treatments-xml/data/03/D1/87/03D187A7DA354E0A3B2CFF14FE72FD6B.xml
2024-06-21 12:22:17 +02:00

846 lines
146 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="144DEE173E349CA677EE9AA5FE202609" ID-CLB-Dataset="6577" ID-DOI="10.5281/zenodo.4502533" ID-GBIF-Dataset="0b64ea7f-8991-4da6-a68d-6475baba8fdd" ID-ISSN="2345-7600" ID-Zenodo-Dep="4502533" ID-ZooBank="819AC644-37BC-43DB-8E11-984D77804AFE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612459355446" checkinUser="carolina" docAuthor="Sørensen, Martin V., Gąsiorowski, Ludwik, Randsø, Phillip V., Sánchez, Nuria &amp; Neves, Ricardo C." docDate="2016" docId="03D187A7DA354E0A3B2CFF14FE72FD6B" docLanguage="en" docName="RafflesBZool.64.3-27.pdf" docOrigin="Raffles Bulletin of Zoology 64" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Centroderes impurus Sørensen &amp; Gąsiorowski &amp; Randsø &amp; Sánchez &amp; Neves 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="10" masterDocId="FFE8FFDFDA374E0D3A0EFA09FF87FA1D" masterDocTitle="First report of kinorhynchs from Singapore, with the description of three new species" masterLastPageNumber="27" masterPageNumber="3" pageNumber="5" updateTime="1698926845134" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="F8E0D14A00B5344AE5F23E3CA7E2D17E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="065FD5EF47C6AFE9A56FD2F3B42FC85F">
<mods:title id="19FEBFF46D1877CF0BB45EC8B755EE51">First report of kinorhynchs from Singapore, with the description of three new species</mods:title>
</mods:titleInfo>
<mods:name id="0F86D4490B0804A5F2971D121CDC87C8" type="personal">
<mods:role id="5513A770FA757DAD69375D5F2B2D5623">
<mods:roleTerm id="94E82C1A0594F021D90C904FC54DA200">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B895244DCD215B028D310B10A395924D">Sørensen, Martin V.</mods:namePart>
</mods:name>
<mods:name id="E80825D2EE9622F39D07BC6721ABCAC4" type="personal">
<mods:role id="2E4DF8FAF325A1EE76B3C8598D3430B9">
<mods:roleTerm id="1D0136E9F0AB6EDEA900C1E215FAE177">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1ECFB2FF0EE6A97E17B204571D77D9A2">Gąsiorowski, Ludwik</mods:namePart>
</mods:name>
<mods:name id="2875690F548E0EFE78558054A90D893C" type="personal">
<mods:role id="1F2060683923DC54D1092A5C275F945A">
<mods:roleTerm id="9A303206799E8B34B5E41B898E804CA0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9D99A649974147182CE75EEB15F6040E">Randsø, Phillip V.</mods:namePart>
</mods:name>
<mods:name id="89862D1A393B8FD67398588B74D435C0" type="personal">
<mods:role id="C2F5C96862EDDFA75CBEAD11994CA595">
<mods:roleTerm id="885A674011EB52A39FE79395449603FA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FC9B7A1460153E0D3F978526D306573C">Sánchez, Nuria</mods:namePart>
</mods:name>
<mods:name id="B1E12ED7B7625DFE1E4F4FE6BD9E681E" type="personal">
<mods:role id="E5167411FE33C03FF5E5ADF50F8EB39E">
<mods:roleTerm id="2A13980DE452DB6927B8A2721F60F021">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7FC543B8A1E8519847C9094458C70876">Neves, Ricardo C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A2D926EC8B15B644D7D3F887C9508B5B">text</mods:typeOfResource>
<mods:relatedItem id="FCBBC12BD3F44F1A62C8CCC16F63F281" type="host">
<mods:titleInfo id="BBE45007C22F2FD50C6EFB420060D03E">
<mods:title id="79D712BD527A417CB3F41050F7C3E757">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="5240C0CA0994BD8DFBB790FB468C6C9E">
<mods:date id="7610476F419A65EAE391D2C7E7703654">2016</mods:date>
<mods:detail id="1497A073C5D84A9726753D85F1796975" type="pubDate">
<mods:number id="EBEDDA9DF8633E9D3723D6008027497B">2016-01-29</mods:number>
</mods:detail>
<mods:detail id="F12B3AAA0E2035A78B38625DED302B29" type="volume">
<mods:number id="4841B1C72711A427DEF7F35865792F68">64</mods:number>
</mods:detail>
<mods:extent id="3280C4C229F4A4123F771FB7D7E3575E" unit="page">
<mods:start id="B208658553A9108FF421C99599FEB736">3</mods:start>
<mods:end id="5FBF2E955BDC0321D3A8D774E3CE0944">27</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5886E695E9E11D262BE6E524711EA77C">journal article</mods:classification>
<mods:identifier id="35C73012D5D6D0B2274692F42C26FD70" type="CLB-Dataset">6577</mods:identifier>
<mods:identifier id="BC96D0195B5F6C2985A4C3C5C26BD93F" type="DOI">10.5281/zenodo.4502533</mods:identifier>
<mods:identifier id="4DFB045FF4A0041691D2CCD96A249E33" type="GBIF-Dataset">0b64ea7f-8991-4da6-a68d-6475baba8fdd</mods:identifier>
<mods:identifier id="2A2AF7EF0D4B772378275499EB933FF0" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="47B583AF1ED5B03885C45F9C37171979" type="Zenodo-Dep">4502533</mods:identifier>
<mods:identifier id="D7010E351039FB156376D5946DE2276A" type="ZooBank">819AC644-37BC-43DB-8E11-984D77804AFE</mods:identifier>
</mods:mods>
<treatment id="03D187A7DA354E0A3B2CFF14FE72FD6B" ID-DOI="http://doi.org/10.5281/zenodo.4776223" ID-GBIF-Taxon="182244072" ID-Zenodo-Dep="4776223" LSID="urn:lsid:plazi:treatment:03D187A7DA354E0A3B2CFF14FE72FD6B" httpUri="http://treatment.plazi.org/id/03D187A7DA354E0A3B2CFF14FE72FD6B" lastPageId="7" lastPageNumber="10" pageId="2" pageNumber="5">
<subSubSection id="C362653ADA354E0F3B2CFF14FDE8FF28" box="[290,623,1309,1333]" pageId="2" pageNumber="5" type="nomenclature">
<paragraph id="8BC736B1DA354E0F3B2CFF14FDE8FF28" blockId="2.[290,623,1309,1365]" box="[290,623,1309,1333]" pageId="2" pageNumber="5">
<heading id="D08F81DDDA354E0F3B2CFF14FDE8FF28" bold="true" box="[290,623,1309,1333]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="5" reason="2">
<emphasis id="B90CEAA3DA354E0F3B2CFF14FDE8FF28" bold="true" box="[290,623,1309,1333]" pageId="2" pageNumber="5">
<taxonomicName id="4C784D32DA354E0F3B2CFF14FD8BFF28" authority="Sørensen &amp; Gąsiorowski &amp; Randsø &amp; Sánchez &amp; Neves, 2016" authorityName="Sørensen &amp; Gąsiorowski &amp; Randsø &amp; Sánchez &amp; Neves" authorityYear="2016" box="[290,524,1309,1333]" class="Kinorhyncha" family="Centroderidae" genus="Centroderes" kingdom="Animalia" order="Cyclorhagida" pageId="2" pageNumber="5" phylum="Cephalorhyncha" rank="species" species="impurus" status="sp. nov.">
<emphasis id="B90CEAA3DA354E0F3B2CFF14FD8BFF28" bold="true" box="[290,524,1309,1333]" italics="true" pageId="2" pageNumber="5">Centroderes impurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57D8DA354E0F3818FF14FDE8FF28" box="[534,623,1309,1333]" pageId="2" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C362653ADA354E0F3B82FF34FD84FF48" box="[396,515,1341,1365]" pageId="2" pageNumber="5" type="description">
<paragraph id="8BC736B1DA354E0F3B82FF34FD84FF48" blockId="2.[290,623,1309,1365]" box="[396,515,1341,1365]" pageId="2" pageNumber="5">
(
<figureCitation id="13432A34DA354E0F3B9AFF34FE7CFF48" box="[404,507,1341,1365]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="4.[135,170,1977,1999]" captionStartId-1="5.[113,148,2063,2085]" captionStartId-2="6.[136,171,1919,1941]" captionStartId-3="8.[136,171,1694,1716]" captionTargetBox-0="[196,1415,193,1962]" captionTargetBox-1="[235,1328,869,2047]" captionTargetBox-2="[279,1332,189,1902]" captionTargetBox-3="[136,1472,297,1678]" captionTargetId-0="figure-8@4.[194,1418,193,1965]" captionTargetId-1="figure-213@5.[235,1330,869,2047]" captionTargetId-2="figure-7@6.[278,1332,189,1904]" captionTargetId-3="figure-8@8.[136,1474,297,1681]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionText-0="Fig. 2. Line art illustration of Centroderes impurus sp. nov. male stage-2, lateral view. Abbreviations: ij, intersegmentary joint line; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral spine; mdp, middorsal placid; mds, middorsal spine; mls, midlateral spine; mlss, midlateral sensory spot; mts, midterminal spine; mvp, midventral process; pdss, paradorsal sensory spot; pf, pectinate fringe; sdss, subdorsal sensory spot; sf, secondary fringe; vms, ventromedial spine; vmss, ventromedial sensory spot. Digits following labels refer to segment numbers." captionText-1="Fig. 3. Light micrographs showing lateral overviews of Centroderes impurus sp. nov. A, holotype, ZRC.MIS.0001, male stage-2. B, paratype, KIN-846, putative J6 or adult stage-1 of unknown gender." captionText-2="Fig. 4. Scanning electron micrographs of male stage-2 (A, B, F, H) and putative J6 or adult stage-1 of unknown gender (CE, G) of Centroderes impurus sp. nov. A, lateral overview. B, ventral overview. C, Lateral view of segment 1. D, Detail showing lateral accessory sensory spot on segment 1. E, Region of tergosternal junctions of segments 4 and 5; dashed circles outline extended cuticular hairs on posterior margin of segment 3. F, Region of tergosternal junction of segments 4 and 5. G, lateroventral view of segments 8 and 9. H, Lateral view of segments 810 in male stage-2. Abbreviations: lvs, lateroventral spine; lvt, lateroventral tube; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; mdcr, middorsal crenulated spine; mlcr, midlateral crenulated spine; mvp, midventral process; s, segment followed by segment number; sf, secondary fringe; vms, ventromedial spine on segment 1." captionText-3="Fig. 5. Light micrographs showing details of trunk morphology in Centroderes impurus sp. nov. A, C, E: holotype, ZRC.MIS.0001, male stage-2; B, D, F: paratype, KIN-846, putative J6 or adult stage-1 of unknown gender. A, Segments 36, lateral view, with dashed circles indicating laterodorsal and lateral accessory sensory spots. B, Segments 13, lateral view, showing ventrolateral tube on segment 2, which is found only in this stage and not in male stage-2. C, Segments 10 and 11, lateral view, showing crenulated spines in middorsal position on segment 10. D, Segments 56, lateral view, showing indications of lateroventral tube on segment 5, which is found only in this stage and not in male stage-2. E, Segment 811, lateral view, showing crenulated spines in midlateral positions on segment 10; note the conspicuous difference between the lateroventral spines on segment 8 and segment 9; dashed circles indicate laterodorsal and lateral accessory sensory spots. F, Segment 811, lateral view, showing acicular spines in middorsal and midlateral positions on segment 10. Abbreviations: lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdac, middorsal acicular spine; mdcr, middorsal crenulated spine; mlac, midlateral acicular spine; mlcr, midlateral crenulated spine; vlt, ventrolateral tube; vms, ventromedial spine." figureDoi-0="http://doi.org/10.5281/zenodo.4502537" figureDoi-1="http://doi.org/10.5281/zenodo.4502541" figureDoi-2="http://doi.org/10.5281/zenodo.4502543" figureDoi-3="http://doi.org/10.5281/zenodo.4502551" httpUri-0="https://zenodo.org/record/4502537/files/figure.png" httpUri-1="https://zenodo.org/record/4502541/files/figure.png" httpUri-2="https://zenodo.org/record/4502543/files/figure.png" httpUri-3="https://zenodo.org/record/4502551/files/figure.png" pageId="2" pageNumber="5">Figs. 25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C362653ADA354E0F3A86FF74FD7AFD68" pageId="2" pageNumber="5" type="materials_examined">
<paragraph id="8BC736B1DA354E0F3A86FF74FD7AFD68" blockId="2.[136,775,1405,1910]" pageId="2" pageNumber="5">
<emphasis id="B90CEAA3DA354E0F3A86FF74FEE1FF88" bold="true" box="[136,358,1405,1429]" pageId="2" pageNumber="5">Material examined.</emphasis>
<materialsCitation id="3B103CECDA354E0F3B63FF74FE0FFC48" ID-GBIF-Occurrence="3114480302" collectingDate="2014-05-16" collectionCode="ZRC" collectorName="Fluoromount G &amp; Lee Kong Chian Natural History" country="Singapore Island" elevation="9" latitude="1.30645" location="Bedok Jetty" longLatPrecision="1" longitude="103.95985" pageId="2" pageNumber="5" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<typeStatus id="54C38813DA354E0F3B63FF74FE57FF88" box="[365,464,1405,1429]" pageId="2" pageNumber="5" type="holotype">Holotype</typeStatus>
adult stage-
<specimenCount id="9D7EFD38DA354E0F385CFF74FD1BFF88" box="[594,668,1405,1429]" pageId="2" pageNumber="5" type="male">2 male</specimenCount>
, collected from sand with mud on
<date id="FFC61071DA354E0F3B85FF94FD9AFFA8" box="[395,541,1437,1462]" pageId="2" pageNumber="5" value="2014-05-16">
<collectingDate id="EF82E999DA354E0F3B85FF94FD9AFFA8" box="[395,541,1437,1462]" pageId="2" pageNumber="5" value="2014-05-16">16 May 2014</collectingDate>
</date>
, at station SI-03 (
<figureCitation id="13432A34DA354E0F38D2FF94FF12FFC8" captionStart="Fig" captionStartId="1.[113,148,1214,1236]" captionTargetBox="[201,1364,189,1199]" captionTargetId="figure-177@1.[201,1364,189,1199]" captionTargetPageId="1" captionText="Fig. 1. Map showing the sampling localities south of Singapore Island. Inset to the lower right shows the Malaya Peninsula and Indonesian Archipelago, with Singapore indicated in the square." figureDoi="http://doi.org/10.5281/zenodo.4502535" httpUri="https://zenodo.org/record/4502535/files/figure.png" pageId="2" pageNumber="5">Fig. 1</figureCitation>
,
<tableCitation id="C6FA030ADA354E0F3AAAFFB4FF72FFC8" box="[164,245,1469,1493]" captionStart="Table 1" captionStartId="1.[113,168,1320,1342]" captionText="Table 1. Summary of data on stations and taxa identified." pageId="2" pageNumber="5">Table 1</tableCitation>
), at
<elevation id="0055D182DA354E0F3B27FFB4FE1EFFC8" box="[297,409,1469,1493]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="5" unit="m" value="9.0">
<quantity id="4C809B54DA354E0F3B27FFB4FED4FFC8" box="[297,339,1469,1493]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="5" unit="m" value="9.0">9 m</quantity>
depth
</elevation>
, between
<location id="8EA7606ADA354E0F3808FFB4FD01FFCB" LSID="urn:lsid:plazi:treatment:03D187A7DA354E0A3B2CFF14FE72FD6B:8EA7606ADA354E0F3808FFB4FD01FFCB" box="[518,646,1469,1494]" country="Singapore Island" latitude="1.30645" longLatPrecision="1" longitude="103.95985" name="Bedok Jetty" pageId="2" pageNumber="5">Bedok Jetty</location>
and Sungei Bedok in the southeast part of
<collectingCountry id="F36F7621DA354E0F3BB3FFD4FDECFFE8" box="[445,619,1501,1525]" name="Singapore Island" pageId="2" pageNumber="5">Singapore Island</collectingCountry>
(
<geoCoordinate id="EE4C5076DA354E0F3876FFD4FC84FFE8" box="[632,771,1501,1525]" degrees="01" direction="north" minutes="18.387" orientation="latitude" pageId="2" pageNumber="5" precision="1" value="1.30645">01°18.387N</geoCoordinate>
,
<geoCoordinate id="EE4C5076DA354E0F3A86FFF4FEA6FC08" box="[136,289,1533,1557]" degrees="103" direction="east" minutes="57.591" orientation="longitude" pageId="2" pageNumber="5" precision="1" value="103.95985">103°57.591E</geoCoordinate>
), mounted in
<collectorName id="268D5367DA354E0F3BBEFFF4FDD0FC08" box="[432,599,1533,1557]" pageId="2" pageNumber="5">Fluoromount G</collectorName>
, deposited at the
<collectorName id="268D5367DA354E0F3A86FC17FE6EFC28" box="[136,489,1565,1590]" pageId="2" pageNumber="5">Lee Kong Chian Natural History</collectorName>
Museum, under catalogue number
<collectionCode id="ED69AE74DA354E0F3AD0FC34FE90FC48" box="[222,279,1597,1621]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="5">ZRC</collectionCode>
.MIS.0001
</materialsCitation>
.
<materialsCitation id="3B103CECDA354E0F3B9EFC37FD7EFD68" ID-GBIF-Occurrence="3114480303" collectingDate="2014-05-16" collectionCode="ZMUC, SEM" collectorName="Fluoromount G" country="Denmark" elevation="9" latitude="1.30645" location="Bedok Jetty" longLatPrecision="1" longitude="103.95985" pageId="2" pageNumber="5" specimenCode="KIN-846, KIN-847" specimenCount="4" specimenCount-male="3" typeStatus="paratype">
<typeStatus id="54C38813DA354E0F3B9EFC37FE70FC4B" box="[400,503,1598,1622]" pageId="2" pageNumber="5" type="paratype">Paratypes</typeStatus>
include two specimens of uncertain sex, probably either adult stage-
<specimenCount id="9D7EFD38DA354E0F3841FC54FD14FC68" box="[591,659,1629,1653]" pageId="2" pageNumber="5" type="male">1 male</specimenCount>
or female, or eventually preadult stages (J6), collected at same locality as holotype, mounted in
<collectorName id="268D5367DA354E0F3B95FC94FDC1FCA8" box="[411,582,1693,1717]" pageId="2" pageNumber="5">Fluoromount G</collectorName>
, and deposited at the Natural History Museum of
<collectingCountry id="F36F7621DA354E0F3BE6FCB4FDC9FCC8" box="[488,590,1725,1749]" name="Denmark" pageId="2" pageNumber="5">Denmark</collectingCountry>
, under catalogue numbers
<collectionCode id="ED69AE74DA354E0F3AE9FCD4FEB4FCE8" box="[231,307,1757,1781]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="2" pageNumber="5">ZMUC</collectionCode>
<specimenCode id="DBDE9ECADA354E0F3B36FCD4FE10FCE8" box="[312,407,1757,1781]" collectionCode="KIN-" pageId="2" pageNumber="5">KIN-846</specimenCode>
and
<specimenCode id="DBDE9ECADA354E0F3BC8FCD4FDAFFCE8" box="[454,552,1757,1781]" collectionCode="KIN-" pageId="2" pageNumber="5">KIN-847</specimenCode>
. Additional, non-type material, includes one adult stage-
<specimenCount id="9D7EFD38DA354E0F3BFAFCF4FDBCFD08" box="[500,571,1789,1813]" pageId="2" pageNumber="5" type="male">2 male</specimenCount>
and two putatively stage-
<specimenCount id="9D7EFD38DA354E0F3AC2FD14FECFFD28" box="[204,328,1821,1845]" pageId="2" pageNumber="5" type="generic">1 specimens</specimenCount>
of uncertain sex, collected at same locality as
<typeStatus id="54C38813DA354E0F3AAAFD34FE83FD48" box="[164,260,1853,1877]" pageId="2" pageNumber="5" type="holotype">holotype</typeStatus>
, and mounted for
<collectionCode id="ED69AE74DA354E0F3BCCFD34FE7BFD48" box="[450,508,1853,1877]" pageId="2" pageNumber="5">SEM</collectionCode>
. The
<collectionCode id="ED69AE74DA354E0F383BFD34FDEBFD48" box="[565,620,1853,1877]" pageId="2" pageNumber="5">SEM</collectionCode>
specimens are very dirty, and contributed only with limited information
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C362653ADA354E0F3A86FD94FBDEF877" pageId="2" pageNumber="5" type="diagnosis">
<paragraph id="8BC736B1DA354E0F3A86FD94FBDEF877" blockId="2.[136,775,1949,2070]" lastBlockId="2.[835,1475,188,618]" pageId="2" pageNumber="5">
<emphasis id="B90CEAA3DA354E0F3A86FD94FF7AFDA8" bold="true" box="[136,253,1949,1973]" pageId="2" pageNumber="5">Diagnosis.</emphasis>
<taxonomicName id="4C784D32DA354E0F3B08FD94FE0DFDA8" authorityName="Zelinka" authorityYear="1907" box="[262,394,1949,1973]" class="Kinorhyncha" family="Centroderidae" genus="Centroderes" kingdom="Animalia" order="Cyclorhagida" pageId="2" pageNumber="5" phylum="Cephalorhyncha" rank="genus">
<emphasis id="B90CEAA3DA354E0F3B08FD94FE0DFDA8" box="[262,394,1949,1973]" italics="true" pageId="2" pageNumber="5">Centroderes</emphasis>
</taxonomicName>
with middorsal acicular spines on segments 19 and 11; long, flexible, ventromedial acicular spines on segment 1; rigid lateroventral acicular spines on segments 8 and 9, with spine on segment 9 being conspicuously stout and robust.Male stage-2 with middorsal and midlateral crenulated spines on segment 10. Putative adult stage-
<specimenCount id="9D7EFD38DA354E0F39C2FB09FBD0FB07" box="[972,1111,256,282]" pageId="2" pageNumber="5" type="generic">1 specimens</specimenCount>
, or alternatively preadults, with middorsal and midlateral acicular spines on segment 10, and with minute tubes present in ventrolateral positions on segment 2 and lateroventral positions on segment 5. Sensory spots present paradorsally on segments 26 and 89, subdorsally on segment 1 (in male stage-2 only) and segment 11 (two pairs), laterodorsally on segments 1 (in putative J6 or stage-
<specimenCount id="9D7EFD38DA354E0F3E73FBE2FB7BF818" box="[1149,1276,491,517]" pageId="2" pageNumber="5" type="generic">1 specimen</specimenCount>
only) and 310, midlaterally on segment 2, in a lateral accessory position on segments 1, 34 and 69 and ventromedially on segments 34 and 610.
