionStartId="8.[167,206,542,561]" captionTargetBox="[190,932,195,518]" captionTargetId="figure-480@8.[187,934,194,519]" captionTargetPageId="8" captionText="Figs. 47–50. Apical margin of terminal sternum of Dendrocellus species. 47) and 49) D. geniculatus (Tibet); 48) and 50) D. crassus (Borneo); 47 and 48 5 males; 49 and 50 5 females. Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461682" httpUri="https://zenodo.org/record/5461682/files/figure.png" pageId="3">Figs. 47, 48</figureCitation>
|
||
), almost straight or widely rounded in female (
|
||
<figureCitation id="13432A2FFFD048072AB6BF65244FFCDA" box="[702,835,872,894]" captionStart="Figs" captionStartId="8.[167,206,542,561]" captionTargetBox="[190,932,195,518]" captionTargetId="figure-480@8.[187,934,194,519]" captionTargetPageId="8" captionText="Figs. 47–50. Apical margin of terminal sternum of Dendrocellus species. 47) and 49) D. geniculatus (Tibet); 48) and 50) D. crassus (Borneo); 47 and 48 5 males; 49 and 50 5 females. Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461682" httpUri="https://zenodo.org/record/5461682/files/figure.png" pageId="3">Figs. 49, 50</figureCitation>
|
||
). Legs slender, protarsomere 3 strongly (
|
||
<figureCitation id="13432A2FFFD0480729D2BF892528FC3E" box="[474,548,900,922]" captionStart="Figs" captionStartId="8.[167,206,1502,1521]" captionTargetBox="[154,967,1068,1479]" captionTargetId="figure-518@8.[152,968,1067,1480]" captionTargetPageId="8" captionText="Figs. 51–56. Tarsomeres 1–4 of right protarsus of Dendrocellus species (dorsal view). 51) and 54) D. viridipennis (Congo); 52) and 55) D. geniculatus (Tibet); 53) and 56) D. crassus (Borneo); 51 to 53 5 males; 54 to 56 5 females. Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461684" httpUri="https://zenodo.org/record/5461684/files/figure.png" pageId="3">Fig. 51</figureCitation>
|
||
) or slightly (
|
||
<figureCitation id="13432A2FFFD048072AC5BF89242CFC3E" box="[717,800,900,922]" captionStart="Figs" captionStartId="8.[167,206,1502,1521]" captionTargetBox="[154,967,1068,1479]" captionTargetId="figure-518@8.[152,968,1067,1480]" captionTargetPageId="8" captionText="Figs. 51–56. Tarsomeres 1–4 of right protarsus of Dendrocellus species (dorsal view). 51) and 54) D. viridipennis (Congo); 52) and 55) D. geniculatus (Tibet); 53) and 56) D. crassus (Borneo); 51 to 53 5 males; 54 to 56 5 females. Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461684" httpUri="https://zenodo.org/record/5461684/files/figure.png" pageId="3">Figs. 52</figureCitation>
|
||
) asymmetrical or symmetrical (
|
||
<figureCitation id="13432A2FFFD048072934BF92269CFC11" box="[316,400,927,949]" captionStart="Figs" captionStartId="8.[167,206,1502,1521]" captionTargetBox="[154,967,1068,1479]" captionTargetId="figure-518@8.[152,968,1067,1480]" captionTargetPageId="8" captionText="Figs. 51–56. Tarsomeres 1–4 of right protarsus of Dendrocellus species (dorsal view). 51) and 54) D. viridipennis (Congo); 52) and 55) D. geniculatus (Tibet); 53) and 56) D. crassus (Borneo); 51 to 53 5 males; 54 to 56 5 females. Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461684" httpUri="https://zenodo.org/record/5461684/files/figure.png" pageId="3">Figs. 53</figureCitation>
|
||
) in male, symmetrical in female (
|
||
<figureCitation id="13432A2FFFD048072AF0BF92247EFC11" box="[760,882,927,949]" captionStart="Figs" captionStartId="8.[167,206,1502,1521]" captionTargetBox="[154,967,1068,1479]" captionTargetId="figure-518@8.[152,968,1067,1480]" captionTargetPageId="8" captionText="Figs. 51–56. Tarsomeres 1–4 of right protarsus of Dendrocellus species (dorsal view). 51) and 54) D. viridipennis (Congo); 52) and 55) D. geniculatus (Tibet); 53) and 56) D. crassus (Borneo); 51 to 53 5 males; 54 to 56 5 females. Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461684" httpUri="https://zenodo.org/record/5461684/files/figure.png" pageId="3">Figs. 54–56</figureCitation>
|
||
), tarsomere 4 bilobed (
|
||
<figureCitation id="13432A2FFFD048072917BFB7269CFC74" box="[287,400,954,976]" captionStart="Figs" captionStartId="8.[167,206,1502,1521]" captionTargetBox="[154,967,1068,1479]" captionTargetId="figure-518@8.[152,968,1067,1480]" captionTargetPageId="8" captionText="Figs. 51–56. Tarsomeres 1–4 of right protarsus of Dendrocellus species (dorsal view). 51) and 54) D. viridipennis (Congo); 52) and 55) D. geniculatus (Tibet); 53) and 56) D. crassus (Borneo); 51 to 53 5 males; 54 to 56 5 females. Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461684" httpUri="https://zenodo.org/record/5461684/files/figure.png" pageId="3">Fig. 51–56</figureCitation>
|
||
); tarsal claws slender, more or less pectinate on inner side (
|
||
<figureCitation id="13432A2FFFD0480728BFBFD8263DFC4F" box="[183,305,981,1003]" captionStart="Figs" captionStartId="9.[196,235,972,991]" captionTargetBox="[176,1003,194,949]" captionTargetId="figure-360@9.[176,1003,194,949]" captionTargetPageId="9" captionText="Figs. 57–72. Tarsal claws of hind tarsi of Dendrocellus species and Drypta species. 57) D. rugicollis (lectotype); 58) D. elegans (male, New Guinea); 59) D. schultzei (lectotype); 60) D. sinicus (holotype); 61) D. inexpectus (holotype); 62) D. javanus (male, Java Occid., paratype); 63) D. nigripennis (holotype); 64) D. geniculatus (male, Tibet); 65) D. australis (male, Congo); 66) D. queenslandicus (holotype); 67) D. confusus (male, Thailand); 68) D. nepalensis (male, Pedong, India); 69) D. coelestinus (syntype); 70) D. micropectinatus (male paratype, Congo); 71) Drypta longicollis Macleay 1825 (male, Thailand); 72) Drypta lineola virgata Chaudoir 1850 (male, Vietnam). Scale line 5 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5461686" httpUri="https://zenodo.org/record/5461686/files/figure.png" pageId="3">Figs. 57–70</figureCitation>
|
||
), very different from those of
|
||
<taxonomicName id="4C784D29FFD048072A65BFD825B8FC4E" box="[621,692,981,1002]" class="Insecta" family="Carabidae" genus="Drypta" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90CEAB8FFD048072A65BFD825B8FC4E" box="[621,692,981,1002]" italics="true" pageId="3">Drypta</emphasis>
|
||
</taxonomicName>
|
||
(claws thick, strongly curved, and smooth on inner side;
|
||
<emphasis id="B90CEAB8FFD0480729C8BFFD26EFFBA1" box="[448,483,1008,1029]" italics="true" pageId="3">e.g.</emphasis>
|
||
,
|
||
<figureCitation id="13432A2FFFD0480729F8BFFD2567FBA2" box="[496,619,1008,1030]" captionStart="Figs" captionStartId="9.[196,235,972,991]" captionTargetBox="[176,1003,194,949]" captionTargetId="figure-360@9.[176,1003,194,949]" captionTargetPageId="9" captionText="Figs. 57–72. Tarsal claws of hind tarsi of Dendrocellus species and Drypta species. 57) D. rugicollis (lectotype); 58) D. elegans (male, New Guinea); 59) D. schultzei (lectotype); 60) D. sinicus (holotype); 61) D. inexpectus (holotype); 62) D. javanus (male, Java Occid., paratype); 63) D. nigripennis (holotype); 64) D. geniculatus (male, Tibet); 65) D. australis (male, Congo); 66) D. queenslandicus (holotype); 67) D. confusus (male, Thailand); 68) D. nepalensis (male, Pedong, India); 69) D. coelestinus (syntype); 70) D. micropectinatus (male paratype, Congo); 71) Drypta longicollis Macleay 1825 (male, Thailand); 72) Drypta lineola virgata Chaudoir 1850 (male, Vietnam). Scale line 5 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5461686" httpUri="https://zenodo.org/record/5461686/files/figure.png" pageId="3">Figs. 71, 72</figureCitation>