</paragraph>
</subSubSection>
<subSubSection id="C362653ADA354E0F394DF89AFA81F916" pageId="2" pageNumber="5" type="etymology">
<paragraph id="8BC736B1DA354E0F394DF89AFA81F916" blockId="2.[835,1475,659,779]" pageId="2" pageNumber="5">
<emphasis id="B90CEAA3DA354E0F394DF89AFC4DF8B6" bold="true" box="[835,970,659,683]" pageId="2" pageNumber="5">Etymology.</emphasis>
The species is named after the Latin word “
<taxonomicName id="4C784D32DA354E0F395EF8BDFC2BF8D6" authorityName="Sørensen &amp; Gąsiorowski &amp; Randsø &amp; Sánchez &amp; Neves" authorityYear="2016" box="[848,940,692,715]" class="Kinorhyncha" family="Centroderidae" genus="Centroderes" kingdom="Animalia" order="Cyclorhagida" pageId="2" pageNumber="5" phylum="Cephalorhyncha" rank="species" species="impurus">
<emphasis id="B90CEAA3DA354E0F395EF8BDFC2BF8D6" box="[848,940,692,715]" italics="true" pageId="2" pageNumber="5">impurus</emphasis>
</taxonomicName>
”, meaning “unclean” or “covered with dirt”, referring to the severe dirt problems that were experienced during the examination of the specimens.
</paragraph>
</subSubSection>
<subSubSection id="C362653ADA354E0A394DF93AFE72FD6B" lastPageId="7" lastPageNumber="10" pageId="2" pageNumber="5" type="description">
<paragraph id="8BC736B1DA354E0F394DF93AFA45F9B6" blockId="2.[835,1476,819,940]" pageId="2" pageNumber="5">
<emphasis id="B90CEAA3DA354E0F394DF93AFC5EF956" bold="true" box="[835,985,819,843]" pageId="2" pageNumber="5">Description.</emphasis>
Adult with head, neck and eleven trunk segments (
<figureCitation id="13432A34DA354E0F39B4F95AFB8EF976" box="[954,1033,851,875]" captionStart="Fig" captionStartId="4.[135,170,1977,1999]" captionTargetBox="[196,1415,193,1962]" captionTargetId="figure-8@4.[194,1418,193,1965]" captionTargetPageId="4" captionText="Fig. 2. Line art illustration of Centroderes impurus sp. nov. male stage-2, lateral view. Abbreviations: ij, intersegmentary joint line; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral spine; mdp, middorsal placid; mds, middorsal spine; mls, midlateral spine; mlss, midlateral sensory spot; mts, midterminal spine; mvp, midventral process; pdss, paradorsal sensory spot; pf, pectinate fringe; sdss, subdorsal sensory spot; sf, secondary fringe; vms, ventromedial spine; vmss, ventromedial sensory spot. Digits following labels refer to segment numbers." figureDoi="http://doi.org/10.5281/zenodo.4502537" httpUri="https://zenodo.org/record/4502537/files/figure.png" pageId="2" pageNumber="5">Figs. 2</figureCitation>
,
<figureCitation id="13432A34DA354E0F3E16F95AFBD9F976" box="[1048,1118,851,875]" captionStart="Fig" captionStartId="5.[113,148,2063,2085]" captionTargetBox="[235,1328,869,2047]" captionTargetId="figure-213@5.[235,1330,869,2047]" captionTargetPageId="5" captionText="Fig. 3. Light micrographs showing lateral overviews of Centroderes impurus sp. nov. A, holotype, ZRC.MIS.0001, male stage-2. B, paratype, KIN-846, putative J6 or adult stage-1 of unknown gender." figureDoi="http://doi.org/10.5281/zenodo.4502541" httpUri="https://zenodo.org/record/4502541/files/figure.png" pageId="2" pageNumber="5">3A, B</figureCitation>
,
<figureCitation id="13432A34DA354E0F3E63F95AFB35F976" box="[1133,1202,851,875]" captionStart="Fig" captionStartId="6.[136,171,1919,1941]" captionTargetBox="[279,1332,189,1902]" captionTargetId="figure-7@6.[278,1332,189,1904]" captionTargetPageId="6" captionText="Fig. 4. Scanning electron micrographs of male stage-2 (A, B, F, H) and putative J6 or adult stage-1 of unknown gender (CE, G) of Centroderes impurus sp. nov. A, lateral overview. B, ventral overview. C, Lateral view of segment 1. D, Detail showing lateral accessory sensory spot on segment 1. E, Region of tergosternal junctions of segments 4 and 5; dashed circles outline extended cuticular hairs on posterior margin of segment 3. F, Region of tergosternal junction of segments 4 and 5. G, lateroventral view of segments 8 and 9. H, Lateral view of segments 810 in male stage-2. Abbreviations: lvs, lateroventral spine; lvt, lateroventral tube; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; mdcr, middorsal crenulated spine; mlcr, midlateral crenulated spine; mvp, midventral process; s, segment followed by segment number; sf, secondary fringe; vms, ventromedial spine on segment 1." figureDoi="http://doi.org/10.5281/zenodo.4502543" httpUri="https://zenodo.org/record/4502543/files/figure.png" pageId="2" pageNumber="5">4A, B</figureCitation>
). For measurements and dimensions, see
<tableCitation id="C6FA030ADA354E0F39FDF97AFBC0F996" box="[1011,1095,883,907]" captionStart="Table 2" captionStartId="3.[113,168,189,211]" captionText="Table 2. Measurements from light microscopy of adult Centroderes impurus sp. nov. (in µm) from station SI-03, Singapore, including individual measures, mean and standard deviation (SD). Abbreviations: LTAS: lateral terminal accessory spine; LTS: lateral terminal spine; LV: lateroventral spine or tube; MD, middorsal spine; ML: midlateral spine; MSW-6: maximum sternal width, measured on segment 6 in this species; MTS: midterminal spine; S: segment lengths; SW-10, standard width, always measured on segment 10; TL: trunk length; VM: ventromedial spine." pageId="2" pageNumber="5">Table 2</tableCitation>
. Distribution of cuticular structures (spines, tubes and sensory spots) is summarised in
<tableCitation id="C6FA030ADA354E0F3F64F99AFA39F9B6" box="[1386,1470,915,939]" captionStart="Table 3" captionStartId="5.[113,167,189,211]" captionText="Table 3. Summary of nature and location of spines, tubes and sensory spots arranged by series in Centroderes impurus sp. nov. Abbreviations: LA: Lateral accessory; LD: laterodorsal; LV: lateroventral; MD: middorsal; ML: midlateral; PD:paradorsal; SD: subdorsal; VL:ventrolateral; VM: ventromedial; a1, condition of character in the putative J6 preadult or adult stage-1; ac, acicular spine; cr, crenulated spines; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mts, midterminal spine; ss, sensory spot; tu, tube; ♂2, condition of character in the male stage-2; -, no structure present." pageId="2" pageNumber="5">Table 3</tableCitation>
.
</paragraph>
<paragraph id="8BC736B1DA354E0F394DF9DAFBB1FEB6" blockId="2.[835,1475,979,1195]" pageId="2" pageNumber="5">The head consists of a retractable mouth cone and an introvert. The mouth cone is equipped with nine outer oral styles, each consisting of two joined units, arranged as one style anterior to each introvert sector, except at the middorsal sector 6. A double fringe consisting of numerous tips is located basally to each style. Scalid arrangement could not be examined in detail.</paragraph>
<paragraph id="8BC736B1DA354E0F394DFEDAFBA1FF76" blockId="2.[835,1475,1235,1387]" pageId="2" pageNumber="5">
The neck has 16 placids that dorsally and laterally alternate in size width between broader (
<quantity id="4C809B54DA354E0F3E7BFEFAFB64FF11" box="[1141,1251,1267,1292]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="2" pageNumber="5" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">1516 µm</quantity>
width) and narrower (
<quantity id="4C809B54DA354E0F3945FF1AFC18FF31" box="[843,927,1299,1324]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="2" pageNumber="5" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">68 µm</quantity>
width) ones; all placids measure
<quantity id="4C809B54DA354E0F3F0AFF1AFAF5FF31" box="[1284,1394,1299,1324]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="2" pageNumber="5" unit="mm" value="16.0" valueMax="17.0" valueMin="15.0">1517 µm</quantity>
length. The midventral placid is broader and flanked by two narrower placids on each side.
</paragraph>
<paragraph id="8BC736B1DA354E0E394DFF9AFDF8FCDD" blockId="2.[835,1475,1427,2061]" lastBlockId="3.[113,752,1639,1728]" lastPageId="3" lastPageNumber="6" pageId="2" pageNumber="5">
Segment 1 consists of one complete cuticular ring with middorsal acicular spine, being short and stout in male stage-2 (
<figureCitation id="13432A34DA354E0F3945FFDAFC27FFF6" box="[843,928,1491,1515]" captionStart="Fig" captionStartId="5.[113,148,2063,2085]" captionTargetBox="[235,1328,869,2047]" captionTargetId="figure-213@5.[235,1330,869,2047]" captionTargetPageId="5" captionText="Fig. 3. Light micrographs showing lateral overviews of Centroderes impurus sp. nov. A, holotype, ZRC.MIS.0001, male stage-2. B, paratype, KIN-846, putative J6 or adult stage-1 of unknown gender." figureDoi="http://doi.org/10.5281/zenodo.4502541" httpUri="https://zenodo.org/record/4502541/files/figure.png" pageId="2" pageNumber="5">Fig. 3A</figureCitation>
), and two slender, elongated spines in ventromedial positions, which extend over two following segments (
<figureCitation id="13432A34DA354E0F3F82FFFAFCD5FC36" captionStart="Fig" captionStartId="4.[135,170,1977,1999]" captionTargetBox="[196,1415,193,1962]" captionTargetId="figure-8@4.[194,1418,193,1965]" captionTargetPageId="4" captionText="Fig. 2. Line art illustration of Centroderes impurus sp. nov. male stage-2, lateral view. Abbreviations: ij, intersegmentary joint line; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral spine; mdp, middorsal placid; mds, middorsal spine; mls, midlateral spine; mlss, midlateral sensory spot; mts, midterminal spine; mvp, midventral process; pdss, paradorsal sensory spot; pf, pectinate fringe; sdss, subdorsal sensory spot; sf, secondary fringe; vms, ventromedial spine; vmss, ventromedial sensory spot. Digits following labels refer to segment numbers." figureDoi="http://doi.org/10.5281/zenodo.4502537" httpUri="https://zenodo.org/record/4502537/files/figure.png" pageId="2" pageNumber="5">Figs. 2</figureCitation>
,
<figureCitation id="13432A34DA354E0F396CFC1AFC03FC36" box="[866,900,1555,1579]" captionStart="Fig" captionStartId="8.[136,171,1694,1716]" captionTargetBox="[136,1472,297,1678]" captionTargetId="figure-8@8.[136,1474,297,1681]" captionTargetPageId="8" captionText="Fig. 5. Light micrographs showing details of trunk morphology in Centroderes impurus sp. nov. A, C, E: holotype, ZRC.MIS.0001, male stage-2; B, D, F: paratype, KIN-846, putative J6 or adult stage-1 of unknown gender. A, Segments 36, lateral view, with dashed circles indicating laterodorsal and lateral accessory sensory spots. B, Segments 13, lateral view, showing ventrolateral tube on segment 2, which is found only in this stage and not in male stage-2. C, Segments 10 and 11, lateral view, showing crenulated spines in middorsal position on segment 10. D, Segments 56, lateral view, showing indications of lateroventral tube on segment 5, which is found only in this stage and not in male stage-2. E, Segment 811, lateral view, showing crenulated spines in midlateral positions on segment 10; note the conspicuous difference between the lateroventral spines on segment 8 and segment 9; dashed circles indicate laterodorsal and lateral accessory sensory spots. F, Segment 811, lateral view, showing acicular spines in middorsal and midlateral positions on segment 10. Abbreviations: lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdac, middorsal acicular spine; mdcr, middorsal crenulated spine; mlac, midlateral acicular spine; mlcr, midlateral crenulated spine; vlt, ventrolateral tube; vms, ventromedial spine." figureDoi="http://doi.org/10.5281/zenodo.4502551" httpUri="https://zenodo.org/record/4502551/files/figure.png" pageId="2" pageNumber="5">5B</figureCitation>
). All spines on this and following segments are at least partly covered with minute cuticular hairs, however, frequently the proximal 1/3 of the surface is smooth. At least until segment 6 or 7, the middorsal spines appear stout in male stage-2, whereas they are more slender and acicular in the putative J6 or stage-
<specimenCount id="9D7EFD38DA354E0F3E7DFCBAFB72FCD6" box="[1139,1269,1715,1739]" pageId="2" pageNumber="5" type="generic">1 specimens</specimenCount>
(compare
<figureCitation id="13432A34DA354E0F3F63FCBAFA45FCD6" box="[1389,1474,1715,1739]" captionStart="Fig" captionStartId="5.[113,148,2063,2085]" captionTargetBox="[235,1328,869,2047]" captionTargetId="figure-213@5.[235,1330,869,2047]" captionTargetPageId="5" captionText="Fig. 3. Light micrographs showing lateral overviews of Centroderes impurus sp. nov. A, holotype, ZRC.MIS.0001, male stage-2. B, paratype, KIN-846, putative J6 or adult stage-1 of unknown gender." figureDoi="http://doi.org/10.5281/zenodo.4502541" httpUri="https://zenodo.org/record/4502541/files/figure.png" pageId="2" pageNumber="5">Fig. 3A</figureCitation>
with 3B). A midventral process is present between the two ventromedial spines (
<figureCitation id="13432A34DA354E0F3E26FCFAFBF4FD16" box="[1064,1139,1779,1803]" captionStart="Fig" captionStartId="4.[135,170,1977,1999]" captionTargetBox="[196,1415,193,1962]" captionTargetId="figure-8@4.[194,1418,193,1965]" captionTargetPageId="4" captionText="Fig. 2. Line art illustration of Centroderes impurus sp. nov. male stage-2, lateral view. Abbreviations: ij, intersegmentary joint line; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral spine; mdp, middorsal placid; mds, middorsal spine; mls, midlateral spine; mlss, midlateral sensory spot; mts, midterminal spine; mvp, midventral process; pdss, paradorsal sensory spot; pf, pectinate fringe; sdss, subdorsal sensory spot; sf, secondary fringe; vms, ventromedial spine; vmss, ventromedial sensory spot. Digits following labels refer to segment numbers." figureDoi="http://doi.org/10.5281/zenodo.4502537" httpUri="https://zenodo.org/record/4502537/files/figure.png" pageId="2" pageNumber="5">Figs. 2</figureCitation>
,
<figureCitation id="13432A34DA354E0F3E71FCFAFB26FD16" box="[1151,1185,1779,1803]" captionStart="Fig" captionStartId="6.[136,171,1919,1941]" captionTargetBox="[279,1332,189,1902]" captionTargetId="figure-7@6.[278,1332,189,1904]" captionTargetPageId="6" captionText="Fig. 4. Scanning electron micrographs of male stage-2 (A, B, F, H) and putative J6 or adult stage-1 of unknown gender (CE, G) of Centroderes impurus sp. nov. A, lateral overview. B, ventral overview. C, Lateral view of segment 1. D, Detail showing lateral accessory sensory spot on segment 1. E, Region of tergosternal junctions of segments 4 and 5; dashed circles outline extended cuticular hairs on posterior margin of segment 3. F, Region of tergosternal junction of segments 4 and 5. G, lateroventral view of segments 8 and 9. H, Lateral view of segments 810 in male stage-2. Abbreviations: lvs, lateroventral spine; lvt, lateroventral tube; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; mdcr, middorsal crenulated spine; mlcr, midlateral crenulated spine; mvp, midventral process; s, segment followed by segment number; sf, secondary fringe; vms, ventromedial spine on segment 1." figureDoi="http://doi.org/10.5281/zenodo.4502543" httpUri="https://zenodo.org/record/4502543/files/figure.png" pageId="2" pageNumber="5">4B</figureCitation>
). Sensory spots are located medially on the segment in either subdorsal (male stage-2) or laterodorsal (putative J6 or stage-
<specimenCount id="9D7EFD38DA354E0F3EC3FD3CFA99FD50" box="[1229,1310,1845,1869]" pageId="2" pageNumber="5" type="adult">1 adults</specimenCount>
) positions, and always in lateral accessory position (
<figureCitation id="13432A34DA354E0F3EC7FD5CFA9AFD70" box="[1225,1309,1877,1901]" captionStart="Fig" captionStartId="6.[136,171,1919,1941]" captionTargetBox="[279,1332,189,1902]" captionTargetId="figure-7@6.[278,1332,189,1904]" captionTargetPageId="6" captionText="Fig. 4. Scanning electron micrographs of male stage-2 (A, B, F, H) and putative J6 or adult stage-1 of unknown gender (CE, G) of Centroderes impurus sp. nov. A, lateral overview. B, ventral overview. C, Lateral view of segment 1. D, Detail showing lateral accessory sensory spot on segment 1. E, Region of tergosternal junctions of segments 4 and 5; dashed circles outline extended cuticular hairs on posterior margin of segment 3. F, Region of tergosternal junction of segments 4 and 5. G, lateroventral view of segments 8 and 9. H, Lateral view of segments 810 in male stage-2. Abbreviations: lvs, lateroventral spine; lvt, lateroventral tube; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; mdcr, middorsal crenulated spine; mlcr, midlateral crenulated spine; mvp, midventral process; s, segment followed by segment number; sf, secondary fringe; vms, ventromedial spine on segment 1." figureDoi="http://doi.org/10.5281/zenodo.4502543" httpUri="https://zenodo.org/record/4502543/files/figure.png" pageId="2" pageNumber="5">Fig. 4C</figureCitation>
). Sensory spots in subdorsal/laterodorsal positions are circular and composed of micropapillae arranged around a central pore (
<figureCitation id="13432A34DA354E0F3F50FD9CFA32FDB0" box="[1374,1461,1941,1965]" captionStart="Fig" captionStartId="6.[136,171,1919,1941]" captionTargetBox="[279,1332,189,1902]" captionTargetId="figure-7@6.[278,1332,189,1904]" captionTargetPageId="6" captionText="Fig. 4. Scanning electron micrographs of male stage-2 (A, B, F, H) and putative J6 or adult stage-1 of unknown gender (CE, G) of Centroderes impurus sp. nov. A, lateral overview. B, ventral overview. C, Lateral view of segment 1. D, Detail showing lateral accessory sensory spot on segment 1. E, Region of tergosternal junctions of segments 4 and 5; dashed circles outline extended cuticular hairs on posterior margin of segment 3. F, Region of tergosternal junction of segments 4 and 5. G, lateroventral view of segments 8 and 9. H, Lateral view of segments 810 in male stage-2. Abbreviations: lvs, lateroventral spine; lvt, lateroventral tube; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; mdcr, middorsal crenulated spine; mlcr, midlateral crenulated spine; mvp, midventral process; s, segment followed by segment number; sf, secondary fringe; vms, ventromedial spine on segment 1." figureDoi="http://doi.org/10.5281/zenodo.4502543" httpUri="https://zenodo.org/record/4502543/files/figure.png" pageId="2" pageNumber="5">Fig. 4C</figureCitation>
); all sensory spots on all the following segments show the same appearance. The sensory spots in the lateral accessory positions differ though, and are more oval with cuticular papillae arranged in three rows and adhering to the cuticle surface (
<figureCitation id="13432A34DA344E0E3AC2FC8EFEA6FC82" box="[204,289,1671,1695]" captionStart="Fig" captionStartId="6.[136,171,1919,1941]" captionTargetBox="[279,1332,189,1902]" captionTargetId="figure-7@6.[278,1332,189,1904]" captionTargetPageId="6" captionText="Fig. 4. Scanning electron micrographs of male stage-2 (A, B, F, H) and putative J6 or adult stage-1 of unknown gender (CE, G) of Centroderes impurus sp. nov. A, lateral overview. B, ventral overview. C, Lateral view of segment 1. D, Detail showing lateral accessory sensory spot on segment 1. E, Region of tergosternal junctions of segments 4 and 5; dashed circles outline extended cuticular hairs on posterior margin of segment 3. F, Region of tergosternal junction of segments 4 and 5. G, lateroventral view of segments 8 and 9. H, Lateral view of segments 810 in male stage-2. Abbreviations: lvs, lateroventral spine; lvt, lateroventral tube; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; mdcr, middorsal crenulated spine; mlcr, midlateral crenulated spine; mvp, midventral process; s, segment followed by segment number; sf, secondary fringe; vms, ventromedial spine on segment 1." figureDoi="http://doi.org/10.5281/zenodo.4502543" httpUri="https://zenodo.org/record/4502543/files/figure.png" pageId="3" pageNumber="6">Fig. 4D</figureCitation>
). Minute, densely distributed cuticular hairs are present on the posterior half of the segment.