|
||
). In Asian species, aedeagus generally thin and long (
|
||
<figureCitation id="13432A2FFFD048072942B806251FFB85" box="[330,531,1035,1057]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="10.[167,206,1115,1134]" captionStartId-1="11.[196,235,1140,1159]" captionTargetBox-0="[147,968,194,1092]" captionTargetBox-1="[176,1003,194,1116]" captionTargetId-0="figure-214@10.[147,974,194,1093]" captionTargetId-1="figure-189@11.[176,1003,194,1117]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Figs. 73–90. Male genitalia of Dendrocellus species. 73) and 74) D. confusus (Thailand); 75) and 76) D. rugicollis (lectotype); 77) and 78) D. coelestinus (Thailand); 79) and 80) D. nepalensis (Pedong, India); 81) and 82) D. kulti (Hubei, China); 83) and 84) D. crassus (Borneo); 85) and 86) D. nigripennis (holotype); 87) and 88) D. geniculatus (Pedong, India); 89) and 90) D. javanus (holotype). Scale lines 5 1.0 mm, scale line A for Figs. 73, 75, 77, 79, 81, 83, 85, 87, 89 (apical part, dorsal view); scale line B for Figs. 74, 76, 78, 80, 82, 84, 86, 88, 90 (left lateral view)." captionText-1="Figs. 91–108. Male genitalia of Dendrocellus species. 91) and 92) D. queenslandicus (holotype); 93) and 94) D. ternatensis (Batchian, Indonesia); 95) and 96) D. elegans (New Guinea); 97) and 98) D. unidentatus (male, Java); 99) and 100) D. inexpectus (holotype); 101) and 102) D. sinicus (holotype); 103) and 104) D. viridipennis (male, Congo); 105) and 106) D. micropectinatus (holotype); 107) and 108) D. australis (male, Congo). Scale lines 5 1.0 mm, scale line A for Figs. 91, 93, 95, 97, 99, 101, 103, 105, 107 (apical part, dorsal view); scale line B for Figs. 92, 94, 96, 98, 100, 102, 104, 106, 108 (left lateral view)." figureDoi-0="http://doi.org/10.5281/zenodo.5461688" figureDoi-1="http://doi.org/10.5281/zenodo.5461690" httpUri-0="https://zenodo.org/record/5461688/files/figure.png" httpUri-1="https://zenodo.org/record/5461690/files/figure.png" pageId="3">Fig. 73, 74, 77–102</figureCitation>
|
||
), with apical lamella wide and long, round or truncate at apex (
|
||
<figureCitation id="13432A2FFFD048072978B82B26E6FB98" box="[368,490,1062,1084]" captionStart="Figs" captionStartId="10.[167,206,1115,1134]" captionTargetBox="[147,968,194,1092]" captionTargetId="figure-214@10.[147,974,194,1093]" captionTargetPageId="10" captionText="Figs. 73–90. Male genitalia of Dendrocellus species. 73) and 74) D. confusus (Thailand); 75) and 76) D. rugicollis (lectotype); 77) and 78) D. coelestinus (Thailand); 79) and 80) D. nepalensis (Pedong, India); 81) and 82) D. kulti (Hubei, China); 83) and 84) D. crassus (Borneo); 85) and 86) D. nigripennis (holotype); 87) and 88) D. geniculatus (Pedong, India); 89) and 90) D. javanus (holotype). Scale lines 5 1.0 mm, scale line A for Figs. 73, 75, 77, 79, 81, 83, 85, 87, 89 (apical part, dorsal view); scale line B for Figs. 74, 76, 78, 80, 82, 84, 86, 88, 90 (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5461688" httpUri="https://zenodo.org/record/5461688/files/figure.png" pageId="3">Figs. 73–86</figureCitation>
|
||
), narrow, digitiform (
|
||
<figureCitation id="13432A2FFFD048072ADDB82B2443FB98" box="[725,847,1062,1084]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="10.[167,206,1115,1134]" captionStartId-1="11.[196,235,1140,1159]" captionTargetBox-0="[147,968,194,1092]" captionTargetBox-1="[176,1003,194,1116]" captionTargetId-0="figure-214@10.[147,974,194,1093]" captionTargetId-1="figure-189@11.[176,1003,194,1117]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Figs. 73–90. Male genitalia of Dendrocellus species. 73) and 74) D. confusus (Thailand); 75) and 76) D. rugicollis (lectotype); 77) and 78) D. coelestinus (Thailand); 79) and 80) D. nepalensis (Pedong, India); 81) and 82) D. kulti (Hubei, China); 83) and 84) D. crassus (Borneo); 85) and 86) D. nigripennis (holotype); 87) and 88) D. geniculatus (Pedong, India); 89) and 90) D. javanus (holotype). Scale lines 5 1.0 mm, scale line A for Figs. 73, 75, 77, 79, 81, 83, 85, 87, 89 (apical part, dorsal view); scale line B for Figs. 74, 76, 78, 80, 82, 84, 86, 88, 90 (left lateral view)." captionText-1="Figs. 91–108. Male genitalia of Dendrocellus species. 91) and 92) D. queenslandicus (holotype); 93) and 94) D. ternatensis (Batchian, Indonesia); 95) and 96) D. elegans (New Guinea); 97) and 98) D. unidentatus (male, Java); 99) and 100) D. inexpectus (holotype); 101) and 102) D. sinicus (holotype); 103) and 104) D. viridipennis (male, Congo); 105) and 106) D. micropectinatus (holotype); 107) and 108) D. australis (male, Congo). Scale lines 5 1.0 mm, scale line A for Figs. 91, 93, 95, 97, 99, 101, 103, 105, 107 (apical part, dorsal view); scale line B for Figs. 92, 94, 96, 98, 100, 102, 104, 106, 108 (left lateral view)." figureDoi-0="http://doi.org/10.5281/zenodo.5461688" figureDoi-1="http://doi.org/10.5281/zenodo.5461690" httpUri-0="https://zenodo.org/record/5461688/files/figure.png" httpUri-1="https://zenodo.org/record/5461690/files/figure.png" pageId="3">Figs. 87–96</figureCitation>
|
||
), or short and round at apex (
|
||
<figureCitation id="13432A2FFFD048072950B84C26D2FBF3" box="[344,478,1089,1111]" captionStart="Figs" captionStartId="11.[196,235,1140,1159]" captionTargetBox="[176,1003,194,1116]" captionTargetId="figure-189@11.[176,1003,194,1117]" captionTargetPageId="11" captionText="Figs. 91–108. Male genitalia of Dendrocellus species. 91) and 92) D. queenslandicus (holotype); 93) and 94) D. ternatensis (Batchian, Indonesia); 95) and 96) D. elegans (New Guinea); 97) and 98) D. unidentatus (male, Java); 99) and 100) D. inexpectus (holotype); 101) and 102) D. sinicus (holotype); 103) and 104) D. viridipennis (male, Congo); 105) and 106) D. micropectinatus (holotype); 107) and 108) D. australis (male, Congo). Scale lines 5 1.0 mm, scale line A for Figs. 91, 93, 95, 97, 99, 101, 103, 105, 107 (apical part, dorsal view); scale line B for Figs. 92, 94, 96, 98, 100, 102, 104, 106, 108 (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5461690" httpUri="https://zenodo.org/record/5461690/files/figure.png" pageId="3">Figs. 97–102</figureCitation>
|
||
); in African species, aedeagus stout, with lamella long or short (
|
||
<figureCitation id="13432A2FFFD048072959B85126EFFBD6" box="[337,483,1116,1138]" captionStart="Figs" captionStartId="11.[196,235,1140,1159]" captionTargetBox="[176,1003,194,1116]" captionTargetId="figure-189@11.[176,1003,194,1117]" captionTargetPageId="11" captionText="Figs. 91–108. Male genitalia of Dendrocellus species. 91) and 92) D. queenslandicus (holotype); 93) and 94) D. ternatensis (Batchian, Indonesia); 95) and 96) D. elegans (New Guinea); 97) and 98) D. unidentatus (male, Java); 99) and 100) D. inexpectus (holotype); 101) and 102) D. sinicus (holotype); 103) and 104) D. viridipennis (male, Congo); 105) and 106) D. micropectinatus (holotype); 107) and 108) D. australis (male, Congo). Scale lines 5 1.0 mm, scale line A for Figs. 91, 93, 95, 97, 99, 101, 103, 105, 107 (apical part, dorsal view); scale line B for Figs. 92, 94, 96, 98, 100, 102, 104, 106, 108 (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5461690" httpUri="https://zenodo.org/record/5461690/files/figure.png" pageId="3">Figs. 103–108</figureCitation>
|
||
), apex round, thickened (
|
||