</paragraph>
<caption id="DF076639DA344E0E3A7FFAB4FEE1FB5B" pageId="3" pageNumber="6">
<paragraph id="8BC736B1DA344E0E3A7FFAB4FEE1FB5B" blockId="3.[113,1452,189,326]" pageId="3" pageNumber="6">
Table 2. Measurements from light microscopy of adult
<taxonomicName id="4C784D32DA344E0E3894FAB4FCEEFACE" authority="Sørensen &amp; Gąsiorowski &amp; Randsø &amp; Sánchez &amp; Neves, 2016" authorityName="Sørensen &amp; Gąsiorowski &amp; Randsø &amp; Sánchez &amp; Neves" authorityYear="2016" box="[666,873,189,211]" class="Kinorhyncha" family="Centroderidae" genus="Centroderes" kingdom="Animalia" order="Cyclorhagida" pageId="3" pageNumber="6" phylum="Cephalorhyncha" rank="species" species="impurus" status="sp. nov.">
<emphasis id="B90CEAA3DA344E0E3894FAB4FCEEFACE" box="[666,873,189,211]" italics="true" pageId="3" pageNumber="6">Centroderes impurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57D8DA344E0E397FFAB7FC47FACE" box="[881,960,190,211]" pageId="3" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
(in µm) from station SI-03, Singapore, including individual measures, mean and standard deviation (SD). Abbreviations: LTAS: lateral terminal accessory spine; LTS: lateral terminal spine; LV: lateroventral spine or tube; MD, middorsal spine; ML: midlateral spine; MSW-6: maximum sternal width, measured on segment 6 in this species; MTS: midterminal spine; S: segment lengths; SW-10, standard width, always measured on segment 10; TL: trunk length; VM: ventromedial spine.
</paragraph>
</caption>
<paragraph id="8BC736B1DA344E0E3A8EFB8FFADCFC3D" pageId="3" pageNumber="6">
<table id="F978C411DA34B1F23A8EFB71FADCFC3D" box="[128,1371,376,1568]" gridcols="6" gridrows="40" pageId="3" pageNumber="6">
<tr id="354834F3DA34B1F23A8EFB71FADCFBB1" box="[128,1371,376,428]" gridrow="0" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFB71FE85FBB1" box="[128,258,376,428]" gridcol="0" gridrow="0" pageId="3" pageNumber="6">
<emphasis id="B90CEAA3DA344E0E3A8EFB8FFF6EFB81" bold="true" box="[128,233,390,412]" pageId="3" pageNumber="6">Character</emphasis>
</th>
<th id="76995D8FDA34B1F23B7CFB71FD89FBB1" box="[370,526,376,428]" gridcol="1" gridrow="0" pageId="3" pageNumber="6">
<emphasis id="B90CEAA3DA344E0E3B7CFB71FE7CFBB7" bold="true" pageId="3" pageNumber="6">ZRC.MIS.0001 ♂ Holotype</emphasis>
</th>
<th id="76995D8FDA34B1F2387EFB71FD4AFBB1" box="[624,717,376,428]" gridcol="2" gridrow="0" pageId="3" pageNumber="6">
<emphasis id="B90CEAA3DA344E0E387CFB71FD4AFBB7" bold="true" pageId="3" pageNumber="6">KIN-846 Paratype</emphasis>
</th>
<th id="76995D8FDA34B1F23941FB71FC2BFBB1" box="[847,940,376,428]" gridcol="3" gridrow="0" pageId="3" pageNumber="6">
<emphasis id="B90CEAA3DA344E0E395FFB71FC2BFBB7" bold="true" pageId="3" pageNumber="6">KIN-847 Paratype</emphasis>
</th>
<th id="76995D8FDA34B1F23E30FB71FBFBFBB1" box="[1086,1148,376,428]" gridcol="4" gridrow="0" pageId="3" pageNumber="6">
<emphasis id="B90CEAA3DA344E0E3E4EFB8EFBFCFB81" bold="true" box="[1088,1147,391,412]" pageId="3" pageNumber="6">Mean</emphasis>
</th>
<th id="76995D8FDA34B1F23F13FB71FADCFBB1" box="[1309,1371,376,428]" gridcol="5" gridrow="0" pageId="3" pageNumber="6">
<emphasis id="B90CEAA3DA344E0E3F23FB8FFACBFB81" bold="true" box="[1325,1356,390,412]" pageId="3" pageNumber="6">SD</emphasis>
</th>
</tr>
<tr id="354834F3DA34B1F23A8EFBCDFADCFBC7" box="[128,1371,452,474]" gridrow="1" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFBCDFE85FBC7" box="[128,258,452,474]" gridcol="0" gridrow="1" pageId="3" pageNumber="6">TL</th>
<td id="76995D8FDA34B1F23B7CFBCDFD89FBC7" box="[370,526,452,474]" gridcol="1" gridrow="1" pageId="3" pageNumber="6">376</td>
<td id="76995D8FDA34B1F2387EFBCDFD4AFBC7" box="[624,717,452,474]" gridcol="2" gridrow="1" pageId="3" pageNumber="6">264</td>
<td id="76995D8FDA34B1F23941FBCDFC2BFBC7" box="[847,940,452,474]" gridcol="3" gridrow="1" pageId="3" pageNumber="6">255</td>
<td id="76995D8FDA34B1F23E30FBCDFBFBFBC7" box="[1086,1148,452,474]" gridcol="4" gridrow="1" pageId="3" pageNumber="6">298</td>
<td id="76995D8FDA34B1F23F13FBCDFADCFBC7" box="[1309,1371,452,474]" gridcol="5" gridrow="1" pageId="3" pageNumber="6">67.41</td>
</tr>
<tr id="354834F3DA34B1F23A8EFBE9FADCFBEB" box="[128,1371,480,502]" gridrow="2" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFBE9FE85FBEB" box="[128,258,480,502]" gridcol="0" gridrow="2" pageId="3" pageNumber="6">MSW-6</th>
<td id="76995D8FDA34B1F23B7CFBE9FD89FBEB" box="[370,526,480,502]" gridcol="1" gridrow="2" pageId="3" pageNumber="6">64</td>
<td id="76995D8FDA34B1F2387EFBE9FD4AFBEB" box="[624,717,480,502]" gridcol="2" gridrow="2" pageId="3" pageNumber="6">62</td>
<td id="76995D8FDA34B1F23941FBE9FC2BFBEB" box="[847,940,480,502]" gridcol="3" gridrow="2" pageId="3" pageNumber="6">57</td>
<td id="76995D8FDA34B1F23E30FBE9FBFBFBEB" box="[1086,1148,480,502]" gridcol="4" gridrow="2" pageId="3" pageNumber="6">61</td>
<td id="76995D8FDA34B1F23F13FBE9FADCFBEB" box="[1309,1371,480,502]" gridcol="5" gridrow="2" pageId="3" pageNumber="6">3.61</td>
</tr>
<tr id="354834F3DA34B1F23A8EFBF4FADCF80E" box="[128,1371,509,531]" gridrow="3" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFBF4FE85F80E" box="[128,258,509,531]" gridcol="0" gridrow="3" pageId="3" pageNumber="6">MSW-6/TL</th>
<td id="76995D8FDA34B1F23B7CFBF4FD89F80E" box="[370,526,509,531]" gridcol="1" gridrow="3" pageId="3" pageNumber="6">17.0%</td>
<td id="76995D8FDA34B1F2387EFBF4FD4AF80E" box="[624,717,509,531]" gridcol="2" gridrow="3" pageId="3" pageNumber="6">23.5%</td>
<td id="76995D8FDA34B1F23941FBF4FC2BF80E" box="[847,940,509,531]" gridcol="3" gridrow="3" pageId="3" pageNumber="6">22.4%</td>
<td id="76995D8FDA34B1F23E30FBF4FBFBF80E" box="[1086,1148,509,531]" gridcol="4" gridrow="3" pageId="3" pageNumber="6">21%</td>
<td id="76995D8FDA34B1F23F13FBF4FADCF80E" box="[1309,1371,509,531]" gridcol="5" gridrow="3" pageId="3" pageNumber="6">3.45%</td>
</tr>
<tr id="354834F3DA34B1F23A8EF813FADCF82D" box="[128,1371,538,560]" gridrow="4" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF813FE85F82D" box="[128,258,538,560]" gridcol="0" gridrow="4" pageId="3" pageNumber="6">SW-10</th>
<td id="76995D8FDA34B1F23B7CF813FD89F82D" box="[370,526,538,560]" gridcol="1" gridrow="4" pageId="3" pageNumber="6">45</td>
<td id="76995D8FDA34B1F2387EF813FD4AF82D" box="[624,717,538,560]" gridcol="2" gridrow="4" pageId="3" pageNumber="6"></td>
<td id="76995D8FDA34B1F23941F813FC2BF82D" box="[847,940,538,560]" gridcol="3" gridrow="4" pageId="3" pageNumber="6">19</td>
<td id="76995D8FDA34B1F23E30F813FBFBF82D" box="[1086,1148,538,560]" gridcol="4" gridrow="4" pageId="3" pageNumber="6">32</td>
<td id="76995D8FDA34B1F23F13F813FADCF82D" box="[1309,1371,538,560]" gridcol="5" gridrow="4" pageId="3" pageNumber="6">18.38</td>
</tr>
<tr id="354834F3DA34B1F23A8EF83EFADCF850" box="[128,1371,567,589]" gridrow="5" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF83EFE85F850" box="[128,258,567,589]" gridcol="0" gridrow="5" pageId="3" pageNumber="6">SW-10/TL</th>
<td id="76995D8FDA34B1F23B7CF83EFD89F850" box="[370,526,567,589]" gridcol="1" gridrow="5" pageId="3" pageNumber="6">12.0%</td>
<td id="76995D8FDA34B1F2387EF83EFD4AF850" box="[624,717,567,589]" gridcol="2" gridrow="5" pageId="3" pageNumber="6"></td>
<td id="76995D8FDA34B1F23941F83EFC2BF850" box="[847,940,567,589]" gridcol="3" gridrow="5" pageId="3" pageNumber="6">7.5%</td>
<td id="76995D8FDA34B1F23E30F83EFBFBF850" box="[1086,1148,567,589]" gridcol="4" gridrow="5" pageId="3" pageNumber="6">9.7%</td>
<td id="76995D8FDA34B1F23F13F83EFADCF850" box="[1309,1371,567,589]" gridcol="5" gridrow="5" pageId="3" pageNumber="6">3.19%</td>
</tr>
<tr id="354834F3DA34B1F23A8EF85DFADCF877" box="[128,1371,596,618]" gridrow="6" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF85DFE85F877" box="[128,258,596,618]" gridcol="0" gridrow="6" pageId="3" pageNumber="6">S1</th>
<td id="76995D8FDA34B1F23B7CF85DFD89F877" box="[370,526,596,618]" gridcol="1" gridrow="6" pageId="3" pageNumber="6">36</td>
<td id="76995D8FDA34B1F2387EF85DFD4AF877" box="[624,717,596,618]" gridcol="2" gridrow="6" pageId="3" pageNumber="6">29</td>
<td id="76995D8FDA34B1F23941F85DFC2BF877" box="[847,940,596,618]" gridcol="3" gridrow="6" pageId="3" pageNumber="6">28</td>
<td id="76995D8FDA34B1F23E30F85DFBFBF877" box="[1086,1148,596,618]" gridcol="4" gridrow="6" pageId="3" pageNumber="6">32</td>
<td id="76995D8FDA34B1F23F13F85DFADCF877" box="[1309,1371,596,618]" gridcol="5" gridrow="6" pageId="3" pageNumber="6">6.08</td>
</tr>
<tr id="354834F3DA34B1F23A8EF879FADCF89B" box="[128,1371,624,646]" gridrow="7" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF879FE85F89B" box="[128,258,624,646]" gridcol="0" gridrow="7" pageId="3" pageNumber="6">S2</th>
<td id="76995D8FDA34B1F23B7CF879FD89F89B" box="[370,526,624,646]" gridcol="1" gridrow="7" pageId="3" pageNumber="6">40</td>
<td id="76995D8FDA34B1F2387EF879FD4AF89B" box="[624,717,624,646]" gridcol="2" gridrow="7" pageId="3" pageNumber="6">24</td>
<td id="76995D8FDA34B1F23941F879FC2BF89B" box="[847,940,624,646]" gridcol="3" gridrow="7" pageId="3" pageNumber="6">24</td>
<td id="76995D8FDA34B1F23E30F879FBFBF89B" box="[1086,1148,624,646]" gridcol="4" gridrow="7" pageId="3" pageNumber="6">28</td>
<td id="76995D8FDA34B1F23F13F879FADCF89B" box="[1309,1371,624,646]" gridcol="5" gridrow="7" pageId="3" pageNumber="6">7.51</td>
</tr>
<tr id="354834F3DA34B1F23A8EF884FADCF8BE" box="[128,1371,653,675]" gridrow="8" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF884FE85F8BE" box="[128,258,653,675]" gridcol="0" gridrow="8" pageId="3" pageNumber="6">S3</th>
<td id="76995D8FDA34B1F23B7CF884FD89F8BE" box="[370,526,653,675]" gridcol="1" gridrow="8" pageId="3" pageNumber="6">42</td>
<td id="76995D8FDA34B1F2387EF884FD4AF8BE" box="[624,717,653,675]" gridcol="2" gridrow="8" pageId="3" pageNumber="6">25</td>
<td id="76995D8FDA34B1F23941F884FC2BF8BE" box="[847,940,653,675]" gridcol="3" gridrow="8" pageId="3" pageNumber="6">24</td>
<td id="76995D8FDA34B1F23E30F884FBFBF8BE" box="[1086,1148,653,675]" gridcol="4" gridrow="8" pageId="3" pageNumber="6">30</td>
<td id="76995D8FDA34B1F23F13F884FADCF8BE" box="[1309,1371,653,675]" gridcol="5" gridrow="8" pageId="3" pageNumber="6">8.96</td>
</tr>
<tr id="354834F3DA34B1F23A8EF8A3FADCF8DD" box="[128,1371,682,704]" gridrow="9" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF8A3FE85F8DD" box="[128,258,682,704]" gridcol="0" gridrow="9" pageId="3" pageNumber="6">S4</th>
<td id="76995D8FDA34B1F23B7CF8A3FD89F8DD" box="[370,526,682,704]" gridcol="1" gridrow="9" pageId="3" pageNumber="6">47</td>
<td id="76995D8FDA34B1F2387EF8A3FD4AF8DD" box="[624,717,682,704]" gridcol="2" gridrow="9" pageId="3" pageNumber="6">27</td>
<td id="76995D8FDA34B1F23941F8A3FC2BF8DD" box="[847,940,682,704]" gridcol="3" gridrow="9" pageId="3" pageNumber="6">27</td>
<td id="76995D8FDA34B1F23E30F8A3FBFBF8DD" box="[1086,1148,682,704]" gridcol="4" gridrow="9" pageId="3" pageNumber="6">32</td>
<td id="76995D8FDA34B1F23F13F8A3FADCF8DD" box="[1309,1371,682,704]" gridcol="5" gridrow="9" pageId="3" pageNumber="6">8.66</td>
</tr>
<tr id="354834F3DA34B1F23A8EF8CEFADCF8C0" box="[128,1371,711,733]" gridrow="10" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF8CEFE85F8C0" box="[128,258,711,733]" gridcol="0" gridrow="10" pageId="3" pageNumber="6">S5</th>
<td id="76995D8FDA34B1F23B7CF8CEFD89F8C0" box="[370,526,711,733]" gridcol="1" gridrow="10" pageId="3" pageNumber="6">50</td>
<td id="76995D8FDA34B1F2387EF8CEFD4AF8C0" box="[624,717,711,733]" gridcol="2" gridrow="10" pageId="3" pageNumber="6">27</td>
<td id="76995D8FDA34B1F23941F8CEFC2BF8C0" box="[847,940,711,733]" gridcol="3" gridrow="10" pageId="3" pageNumber="6">27</td>
<td id="76995D8FDA34B1F23E30F8CEFBFBF8C0" box="[1086,1148,711,733]" gridcol="4" gridrow="10" pageId="3" pageNumber="6">34</td>
<td id="76995D8FDA34B1F23F13F8CEFADCF8C0" box="[1309,1371,711,733]" gridcol="5" gridrow="10" pageId="3" pageNumber="6">11.55</td>
</tr>
<tr id="354834F3DA34B1F23A8EF8EDFADCF8E7" box="[128,1371,740,762]" gridrow="11" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF8EDFE85F8E7" box="[128,258,740,762]" gridcol="0" gridrow="11" pageId="3" pageNumber="6">S6</th>
<td id="76995D8FDA34B1F23B7CF8EDFD89F8E7" box="[370,526,740,762]" gridcol="1" gridrow="11" pageId="3" pageNumber="6">52</td>
<td id="76995D8FDA34B1F2387EF8EDFD4AF8E7" box="[624,717,740,762]" gridcol="2" gridrow="11" pageId="3" pageNumber="6">32</td>
<td id="76995D8FDA34B1F23941F8EDFC2BF8E7" box="[847,940,740,762]" gridcol="3" gridrow="11" pageId="3" pageNumber="6">28</td>
<td id="76995D8FDA34B1F23E30F8EDFBFBF8E7" box="[1086,1148,740,762]" gridcol="4" gridrow="11" pageId="3" pageNumber="6">35</td>
<td id="76995D8FDA34B1F23F13F8EDFADCF8E7" box="[1309,1371,740,762]" gridcol="5" gridrow="11" pageId="3" pageNumber="6">9.45</td>
</tr>
<tr id="354834F3DA34B1F23A8EF909FADCF90B" box="[128,1371,768,790]" gridrow="12" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF909FE85F90B" box="[128,258,768,790]" gridcol="0" gridrow="12" pageId="3" pageNumber="6">S7</th>
<td id="76995D8FDA34B1F23B7CF909FD89F90B" box="[370,526,768,790]" gridcol="1" gridrow="12" pageId="3" pageNumber="6">55</td>
<td id="76995D8FDA34B1F2387EF909FD4AF90B" box="[624,717,768,790]" gridcol="2" gridrow="12" pageId="3" pageNumber="6">28</td>
<td id="76995D8FDA34B1F23941F909FC2BF90B" box="[847,940,768,790]" gridcol="3" gridrow="12" pageId="3" pageNumber="6">27</td>
<td id="76995D8FDA34B1F23E30F909FBFBF90B" box="[1086,1148,768,790]" gridcol="4" gridrow="12" pageId="3" pageNumber="6">34</td>
<td id="76995D8FDA34B1F23F13F909FADCF90B" box="[1309,1371,768,790]" gridcol="5" gridrow="12" pageId="3" pageNumber="6">10.69</td>
</tr>
<tr id="354834F3DA34B1F23A8EF914FADCF92E" box="[128,1371,797,819]" gridrow="13" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF914FE85F92E" box="[128,258,797,819]" gridcol="0" gridrow="13" pageId="3" pageNumber="6">S8</th>
<td id="76995D8FDA34B1F23B7CF914FD89F92E" box="[370,526,797,819]" gridcol="1" gridrow="13" pageId="3" pageNumber="6">58</td>
<td id="76995D8FDA34B1F2387EF914FD4AF92E" box="[624,717,797,819]" gridcol="2" gridrow="13" pageId="3" pageNumber="6">32</td>
<td id="76995D8FDA34B1F23941F914FC2BF92E" box="[847,940,797,819]" gridcol="3" gridrow="13" pageId="3" pageNumber="6">28</td>
<td id="76995D8FDA34B1F23E30F914FBFBF92E" box="[1086,1148,797,819]" gridcol="4" gridrow="13" pageId="3" pageNumber="6">37</td>
<td id="76995D8FDA34B1F23F13F914FADCF92E" box="[1309,1371,797,819]" gridcol="5" gridrow="13" pageId="3" pageNumber="6">11.72</td>
</tr>
<tr id="354834F3DA34B1F23A8EF933FADCF94D" box="[128,1371,826,848]" gridrow="14" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF933FE85F94D" box="[128,258,826,848]" gridcol="0" gridrow="14" pageId="3" pageNumber="6">S9</th>
<td id="76995D8FDA34B1F23B7CF933FD89F94D" box="[370,526,826,848]" gridcol="1" gridrow="14" pageId="3" pageNumber="6">58</td>
<td id="76995D8FDA34B1F2387EF933FD4AF94D" box="[624,717,826,848]" gridcol="2" gridrow="14" pageId="3" pageNumber="6">28</td>
<td id="76995D8FDA34B1F23941F933FC2BF94D" box="[847,940,826,848]" gridcol="3" gridrow="14" pageId="3" pageNumber="6">24</td>
<td id="76995D8FDA34B1F23E30F933FBFBF94D" box="[1086,1148,826,848]" gridcol="4" gridrow="14" pageId="3" pageNumber="6">34</td>
<td id="76995D8FDA34B1F23F13F933FADCF94D" box="[1309,1371,826,848]" gridcol="5" gridrow="14" pageId="3" pageNumber="6">13.43</td>
</tr>
<tr id="354834F3DA34B1F23A8EF95EFADCF970" box="[128,1371,855,877]" gridrow="15" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF95EFE85F970" box="[128,258,855,877]" gridcol="0" gridrow="15" pageId="3" pageNumber="6">S10</th>
<td id="76995D8FDA34B1F23B7CF95EFD89F970" box="[370,526,855,877]" gridcol="1" gridrow="15" pageId="3" pageNumber="6">39</td>
<td id="76995D8FDA34B1F2387EF95EFD4AF970" box="[624,717,855,877]" gridcol="2" gridrow="15" pageId="3" pageNumber="6">24</td>
<td id="76995D8FDA34B1F23941F95EFC2BF970" box="[847,940,855,877]" gridcol="3" gridrow="15" pageId="3" pageNumber="6">20</td>
<td id="76995D8FDA34B1F23E30F95EFBFBF970" box="[1086,1148,855,877]" gridcol="4" gridrow="15" pageId="3" pageNumber="6">28</td>
<td id="76995D8FDA34B1F23F13F95EFADCF970" box="[1309,1371,855,877]" gridcol="5" gridrow="15" pageId="3" pageNumber="6">10.58</td>
</tr>
<tr id="354834F3DA34B1F23A8EF97DFADCF997" box="[128,1371,884,906]" gridrow="16" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF97DFE85F997" box="[128,258,884,906]" gridcol="0" gridrow="16" pageId="3" pageNumber="6">S11</th>
<td id="76995D8FDA34B1F23B7CF97DFD89F997" box="[370,526,884,906]" gridcol="1" gridrow="16" pageId="3" pageNumber="6">28</td>
<td id="76995D8FDA34B1F2387EF97DFD4AF997" box="[624,717,884,906]" gridcol="2" gridrow="16" pageId="3" pageNumber="6">21</td>
<td id="76995D8FDA34B1F23941F97DFC2BF997" box="[847,940,884,906]" gridcol="3" gridrow="16" pageId="3" pageNumber="6">17</td>
<td id="76995D8FDA34B1F23E30F97DFBFBF997" box="[1086,1148,884,906]" gridcol="4" gridrow="16" pageId="3" pageNumber="6">22</td>
<td id="76995D8FDA34B1F23F13F97DFADCF997" box="[1309,1371,884,906]" gridcol="5" gridrow="16" pageId="3" pageNumber="6">5.57</td>
</tr>
<tr id="354834F3DA34B1F23A8EF999FADCF9BB" box="[128,1371,912,934]" gridrow="17" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF999FE85F9BB" box="[128,258,912,934]" gridcol="0" gridrow="17" pageId="3" pageNumber="6">MD1</th>
<td id="76995D8FDA34B1F23B7CF999FD89F9BB" box="[370,526,912,934]" gridcol="1" gridrow="17" pageId="3" pageNumber="6">16</td>
<td id="76995D8FDA34B1F2387EF999FD4AF9BB" box="[624,717,912,934]" gridcol="2" gridrow="17" pageId="3" pageNumber="6">17</td>
<td id="76995D8FDA34B1F23941F999FC2BF9BB" box="[847,940,912,934]" gridcol="3" gridrow="17" pageId="3" pageNumber="6">15</td>
<td id="76995D8FDA34B1F23E30F999FBFBF9BB" box="[1086,1148,912,934]" gridcol="4" gridrow="17" pageId="3" pageNumber="6">16</td>
<td id="76995D8FDA34B1F23F13F999FADCF9BB" box="[1309,1371,912,934]" gridcol="5" gridrow="17" pageId="3" pageNumber="6">1.00</td>
</tr>
<tr id="354834F3DA34B1F23A8EF9A4FADCF9DE" box="[128,1371,941,963]" gridrow="18" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF9A4FE85F9DE" box="[128,258,941,963]" gridcol="0" gridrow="18" pageId="3" pageNumber="6">MD2</th>
<td id="76995D8FDA34B1F23B7CF9A4FD89F9DE" box="[370,526,941,963]" gridcol="1" gridrow="18" pageId="3" pageNumber="6">16</td>
<td id="76995D8FDA34B1F2387EF9A4FD4AF9DE" box="[624,717,941,963]" gridcol="2" gridrow="18" pageId="3" pageNumber="6">16</td>
<td id="76995D8FDA34B1F23941F9A4FC2BF9DE" box="[847,940,941,963]" gridcol="3" gridrow="18" pageId="3" pageNumber="6">17</td>
<td id="76995D8FDA34B1F23E30F9A4FBFBF9DE" box="[1086,1148,941,963]" gridcol="4" gridrow="18" pageId="3" pageNumber="6">16</td>
<td id="76995D8FDA34B1F23F13F9A4FADCF9DE" box="[1309,1371,941,963]" gridcol="5" gridrow="18" pageId="3" pageNumber="6">0.58</td>
</tr>
<tr id="354834F3DA34B1F23A8EF9C3FADCF9FD" box="[128,1371,970,992]" gridrow="19" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF9C3FE85F9FD" box="[128,258,970,992]" gridcol="0" gridrow="19" pageId="3" pageNumber="6">MD3</th>
<td id="76995D8FDA34B1F23B7CF9C3FD89F9FD" box="[370,526,970,992]" gridcol="1" gridrow="19" pageId="3" pageNumber="6">16</td>
<td id="76995D8FDA34B1F2387EF9C3FD4AF9FD" box="[624,717,970,992]" gridcol="2" gridrow="19" pageId="3" pageNumber="6">21</td>
<td id="76995D8FDA34B1F23941F9C3FC2BF9FD" box="[847,940,970,992]" gridcol="3" gridrow="19" pageId="3" pageNumber="6">18</td>
<td id="76995D8FDA34B1F23E30F9C3FBFBF9FD" box="[1086,1148,970,992]" gridcol="4" gridrow="19" pageId="3" pageNumber="6">18</td>
<td id="76995D8FDA34B1F23F13F9C3FADCF9FD" box="[1309,1371,970,992]" gridcol="5" gridrow="19" pageId="3" pageNumber="6">2.52</td>
</tr>
<tr id="354834F3DA34B1F23A8EF9EEFADCF9E0" box="[128,1371,999,1021]" gridrow="20" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EF9EEFE85F9E0" box="[128,258,999,1021]" gridcol="0" gridrow="20" pageId="3" pageNumber="6">MD4</th>
<td id="76995D8FDA34B1F23B7CF9EEFD89F9E0" box="[370,526,999,1021]" gridcol="1" gridrow="20" pageId="3" pageNumber="6">16</td>
<td id="76995D8FDA34B1F2387EF9EEFD4AF9E0" box="[624,717,999,1021]" gridcol="2" gridrow="20" pageId="3" pageNumber="6">20</td>
<td id="76995D8FDA34B1F23941F9EEFC2BF9E0" box="[847,940,999,1021]" gridcol="3" gridrow="20" pageId="3" pageNumber="6">21</td>
<td id="76995D8FDA34B1F23E30F9EEFBFBF9E0" box="[1086,1148,999,1021]" gridcol="4" gridrow="20" pageId="3" pageNumber="6">19</td>
<td id="76995D8FDA34B1F23F13F9EEFADCF9E0" box="[1309,1371,999,1021]" gridcol="5" gridrow="20" pageId="3" pageNumber="6">2.65</td>
</tr>
<tr id="354834F3DA34B1F23A8EFE0DFADCFE07" box="[128,1371,1028,1050]" gridrow="21" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFE0DFE85FE07" box="[128,258,1028,1050]" gridcol="0" gridrow="21" pageId="3" pageNumber="6">MD5</th>
<td id="76995D8FDA34B1F23B7CFE0DFD89FE07" box="[370,526,1028,1050]" gridcol="1" gridrow="21" pageId="3" pageNumber="6">19</td>
<td id="76995D8FDA34B1F2387EFE0DFD4AFE07" box="[624,717,1028,1050]" gridcol="2" gridrow="21" pageId="3" pageNumber="6">23</td>
<td id="76995D8FDA34B1F23941FE0DFC2BFE07" box="[847,940,1028,1050]" gridcol="3" gridrow="21" pageId="3" pageNumber="6">23</td>
<td id="76995D8FDA34B1F23E30FE0DFBFBFE07" box="[1086,1148,1028,1050]" gridcol="4" gridrow="21" pageId="3" pageNumber="6">22</td>
<td id="76995D8FDA34B1F23F13FE0DFADCFE07" box="[1309,1371,1028,1050]" gridcol="5" gridrow="21" pageId="3" pageNumber="6">2.31</td>
</tr>
<tr id="354834F3DA34B1F23A8EFE29FADCFE2B" box="[128,1371,1056,1078]" gridrow="22" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFE29FE85FE2B" box="[128,258,1056,1078]" gridcol="0" gridrow="22" pageId="3" pageNumber="6">MD6</th>
<td id="76995D8FDA34B1F23B7CFE29FD89FE2B" box="[370,526,1056,1078]" gridcol="1" gridrow="22" pageId="3" pageNumber="6">19</td>
<td id="76995D8FDA34B1F2387EFE29FD4AFE2B" box="[624,717,1056,1078]" gridcol="2" gridrow="22" pageId="3" pageNumber="6">28</td>
<td id="76995D8FDA34B1F23941FE29FC2BFE2B" box="[847,940,1056,1078]" gridcol="3" gridrow="22" pageId="3" pageNumber="6">26</td>
<td id="76995D8FDA34B1F23E30FE29FBFBFE2B" box="[1086,1148,1056,1078]" gridcol="4" gridrow="22" pageId="3" pageNumber="6">24</td>
<td id="76995D8FDA34B1F23F13FE29FADCFE2B" box="[1309,1371,1056,1078]" gridcol="5" gridrow="22" pageId="3" pageNumber="6">4.73</td>
</tr>
<tr id="354834F3DA34B1F23A8EFE34FADCFE4E" box="[128,1371,1085,1107]" gridrow="23" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFE34FE85FE4E" box="[128,258,1085,1107]" gridcol="0" gridrow="23" pageId="3" pageNumber="6">MD7</th>
<td id="76995D8FDA34B1F23B7CFE34FD89FE4E" box="[370,526,1085,1107]" gridcol="1" gridrow="23" pageId="3" pageNumber="6">23</td>
<td id="76995D8FDA34B1F2387EFE34FD4AFE4E" box="[624,717,1085,1107]" gridcol="2" gridrow="23" pageId="3" pageNumber="6">32</td>
<td id="76995D8FDA34B1F23941FE34FC2BFE4E" box="[847,940,1085,1107]" gridcol="3" gridrow="23" pageId="3" pageNumber="6">28</td>
<td id="76995D8FDA34B1F23E30FE34FBFBFE4E" box="[1086,1148,1085,1107]" gridcol="4" gridrow="23" pageId="3" pageNumber="6">28</td>
<td id="76995D8FDA34B1F23F13FE34FADCFE4E" box="[1309,1371,1085,1107]" gridcol="5" gridrow="23" pageId="3" pageNumber="6">4.51</td>
</tr>
<tr id="354834F3DA34B1F23A8EFE53FADCFE6D" box="[128,1371,1114,1136]" gridrow="24" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFE53FE85FE6D" box="[128,258,1114,1136]" gridcol="0" gridrow="24" pageId="3" pageNumber="6">MD8</th>
<td id="76995D8FDA34B1F23B7CFE53FD89FE6D" box="[370,526,1114,1136]" gridcol="1" gridrow="24" pageId="3" pageNumber="6">30</td>
<td id="76995D8FDA34B1F2387EFE53FD4AFE6D" box="[624,717,1114,1136]" gridcol="2" gridrow="24" pageId="3" pageNumber="6">37</td>
<td id="76995D8FDA34B1F23941FE53FC2BFE6D" box="[847,940,1114,1136]" gridcol="3" gridrow="24" pageId="3" pageNumber="6">33</td>
<td id="76995D8FDA34B1F23E30FE53FBFBFE6D" box="[1086,1148,1114,1136]" gridcol="4" gridrow="24" pageId="3" pageNumber="6">33</td>
<td id="76995D8FDA34B1F23F13FE53FADCFE6D" box="[1309,1371,1114,1136]" gridcol="5" gridrow="24" pageId="3" pageNumber="6">3.51</td>
</tr>
<tr id="354834F3DA34B1F23A8EFE7EFADCFE90" box="[128,1371,1143,1165]" gridrow="25" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFE7EFE85FE90" box="[128,258,1143,1165]" gridcol="0" gridrow="25" pageId="3" pageNumber="6">MD9</th>
<td id="76995D8FDA34B1F23B7CFE7EFD89FE90" box="[370,526,1143,1165]" gridcol="1" gridrow="25" pageId="3" pageNumber="6">33</td>
<td id="76995D8FDA34B1F2387EFE7EFD4AFE90" box="[624,717,1143,1165]" gridcol="2" gridrow="25" pageId="3" pageNumber="6">37</td>
<td id="76995D8FDA34B1F23941FE7EFC2BFE90" box="[847,940,1143,1165]" gridcol="3" gridrow="25" pageId="3" pageNumber="6">36</td>
<td id="76995D8FDA34B1F23E30FE7EFBFBFE90" box="[1086,1148,1143,1165]" gridcol="4" gridrow="25" pageId="3" pageNumber="6">35</td>
<td id="76995D8FDA34B1F23F13FE7EFADCFE90" box="[1309,1371,1143,1165]" gridcol="5" gridrow="25" pageId="3" pageNumber="6">2.08</td>
</tr>
<tr id="354834F3DA34B1F23A8EFE9DFADCFEB4" box="[128,1371,1172,1193]" gridrow="26" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFE9DFE85FEB4" box="[128,258,1172,1193]" gridcol="0" gridrow="26" pageId="3" pageNumber="6">MD10</th>
<td id="76995D8FDA34B1F23B7CFE9DFD89FEB4" box="[370,526,1172,1193]" gridcol="1" gridrow="26" pageId="3" pageNumber="6">28</td>
<td id="76995D8FDA34B1F2387EFE9DFD4AFEB4" box="[624,717,1172,1193]" gridcol="2" gridrow="26" pageId="3" pageNumber="6">48</td>
<td id="76995D8FDA34B1F23941FE9DFC2BFEB4" box="[847,940,1172,1193]" gridcol="3" gridrow="26" pageId="3" pageNumber="6">47</td>
<td id="76995D8FDA34B1F23E30FE9DFBFBFEB4" box="[1086,1148,1172,1193]" gridcol="4" gridrow="26" pageId="3" pageNumber="6">41</td>
<td id="76995D8FDA34B1F23F13FE9DFADCFEB4" box="[1309,1371,1172,1193]" gridcol="5" gridrow="26" pageId="3" pageNumber="6">11.27</td>
</tr>
<tr id="354834F3DA34B1F23A8EFEB9FADCFEDB" box="[128,1371,1200,1222]" gridrow="27" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFEB9FE85FEDB" box="[128,258,1200,1222]" gridcol="0" gridrow="27" pageId="3" pageNumber="6">MD11</th>
<td id="76995D8FDA34B1F23B7CFEB9FD89FEDB" box="[370,526,1200,1222]" gridcol="1" gridrow="27" pageId="3" pageNumber="6">34</td>
<td id="76995D8FDA34B1F2387EFEB9FD4AFEDB" box="[624,717,1200,1222]" gridcol="2" gridrow="27" pageId="3" pageNumber="6">41</td>
<td id="76995D8FDA34B1F23941FEB9FC2BFEDB" box="[847,940,1200,1222]" gridcol="3" gridrow="27" pageId="3" pageNumber="6">50</td>
<td id="76995D8FDA34B1F23E30FEB9FBFBFEDB" box="[1086,1148,1200,1222]" gridcol="4" gridrow="27" pageId="3" pageNumber="6">42</td>
<td id="76995D8FDA34B1F23F13FEB9FADCFEDB" box="[1309,1371,1200,1222]" gridcol="5" gridrow="27" pageId="3" pageNumber="6">8.02</td>
</tr>
<tr id="354834F3DA34B1F23A8EFEC4FADCFEFE" box="[128,1371,1229,1251]" gridrow="28" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFEC4FE85FEFE" box="[128,258,1229,1251]" gridcol="0" gridrow="28" pageId="3" pageNumber="6">VM1</th>
<td id="76995D8FDA34B1F23B7CFEC4FD89FEFE" box="[370,526,1229,1251]" gridcol="1" gridrow="28" pageId="3" pageNumber="6">62</td>
<td id="76995D8FDA34B1F2387EFEC4FD4AFEFE" box="[624,717,1229,1251]" gridcol="2" gridrow="28" pageId="3" pageNumber="6">53</td>
<td id="76995D8FDA34B1F23941FEC4FC2BFEFE" box="[847,940,1229,1251]" gridcol="3" gridrow="28" pageId="3" pageNumber="6">52</td>
<td id="76995D8FDA34B1F23E30FEC4FBFBFEFE" box="[1086,1148,1229,1251]" gridcol="4" gridrow="28" pageId="3" pageNumber="6">56</td>
<td id="76995D8FDA34B1F23F13FEC4FADCFEFE" box="[1309,1371,1229,1251]" gridcol="5" gridrow="28" pageId="3" pageNumber="6">5.51</td>
</tr>
<tr id="354834F3DA34B1F23A8EFEE3FADCFF1D" box="[128,1371,1258,1280]" gridrow="29" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFEE3FE85FF1D" box="[128,258,1258,1280]" gridcol="0" gridrow="29" pageId="3" pageNumber="6">LV2</th>
<td id="76995D8FDA34B1F23B7CFEE3FD89FF1D" box="[370,526,1258,1280]" gridcol="1" gridrow="29" pageId="3" pageNumber="6"></td>
<td id="76995D8FDA34B1F2387EFEE3FD4AFF1D" box="[624,717,1258,1280]" gridcol="2" gridrow="29" pageId="3" pageNumber="6">8</td>
<td id="76995D8FDA34B1F23941FEE3FC2BFF1D" box="[847,940,1258,1280]" gridcol="3" gridrow="29" pageId="3" pageNumber="6"></td>
<td id="76995D8FDA34B1F23E30FEE3FBFBFF1D" box="[1086,1148,1258,1280]" gridcol="4" gridrow="29" pageId="3" pageNumber="6">8</td>
<td id="76995D8FDA34B1F23F13FEE3FADCFF1D" box="[1309,1371,1258,1280]" gridcol="5" gridrow="29" pageId="3" pageNumber="6"></td>
</tr>
<tr id="354834F3DA34B1F23A8EFF0EFADCFF00" box="[128,1371,1287,1309]" gridrow="30" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFF0EFE85FF00" box="[128,258,1287,1309]" gridcol="0" gridrow="30" pageId="3" pageNumber="6">LV5</th>
<td id="76995D8FDA34B1F23B7CFF0EFD89FF00" box="[370,526,1287,1309]" gridcol="1" gridrow="30" pageId="3" pageNumber="6"></td>
<td id="76995D8FDA34B1F2387EFF0EFD4AFF00" box="[624,717,1287,1309]" gridcol="2" gridrow="30" pageId="3" pageNumber="6">9</td>
<td id="76995D8FDA34B1F23941FF0EFC2BFF00" box="[847,940,1287,1309]" gridcol="3" gridrow="30" pageId="3" pageNumber="6">10</td>
<td id="76995D8FDA34B1F23E30FF0EFBFBFF00" box="[1086,1148,1287,1309]" gridcol="4" gridrow="30" pageId="3" pageNumber="6">10</td>
<td id="76995D8FDA34B1F23F13FF0EFADCFF00" box="[1309,1371,1287,1309]" gridcol="5" gridrow="30" pageId="3" pageNumber="6">0.71</td>
</tr>
<tr id="354834F3DA34B1F23A8EFF2DFADCFF27" box="[128,1371,1316,1338]" gridrow="31" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFF2DFE85FF27" box="[128,258,1316,1338]" gridcol="0" gridrow="31" pageId="3" pageNumber="6">LV8</th>
<td id="76995D8FDA34B1F23B7CFF2DFD89FF27" box="[370,526,1316,1338]" gridcol="1" gridrow="31" pageId="3" pageNumber="6">18</td>
<td id="76995D8FDA34B1F2387EFF2DFD4AFF27" box="[624,717,1316,1338]" gridcol="2" gridrow="31" pageId="3" pageNumber="6">17</td>
<td id="76995D8FDA34B1F23941FF2DFC2BFF27" box="[847,940,1316,1338]" gridcol="3" gridrow="31" pageId="3" pageNumber="6">17</td>
<td id="76995D8FDA34B1F23E30FF2DFBFBFF27" box="[1086,1148,1316,1338]" gridcol="4" gridrow="31" pageId="3" pageNumber="6">17</td>
<td id="76995D8FDA34B1F23F13FF2DFADCFF27" box="[1309,1371,1316,1338]" gridcol="5" gridrow="31" pageId="3" pageNumber="6">0,58</td>
</tr>
<tr id="354834F3DA34B1F23A8EFF49FADCFF4B" box="[128,1371,1344,1366]" gridrow="32" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFF49FE85FF4B" box="[128,258,1344,1366]" gridcol="0" gridrow="32" pageId="3" pageNumber="6">LV9</th>
<td id="76995D8FDA34B1F23B7CFF49FD89FF4B" box="[370,526,1344,1366]" gridcol="1" gridrow="32" pageId="3" pageNumber="6">39</td>
<td id="76995D8FDA34B1F2387EFF49FD4AFF4B" box="[624,717,1344,1366]" gridcol="2" gridrow="32" pageId="3" pageNumber="6">28</td>
<td id="76995D8FDA34B1F23941FF49FC2BFF4B" box="[847,940,1344,1366]" gridcol="3" gridrow="32" pageId="3" pageNumber="6">27</td>
<td id="76995D8FDA34B1F23E30FF49FBFBFF4B" box="[1086,1148,1344,1366]" gridcol="4" gridrow="32" pageId="3" pageNumber="6">31</td>
<td id="76995D8FDA34B1F23F13FF49FADCFF4B" box="[1309,1371,1344,1366]" gridcol="5" gridrow="32" pageId="3" pageNumber="6">6.66</td>
</tr>
<tr id="354834F3DA34B1F23A8EFF54FADCFF6E" box="[128,1371,1373,1395]" gridrow="33" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFF54FE85FF6E" box="[128,258,1373,1395]" gridcol="0" gridrow="33" pageId="3" pageNumber="6">ML10</th>
<td id="76995D8FDA34B1F23B7CFF54FD89FF6E" box="[370,526,1373,1395]" gridcol="1" gridrow="33" pageId="3" pageNumber="6">24</td>
<td id="76995D8FDA34B1F2387EFF54FD4AFF6E" box="[624,717,1373,1395]" gridcol="2" gridrow="33" pageId="3" pageNumber="6">24</td>
<td id="76995D8FDA34B1F23941FF54FC2BFF6E" box="[847,940,1373,1395]" gridcol="3" gridrow="33" pageId="3" pageNumber="6">23</td>
<td id="76995D8FDA34B1F23E30FF54FBFBFF6E" box="[1086,1148,1373,1395]" gridcol="4" gridrow="33" pageId="3" pageNumber="6">24</td>
<td id="76995D8FDA34B1F23F13FF54FADCFF6E" box="[1309,1371,1373,1395]" gridcol="5" gridrow="33" pageId="3" pageNumber="6">0.58</td>
</tr>
<tr id="354834F3DA34B1F23A8EFF73FADCFF8D" box="[128,1371,1402,1424]" gridrow="34" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFF73FE85FF8D" box="[128,258,1402,1424]" gridcol="0" gridrow="34" pageId="3" pageNumber="6">LTAS</th>
<td id="76995D8FDA34B1F23B7CFF73FD89FF8D" box="[370,526,1402,1424]" gridcol="1" gridrow="34" pageId="3" pageNumber="6">(&gt;42)</td>
<td id="76995D8FDA34B1F2387EFF73FD4AFF8D" box="[624,717,1402,1424]" gridcol="2" gridrow="34" pageId="3" pageNumber="6">59</td>
<td id="76995D8FDA34B1F23941FF73FC2BFF8D" box="[847,940,1402,1424]" gridcol="3" gridrow="34" pageId="3" pageNumber="6">60</td>
<td id="76995D8FDA34B1F23E30FF73FBFBFF8D" box="[1086,1148,1402,1424]" gridcol="4" gridrow="34" pageId="3" pageNumber="6">60</td>
<td id="76995D8FDA34B1F23F13FF73FADCFF8D" box="[1309,1371,1402,1424]" gridcol="5" gridrow="34" pageId="3" pageNumber="6">0.71</td>
</tr>
<tr id="354834F3DA34B1F23A8EFF9EFADCFFB0" box="[128,1371,1431,1453]" gridrow="35" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFF9EFE85FFB0" box="[128,258,1431,1453]" gridcol="0" gridrow="35" pageId="3" pageNumber="6">LTS</th>
<td id="76995D8FDA34B1F23B7CFF9EFD89FFB0" box="[370,526,1431,1453]" gridcol="1" gridrow="35" pageId="3" pageNumber="6">44</td>
<td id="76995D8FDA34B1F2387EFF9EFD4AFFB0" box="[624,717,1431,1453]" gridcol="2" gridrow="35" pageId="3" pageNumber="6">44</td>
<td id="76995D8FDA34B1F23941FF9EFC2BFFB0" box="[847,940,1431,1453]" gridcol="3" gridrow="35" pageId="3" pageNumber="6">45</td>
<td id="76995D8FDA34B1F23E30FF9EFBFBFFB0" box="[1086,1148,1431,1453]" gridcol="4" gridrow="35" pageId="3" pageNumber="6">44</td>
<td id="76995D8FDA34B1F23F13FF9EFADCFFB0" box="[1309,1371,1431,1453]" gridcol="5" gridrow="35" pageId="3" pageNumber="6">0.58</td>
</tr>
<tr id="354834F3DA34B1F23A8EFFBDFADCFFD7" box="[128,1371,1460,1482]" gridrow="36" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFFBDFE85FFD7" box="[128,258,1460,1482]" gridcol="0" gridrow="36" pageId="3" pageNumber="6">LTS/LTAS</th>
<td id="76995D8FDA34B1F23B7CFFBDFD89FFD7" box="[370,526,1460,1482]" gridcol="1" gridrow="36" pageId="3" pageNumber="6"></td>
<td id="76995D8FDA34B1F2387EFFBDFD4AFFD7" box="[624,717,1460,1482]" gridcol="2" gridrow="36" pageId="3" pageNumber="6">74.6%</td>
<td id="76995D8FDA34B1F23941FFBDFC2BFFD7" box="[847,940,1460,1482]" gridcol="3" gridrow="36" pageId="3" pageNumber="6">75.0%</td>
<td id="76995D8FDA34B1F23E30FFBDFBFBFFD7" box="[1086,1148,1460,1482]" gridcol="4" gridrow="36" pageId="3" pageNumber="6">74.8%</td>
<td id="76995D8FDA34B1F23F13FFBDFADCFFD7" box="[1309,1371,1460,1482]" gridcol="5" gridrow="36" pageId="3" pageNumber="6">0.30%</td>
</tr>
<tr id="354834F3DA34B1F23A8EFFD9FADCFFFB" box="[128,1371,1488,1510]" gridrow="37" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFFD9FE85FFFB" box="[128,258,1488,1510]" gridcol="0" gridrow="37" pageId="3" pageNumber="6">MD11/LTAS</th>
<td id="76995D8FDA34B1F23B7CFFD9FD89FFFB" box="[370,526,1488,1510]" gridcol="1" gridrow="37" pageId="3" pageNumber="6"></td>
<td id="76995D8FDA34B1F2387EFFD9FD4AFFFB" box="[624,717,1488,1510]" gridcol="2" gridrow="37" pageId="3" pageNumber="6">69.5%</td>
<td id="76995D8FDA34B1F23941FFD9FC2BFFFB" box="[847,940,1488,1510]" gridcol="3" gridrow="37" pageId="3" pageNumber="6">83.3%</td>
<td id="76995D8FDA34B1F23E30FFD9FBFBFFFB" box="[1086,1148,1488,1510]" gridcol="4" gridrow="37" pageId="3" pageNumber="6">76.4%</td>
<td id="76995D8FDA34B1F23F13FFD9FADCFFFB" box="[1309,1371,1488,1510]" gridcol="5" gridrow="37" pageId="3" pageNumber="6">9.79%</td>
</tr>
<tr id="354834F3DA34B1F23A8EFFE4FADCFC1E" box="[128,1371,1517,1539]" gridrow="38" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFFE4FE85FC1E" box="[128,258,1517,1539]" gridcol="0" gridrow="38" pageId="3" pageNumber="6">MTS</th>
<td id="76995D8FDA34B1F23B7CFFE4FD89FC1E" box="[370,526,1517,1539]" gridcol="1" gridrow="38" pageId="3" pageNumber="6">(&gt;85)</td>
<td id="76995D8FDA34B1F2387EFFE4FD4AFC1E" box="[624,717,1517,1539]" gridcol="2" gridrow="38" pageId="3" pageNumber="6">120</td>
<td id="76995D8FDA34B1F23941FFE4FC2BFC1E" box="[847,940,1517,1539]" gridcol="3" gridrow="38" pageId="3" pageNumber="6">93</td>
<td id="76995D8FDA34B1F23E30FFE4FBFBFC1E" box="[1086,1148,1517,1539]" gridcol="4" gridrow="38" pageId="3" pageNumber="6">107</td>
<td id="76995D8FDA34B1F23F13FFE4FADCFC1E" box="[1309,1371,1517,1539]" gridcol="5" gridrow="38" pageId="3" pageNumber="6">19.09</td>
</tr>
<tr id="354834F3DA34B1F23A8EFC03FADCFC3D" box="[128,1371,1546,1568]" gridrow="39" pageId="3" pageNumber="6">
<th id="76995D8FDA34B1F23A8EFC03FE85FC3D" box="[128,258,1546,1568]" gridcol="0" gridrow="39" pageId="3" pageNumber="6">MTS/TL</th>
<td id="76995D8FDA34B1F23B7CFC03FD89FC3D" box="[370,526,1546,1568]" gridcol="1" gridrow="39" pageId="3" pageNumber="6"></td>
<td id="76995D8FDA34B1F2387EFC03FD4AFC3D" box="[624,717,1546,1568]" gridcol="2" gridrow="39" pageId="3" pageNumber="6">45.5%</td>
<td id="76995D8FDA34B1F23941FC03FC2BFC3D" box="[847,940,1546,1568]" gridcol="3" gridrow="39" pageId="3" pageNumber="6">36.5%</td>
<td id="76995D8FDA34B1F23E30FC03FBFBFC3D" box="[1086,1148,1546,1568]" gridcol="4" gridrow="39" pageId="3" pageNumber="6">41.0%</td>
<td id="76995D8FDA34B1F23F13FC03FADCFC3D" box="[1309,1371,1546,1568]" gridcol="5" gridrow="39" pageId="3" pageNumber="6">6.35%</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC736B1DA344E0E3A7FFCEEFBE0FD62" blockId="3.[113,753,1767,2112]" lastBlockId="3.[812,1452,1639,1920]" pageId="3" pageNumber="6">
Segment 2 and all remaining segments consist of one tergal and two sternal plates. A middorsal spine is present in all specimens. Furthermore, minute ventrolateral tubes were observed in the putative J6 or stage-
<specimenCount id="9D7EFD38DA344E0E3BF1FD4EFDF8FD42" box="[511,639,1863,1887]" pageId="3" pageNumber="6" type="generic">1 specimens</specimenCount>
(
<figureCitation id="13432A34DA344E0E3881FD4EFD63FD42" box="[655,740,1863,1887]" captionStart="Fig" captionStartId="8.[136,171,1694,1716]" captionTargetBox="[136,1472,297,1678]" captionTargetId="figure-8@8.[136,1474,297,1681]" captionTargetPageId="8" captionText="Fig. 5. Light micrographs showing details of trunk morphology in Centroderes impurus sp. nov. A, C, E: holotype, ZRC.MIS.0001, male stage-2; B, D, F: paratype, KIN-846, putative J6 or adult stage-1 of unknown gender. A, Segments 36, lateral view, with dashed circles indicating laterodorsal and lateral accessory sensory spots. B, Segments 13, lateral view, showing ventrolateral tube on segment 2, which is found only in this stage and not in male stage-2. C, Segments 10 and 11, lateral view, showing crenulated spines in middorsal position on segment 10. D, Segments 56, lateral view, showing indications of lateroventral tube on segment 5, which is found only in this stage and not in male stage-2. E, Segment 811, lateral view, showing crenulated spines in midlateral positions on segment 10; note the conspicuous difference between the lateroventral spines on segment 8 and segment 9; dashed circles indicate laterodorsal and lateral accessory sensory spots. F, Segment 811, lateral view, showing acicular spines in middorsal and midlateral positions on segment 10. Abbreviations: lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdac, middorsal acicular spine; mdcr, middorsal crenulated spine; mlac, midlateral acicular spine; mlcr, midlateral crenulated spine; vlt, ventrolateral tube; vms, ventromedial spine." figureDoi="http://doi.org/10.5281/zenodo.4502551" httpUri="https://zenodo.org/record/4502551/files/figure.png" pageId="3" pageNumber="6">Fig. 5B</figureCitation>
). In male stage-2 no such structure was observed neither in LM nor in SEM; in the single specimen examined with SEM it was evident that the position where the tube would attach was filled with densely arranged prominent hairs. Similar hairs were spotted on other segments as well (
<figureCitation id="13432A34DA344E0E38CBFDEEFF16F202" captionStart="Fig" captionStartId="6.[136,171,1919,1941]" captionTargetBox="[279,1332,189,1902]" captionTargetId="figure-7@6.[278,1332,189,1904]" captionTargetPageId="6" captionText="Fig. 4. Scanning electron micrographs of male stage-2 (A, B, F, H) and putative J6 or adult stage-1 of unknown gender (CE, G) of Centroderes impurus sp. nov. A, lateral overview. B, ventral overview. C, Lateral view of segment 1. D, Detail showing lateral accessory sensory spot on segment 1. E, Region of tergosternal junctions of segments 4 and 5; dashed circles outline extended cuticular hairs on posterior margin of segment 3. F, Region of tergosternal junction of segments 4 and 5. G, lateroventral view of segments 8 and 9. H, Lateral view of segments 810 in male stage-2. Abbreviations: lvs, lateroventral spine; lvt, lateroventral tube; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; mdcr, middorsal crenulated spine; mlcr, midlateral crenulated spine; mvp, midventral process; s, segment followed by segment number; sf, secondary fringe; vms, ventromedial spine on segment 1." figureDoi="http://doi.org/10.5281/zenodo.4502543" httpUri="https://zenodo.org/record/4502543/files/figure.png" pageId="3" pageNumber="6">Fig. 4F</figureCitation>
). Sensory spots are present in paradorsal and midlateral position. The paradorsal sensory spots are located very close to the posterior segment margin, next to the base of the middorsal spine. Secondary pectinate fringe consisting of minute cuticular hairs present on anterior part of segment; the fringe may be partially covered by the posterior margin of preceding segment. Additional hairs are densely and evenly distributed over the posterior half of the segment. Hairs in the tergosternal junction region are distinctly longer, especially anteriorly. A similar arrangement of hairs is present on segments 310 (
<figureCitation id="13432A34DA344E0E39D1FD6EFBABFD62" box="[991,1068,1895,1919]" captionStart="Fig" captionStartId="4.[135,170,1977,1999]" captionTargetBox="[196,1415,193,1962]" captionTargetId="figure-8@4.[194,1418,193,1965]" captionTargetPageId="4" captionText="Fig. 2. Line art illustration of Centroderes impurus sp. nov. male stage-2, lateral view. Abbreviations: ij, intersegmentary joint line; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral spine; mdp, middorsal placid; mds, middorsal spine; mls, midlateral spine; mlss, midlateral sensory spot; mts, midterminal spine; mvp, midventral process; pdss, paradorsal sensory spot; pf, pectinate fringe; sdss, subdorsal sensory spot; sf, secondary fringe; vms, ventromedial spine; vmss, ventromedial sensory spot. Digits following labels refer to segment numbers." figureDoi="http://doi.org/10.5281/zenodo.4502537" httpUri="https://zenodo.org/record/4502537/files/figure.png" pageId="3" pageNumber="6">
Figs.
<geoCoordinate id="EE4C5076DA344E0E3E13FD6EFBABFD62" box="[1053,1068,1895,1919]" degrees="2, 4" direction="east" orientation="longitude" pageId="3" pageNumber="6" precision="5555" value="2.4">2</geoCoordinate>
</figureCitation>
,
<figureCitation id="13432A34DA344E0E3E34FD6EFBDBFD62" box="[1082,1116,1895,1919]" captionStart="Fig" captionStartId="6.[136,171,1919,1941]" captionTargetBox="[279,1332,189,1902]" captionTargetId="figure-7@6.[278,1332,189,1904]" captionTargetPageId="6" captionText="Fig. 4. Scanning electron micrographs of male stage-2 (A, B, F, H) and putative J6 or adult stage-1 of unknown gender (CE, G) of Centroderes impurus sp. nov. A, lateral overview. B, ventral overview. C, Lateral view of segment 1. D, Detail showing lateral accessory sensory spot on segment 1. E, Region of tergosternal junctions of segments 4 and 5; dashed circles outline extended cuticular hairs on posterior margin of segment 3. F, Region of tergosternal junction of segments 4 and 5. G, lateroventral view of segments 8 and 9. H, Lateral view of segments 810 in male stage-2. Abbreviations: lvs, lateroventral spine; lvt, lateroventral tube; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; mdcr, middorsal crenulated spine; mlcr, midlateral crenulated spine; mvp, midventral process; s, segment followed by segment number; sf, secondary fringe; vms, ventromedial spine on segment 1." figureDoi="http://doi.org/10.5281/zenodo.4502543" httpUri="https://zenodo.org/record/4502543/files/figure.png" pageId="3" pageNumber="6">4E</figureCitation>
).
</paragraph>
<paragraph id="8BC736B1DA344E0E3923FDAEFBA5F23D" blockId="3.[813,1451,1959,2080]" pageId="3" pageNumber="6">
Segments 3 and 4 with middorsal acicular spine. Sensory spots are present in paradorsal, laterodorsal, lateral accessory and ventromedial positions (
<figureCitation id="13432A34DA344E0E3E51FDEEFB2CFDE2" box="[1119,1195,2023,2047]" captionStart="Fig" captionStartId="4.[135,170,1977,1999]" captionTargetBox="[196,1415,193,1962]" captionTargetId="figure-8@4.[194,1418,193,1965]" captionTargetPageId="4" captionText="Fig. 2. Line art illustration of Centroderes impurus sp. nov. male stage-2, lateral view. Abbreviations: ij, intersegmentary joint line; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral spine; mdp, middorsal placid; mds, middorsal spine; mls, midlateral spine; mlss, midlateral sensory spot; mts, midterminal spine; mvp, midventral process; pdss, paradorsal sensory spot; pf, pectinate fringe; sdss, subdorsal sensory spot; sf, secondary fringe; vms, ventromedial spine; vmss, ventromedial sensory spot. Digits following labels refer to segment numbers." figureDoi="http://doi.org/10.5281/zenodo.4502537" httpUri="https://zenodo.org/record/4502537/files/figure.png" pageId="3" pageNumber="6">Figs. 2</figureCitation>
,
<figureCitation id="13432A34DA344E0E3EB9FDEEFB5BFDE2" box="[1207,1244,2023,2047]" captionStart="Fig" captionStartId="8.[136,171,1694,1716]" captionTargetBox="[136,1472,297,1678]" captionTargetId="figure-8@8.[136,1474,297,1681]" captionTargetPageId="8" captionText="Fig. 5. Light micrographs showing details of trunk morphology in Centroderes impurus sp. nov. A, C, E: holotype, ZRC.MIS.0001, male stage-2; B, D, F: paratype, KIN-846, putative J6 or adult stage-1 of unknown gender. A, Segments 36, lateral view, with dashed circles indicating laterodorsal and lateral accessory sensory spots. B, Segments 13, lateral view, showing ventrolateral tube on segment 2, which is found only in this stage and not in male stage-2. C, Segments 10 and 11, lateral view, showing crenulated spines in middorsal position on segment 10. D, Segments 56, lateral view, showing indications of lateroventral tube on segment 5, which is found only in this stage and not in male stage-2. E, Segment 811, lateral view, showing crenulated spines in midlateral positions on segment 10; note the conspicuous difference between the lateroventral spines on segment 8 and segment 9; dashed circles indicate laterodorsal and lateral accessory sensory spots. F, Segment 811, lateral view, showing acicular spines in middorsal and midlateral positions on segment 10. Abbreviations: lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdac, middorsal acicular spine; mdcr, middorsal crenulated spine; mlac, midlateral acicular spine; mlcr, midlateral crenulated spine; vlt, ventrolateral tube; vms, ventromedial spine." figureDoi="http://doi.org/10.5281/zenodo.4502551" httpUri="https://zenodo.org/record/4502551/files/figure.png" pageId="3" pageNumber="6">5A</figureCitation>
). Cuticular hairs as on preceding segment.
</paragraph>
<caption id="DF076639DA334E093A89FDB0FB03F25E" ID-DOI="http://doi.org/10.5281/zenodo.4502537" ID-Zenodo-Dep="4502537" httpUri="https://zenodo.org/record/4502537/files/figure.png" pageId="4" pageNumber="7" startId="4.[135,170,1977,1999]" targetBox="[196,1415,193,1962]" targetPageId="4">
<paragraph id="8BC736B1DA334E093A89FDB0FB03F25E" blockId="4.[135,1474,1977,2115]" pageId="4" pageNumber="7">
Fig. 2. Line art illustration of
<taxonomicName id="4C784D32DA334E093BB2FDB0FD09FDD2" authority="Sørensen &amp; Gąsiorowski &amp; Randsø &amp; Sánchez &amp; Neves, 2016" authorityName="Sørensen &amp; Gąsiorowski &amp; Randsø &amp; Sánchez &amp; Neves" authorityYear="2016" box="[444,654,1977,1999]" class="Kinorhyncha" family="Centroderidae" genus="Centroderes" kingdom="Animalia" order="Cyclorhagida" pageId="4" pageNumber="7" phylum="Cephalorhyncha" rank="species" species="impurus" status="sp. nov.">
<emphasis id="B90CEAA3DA334E093BB2FDB0FD09FDD2" box="[444,654,1977,1999]" italics="true" pageId="4" pageNumber="7">Centroderes impurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57D8DA334E093899FDB3FD60FDD2" box="[663,743,1978,1999]" pageId="4" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
male stage-2, lateral view. Abbreviations: ij, intersegmentary joint line; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral spine; mdp, middorsal placid; mds, middorsal spine; mls, midlateral spine; mlss, midlateral sensory spot; mts, midterminal spine; mvp, midventral process; pdss, paradorsal sensory spot; pf, pectinate fringe; sdss, subdorsal sensory spot; sf, secondary fringe; vms, ventromedial spine; vmss, ventromedial sensory spot. Digits following labels refer to segment numbers.
</paragraph>
</caption>
<caption id="DF076639DA324E083A7FFAB4FD9FFB5B" pageId="5" pageNumber="8">
<paragraph id="8BC736B1DA324E083A7FFAB4FD9FFB5B" blockId="5.[112,1452,189,326]" pageId="5" pageNumber="8">
Table 3. Summary of nature and location of spines, tubes and sensory spots arranged by series in
<taxonomicName id="4C784D32DA324E083E0EFAB4FB4DFACE" authority="Sørensen &amp; Gąsiorowski &amp; Randsø &amp; Sánchez &amp; Neves, 2016" authorityName="Sørensen &amp; Gąsiorowski &amp; Randsø &amp; Sánchez &amp; Neves" authorityYear="2016" box="[1024,1226,189,211]" class="Kinorhyncha" family="Centroderidae" genus="Centroderes" kingdom="Animalia" order="Cyclorhagida" pageId="5" pageNumber="8" phylum="Cephalorhyncha" rank="species" species="impurus" status="sp. nov.">
<emphasis id="B90CEAA3DA324E083E0EFAB4FB4DFACE" box="[1024,1226,189,211]" italics="true" pageId="5" pageNumber="8">Centroderes impurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57D8DA324E083EC0FAB7FA9EFACE" box="[1230,1305,190,211]" pageId="5" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviations: LA: Lateral accessory; LD: laterodorsal; LV: lateroventral; MD: middorsal; ML: midlateral; PD: paradorsal; SD: subdorsal; VL: ventrolateral; VM: ventromedial; a1, condition of character in the putative J6 preadult or adult stage-1; ac, acicular spine; cr, crenulated spines; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mts, midterminal spine; ss, sensory spot; tu, tube; ♂2, condition of character in the male stage-2; -, no structure present.
</paragraph>
</caption>
<paragraph id="8BC736B1DA324E083A82FB71FBDFF902" pageId="5" pageNumber="8">
<table id="F978C411DA32B1F23A87FB71FA15F902" box="[137,1426,376,799]" gridcols="10" gridrows="12" pageId="5" pageNumber="8">
<tr id="354834F3DA32B1F23A87FB71FA15FBB6" box="[137,1426,376,427]" gridrow="0" pageId="5" pageNumber="8">
<th id="76995D8FDA32B1F23A87FB71FF66FBB6" box="[137,225,376,427]" gridcol="0" gridrow="0" pageId="5" pageNumber="8">
<emphasis id="B90CEAA3DA324E083A82FB71FF66FBB6" bold="true" pageId="5" pageNumber="8">Position Segment</emphasis>
</th>
<th id="76995D8FDA32B1F23B05FB71FE11FBB6" box="[267,406,376,427]" gridcol="1" gridrow="0" pageId="5" pageNumber="8">
<emphasis id="B90CEAA3DA324E083B32FB8EFEE3FB81" bold="true" box="[316,356,391,412]" pageId="5" pageNumber="8">MD</emphasis>
</th>
<th id="76995D8FDA32B1F23BCBFB71FE62FBB6" box="[453,485,376,427]" gridcol="2" gridrow="0" pageId="5" pageNumber="8">
<emphasis id="B90CEAA3DA324E083BCBFB8EFE62FB81" bold="true" box="[453,485,391,412]" pageId="5" pageNumber="8">PD</emphasis>
</th>
<th id="76995D8FDA32B1F23812FB71FDDAFBB6" box="[540,605,376,427]" gridcol="3" gridrow="0" pageId="5" pageNumber="8">
<emphasis id="B90CEAA3DA324E083823FB8FFDCBFB81" bold="true" box="[557,588,390,412]" pageId="5" pageNumber="8">SD</emphasis>
</th>
<th id="76995D8FDA32B1F2389BFB71FD48FBB6" box="[661,719,376,427]" gridcol="4" gridrow="0" pageId="5" pageNumber="8">
<emphasis id="B90CEAA3DA324E0838AFFB8EFD45FB81" bold="true" box="[673,706,391,412]" pageId="5" pageNumber="8">LD</emphasis>
</th>
<th id="76995D8FDA32B1F238F0FB71FC0EFBB6" box="[766,905,376,427]" gridcol="5" gridrow="0" pageId="5" pageNumber="8">
<emphasis id="B90CEAA3DA324E08393EFB8EFCD0FB81" bold="true" box="[816,855,391,412]" pageId="5" pageNumber="8">ML</emphasis>
</th>
<th id="76995D8FDA32B1F239B9FB71FC5FFBB6" box="[951,984,376,427]" gridcol="6" gridrow="0" pageId="5" pageNumber="8">
<emphasis id="B90CEAA3DA324E0839B9FB8FFC5FFB86" bold="true" box="[951,984,390,411]" pageId="5" pageNumber="8">LA</emphasis>
</th>
<th id="76995D8FDA32B1F23E07FB71FB09FBB6" box="[1033,1166,376,427]" gridcol="7" gridrow="0" pageId="5" pageNumber="8">
<emphasis id="B90CEAA3DA324E083E35FB8EFBDBFB81" bold="true" box="[1083,1116,391,412]" pageId="5" pageNumber="8">LV</emphasis>
</th>
<th id="76995D8FDA32B1F23EB9FB71FABBFBB6" box="[1207,1340,376,427]" gridcol="8" gridrow="0" pageId="5" pageNumber="8">
<emphasis id="B90CEAA3DA324E083EE7FB8EFA8DFB81" bold="true" box="[1257,1290,391,412]" pageId="5" pageNumber="8">VL</emphasis>
</th>
<th id="76995D8FDA32B1F23F64FB71FA15FBB6" box="[1386,1426,376,427]" gridcol="9" gridrow="0" pageId="5" pageNumber="8">
<emphasis id="B90CEAA3DA324E083F64FB8EFA15FB81" bold="true" box="[1386,1426,391,412]" pageId="5" pageNumber="8">VM</emphasis>
</th>
</tr>
<tr id="354834F3DA32B1F23A87FBCDFA15FBC7" box="[137,1426,452,474]" gridrow="1" pageId="5" pageNumber="8" rowspan-2="1" rowspan-5="1" rowspan-7="1" rowspan-8="1">
<th id="76995D8FDA32B1F23A87FBCDFF66FBC7" box="[137,225,452,474]" gridcol="0" gridrow="1" pageId="5" pageNumber="8">1</th>
<td id="76995D8FDA32B1F23B05FBCDFE11FBC7" box="[267,406,452,474]" gridcol="1" gridrow="1" pageId="5" pageNumber="8">ac</td>
<td id="76995D8FDA32B1F23812FBCDFDDAFBC7" box="[540,605,452,474]" gridcol="3" gridrow="1" pageId="5" pageNumber="8">ss(♂2)</td>
<td id="76995D8FDA32B1F2389BFBCDFD48FBC7" box="[661,719,452,474]" gridcol="4" gridrow="1" pageId="5" pageNumber="8">ss(a1)</td>
<td id="76995D8FDA32B1F239B9FBCDFC5FFBC7" box="[951,984,452,474]" gridcol="6" gridrow="1" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F23F64FBCDFA15FBC7" box="[1386,1426,452,474]" gridcol="9" gridrow="1" pageId="5" pageNumber="8">ac</td>
</tr>
<tr id="354834F3DA32B1F23A87FBE2FA15F81C" box="[137,1426,491,513]" gridrow="2" pageId="5" pageNumber="8" rowspan-3="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="76995D8FDA32B1F23A87FBE2FF66F81C" box="[137,225,491,513]" gridcol="0" gridrow="2" pageId="5" pageNumber="8">2</th>
<td id="76995D8FDA32B1F23B05FBE2FE11F81C" box="[267,406,491,513]" gridcol="1" gridrow="2" pageId="5" pageNumber="8">ac</td>
<td id="76995D8FDA32B1F23BCBFBE2FE62F81C" box="[453,485,491,513]" gridcol="2" gridrow="2" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F238F0FBE2FC0EF81C" box="[766,905,491,513]" gridcol="5" gridrow="2" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F23EB9FBE2FABBF81C" box="[1207,1340,491,513]" gridcol="8" gridrow="2" pageId="5" pageNumber="8">tu(a1); - (♂2)</td>
</tr>
<tr id="354834F3DA32B1F23A87F802FA15F83D" box="[137,1426,523,544]" gridrow="3" pageId="5" pageNumber="8" rowspan-3="1" rowspan-5="1" rowspan-7="1" rowspan-8="1">
<th id="76995D8FDA32B1F23A87F802FF66F83D" box="[137,225,523,544]" gridcol="0" gridrow="3" pageId="5" pageNumber="8">3</th>
<td id="76995D8FDA32B1F23B05F802FE11F83D" box="[267,406,523,544]" gridcol="1" gridrow="3" pageId="5" pageNumber="8">ac</td>
<td id="76995D8FDA32B1F23BCBF802FE62F83D" box="[453,485,523,544]" gridcol="2" gridrow="3" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F2389BF802FD48F83D" box="[661,719,523,544]" gridcol="4" gridrow="3" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F239B9F802FC5FF83D" box="[951,984,523,544]" gridcol="6" gridrow="3" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F23F64F802FA15F83D" box="[1386,1426,523,544]" gridcol="9" gridrow="3" pageId="5" pageNumber="8">ss</td>
</tr>
<tr id="354834F3DA32B1F23A87F822FA15F85D" box="[137,1426,555,576]" gridrow="4" pageId="5" pageNumber="8" rowspan-3="1" rowspan-5="1" rowspan-7="1" rowspan-8="1">
<th id="76995D8FDA32B1F23A87F822FF66F85D" box="[137,225,555,576]" gridcol="0" gridrow="4" pageId="5" pageNumber="8">4</th>
<td id="76995D8FDA32B1F23B05F822FE11F85D" box="[267,406,555,576]" gridcol="1" gridrow="4" pageId="5" pageNumber="8">ac</td>
<td id="76995D8FDA32B1F23BCBF822FE62F85D" box="[453,485,555,576]" gridcol="2" gridrow="4" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F2389BF822FD48F85D" box="[661,719,555,576]" gridcol="4" gridrow="4" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F239B9F822FC5FF85D" box="[951,984,555,576]" gridcol="6" gridrow="4" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F23F64F822FA15F85D" box="[1386,1426,555,576]" gridcol="9" gridrow="4" pageId="5" pageNumber="8">ss</td>
</tr>
<tr id="354834F3DA32B1F23A87F843FA15F87D" box="[137,1426,586,608]" gridrow="5" pageId="5" pageNumber="8" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-8="1" rowspan-9="1">
<th id="76995D8FDA32B1F23A87F843FF66F87D" box="[137,225,586,608]" gridcol="0" gridrow="5" pageId="5" pageNumber="8">5</th>
<td id="76995D8FDA32B1F23B05F843FE11F87D" box="[267,406,586,608]" gridcol="1" gridrow="5" pageId="5" pageNumber="8">ac</td>
<td id="76995D8FDA32B1F23BCBF843FE62F87D" box="[453,485,586,608]" gridcol="2" gridrow="5" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F2389BF843FD48F87D" box="[661,719,586,608]" gridcol="4" gridrow="5" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F23E07F843FB09F87D" box="[1033,1166,586,608]" gridcol="7" gridrow="5" pageId="5" pageNumber="8">tu(a1); - (♂2)</td>
</tr>
<tr id="354834F3DA32B1F23A87F863FA15F89D" box="[137,1426,618,640]" gridrow="6" pageId="5" pageNumber="8" rowspan-3="1" rowspan-5="1" rowspan-7="1" rowspan-8="1">
<th id="76995D8FDA32B1F23A87F863FF66F89D" box="[137,225,618,640]" gridcol="0" gridrow="6" pageId="5" pageNumber="8">6</th>
<td id="76995D8FDA32B1F23B05F863FE11F89D" box="[267,406,618,640]" gridcol="1" gridrow="6" pageId="5" pageNumber="8">ac</td>
<td id="76995D8FDA32B1F23BCBF863FE62F89D" box="[453,485,618,640]" gridcol="2" gridrow="6" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F2389BF863FD48F89D" box="[661,719,618,640]" gridcol="4" gridrow="6" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F239B9F863FC5FF89D" box="[951,984,618,640]" gridcol="6" gridrow="6" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F23F64F863FA15F89D" box="[1386,1426,618,640]" gridcol="9" gridrow="6" pageId="5" pageNumber="8">ss</td>
</tr>
<tr id="354834F3DA32B1F23A87F883FA15F882" box="[137,1426,650,671]" gridrow="7" pageId="5" pageNumber="8" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-7="1" rowspan-8="1">
<th id="76995D8FDA32B1F23A87F883FF66F882" box="[137,225,650,671]" gridcol="0" gridrow="7" pageId="5" pageNumber="8">7</th>
<td id="76995D8FDA32B1F23B05F883FE11F882" box="[267,406,650,671]" gridcol="1" gridrow="7" pageId="5" pageNumber="8">ac</td>
<td id="76995D8FDA32B1F2389BF883FD48F882" box="[661,719,650,671]" gridcol="4" gridrow="7" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F239B9F883FC5FF882" box="[951,984,650,671]" gridcol="6" gridrow="7" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F23F64F883FA15F882" box="[1386,1426,650,671]" gridcol="9" gridrow="7" pageId="5" pageNumber="8">ss</td>
</tr>
<tr id="354834F3DA32B1F23A87F8A3FA15F8A2" box="[137,1426,682,703]" gridrow="8" pageId="5" pageNumber="8" rowspan-3="1" rowspan-5="1" rowspan-8="1">
<th id="76995D8FDA32B1F23A87F8A3FF66F8A2" box="[137,225,682,703]" gridcol="0" gridrow="8" pageId="5" pageNumber="8">8</th>
<td id="76995D8FDA32B1F23B05F8A3FE11F8A2" box="[267,406,682,703]" gridcol="1" gridrow="8" pageId="5" pageNumber="8">ac</td>
<td id="76995D8FDA32B1F23BCBF8A3FE62F8A2" box="[453,485,682,703]" gridcol="2" gridrow="8" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F2389BF8A3FD48F8A2" box="[661,719,682,703]" gridcol="4" gridrow="8" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F239B9F8A3FC5FF8A2" box="[951,984,682,703]" gridcol="6" gridrow="8" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F23E07F8A3FB09F8A2" box="[1033,1166,682,703]" gridcol="7" gridrow="8" pageId="5" pageNumber="8">ac</td>
<td id="76995D8FDA32B1F23F64F8A3FA15F8A2" box="[1386,1426,682,703]" gridcol="9" gridrow="8" pageId="5" pageNumber="8">ss</td>
</tr>
<tr id="354834F3DA32B1F23A87F8C0FA15F8C2" box="[137,1426,713,735]" gridrow="9" pageId="5" pageNumber="8" rowspan-3="1" rowspan-5="1" rowspan-8="1">
<th id="76995D8FDA32B1F23A87F8C0FF66F8C2" box="[137,225,713,735]" gridcol="0" gridrow="9" pageId="5" pageNumber="8">9</th>
<td id="76995D8FDA32B1F23B05F8C0FE11F8C2" box="[267,406,713,735]" gridcol="1" gridrow="9" pageId="5" pageNumber="8">ac</td>
<td id="76995D8FDA32B1F23BCBF8C0FE62F8C2" box="[453,485,713,735]" gridcol="2" gridrow="9" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F2389BF8C0FD48F8C2" box="[661,719,713,735]" gridcol="4" gridrow="9" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F239B9F8C0FC5FF8C2" box="[951,984,713,735]" gridcol="6" gridrow="9" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F23E07F8C0FB09F8C2" box="[1033,1166,713,735]" gridcol="7" gridrow="9" pageId="5" pageNumber="8">robust ac</td>
<td id="76995D8FDA32B1F23F64F8C0FA15F8C2" box="[1386,1426,713,735]" gridcol="9" gridrow="9" pageId="5" pageNumber="8">ss</td>
</tr>
<tr id="354834F3DA32B1F23A87F8E0FA15F8E2" box="[137,1426,745,767]" gridrow="10" pageId="5" pageNumber="8" rowspan-2="1" rowspan-3="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76995D8FDA32B1F23A87F8E0FF66F8E2" box="[137,225,745,767]" gridcol="0" gridrow="10" pageId="5" pageNumber="8">10</th>
<td id="76995D8FDA32B1F23B05F8E0FE11F8E2" box="[267,406,745,767]" gridcol="1" gridrow="10" pageId="5" pageNumber="8">ac(a1); cr(♂2)</td>
<td id="76995D8FDA32B1F2389BF8E0FD48F8E2" box="[661,719,745,767]" gridcol="4" gridrow="10" pageId="5" pageNumber="8">ss</td>
<td id="76995D8FDA32B1F238F0F8E0FC0EF8E2" box="[766,905,745,767]" gridcol="5" gridrow="10" pageId="5" pageNumber="8">ac(a1); cr(♂2)</td>
<td id="76995D8FDA32B1F23F64F8E0FA15F8E2" box="[1386,1426,745,767]" gridcol="9" gridrow="10" pageId="5" pageNumber="8">ss</td>
</tr>
<tr id="354834F3DA32B1F23A87F900FA15F902" box="[137,1426,777,799]" gridrow="11" pageId="5" pageNumber="8" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-8="1" rowspan-9="1">
<th id="76995D8FDA32B1F23A87F900FF66F902" box="[137,225,777,799]" gridcol="0" gridrow="11" pageId="5" pageNumber="8">11</th>
<td id="76995D8FDA32B1F23B05F900FE11F902" box="[267,406,777,799]" gridcol="1" gridrow="11" pageId="5" pageNumber="8">ac, mts</td>
<td id="76995D8FDA32B1F23812F900FDDAF902" box="[540,605,777,799]" gridcol="3" gridrow="11" pageId="5" pageNumber="8">ss, ss</td>
<td id="76995D8FDA32B1F239B9F900FC5FF902" box="[951,984,777,799]" gridcol="6" gridrow="11" pageId="5" pageNumber="8">ltas</td>
<td id="76995D8FDA32B1F23E07F900FB09F902" box="[1033,1166,777,799]" gridcol="7" gridrow="11" pageId="5" pageNumber="8">lts</td>
</tr>
</table>
</paragraph>
<caption id="DF076639DA324E083A7FF206FC89F25F" ID-DOI="http://doi.org/10.5281/zenodo.4502541" ID-Zenodo-Dep="4502541" httpUri="https://zenodo.org/record/4502541/files/figure.png" pageId="5" pageNumber="8" startId="5.[113,148,2063,2085]" targetBox="[235,1328,869,2047]" targetPageId="5">
<paragraph id="8BC736B1DA324E083A7FF206FC89F25F" blockId="5.[113,1451,2063,2114]" pageId="5" pageNumber="8">
Fig. 3. Light micrographs showing lateral overviews of
<taxonomicName id="4C784D32DA324E0838A7F206FCFDF238" authority="Sørensen &amp; Gąsiorowski &amp; Randsø &amp; Sánchez &amp; Neves, 2016" authorityName="Sørensen &amp; Gąsiorowski &amp; Randsø &amp; Sánchez &amp; Neves" authorityYear="2016" box="[681,890,2063,2085]" class="Kinorhyncha" family="Centroderidae" genus="Centroderes" kingdom="Animalia" order="Cyclorhagida" pageId="5" pageNumber="8" phylum="Cephalorhyncha" rank="species" species="impurus" status="sp. nov.">
<emphasis id="B90CEAA3DA324E0838A7F206FCFDF238" box="[681,890,2063,2085]" italics="true" pageId="5" pageNumber="8">Centroderes impurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57D8DA324E08398DF219FC54F238" box="[899,979,2064,2085]" pageId="5" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
A, holotype, ZRC.MIS.0001, male stage-2. B, paratype, KIN-846, putative J6 or adult stage-1 of unknown gender.
</paragraph>
</caption>
<caption id="DF076639DA314E0B3A86FD76FB83F25C" ID-DOI="http://doi.org/10.5281/zenodo.4502543" ID-Zenodo-Dep="4502543" httpUri="https://zenodo.org/record/4502543/files/figure.png" pageId="6" pageNumber="9" startId="6.[136,171,1919,1941]" targetBox="[279,1332,189,1902]" targetPageId="6">
<paragraph id="8BC736B1DA314E0B3A86FD76FB83F25C" blockId="6.[136,1474,1919,2114]" pageId="6" pageNumber="9">
Fig. 4. Scanning electron micrographs of male stage-2 (A, B, F, H) and putative J6 or adult stage-1 of unknown gender (CE, G) of
<taxonomicName id="4C784D32DA314E0B3A86FD95FED3FDAF" authority="Sørensen &amp; Gąsiorowski &amp; Randsø &amp; Sánchez &amp; Neves, 2016" authorityName="Sørensen &amp; Gąsiorowski &amp; Randsø &amp; Sánchez &amp; Neves" authorityYear="2016" box="[136,340,1948,1970]" class="Kinorhyncha" family="Centroderidae" genus="Centroderes" kingdom="Animalia" order="Cyclorhagida" pageId="6" pageNumber="9" phylum="Cephalorhyncha" rank="species" species="impurus" status="sp. nov.">
<emphasis id="B90CEAA3DA314E0B3A86FD95FED3FDAF" box="[136,340,1948,1970]" italics="true" pageId="6" pageNumber="9">Centroderes impurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57D8DA314E0B3B55FD94FE2FFDAF" box="[347,424,1949,1970]" pageId="6" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
A, lateral overview. B, ventral overview. C, Lateral view of segment 1. D, Detail showing lateral accessory sensory spot on segment 1. E, Region of tergosternal junctions of segments 4 and 5; dashed circles outline extended cuticular hairs on posterior margin of segment 3. F, Region of tergosternal junction of segments 4 and 5. G, lateroventral view of segments 8 and 9. H, Lateral view of segments 810 in male stage-2. Abbreviations: lvs, lateroventral spine; lvt, lateroventral tube; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; mdcr, middorsal crenulated spine; mlcr, midlateral crenulated spine; mvp, midventral process; s, segment followed by segment number; sf, secondary fringe; vms, ventromedial spine on segment 1.
</paragraph>
</caption>
<paragraph id="8BC736B1DA304E0A3A7CFAB4FE44FB8B" blockId="7.[114,754,189,406]" pageId="7" pageNumber="10">
Segment 5 with middorsal acicular spine. Minute lateroventral tubes are present in putative J6 or stage-
<specimenCount id="9D7EFD38DA304E0A3822FAD4FD2CFAE8" box="[556,683,221,245]" pageId="7" pageNumber="10" type="generic">1 specimens</specimenCount>
(
<figureCitation id="13432A34DA304E0A38B5FAD4FF15FB08" captionStart="Fig" captionStartId="6.[136,171,1919,1941]" captionTargetBox="[279,1332,189,1902]" captionTargetId="figure-7@6.[278,1332,189,1904]" captionTargetPageId="6" captionText="Fig. 4. Scanning electron micrographs of male stage-2 (A, B, F, H) and putative J6 or adult stage-1 of unknown gender (CE, G) of Centroderes impurus sp. nov. A, lateral overview. B, ventral overview. C, Lateral view of segment 1. D, Detail showing lateral accessory sensory spot on segment 1. E, Region of tergosternal junctions of segments 4 and 5; dashed circles outline extended cuticular hairs on posterior margin of segment 3. F, Region of tergosternal junction of segments 4 and 5. G, lateroventral view of segments 8 and 9. H, Lateral view of segments 810 in male stage-2. Abbreviations: lvs, lateroventral spine; lvt, lateroventral tube; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; mdcr, middorsal crenulated spine; mlcr, midlateral crenulated spine; mvp, midventral process; s, segment followed by segment number; sf, secondary fringe; vms, ventromedial spine on segment 1." figureDoi="http://doi.org/10.5281/zenodo.4502543" httpUri="https://zenodo.org/record/4502543/files/figure.png" pageId="7" pageNumber="10">Figs. 4E</figureCitation>
,
<figureCitation id="13432A34DA304E0A3A93FAF4FF46FB08" box="[157,193,253,277]" captionStart="Fig" captionStartId="8.[136,171,1694,1716]" captionTargetBox="[136,1472,297,1678]" captionTargetId="figure-8@8.[136,1474,297,1681]" captionTargetPageId="8" captionText="Fig. 5. Light micrographs showing details of trunk morphology in Centroderes impurus sp. nov. A, C, E: holotype, ZRC.MIS.0001, male stage-2; B, D, F: paratype, KIN-846, putative J6 or adult stage-1 of unknown gender. A, Segments 36, lateral view, with dashed circles indicating laterodorsal and lateral accessory sensory spots. B, Segments 13, lateral view, showing ventrolateral tube on segment 2, which is found only in this stage and not in male stage-2. C, Segments 10 and 11, lateral view, showing crenulated spines in middorsal position on segment 10. D, Segments 56, lateral view, showing indications of lateroventral tube on segment 5, which is found only in this stage and not in male stage-2. E, Segment 811, lateral view, showing crenulated spines in midlateral positions on segment 10; note the conspicuous difference between the lateroventral spines on segment 8 and segment 9; dashed circles indicate laterodorsal and lateral accessory sensory spots. F, Segment 811, lateral view, showing acicular spines in middorsal and midlateral positions on segment 10. Abbreviations: lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdac, middorsal acicular spine; mdcr, middorsal crenulated spine; mlac, midlateral acicular spine; mlcr, midlateral crenulated spine; vlt, ventrolateral tube; vms, ventromedial spine." figureDoi="http://doi.org/10.5281/zenodo.4502551" httpUri="https://zenodo.org/record/4502551/files/figure.png" pageId="7" pageNumber="10">5D</figureCitation>
). Stage-
<specimenCount id="9D7EFD38DA304E0A3B17FAF4FE1BFB08" box="[281,412,253,277]" pageId="7" pageNumber="10" type="generic">2 specimens</specimenCount>
without such tubes, and instead with densely arranged prominent hairs in the areas where the tubes would attach (
<figureCitation id="13432A34DA304E0A3B5CFB34FE21FB48" box="[338,422,317,341]" captionStart="Fig" captionStartId="6.[136,171,1919,1941]" captionTargetBox="[279,1332,189,1902]" captionTargetId="figure-7@6.[278,1332,189,1904]" captionTargetPageId="6" captionText="Fig. 4. Scanning electron micrographs of male stage-2 (A, B, F, H) and putative J6 or adult stage-1 of unknown gender (CE, G) of Centroderes impurus sp. nov. A, lateral overview. B, ventral overview. C, Lateral view of segment 1. D, Detail showing lateral accessory sensory spot on segment 1. E, Region of tergosternal junctions of segments 4 and 5; dashed circles outline extended cuticular hairs on posterior margin of segment 3. F, Region of tergosternal junction of segments 4 and 5. G, lateroventral view of segments 8 and 9. H, Lateral view of segments 810 in male stage-2. Abbreviations: lvs, lateroventral spine; lvt, lateroventral tube; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; mdcr, middorsal crenulated spine; mlcr, midlateral crenulated spine; mvp, midventral process; s, segment followed by segment number; sf, secondary fringe; vms, ventromedial spine on segment 1." figureDoi="http://doi.org/10.5281/zenodo.4502543" httpUri="https://zenodo.org/record/4502543/files/figure.png" pageId="7" pageNumber="10">Fig. 4F</figureCitation>
). Sensory spots are present in paradorsal and laterodorsal positions (
<figureCitation id="13432A34DA304E0A380AFB54FDC9FB68" box="[516,590,349,373]" captionStart="Fig" captionStartId="4.[135,170,1977,1999]" captionTargetBox="[196,1415,193,1962]" captionTargetId="figure-8@4.[194,1418,193,1965]" captionTargetPageId="4" captionText="Fig. 2. Line art illustration of Centroderes impurus sp. nov. male stage-2, lateral view. Abbreviations: ij, intersegmentary joint line; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral spine; mdp, middorsal placid; mds, middorsal spine; mls, midlateral spine; mlss, midlateral sensory spot; mts, midterminal spine; mvp, midventral process; pdss, paradorsal sensory spot; pf, pectinate fringe; sdss, subdorsal sensory spot; sf, secondary fringe; vms, ventromedial spine; vmss, ventromedial sensory spot. Digits following labels refer to segment numbers." figureDoi="http://doi.org/10.5281/zenodo.4502537" httpUri="https://zenodo.org/record/4502537/files/figure.png" pageId="7" pageNumber="10">Figs. 2</figureCitation>
,
<figureCitation id="13432A34DA304E0A3856FB54FDFAFB68" box="[600,637,349,373]" captionStart="Fig" captionStartId="8.[136,171,1694,1716]" captionTargetBox="[136,1472,297,1678]" captionTargetId="figure-8@8.[136,1474,297,1681]" captionTargetPageId="8" captionText="Fig. 5. Light micrographs showing details of trunk morphology in Centroderes impurus sp. nov. A, C, E: holotype, ZRC.MIS.0001, male stage-2; B, D, F: paratype, KIN-846, putative J6 or adult stage-1 of unknown gender. A, Segments 36, lateral view, with dashed circles indicating laterodorsal and lateral accessory sensory spots. B, Segments 13, lateral view, showing ventrolateral tube on segment 2, which is found only in this stage and not in male stage-2. C, Segments 10 and 11, lateral view, showing crenulated spines in middorsal position on segment 10. D, Segments 56, lateral view, showing indications of lateroventral tube on segment 5, which is found only in this stage and not in male stage-2. E, Segment 811, lateral view, showing crenulated spines in midlateral positions on segment 10; note the conspicuous difference between the lateroventral spines on segment 8 and segment 9; dashed circles indicate laterodorsal and lateral accessory sensory spots. F, Segment 811, lateral view, showing acicular spines in middorsal and midlateral positions on segment 10. Abbreviations: lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdac, middorsal acicular spine; mdcr, middorsal crenulated spine; mlac, midlateral acicular spine; mlcr, midlateral crenulated spine; vlt, ventrolateral tube; vms, ventromedial spine." figureDoi="http://doi.org/10.5281/zenodo.4502551" httpUri="https://zenodo.org/record/4502551/files/figure.png" pageId="7" pageNumber="10">5A</figureCitation>
). Cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="8BC736B1DA304E0A3A7CFBB4FEC2F82B" blockId="7.[114,754,445,566]" pageId="7" pageNumber="10">
Segment 6 with middorsal acicular spine. Sensory spots present in paradorsal, laterodorsal, lateral accessory and ventromedial positions (
<figureCitation id="13432A34DA304E0A3B74FBF4FE40F808" box="[378,455,509,533]" captionStart="Fig" captionStartId="4.[135,170,1977,1999]" captionTargetBox="[196,1415,193,1962]" captionTargetId="figure-8@4.[194,1418,193,1965]" captionTargetPageId="4" captionText="Fig. 2. Line art illustration of Centroderes impurus sp. nov. male stage-2, lateral view. Abbreviations: ij, intersegmentary joint line; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral spine; mdp, middorsal placid; mds, middorsal spine; mls, midlateral spine; mlss, midlateral sensory spot; mts, midterminal spine; mvp, midventral process; pdss, paradorsal sensory spot; pf, pectinate fringe; sdss, subdorsal sensory spot; sf, secondary fringe; vms, ventromedial spine; vmss, ventromedial sensory spot. Digits following labels refer to segment numbers." figureDoi="http://doi.org/10.5281/zenodo.4502537" httpUri="https://zenodo.org/record/4502537/files/figure.png" pageId="7" pageNumber="10">Figs. 2</figureCitation>
,
<figureCitation id="13432A34DA304E0A3BDBFBF4FE7DF808" box="[469,506,509,533]" captionStart="Fig" captionStartId="8.[136,171,1694,1716]" captionTargetBox="[136,1472,297,1678]" captionTargetId="figure-8@8.[136,1474,297,1681]" captionTargetPageId="8" captionText="Fig. 5. Light micrographs showing details of trunk morphology in Centroderes impurus sp. nov. A, C, E: holotype, ZRC.MIS.0001, male stage-2; B, D, F: paratype, KIN-846, putative J6 or adult stage-1 of unknown gender. A, Segments 36, lateral view, with dashed circles indicating laterodorsal and lateral accessory sensory spots. B, Segments 13, lateral view, showing ventrolateral tube on segment 2, which is found only in this stage and not in male stage-2. C, Segments 10 and 11, lateral view, showing crenulated spines in middorsal position on segment 10. D, Segments 56, lateral view, showing indications of lateroventral tube on segment 5, which is found only in this stage and not in male stage-2. E, Segment 811, lateral view, showing crenulated spines in midlateral positions on segment 10; note the conspicuous difference between the lateroventral spines on segment 8 and segment 9; dashed circles indicate laterodorsal and lateral accessory sensory spots. F, Segment 811, lateral view, showing acicular spines in middorsal and midlateral positions on segment 10. Abbreviations: lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdac, middorsal acicular spine; mdcr, middorsal crenulated spine; mlac, midlateral acicular spine; mlcr, midlateral crenulated spine; vlt, ventrolateral tube; vms, ventromedial spine." figureDoi="http://doi.org/10.5281/zenodo.4502551" httpUri="https://zenodo.org/record/4502551/files/figure.png" pageId="7" pageNumber="10">5A</figureCitation>
). Cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="8BC736B1DA304E0A3A7CF854FF54F94B" blockId="7.[114,754,605,854]" pageId="7" pageNumber="10">
Segment 7 with middorsal acicular spine. Sensory spots present in laterodorsal, lateral accessory and ventromedial positions. On this and the following segments, the laterodorsal sensory spots are located slightly closer to the midlateral position. Putative J6 or stage-
<specimenCount id="9D7EFD38DA304E0A3BA4F8D4FDA1F8E8" box="[426,550,733,757]" pageId="7" pageNumber="10" type="generic">1 specimens</specimenCount>
show no indication of female specific glands, as observed in other
<taxonomicName id="4C784D32DA304E0A3863F8F4FD76F908" authorityName="Zelinka" authorityYear="1907" box="[621,753,765,789]" class="Kinorhyncha" family="Centroderidae" genus="Centroderes" kingdom="Animalia" order="Cyclorhagida" pageId="7" pageNumber="10" phylum="Cephalorhyncha" rank="genus">
<emphasis id="B90CEAA3DA304E0A3863F8F4FD76F908" box="[621,753,765,789]" italics="true" pageId="7" pageNumber="10">Centroderes</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFE94B40DA304E0A3AA8F914FE0CF928" author="Neuhaus B &amp; Pardos F &amp; Sorensen MV &amp; Higgins RP" box="[166,395,797,821]" pageId="7" pageNumber="10" pagination="1 - 69" refId="ref15114" refString="Neuhaus B, Pardos F, Sorensen MV &amp; Higgins RP (2014) New species of Centroderes (Kinorhyncha: Cyclorhagida) from the Northwest Atlantic Ocean, life cycle, and ground pattern of the genus. Zootaxa, 3901: 1 - 69." type="journal article" year="2014">Neuhaus et al., 2014</bibRefCitation>
). Cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="8BC736B1DA304E0A3A7CF974FF54FE4B" blockId="7.[114,755,893,1110]" pageId="7" pageNumber="10">
Segment 8 with prominent, long middorsal acicular spine (
<figureCitation id="13432A34DA304E0A3A74F994FF40F9A8" box="[122,199,925,949]" captionStart="Fig" captionStartId="4.[135,170,1977,1999]" captionTargetBox="[196,1415,193,1962]" captionTargetId="figure-8@4.[194,1418,193,1965]" captionTargetPageId="4" captionText="Fig. 2. Line art illustration of Centroderes impurus sp. nov. male stage-2, lateral view. Abbreviations: ij, intersegmentary joint line; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral spine; mdp, middorsal placid; mds, middorsal spine; mls, midlateral spine; mlss, midlateral sensory spot; mts, midterminal spine; mvp, midventral process; pdss, paradorsal sensory spot; pf, pectinate fringe; sdss, subdorsal sensory spot; sf, secondary fringe; vms, ventromedial spine; vmss, ventromedial sensory spot. Digits following labels refer to segment numbers." figureDoi="http://doi.org/10.5281/zenodo.4502537" httpUri="https://zenodo.org/record/4502537/files/figure.png" pageId="7" pageNumber="10">Figs. 2</figureCitation>
,
<figureCitation id="13432A34DA304E0A3ADAF994FF7FF9A8" box="[212,248,925,949]" captionStart="Fig" captionStartId="5.[113,148,2063,2085]" captionTargetBox="[235,1328,869,2047]" captionTargetId="figure-213@5.[235,1330,869,2047]" captionTargetPageId="5" captionText="Fig. 3. Light micrographs showing lateral overviews of Centroderes impurus sp. nov. A, holotype, ZRC.MIS.0001, male stage-2. B, paratype, KIN-846, putative J6 or adult stage-1 of unknown gender." figureDoi="http://doi.org/10.5281/zenodo.4502541" httpUri="https://zenodo.org/record/4502541/files/figure.png" pageId="7" pageNumber="10">3A</figureCitation>
). Small acicular spines present in lateroventral positions (
<figureCitation id="13432A34DA304E0A3AFEF9B4FEDBF9C8" box="[240,348,957,981]" captionStart="Fig" captionStartId="6.[136,171,1919,1941]" captionTargetBox="[279,1332,189,1902]" captionTargetId="figure-7@6.[278,1332,189,1904]" captionTargetPageId="6" captionText="Fig. 4. Scanning electron micrographs of male stage-2 (A, B, F, H) and putative J6 or adult stage-1 of unknown gender (CE, G) of Centroderes impurus sp. nov. A, lateral overview. B, ventral overview. C, Lateral view of segment 1. D, Detail showing lateral accessory sensory spot on segment 1. E, Region of tergosternal junctions of segments 4 and 5; dashed circles outline extended cuticular hairs on posterior margin of segment 3. F, Region of tergosternal junction of segments 4 and 5. G, lateroventral view of segments 8 and 9. H, Lateral view of segments 810 in male stage-2. Abbreviations: lvs, lateroventral spine; lvt, lateroventral tube; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; mdcr, middorsal crenulated spine; mlcr, midlateral crenulated spine; mvp, midventral process; s, segment followed by segment number; sf, secondary fringe; vms, ventromedial spine on segment 1." figureDoi="http://doi.org/10.5281/zenodo.4502543" httpUri="https://zenodo.org/record/4502543/files/figure.png" pageId="7" pageNumber="10">Figs. 4G</figureCitation>
,
<figureCitation id="13432A34DA304E0A3B62F9B4FE35F9C8" box="[364,434,957,981]" captionStart="Fig" captionStartId="8.[136,171,1694,1716]" captionTargetBox="[136,1472,297,1678]" captionTargetId="figure-8@8.[136,1474,297,1681]" captionTargetPageId="8" captionText="Fig. 5. Light micrographs showing details of trunk morphology in Centroderes impurus sp. nov. A, C, E: holotype, ZRC.MIS.0001, male stage-2; B, D, F: paratype, KIN-846, putative J6 or adult stage-1 of unknown gender. A, Segments 36, lateral view, with dashed circles indicating laterodorsal and lateral accessory sensory spots. B, Segments 13, lateral view, showing ventrolateral tube on segment 2, which is found only in this stage and not in male stage-2. C, Segments 10 and 11, lateral view, showing crenulated spines in middorsal position on segment 10. D, Segments 56, lateral view, showing indications of lateroventral tube on segment 5, which is found only in this stage and not in male stage-2. E, Segment 811, lateral view, showing crenulated spines in midlateral positions on segment 10; note the conspicuous difference between the lateroventral spines on segment 8 and segment 9; dashed circles indicate laterodorsal and lateral accessory sensory spots. F, Segment 811, lateral view, showing acicular spines in middorsal and midlateral positions on segment 10. Abbreviations: lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdac, middorsal acicular spine; mdcr, middorsal crenulated spine; mlac, midlateral acicular spine; mlcr, midlateral crenulated spine; vlt, ventrolateral tube; vms, ventromedial spine." figureDoi="http://doi.org/10.5281/zenodo.4502551" httpUri="https://zenodo.org/record/4502551/files/figure.png" pageId="7" pageNumber="10">5E, F</figureCitation>
). Sensory spots present in paradorsal, laterodorsal, lateral accessory and ventromedial position. Putative J6 or stage-
<specimenCount id="9D7EFD38DA304E0A3BA4F9F4FDA1FE08" box="[426,550,1021,1045]" pageId="7" pageNumber="10" type="generic">1 specimens</specimenCount>
show no indication of female specific glands. Cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="8BC736B1DA304E0A3A7CFE74FF54FF6B" blockId="7.[114,754,1149,1398]" pageId="7" pageNumber="10">
Segment 9 with prominent, long middorsal acicular spine (
<figureCitation id="13432A34DA304E0A3A74FE94FF45FEA8" box="[122,194,1181,1205]" captionStart="Fig" captionStartId="4.[135,170,1977,1999]" captionTargetBox="[196,1415,193,1962]" captionTargetId="figure-8@4.[194,1418,193,1965]" captionTargetPageId="4" captionText="Fig. 2. Line art illustration of Centroderes impurus sp. nov. male stage-2, lateral view. Abbreviations: ij, intersegmentary joint line; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral spine; mdp, middorsal placid; mds, middorsal spine; mls, midlateral spine; mlss, midlateral sensory spot; mts, midterminal spine; mvp, midventral process; pdss, paradorsal sensory spot; pf, pectinate fringe; sdss, subdorsal sensory spot; sf, secondary fringe; vms, ventromedial spine; vmss, ventromedial sensory spot. Digits following labels refer to segment numbers." figureDoi="http://doi.org/10.5281/zenodo.4502537" httpUri="https://zenodo.org/record/4502537/files/figure.png" pageId="7" pageNumber="10">Figs. 2</figureCitation>
,
<figureCitation id="13432A34DA304E0A3AC5FE94FF68FEA8" box="[203,239,1181,1205]" captionStart="Fig" captionStartId="5.[113,148,2063,2085]" captionTargetBox="[235,1328,869,2047]" captionTargetId="figure-213@5.[235,1330,869,2047]" captionTargetPageId="5" captionText="Fig. 3. Light micrographs showing lateral overviews of Centroderes impurus sp. nov. A, holotype, ZRC.MIS.0001, male stage-2. B, paratype, KIN-846, putative J6 or adult stage-1 of unknown gender." figureDoi="http://doi.org/10.5281/zenodo.4502541" httpUri="https://zenodo.org/record/4502541/files/figure.png" pageId="7" pageNumber="10">3A</figureCitation>
). Conspicuously robust acicular spines are present in lateroventral positions; these spines can be twice as long as the corresponding spines on segment 8 (
<figureCitation id="13432A34DA304E0A384DFED4FD23FEE8" box="[579,676,1245,1269]" captionStart="Fig" captionStartId="6.[136,171,1919,1941]" captionTargetBox="[279,1332,189,1902]" captionTargetId="figure-7@6.[278,1332,189,1904]" captionTargetPageId="6" captionText="Fig. 4. Scanning electron micrographs of male stage-2 (A, B, F, H) and putative J6 or adult stage-1 of unknown gender (CE, G) of Centroderes impurus sp. nov. A, lateral overview. B, ventral overview. C, Lateral view of segment 1. D, Detail showing lateral accessory sensory spot on segment 1. E, Region of tergosternal junctions of segments 4 and 5; dashed circles outline extended cuticular hairs on posterior margin of segment 3. F, Region of tergosternal junction of segments 4 and 5. G, lateroventral view of segments 8 and 9. H, Lateral view of segments 810 in male stage-2. Abbreviations: lvs, lateroventral spine; lvt, lateroventral tube; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; mdcr, middorsal crenulated spine; mlcr, midlateral crenulated spine; mvp, midventral process; s, segment followed by segment number; sf, secondary fringe; vms, ventromedial spine on segment 1." figureDoi="http://doi.org/10.5281/zenodo.4502543" httpUri="https://zenodo.org/record/4502543/files/figure.png" pageId="7" pageNumber="10">Figs. 4G</figureCitation>
,
<figureCitation id="13432A34DA304E0A38BEFED4FD6AFEE8" box="[688,749,1245,1269]" captionStart="Fig" captionStartId="8.[136,171,1694,1716]" captionTargetBox="[136,1472,297,1678]" captionTargetId="figure-8@8.[136,1474,297,1681]" captionTargetPageId="8" captionText="Fig. 5. Light micrographs showing details of trunk morphology in Centroderes impurus sp. nov. A, C, E: holotype, ZRC.MIS.0001, male stage-2; B, D, F: paratype, KIN-846, putative J6 or adult stage-1 of unknown gender. A, Segments 36, lateral view, with dashed circles indicating laterodorsal and lateral accessory sensory spots. B, Segments 13, lateral view, showing ventrolateral tube on segment 2, which is found only in this stage and not in male stage-2. C, Segments 10 and 11, lateral view, showing crenulated spines in middorsal position on segment 10. D, Segments 56, lateral view, showing indications of lateroventral tube on segment 5, which is found only in this stage and not in male stage-2. E, Segment 811, lateral view, showing crenulated spines in midlateral positions on segment 10; note the conspicuous difference between the lateroventral spines on segment 8 and segment 9; dashed circles indicate laterodorsal and lateral accessory sensory spots. F, Segment 811, lateral view, showing acicular spines in middorsal and midlateral positions on segment 10. Abbreviations: lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdac, middorsal acicular spine; mdcr, middorsal crenulated spine; mlac, midlateral acicular spine; mlcr, midlateral crenulated spine; vlt, ventrolateral tube; vms, ventromedial spine." figureDoi="http://doi.org/10.5281/zenodo.4502551" httpUri="https://zenodo.org/record/4502551/files/figure.png" pageId="7" pageNumber="10">5E, F</figureCitation>
, see also
<tableCitation id="C6FA030ADA304E0A3ADAFEF4FEAEFF08" box="[212,297,1277,1301]" captionStart="Table 2" captionStartId="3.[113,168,189,211]" captionText="Table 2. Measurements from light microscopy of adult Centroderes impurus sp. nov. (in µm) from station SI-03, Singapore, including individual measures, mean and standard deviation (SD). Abbreviations: LTAS: lateral terminal accessory spine; LTS: lateral terminal spine; LV: lateroventral spine or tube; MD, middorsal spine; ML: midlateral spine; MSW-6: maximum sternal width, measured on segment 6 in this species; MTS: midterminal spine; S: segment lengths; SW-10, standard width, always measured on segment 10; TL: trunk length; VM: ventromedial spine." pageId="7" pageNumber="10">Table 2</tableCitation>
), especially in the male stage-2. Sensory spots present in paradorsal, laterodorsal, lateral accessory and ventromedial position. Cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="8BC736B1DA304E0A3A7CFF94FEEFFC4B" blockId="7.[114,754,1437,1622]" pageId="7" pageNumber="10">
Segment 10 with middorsal and midlateral acicular spines in putative J6 or stage-
<specimenCount id="9D7EFD38DA304E0A3B88FFB4FD89FFC8" box="[390,526,1469,1493]" pageId="7" pageNumber="10" type="generic">1 specimens</specimenCount>
(
<figureCitation id="13432A34DA304E0A382CFFB4FD00FFC8" box="[546,647,1469,1493]" captionStart="Fig" captionStartId="5.[113,148,2063,2085]" captionTargetBox="[235,1328,869,2047]" captionTargetId="figure-213@5.[235,1330,869,2047]" captionTargetPageId="5" captionText="Fig. 3. Light micrographs showing lateral overviews of Centroderes impurus sp. nov. A, holotype, ZRC.MIS.0001, male stage-2. B, paratype, KIN-846, putative J6 or adult stage-1 of unknown gender." figureDoi="http://doi.org/10.5281/zenodo.4502541" httpUri="https://zenodo.org/record/4502541/files/figure.png" pageId="7" pageNumber="10">Figs. 3B</figureCitation>
,
<figureCitation id="13432A34DA304E0A3896FFB4FD31FFC8" box="[664,694,1469,1493]" captionStart="Fig" captionStartId="8.[136,171,1694,1716]" captionTargetBox="[136,1472,297,1678]" captionTargetId="figure-8@8.[136,1474,297,1681]" captionTargetPageId="8" captionText="Fig. 5. Light micrographs showing details of trunk morphology in Centroderes impurus sp. nov. A, C, E: holotype, ZRC.MIS.0001, male stage-2; B, D, F: paratype, KIN-846, putative J6 or adult stage-1 of unknown gender. A, Segments 36, lateral view, with dashed circles indicating laterodorsal and lateral accessory sensory spots. B, Segments 13, lateral view, showing ventrolateral tube on segment 2, which is found only in this stage and not in male stage-2. C, Segments 10 and 11, lateral view, showing crenulated spines in middorsal position on segment 10. D, Segments 56, lateral view, showing indications of lateroventral tube on segment 5, which is found only in this stage and not in male stage-2. E, Segment 811, lateral view, showing crenulated spines in midlateral positions on segment 10; note the conspicuous difference between the lateroventral spines on segment 8 and segment 9; dashed circles indicate laterodorsal and lateral accessory sensory spots. F, Segment 811, lateral view, showing acicular spines in middorsal and midlateral positions on segment 10. Abbreviations: lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdac, middorsal acicular spine; mdcr, middorsal crenulated spine; mlac, midlateral acicular spine; mlcr, midlateral crenulated spine; vlt, ventrolateral tube; vms, ventromedial spine." figureDoi="http://doi.org/10.5281/zenodo.4502551" httpUri="https://zenodo.org/record/4502551/files/figure.png" pageId="7" pageNumber="10">5F</figureCitation>
) and with middorsal and midlateral crenulated spines in stage-
<specimenCount id="9D7EFD38DA304E0A38EBFFD4FF36FC08" pageId="7" pageNumber="10" type="male">2 males</specimenCount>
(
<figureCitation id="13432A34DA304E0A3ACAFFF4FE94FC08" box="[196,275,1533,1557]" captionStart="Fig" captionStartId="4.[135,170,1977,1999]" captionTargetBox="[196,1415,193,1962]" captionTargetId="figure-8@4.[194,1418,193,1965]" captionTargetPageId="4" captionText="Fig. 2. Line art illustration of Centroderes impurus sp. nov. male stage-2, lateral view. Abbreviations: ij, intersegmentary joint line; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral spine; mdp, middorsal placid; mds, middorsal spine; mls, midlateral spine; mlss, midlateral sensory spot; mts, midterminal spine; mvp, midventral process; pdss, paradorsal sensory spot; pf, pectinate fringe; sdss, subdorsal sensory spot; sf, secondary fringe; vms, ventromedial spine; vmss, ventromedial sensory spot. Digits following labels refer to segment numbers." figureDoi="http://doi.org/10.5281/zenodo.4502537" httpUri="https://zenodo.org/record/4502537/files/figure.png" pageId="7" pageNumber="10">Figs. 2</figureCitation>
,
<figureCitation id="13432A34DA304E0A3B2DFFF4FEC2FC08" box="[291,325,1533,1557]" captionStart="Fig" captionStartId="5.[113,148,2063,2085]" captionTargetBox="[235,1328,869,2047]" captionTargetId="figure-213@5.[235,1330,869,2047]" captionTargetPageId="5" captionText="Fig. 3. Light micrographs showing lateral overviews of Centroderes impurus sp. nov. A, holotype, ZRC.MIS.0001, male stage-2. B, paratype, KIN-846, putative J6 or adult stage-1 of unknown gender." figureDoi="http://doi.org/10.5281/zenodo.4502541" httpUri="https://zenodo.org/record/4502541/files/figure.png" pageId="7" pageNumber="10">3A</figureCitation>
,
<figureCitation id="13432A34DA304E0A3B5BFFF4FEFFFC08" box="[341,376,1533,1557]" captionStart="Fig" captionStartId="6.[136,171,1919,1941]" captionTargetBox="[279,1332,189,1902]" captionTargetId="figure-7@6.[278,1332,189,1904]" captionTargetPageId="6" captionText="Fig. 4. Scanning electron micrographs of male stage-2 (A, B, F, H) and putative J6 or adult stage-1 of unknown gender (CE, G) of Centroderes impurus sp. nov. A, lateral overview. B, ventral overview. C, Lateral view of segment 1. D, Detail showing lateral accessory sensory spot on segment 1. E, Region of tergosternal junctions of segments 4 and 5; dashed circles outline extended cuticular hairs on posterior margin of segment 3. F, Region of tergosternal junction of segments 4 and 5. G, lateroventral view of segments 8 and 9. H, Lateral view of segments 810 in male stage-2. Abbreviations: lvs, lateroventral spine; lvt, lateroventral tube; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; mdcr, middorsal crenulated spine; mlcr, midlateral crenulated spine; mvp, midventral process; s, segment followed by segment number; sf, secondary fringe; vms, ventromedial spine on segment 1." figureDoi="http://doi.org/10.5281/zenodo.4502543" httpUri="https://zenodo.org/record/4502543/files/figure.png" pageId="7" pageNumber="10">4H</figureCitation>
,
<figureCitation id="13432A34DA304E0A3B88FFF4FE4CFC08" box="[390,459,1533,1557]" captionStart="Fig" captionStartId="8.[136,171,1694,1716]" captionTargetBox="[136,1472,297,1678]" captionTargetId="figure-8@8.[136,1474,297,1681]" captionTargetPageId="8" captionText="Fig. 5. Light micrographs showing details of trunk morphology in Centroderes impurus sp. nov. A, C, E: holotype, ZRC.MIS.0001, male stage-2; B, D, F: paratype, KIN-846, putative J6 or adult stage-1 of unknown gender. A, Segments 36, lateral view, with dashed circles indicating laterodorsal and lateral accessory sensory spots. B, Segments 13, lateral view, showing ventrolateral tube on segment 2, which is found only in this stage and not in male stage-2. C, Segments 10 and 11, lateral view, showing crenulated spines in middorsal position on segment 10. D, Segments 56, lateral view, showing indications of lateroventral tube on segment 5, which is found only in this stage and not in male stage-2. E, Segment 811, lateral view, showing crenulated spines in midlateral positions on segment 10; note the conspicuous difference between the lateroventral spines on segment 8 and segment 9; dashed circles indicate laterodorsal and lateral accessory sensory spots. F, Segment 811, lateral view, showing acicular spines in middorsal and midlateral positions on segment 10. Abbreviations: lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdac, middorsal acicular spine; mdcr, middorsal crenulated spine; mlac, midlateral acicular spine; mlcr, midlateral crenulated spine; vlt, ventrolateral tube; vms, ventromedial spine." figureDoi="http://doi.org/10.5281/zenodo.4502551" httpUri="https://zenodo.org/record/4502551/files/figure.png" pageId="7" pageNumber="10">5C, E</figureCitation>
). Sensory spots present in laterodorsal and ventromedial positions. Cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="8BC736B1DA304E0A3A7CFC74FE72FD6B" blockId="7.[114,754,1661,1910]" pageId="7" pageNumber="10">
Segment 11 with middorsal, lateral terminal, lateral terminal accessory, and midterminal spines (
<figureCitation id="13432A34DA304E0A3BF7FC94FDC0FCA8" box="[505,583,1693,1717]" captionStart="Fig" captionStartId="4.[135,170,1977,1999]" captionTargetBox="[196,1415,193,1962]" captionTargetId="figure-8@4.[194,1418,193,1965]" captionTargetPageId="4" captionText="Fig. 2. Line art illustration of Centroderes impurus sp. nov. male stage-2, lateral view. Abbreviations: ij, intersegmentary joint line; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral spine; mdp, middorsal placid; mds, middorsal spine; mls, midlateral spine; mlss, midlateral sensory spot; mts, midterminal spine; mvp, midventral process; pdss, paradorsal sensory spot; pf, pectinate fringe; sdss, subdorsal sensory spot; sf, secondary fringe; vms, ventromedial spine; vmss, ventromedial sensory spot. Digits following labels refer to segment numbers." figureDoi="http://doi.org/10.5281/zenodo.4502537" httpUri="https://zenodo.org/record/4502537/files/figure.png" pageId="7" pageNumber="10">Figs. 2</figureCitation>
,
<figureCitation id="13432A34DA304E0A3858FC94FDE2FCA8" box="[598,613,1693,1717]" captionStart="Fig" captionStartId="5.[113,148,2063,2085]" captionTargetBox="[235,1328,869,2047]" captionTargetId="figure-213@5.[235,1330,869,2047]" captionTargetPageId="5" captionText="Fig. 3. Light micrographs showing lateral overviews of Centroderes impurus sp. nov. A, holotype, ZRC.MIS.0001, male stage-2. B, paratype, KIN-846, putative J6 or adult stage-1 of unknown gender." figureDoi="http://doi.org/10.5281/zenodo.4502541" httpUri="https://zenodo.org/record/4502541/files/figure.png" pageId="7" pageNumber="10">3</figureCitation>
,
<figureCitation id="13432A34DA304E0A387DFC94FD3FFCA8" box="[627,696,1693,1717]" captionStart="Fig" captionStartId="6.[136,171,1919,1941]" captionTargetBox="[279,1332,189,1902]" captionTargetId="figure-7@6.[278,1332,189,1904]" captionTargetPageId="6" captionText="Fig. 4. Scanning electron micrographs of male stage-2 (A, B, F, H) and putative J6 or adult stage-1 of unknown gender (CE, G) of Centroderes impurus sp. nov. A, lateral overview. B, ventral overview. C, Lateral view of segment 1. D, Detail showing lateral accessory sensory spot on segment 1. E, Region of tergosternal junctions of segments 4 and 5; dashed circles outline extended cuticular hairs on posterior margin of segment 3. F, Region of tergosternal junction of segments 4 and 5. G, lateroventral view of segments 8 and 9. H, Lateral view of segments 810 in male stage-2. Abbreviations: lvs, lateroventral spine; lvt, lateroventral tube; lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; mdcr, middorsal crenulated spine; mlcr, midlateral crenulated spine; mvp, midventral process; s, segment followed by segment number; sf, secondary fringe; vms, ventromedial spine on segment 1." figureDoi="http://doi.org/10.5281/zenodo.4502543" httpUri="https://zenodo.org/record/4502543/files/figure.png" pageId="7" pageNumber="10">4A, B</figureCitation>
,
<figureCitation id="13432A34DA304E0A38CBFC94FD62FCA8" box="[709,741,1693,1717]" captionStart="Fig" captionStartId="8.[136,171,1694,1716]" captionTargetBox="[136,1472,297,1678]" captionTargetId="figure-8@8.[136,1474,297,1681]" captionTargetPageId="8" captionText="Fig. 5. Light micrographs showing details of trunk morphology in Centroderes impurus sp. nov. A, C, E: holotype, ZRC.MIS.0001, male stage-2; B, D, F: paratype, KIN-846, putative J6 or adult stage-1 of unknown gender. A, Segments 36, lateral view, with dashed circles indicating laterodorsal and lateral accessory sensory spots. B, Segments 13, lateral view, showing ventrolateral tube on segment 2, which is found only in this stage and not in male stage-2. C, Segments 10 and 11, lateral view, showing crenulated spines in middorsal position on segment 10. D, Segments 56, lateral view, showing indications of lateroventral tube on segment 5, which is found only in this stage and not in male stage-2. E, Segment 811, lateral view, showing crenulated spines in midlateral positions on segment 10; note the conspicuous difference between the lateroventral spines on segment 8 and segment 9; dashed circles indicate laterodorsal and lateral accessory sensory spots. F, Segment 811, lateral view, showing acicular spines in middorsal and midlateral positions on segment 10. Abbreviations: lass, lateral accessory sensory spot; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdac, middorsal acicular spine; mdcr, middorsal crenulated spine; mlac, midlateral acicular spine; mlcr, midlateral crenulated spine; vlt, ventrolateral tube; vms, ventromedial spine." figureDoi="http://doi.org/10.5281/zenodo.4502551" httpUri="https://zenodo.org/record/4502551/files/figure.png" pageId="7" pageNumber="10">5F</figureCitation>
). The midterminal spine is considerably longer than the other spines on this segment (
<tableCitation id="C6FA030ADA304E0A3B8EFCD4FE5EFCE8" box="[384,473,1757,1781]" captionStart="Table 2" captionStartId="3.[113,168,189,211]" captionText="Table 2. Measurements from light microscopy of adult Centroderes impurus sp. nov. (in µm) from station SI-03, Singapore, including individual measures, mean and standard deviation (SD). Abbreviations: LTAS: lateral terminal accessory spine; LTS: lateral terminal spine; LV: lateroventral spine or tube; MD, middorsal spine; ML: midlateral spine; MSW-6: maximum sternal width, measured on segment 6 in this species; MTS: midterminal spine; S: segment lengths; SW-10, standard width, always measured on segment 10; TL: trunk length; VM: ventromedial spine." pageId="7" pageNumber="10">Table 2</tableCitation>
). Two pairs of subdorsal sensory spots, one pair being more anterior than the other, are present. The whole cuticular surface is covered with minute cuticular hairs which turn slightly longer and denser in the terminal part of the segment.
</paragraph>
</subSubSection>
</treatment>
</document>