<figureCitation id="13432A2FFFD048072AF2B8512483FBD6" box="[762,911,1116,1138]" captionStart="Figs" captionStartId="11.[196,235,1140,1159]" captionTargetBox="[176,1003,194,1116]" captionTargetId="figure-189@11.[176,1003,194,1117]" captionTargetPageId="11" captionText="Figs. 91–108. Male genitalia of Dendrocellus species. 91) and 92) D. queenslandicus (holotype); 93) and 94) D. ternatensis (Batchian, Indonesia); 95) and 96) D. elegans (New Guinea); 97) and 98) D. unidentatus (male, Java); 99) and 100) D. inexpectus (holotype); 101) and 102) D. sinicus (holotype); 103) and 104) D. viridipennis (male, Congo); 105) and 106) D. micropectinatus (holotype); 107) and 108) D. australis (male, Congo). Scale lines 5 1.0 mm, scale line A for Figs. 91, 93, 95, 97, 99, 101, 103, 105, 107 (apical part, dorsal view); scale line B for Figs. 92, 94, 96, 98, 100, 102, 104, 106, 108 (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5461690" httpUri="https://zenodo.org/record/5461690/files/figure.png" pageId="3">Figs. 104, 108</figureCitation>
|
||
) or thin (
|
||
<figureCitation id="13432A2FFFD0480728BFB87A261CFB29" box="[183,272,1143,1165]" captionStart="Figs" captionStartId="11.[196,235,1140,1159]" captionTargetBox="[176,1003,194,1116]" captionTargetId="figure-189@11.[176,1003,194,1117]" captionTargetPageId="11" captionText="Figs. 91–108. Male genitalia of Dendrocellus species. 91) and 92) D. queenslandicus (holotype); 93) and 94) D. ternatensis (Batchian, Indonesia); 95) and 96) D. elegans (New Guinea); 97) and 98) D. unidentatus (male, Java); 99) and 100) D. inexpectus (holotype); 101) and 102) D. sinicus (holotype); 103) and 104) D. viridipennis (male, Congo); 105) and 106) D. micropectinatus (holotype); 107) and 108) D. australis (male, Congo). Scale lines 5 1.0 mm, scale line A for Figs. 91, 93, 95, 97, 99, 101, 103, 105, 107 (apical part, dorsal view); scale line B for Figs. 92, 94, 96, 98, 100, 102, 104, 106, 108 (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5461690" httpUri="https://zenodo.org/record/5461690/files/figure.png" pageId="3">Fig. 106</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3626521FFD0480728CFB89E2518FB7B" pageId="3" type="biology_ecology">
|
||
<paragraph id="8BC736AAFFD0480728CFB89E2518FB7B" blockId="3.[175,1005,194,1275]" pageId="3">
|
||
<emphasis id="B90CEAB8FFD0480728CFB89E2616FB0D" bold="true" box="[199,282,1171,1193]" pageId="3">Biology.</emphasis>
|
||
These beetles live on vegetation, but adults of some species (
|
||
<emphasis id="B90CEAB8FFD048072BA9B89E24C8FB0C" box="[929,964,1171,1192]" italics="true" pageId="3">e.g.</emphasis>
|
||
,
|
||
<taxonomicName id="4C784D29FFD048072BDBB89E2662FB60" authority="(Hope))" authorityName="Hope, Kryzhanovskij" authorityYear="1831" baseAuthorityName="Hope" class="Insecta" family="Carabidae" genus="Desera" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="nepalensis">
|
||
<emphasis id="B90CEAB8FFD048072BDBB89E261FFB67" italics="true" pageId="3">D. nepalensis</emphasis>
|
||
(Hope))
|
||
</taxonomicName>
|
||
also hide under shrub debris or inside decayed tree trunks. Adults are attracted to light traps.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3626521FFD0480728CFB8E8256BFB5F" box="[199,615,1253,1275]" pageId="3" type="distribution">
|
||
<paragraph id="8BC736AAFFD0480728CFB8E8256BFB5F" blockId="3.[175,1005,194,1275]" box="[199,615,1253,1275]" pageId="3">
|
||
<emphasis id="B90CEAB8FFD0480728CFB8E8264AFB5F" bold="true" box="[199,326,1253,1275]" pageId="3">Distribution.</emphasis>
|
||
Asia,
|
||
<collectingCountry id="F36F763AFFD048072982B8E826E5FB5F" box="[394,489,1253,1275]" name="Australia" pageId="3">Australia</collectingCountry>
|
||
and Africa.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |