treatments-xml/data/03/AF/A4/03AFA402FFF2FFD68F8250797B862D85.xml

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<mods:title id="2D7460F4C2D223C9091B38A09A52A657">Description of the larvae of Drusus ramae Marinković- Gospodnetić and Drusus medianus Marinković- Gospodnetić (Trichoptera: Limnephilidae) with some genetic, distributional, ecological, faunal and conservation notes</mods:title>
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<treatment id="03AFA402FFF2FFD68F8250797B862D85" ID-DOI="http://doi.org/10.5281/zenodo.6201450" ID-GBIF-Taxon="134090652" ID-Zenodo-Dep="6201450" LSID="urn:lsid:plazi:treatment:03AFA402FFF2FFD68F8250797B862D85" httpUri="http://treatment.plazi.org/id/03AFA402FFF2FFD68F8250797B862D85" lastPageId="16" lastPageNumber="17" pageId="12" pageNumber="13">
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<subSubSection id="C31C469FFFF2FFCA8F8250797AAF2D8F" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BB91514FFF2FFCA8F8250797AAF2D8F" blockId="12.[151,1437,1383,1447]" pageId="12" pageNumber="13">
<heading id="D0F1A278FFF2FFCA8F8250797AAF2D8F" bold="true" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="B972C906FFF2FFCA8F8250797AAF2D8F" bold="true" pageId="12" pageNumber="13">
Morphological separation of larvae of 
<emphasis id="B972C906FFF2FFCA8D8E50797D4F2DA8" bold="true" box="[667,850,1383,1409]" italics="true" pageId="12" pageNumber="13">Drusus ramae</emphasis>
and 
<emphasis id="B972C906FFF2FFCA8C8050797A652DA8" bold="true" box="[917,1144,1383,1409]" italics="true" pageId="12" pageNumber="13">Drusus medianus</emphasis>
from other European 
<taxonomicName id="4C066E97FFF2FFCA8F8250927F142D8F" ID-CoL="JGV" authorityName="Banks" authorityYear="1916" box="[151,265,1420,1446]" class="Insecta" family="Limnephilidae" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subFamily" subFamily="Drusinae">Drusinae</taxonomicName>
and identification key for known 
<taxonomicName id="4C066E97FFF2FFCA8DA550937D152D8E" box="[688,776,1421,1447]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B972C906FFF2FFCA8DA550937D152D8E" bold="true" box="[688,776,1421,1447]" italics="true" pageId="12" pageNumber="13">Drusus</emphasis>
</taxonomicName>
larvae from the Balkan Peninsula
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31C469FFFF2FFC88FD350C97A57299E" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8BB91514FFF2FFCA8FD350C97A8D2E70" blockId="12.[151,1437,1495,2033]" pageId="12" pageNumber="13">
Larvae of only 2 endemic species from the Balkan Peninsula have been described previously, 
<taxonomicName id="4C066E97FFF2FFCA8A1950C67B812DD8" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[1292,1436,1496,1521]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="croaticus">
<emphasis id="B972C906FFF2FFCA8A1950C67B812DD8" box="[1292,1436,1496,1521]" italics="true" pageId="12" pageNumber="13">D. croaticus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF2FFCA8FDE50E47D202E3D" authority="(Kucinic et al. 2008)" authorityName="Kucinic et al." authorityYear="2008" box="[203,829,1529,1556]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="septentrionis">
<emphasis id="B972C906FFF2FFCA8FDE50E47C292E3A" box="[203,564,1530,1555]" italics="true" pageId="12" pageNumber="13">D. radovanovici septentrionis</emphasis>
(
<bibRefCitation id="EF9768E5FFF2FFCA8D5050E47D292E3D" author="Kucinic" box="[581,820,1529,1556]" pageId="12" pageNumber="13" refString="Kucinic, M., Previsic, A., Gottstein, S., Hrasovec, B., Stanic- Kostroman, S., Pernek, M. &amp; Delic, A. (2008) Description of the larvae of Drusus radovanovici septentrionis Marinkovic- Gospodnetic, 1976 and Drusus croaticus Marinkovic- Gospodnetic, 1971 (Trichoptera: Limnephilidae) from Bosnia and Herzegovina, and Croatia. Zootaxa, 1783, 1 - 17." type="journal article" year="2008">Kučinić et al. 2008</bibRefCitation>
)
</taxonomicName>
. 
<taxonomicName id="4C066E97FFF2FFCA8C5C50E47A0A2E3A" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[841,1047,1530,1555]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="croaticus">
<emphasis id="B972C906FFF2FFCA8C5C50E47A0A2E3A" box="[841,1047,1530,1555]" italics="true" pageId="12" pageNumber="13">Drusus croaticus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C066E97FFF2FFCA8B3250E47A842E3A" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[1063,1177,1530,1555]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF2FFCA8B3250E47A842E3A" box="[1063,1177,1530,1555]" italics="true" pageId="12" pageNumber="13">D. ramae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C066E97FFF2FFCA8BBF50E47B5E2E3A" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[1194,1347,1530,1555]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF2FFCA8BBF50E47B5E2E3A" box="[1194,1347,1530,1555]" italics="true" pageId="12" pageNumber="13">D. medianus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF2FFCA8A6A50E47FCD2E1F" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="septentrionis">
<emphasis id="B972C906FFF2FFCA8A6A50E47FCD2E1F" italics="true" pageId="12" pageNumber="13">D. radovanovici septentrionis</emphasis>
</taxonomicName>
larvae are morphologically very distinct, especially regarding features of the head (
<figureCitation id="133D0991FFF2FFCA8F8A53217F3B2E70" box="[159,294,1599,1625]" captionStart-0="FIGURES 33 – 36. 33" captionStart-1="FIGURES 37 – 40. 37" captionStartId-0="11.[151,269,1363,1387]" captionStartId-1="12.[151,269,1191,1215]" captionTargetBox-0="[151,1436,370,1342]" captionTargetBox-1="[151,1436,197,1167]" captionTargetId-0="figure@11.[151,1436,370,1342]" captionTargetId-1="figure@12.[151,1436,197,1167]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURES 33 – 36. 33, Drusus croaticus Marinković- Gospodnetić, 1971: head and pronotum protruding from case, right lateral view; 34, Drusus ramae Marinković- Gospodnetić, 1971: head and pronotum, right lateral view; 35, Drusus medianus Marinković- Gospodnetić, 1976: head and pronotum, right lateral view; 36, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: head and pronotum, right lateral view." captionText-1="FIGURES 37 – 40. 37, Drusus croaticus Marinković- Gospodnetić, 1971: posterior part of the head, right lateral view; 38, Drusus ramae Marinković- Gospodnetić, 1971: posterior part of the head, right lateral view; 39, Drusus medianus Marinković- Gospodnetić, 1976: parietal part of the head at eye level, right lateral view; 40, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: parietal part of the head at eye level, right lateral view." httpUri-0="https://zenodo.org/record/1051922/files/figure.png" httpUri-1="https://zenodo.org/record/1051924/files/figure.png" pageId="12" pageNumber="13">Figs 33–40</figureCitation>
, 
<figureCitation id="133D0991FFF2FFCA8E2753217F4E2E70" box="[306,339,1599,1625]" captionStart="FIGURES 41 – 45. 41" captionStartId="13.[151,269,1233,1257]" captionTargetBox="[151,1436,197,1206]" captionTargetId="figure@13.[151,1436,197,1209]" captionTargetPageId="13" captionText="FIGURES 41 – 45. 41, Drusus croaticus Marinković- Gospodnetić, 1971: pronotum, right lateral view; 42, Drusus ramae Marinković- Gospodnetić, 1971: pronotum, right lateral view; 43, Drusus medianus Marinković- Gospodnetić, 1976: pronotum, right lateral view; 44, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: pronotum, right lateral view; 45, Drusus medianus Marinković- Gospodnetić, 1976 (left), and D. radovanovici septentrionis Marinković- Gospodnetić, 1976 (right): parietal part of the head at eye level showing spinules, right lateral view." httpUri="https://zenodo.org/record/1051927/files/figure.png" pageId="12" pageNumber="13">45</figureCitation>
), pronotum (
<figureCitation id="133D0991FFF2FFCA8EFF53217C6C2E70" box="[490,625,1599,1625]" captionStart="FIGURES 33 – 36. 33" captionStartId="11.[151,269,1363,1387]" captionTargetBox="[151,1436,370,1342]" captionTargetId="figure@11.[151,1436,370,1342]" captionTargetPageId="11" captionText="FIGURES 33 – 36. 33, Drusus croaticus Marinković- Gospodnetić, 1971: head and pronotum protruding from case, right lateral view; 34, Drusus ramae Marinković- Gospodnetić, 1971: head and pronotum, right lateral view; 35, Drusus medianus Marinković- Gospodnetić, 1976: head and pronotum, right lateral view; 36, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: head and pronotum, right lateral view." httpUri="https://zenodo.org/record/1051922/files/figure.png" pageId="12" pageNumber="13">Figs 33–36</figureCitation>
, 
<figureCitation id="133D0991FFF2FFCA8D6853217CD52E70" box="[637,712,1599,1625]" captionStart="FIGURES 41 – 45. 41" captionStartId="13.[151,269,1233,1257]" captionTargetBox="[151,1436,197,1206]" captionTargetId="figure@13.[151,1436,197,1209]" captionTargetPageId="13" captionText="FIGURES 41 – 45. 41, Drusus croaticus Marinković- Gospodnetić, 1971: pronotum, right lateral view; 42, Drusus ramae Marinković- Gospodnetić, 1971: pronotum, right lateral view; 43, Drusus medianus Marinković- Gospodnetić, 1976: pronotum, right lateral view; 44, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: pronotum, right lateral view; 45, Drusus medianus Marinković- Gospodnetić, 1976 (left), and D. radovanovici septentrionis Marinković- Gospodnetić, 1976 (right): parietal part of the head at eye level showing spinules, right lateral view." httpUri="https://zenodo.org/record/1051927/files/figure.png" pageId="12" pageNumber="13">41–44</figureCitation>
) and metanotal sclerites (
<figureCitation id="133D0991FFF2FFCA8CE153217A992E70" box="[1012,1156,1599,1625]" captionStart="FIGURES 46 – 53" captionStartId="15.[151,272,1869,1893]" captionTargetBox="[151,1424,197,1846]" captionTargetId="figure@15.[151,1436,197,1846]" captionTargetPageId="15" captionText="FIGURES 46 – 53. Metanota of Drusus spp., showing anteromedian (sa 1) and posteromedian (sa 2) sclerites. 46 – 47, Drusus ramae Marinković- Gospodnetić, 1976; 48 – 49, Drusus medianus Marinković- Gospodnetić, 1971; 50 – 51, D. radovanovici septentrionis Marinković- Gospodnetić, 1976; 52 – 53, Drusus croaticus Marinković- Gospodnetić, 1971." httpUri="https://zenodo.org/record/1051929/files/figure.png" pageId="12" pageNumber="13">Figs. 46–53</figureCitation>
).
</paragraph>
<paragraph id="8BB91514FFF2FFCA8FD3537C7AA82EEE" blockId="12.[151,1437,1495,2033]" pageId="12" pageNumber="13">
Larvae of 
<taxonomicName id="4C066E97FFF2FFCA8E54537C7FAC2E52" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[321,433,1634,1659]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF2FFCA8E54537C7FAC2E52" box="[321,433,1634,1659]" italics="true" pageId="12" pageNumber="13">D. ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF2FFCA8EF8537C7C9F2E52" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[493,642,1634,1659]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF2FFCA8EF8537C7C9F2E52" box="[493,642,1634,1659]" italics="true" pageId="12" pageNumber="13">D. medianus</emphasis>
</taxonomicName>
are easily distinguished from each other by the markedly different shape of the pronotum (
<figureCitation id="133D0991FFF2FFCA8EA953997C572E88" box="[444,586,1671,1697]" captionStart="FIGURES 33 – 36. 33" captionStartId="11.[151,269,1363,1387]" captionTargetBox="[151,1436,370,1342]" captionTargetId="figure@11.[151,1436,370,1342]" captionTargetPageId="11" captionText="FIGURES 33 – 36. 33, Drusus croaticus Marinković- Gospodnetić, 1971: head and pronotum protruding from case, right lateral view; 34, Drusus ramae Marinković- Gospodnetić, 1971: head and pronotum, right lateral view; 35, Drusus medianus Marinković- Gospodnetić, 1976: head and pronotum, right lateral view; 36, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: head and pronotum, right lateral view." httpUri="https://zenodo.org/record/1051922/files/figure.png" pageId="12" pageNumber="13">Figs 34–35</figureCitation>
, 
<figureCitation id="133D0991FFF2FFCA8D4D53997CB42E88" box="[600,681,1671,1697]" captionStart="FIGURES 41 – 45. 41" captionStartId="13.[151,269,1233,1257]" captionTargetBox="[151,1436,197,1206]" captionTargetId="figure@13.[151,1436,197,1209]" captionTargetPageId="13" captionText="FIGURES 41 – 45. 41, Drusus croaticus Marinković- Gospodnetić, 1971: pronotum, right lateral view; 42, Drusus ramae Marinković- Gospodnetić, 1971: pronotum, right lateral view; 43, Drusus medianus Marinković- Gospodnetić, 1976: pronotum, right lateral view; 44, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: pronotum, right lateral view; 45, Drusus medianus Marinković- Gospodnetić, 1976 (left), and D. radovanovici septentrionis Marinković- Gospodnetić, 1976 (right): parietal part of the head at eye level showing spinules, right lateral view." httpUri="https://zenodo.org/record/1051927/files/figure.png" pageId="12" pageNumber="13">42–43</figureCitation>
). Moreover, larvae of 
<taxonomicName id="4C066E97FFF2FFCA8CAE53967A4B2E88" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[955,1110,1672,1697]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF2FFCA8CAE53967A4B2E88" box="[955,1110,1672,1697]" italics="true" pageId="12" pageNumber="13">D. medianus</emphasis>
</taxonomicName>
have spinules on the head beside the eyes (
<figureCitation id="133D0991FFF2FFCA8E4D53B37FFB2EEE" box="[344,486,1709,1735]" captionStart="FIGURES 16 – 21" captionStartId="8.[151,270,1652,1676]" captionTargetBox="[151,1436,197,1629]" captionTargetId="figure@8.[151,1436,197,1629]" captionTargetPageId="8" captionText="FIGURES 16 – 21. Drusus medianus Marinković- Gospodnetić, 1976. 16, larva in its case, right lateral view; 17, larva, right lateral view; 18, head, pronotum and mesonotum, right lateral view; 19, head with spinules above eye, right lateral view; 20, area with spinules, right lateral view; 21, spinules, right lateral view." httpUri="https://zenodo.org/record/1051916/files/figure.png" pageId="12" pageNumber="13">Figs. 19–21</figureCitation>
, 
<figureCitation id="133D0991FFF2FFCA8EE753B37C0E2EEE" box="[498,531,1709,1735]" captionStart="FIGURES 41 – 45. 41" captionStartId="13.[151,269,1233,1257]" captionTargetBox="[151,1436,197,1206]" captionTargetId="figure@13.[151,1436,197,1209]" captionTargetPageId="13" captionText="FIGURES 41 – 45. 41, Drusus croaticus Marinković- Gospodnetić, 1971: pronotum, right lateral view; 42, Drusus ramae Marinković- Gospodnetić, 1971: pronotum, right lateral view; 43, Drusus medianus Marinković- Gospodnetić, 1976: pronotum, right lateral view; 44, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: pronotum, right lateral view; 45, Drusus medianus Marinković- Gospodnetić, 1976 (left), and D. radovanovici septentrionis Marinković- Gospodnetić, 1976 (right): parietal part of the head at eye level showing spinules, right lateral view." httpUri="https://zenodo.org/record/1051927/files/figure.png" pageId="12" pageNumber="13">45</figureCitation>
), while larvae of 
<taxonomicName id="4C066E97FFF2FFCA8DCA53B37D532EEF" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[735,846,1709,1734]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF2FFCA8DCA53B37D532EEF" box="[735,846,1709,1734]" italics="true" pageId="12" pageNumber="13">D. ramae</emphasis>
</taxonomicName>
lack this feature (
<figureCitation id="133D0991FFF2FFCA8B0A53B37A672EEE" box="[1055,1146,1709,1735]" captionStart="FIGURES 33 – 36. 33" captionStartId="11.[151,269,1363,1387]" captionTargetBox="[151,1436,370,1342]" captionTargetId="figure@11.[151,1436,370,1342]" captionTargetPageId="11" captionText="FIGURES 33 – 36. 33, Drusus croaticus Marinković- Gospodnetić, 1971: head and pronotum protruding from case, right lateral view; 34, Drusus ramae Marinković- Gospodnetić, 1971: head and pronotum, right lateral view; 35, Drusus medianus Marinković- Gospodnetić, 1976: head and pronotum, right lateral view; 36, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: head and pronotum, right lateral view." httpUri="https://zenodo.org/record/1051922/files/figure.png" pageId="12" pageNumber="13">Figs 34</figureCitation>
, 
<figureCitation id="133D0991FFF2FFCA8B9253B37AB52EEE" box="[1159,1192,1709,1735]" captionStart="FIGURES 37 – 40. 37" captionStartId="12.[151,269,1191,1215]" captionTargetBox="[151,1436,197,1167]" captionTargetId="figure@12.[151,1436,197,1167]" captionTargetPageId="12" captionText="FIGURES 37 – 40. 37, Drusus croaticus Marinković- Gospodnetić, 1971: posterior part of the head, right lateral view; 38, Drusus ramae Marinković- Gospodnetić, 1971: posterior part of the head, right lateral view; 39, Drusus medianus Marinković- Gospodnetić, 1976: parietal part of the head at eye level, right lateral view; 40, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: parietal part of the head at eye level, right lateral view." httpUri="https://zenodo.org/record/1051924/files/figure.png" pageId="12" pageNumber="13">38</figureCitation>
).
</paragraph>
<paragraph id="8BB91514FFF2FFCA8FD353CC7D702FD8" blockId="12.[151,1437,1495,2033]" pageId="12" pageNumber="13">
The larva of 
<taxonomicName id="4C066E97FFF2FFCA8E7353CC7FE42EC2" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[358,505,1746,1771]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="croaticus">
<emphasis id="B972C906FFF2FFCA8E7353CC7FE42EC2" box="[358,505,1746,1771]" italics="true" pageId="12" pageNumber="13">D. croaticus</emphasis>
</taxonomicName>
is easily distinguished from the larvae of 
<taxonomicName id="4C066E97FFF2FFCA8CE853CC7A722EC2" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[1021,1135,1746,1771]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF2FFCA8CE853CC7A722EC2" box="[1021,1135,1746,1771]" italics="true" pageId="12" pageNumber="13">D. ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF2FFCA8BBA53CC7B5A2EC2" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[1199,1351,1746,1771]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF2FFCA8BBA53CC7B5A2EC2" box="[1199,1351,1746,1771]" italics="true" pageId="12" pageNumber="13">D. medianus</emphasis>
</taxonomicName>
by the following characters: the shape of pronotum (
<figureCitation id="133D0991FFF2FFCA8DA453E97D272F38" box="[689,826,1783,1809]" captionStart="FIGURES 33 – 36. 33" captionStartId="11.[151,269,1363,1387]" captionTargetBox="[151,1436,370,1342]" captionTargetId="figure@11.[151,1436,370,1342]" captionTargetPageId="11" captionText="FIGURES 33 – 36. 33, Drusus croaticus Marinković- Gospodnetić, 1971: head and pronotum protruding from case, right lateral view; 34, Drusus ramae Marinković- Gospodnetić, 1971: head and pronotum, right lateral view; 35, Drusus medianus Marinković- Gospodnetić, 1976: head and pronotum, right lateral view; 36, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: head and pronotum, right lateral view." httpUri="https://zenodo.org/record/1051922/files/figure.png" pageId="12" pageNumber="13">Figs 33–35</figureCitation>
, 
<figureCitation id="133D0991FFF2FFCA8C5353E97D8C2F38" box="[838,913,1783,1809]" captionStart="FIGURES 41 – 45. 41" captionStartId="13.[151,269,1233,1257]" captionTargetBox="[151,1436,197,1206]" captionTargetId="figure@13.[151,1436,197,1209]" captionTargetPageId="13" captionText="FIGURES 41 – 45. 41, Drusus croaticus Marinković- Gospodnetić, 1971: pronotum, right lateral view; 42, Drusus ramae Marinković- Gospodnetić, 1971: pronotum, right lateral view; 43, Drusus medianus Marinković- Gospodnetić, 1976: pronotum, right lateral view; 44, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: pronotum, right lateral view; 45, Drusus medianus Marinković- Gospodnetić, 1976 (left), and D. radovanovici septentrionis Marinković- Gospodnetić, 1976 (right): parietal part of the head at eye level showing spinules, right lateral view." httpUri="https://zenodo.org/record/1051927/files/figure.png" pageId="12" pageNumber="13">41–43</figureCitation>
) (
<taxonomicName id="4C066E97FFF2FFCA8CBE53E67A262F38" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[939,1083,1784,1809]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="croaticus">
<emphasis id="B972C906FFF2FFCA8CBE53E67A262F38" box="[939,1083,1784,1809]" italics="true" pageId="12" pageNumber="13">D. croaticus</emphasis>
</taxonomicName>
has a more rounded posterior part of the pronotum and a flattened anterior part), absence of spinules on the head, and absence of prominent black median setae at the anterior border of the pronotum in 
<taxonomicName id="4C066E97FFF2FFCA8C4D525C7DFA2F72" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[856,999,1858,1883]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="croaticus">
<emphasis id="B972C906FFF2FFCA8C4D525C7DFA2F72" box="[856,999,1858,1883]" italics="true" pageId="12" pageNumber="13">D. croaticus</emphasis>
</taxonomicName>
(
<figureCitation id="133D0991FFF2FFCA8CE3525C7A4C2F75" box="[1014,1105,1858,1884]" captionStart="FIGURES 33 – 36. 33" captionStartId="11.[151,269,1363,1387]" captionTargetBox="[151,1436,370,1342]" captionTargetId="figure@11.[151,1436,370,1342]" captionTargetPageId="11" captionText="FIGURES 33 – 36. 33, Drusus croaticus Marinković- Gospodnetić, 1971: head and pronotum protruding from case, right lateral view; 34, Drusus ramae Marinković- Gospodnetić, 1971: head and pronotum, right lateral view; 35, Drusus medianus Marinković- Gospodnetić, 1976: head and pronotum, right lateral view; 36, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: head and pronotum, right lateral view." httpUri="https://zenodo.org/record/1051922/files/figure.png" pageId="12" pageNumber="13">Figs 33</figureCitation>
, 
<figureCitation id="133D0991FFF2FFCA8B49525C7A612F75" box="[1116,1148,1858,1884]" captionStart="FIGURES 37 – 40. 37" captionStartId="12.[151,269,1191,1215]" captionTargetBox="[151,1436,197,1167]" captionTargetId="figure@12.[151,1436,197,1167]" captionTargetPageId="12" captionText="FIGURES 37 – 40. 37, Drusus croaticus Marinković- Gospodnetić, 1971: posterior part of the head, right lateral view; 38, Drusus ramae Marinković- Gospodnetić, 1971: posterior part of the head, right lateral view; 39, Drusus medianus Marinković- Gospodnetić, 1976: parietal part of the head at eye level, right lateral view; 40, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: parietal part of the head at eye level, right lateral view." httpUri="https://zenodo.org/record/1051924/files/figure.png" pageId="12" pageNumber="13">37</figureCitation>
, 
<figureCitation id="133D0991FFF2FFCA8B9D525C7AB42F75" box="[1160,1193,1858,1884]" captionStart="FIGURES 41 – 45. 41" captionStartId="13.[151,269,1233,1257]" captionTargetBox="[151,1436,197,1206]" captionTargetId="figure@13.[151,1436,197,1209]" captionTargetPageId="13" captionText="FIGURES 41 – 45. 41, Drusus croaticus Marinković- Gospodnetić, 1971: pronotum, right lateral view; 42, Drusus ramae Marinković- Gospodnetić, 1971: pronotum, right lateral view; 43, Drusus medianus Marinković- Gospodnetić, 1976: pronotum, right lateral view; 44, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: pronotum, right lateral view; 45, Drusus medianus Marinković- Gospodnetić, 1976 (left), and D. radovanovici septentrionis Marinković- Gospodnetić, 1976 (right): parietal part of the head at eye level showing spinules, right lateral view." httpUri="https://zenodo.org/record/1051927/files/figure.png" pageId="12" pageNumber="13">41</figureCitation>
). Similarly, the larva of 
<taxonomicName id="4C066E97FFF2FFCA8FAF52767C222FA8" box="[186,575,1896,1921]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="serptentrionis">
<emphasis id="B972C906FFF2FFCA8FAF52767C222FA8" box="[186,575,1896,1921]" italics="true" pageId="12" pageNumber="13">D. radovanovici serptentrionis</emphasis>
</taxonomicName>
is easily distinguished from the larva of 
<taxonomicName id="4C066E97FFF2FFCA8B5E52767ADE2FA8" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[1099,1219,1896,1921]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF2FFCA8B5E52767ADE2FA8" box="[1099,1219,1896,1921]" italics="true" pageId="12" pageNumber="13">D. ramae</emphasis>
</taxonomicName>
by the following characters: the shape of the pronotum (
<taxonomicName id="4C066E97FFF2FFCA8D7352937CC82F8F" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[614,725,1933,1958]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF2FFCA8D7352937CC82F8F" box="[614,725,1933,1958]" italics="true" pageId="12" pageNumber="13">D. ramae</emphasis>
</taxonomicName>
has two prominent acute humps lacking in 
<taxonomicName id="4C066E97FFF2FFCA8BC952937F282FE2" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="septentrionis">
<emphasis id="B972C906FFF2FFCA8BC952937F282FE2" italics="true" pageId="12" pageNumber="13">D. radovanovici septentrionis</emphasis>
</taxonomicName>
) (
<figureCitation id="133D0991FFF2FFCA8E4452AC7FC22FE5" box="[337,479,1970,1996]" captionStart="FIGURES 33 – 36. 33" captionStartId="11.[151,269,1363,1387]" captionTargetBox="[151,1436,370,1342]" captionTargetId="figure@11.[151,1436,370,1342]" captionTargetPageId="11" captionText="FIGURES 33 – 36. 33, Drusus croaticus Marinković- Gospodnetić, 1971: head and pronotum protruding from case, right lateral view; 34, Drusus ramae Marinković- Gospodnetić, 1971: head and pronotum, right lateral view; 35, Drusus medianus Marinković- Gospodnetić, 1976: head and pronotum, right lateral view; 36, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: head and pronotum, right lateral view." httpUri="https://zenodo.org/record/1051922/files/figure.png" pageId="12" pageNumber="13">Figs 34, 36</figureCitation>
, 
<figureCitation id="133D0991FFF2FFCA8EF852AC7C212FE5" box="[493,572,1970,1996]" captionStart="FIGURES 41 – 45. 41" captionStartId="13.[151,269,1233,1257]" captionTargetBox="[151,1436,197,1206]" captionTargetId="figure@13.[151,1436,197,1209]" captionTargetPageId="13" captionText="FIGURES 41 – 45. 41, Drusus croaticus Marinković- Gospodnetić, 1971: pronotum, right lateral view; 42, Drusus ramae Marinković- Gospodnetić, 1971: pronotum, right lateral view; 43, Drusus medianus Marinković- Gospodnetić, 1976: pronotum, right lateral view; 44, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: pronotum, right lateral view; 45, Drusus medianus Marinković- Gospodnetić, 1976 (left), and D. radovanovici septentrionis Marinković- Gospodnetić, 1976 (right): parietal part of the head at eye level showing spinules, right lateral view." httpUri="https://zenodo.org/record/1051927/files/figure.png" pageId="12" pageNumber="13">42, 44</figureCitation>
) and presence/absence of head spinules beside the eyes (present in 
<taxonomicName id="4C066E97FFF2FFCA8A6A52AC7FCD2FD8" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="septentrionis">
<emphasis id="B972C906FFF2FFCA8A6A52AC7FCD2FD8" italics="true" pageId="12" pageNumber="13">D. radovanovici septentrionis</emphasis>
</taxonomicName>
, absent in 
<taxonomicName id="4C066E97FFF2FFCA8D5A52C67CA32FD8" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[591,702,2008,2033]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF2FFCA8D5A52C67CA32FD8" box="[591,702,2008,2033]" italics="true" pageId="12" pageNumber="13">D. ramae</emphasis>
</taxonomicName>
) (
<figureCitation id="133D0991FFF2FFCA8DC252C97D2F2FD8" box="[727,818,2007,2033]" captionStart="FIGURES 33 – 36. 33" captionStartId="11.[151,269,1363,1387]" captionTargetBox="[151,1436,370,1342]" captionTargetId="figure@11.[151,1436,370,1342]" captionTargetPageId="11" captionText="FIGURES 33 – 36. 33, Drusus croaticus Marinković- Gospodnetić, 1971: head and pronotum protruding from case, right lateral view; 34, Drusus ramae Marinković- Gospodnetić, 1971: head and pronotum, right lateral view; 35, Drusus medianus Marinković- Gospodnetić, 1976: head and pronotum, right lateral view; 36, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: head and pronotum, right lateral view." httpUri="https://zenodo.org/record/1051922/files/figure.png" pageId="12" pageNumber="13">Figs 34</figureCitation>
, 
<figureCitation id="133D0991FFF2FFCA8C2B52C97D422FD8" box="[830,863,2007,2033]" captionStart="FIGURES 37 – 40. 37" captionStartId="12.[151,269,1191,1215]" captionTargetBox="[151,1436,197,1167]" captionTargetId="figure@12.[151,1436,197,1167]" captionTargetPageId="12" captionText="FIGURES 37 – 40. 37, Drusus croaticus Marinković- Gospodnetić, 1971: posterior part of the head, right lateral view; 38, Drusus ramae Marinković- Gospodnetić, 1971: posterior part of the head, right lateral view; 39, Drusus medianus Marinković- Gospodnetić, 1976: parietal part of the head at eye level, right lateral view; 40, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: parietal part of the head at eye level, right lateral view." httpUri="https://zenodo.org/record/1051924/files/figure.png" pageId="12" pageNumber="13">38</figureCitation>
).
</paragraph>
<caption id="DF79459CFFF3FFCB8F8251CF7AA22D41" httpUri="https://zenodo.org/record/1051927/files/figure.png" pageId="13" pageNumber="14" targetBox="[151,1436,197,1206]" targetPageId="13">
<paragraph id="8BB91514FFF3FFCB8F8251CF7AA22D41" blockId="13.[151,1437,1233,1385]" pageId="13" pageNumber="14">
<emphasis id="B972C906FFF3FFCB8F8251CF7F4A2CC0" bold="true" box="[151,343,1233,1257]" pageId="13" pageNumber="14">FIGURES 41–45</emphasis>
. 41, 
<taxonomicName id="4C066E97FFF3FFCB8E9E51CC7C5C2CC0" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[395,577,1234,1257]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="croaticus">
<emphasis id="B972C906FFF3FFCB8E9E51CC7C5C2CC0" box="[395,577,1234,1257]" italics="true" pageId="13" pageNumber="14">Drusus croaticus</emphasis>
</taxonomicName>
Marinković- Gospodnetić, 1971: pronotum, right lateral view; 42, 
<taxonomicName id="4C066E97FFF3FFCB8A1151CC7B812CC0" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[1284,1436,1234,1257]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF3FFCB8A1151CC7B812CC0" box="[1284,1436,1234,1257]" italics="true" pageId="13" pageNumber="14">Drusus ramae</emphasis>
</taxonomicName>
Marinković- Gospodnetić, 1971: pronotum, right lateral view; 43, 
<taxonomicName id="4C066E97FFF3FFCB8C6751EC7A2F2D20" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[882,1074,1266,1289]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF3FFCB8C6751EC7A2F2D20" box="[882,1074,1266,1289]" italics="true" pageId="13" pageNumber="14">Drusus medianus</emphasis>
</taxonomicName>
Marinković- Gospodnetić, 1976: pronotum, right lateral view; 44, 
<taxonomicName id="4C066E97FFF3FFCB8EEB500C7D5D2D00" box="[510,832,1298,1321]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="septentrionis">
<emphasis id="B972C906FFF3FFCB8EEB500C7D5D2D00" box="[510,832,1298,1321]" italics="true" pageId="13" pageNumber="14">D. radovanovici septentrionis</emphasis>
</taxonomicName>
Marinković- Gospodnetić, 1976: pronotum, right lateral view; 45, 
<taxonomicName id="4C066E97FFF3FFCB8E1D502C7FD12D60" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[264,460,1330,1353]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF3FFCB8E1D502C7FD12D60" box="[264,460,1330,1353]" italics="true" pageId="13" pageNumber="14">Drusus medianus</emphasis>
</taxonomicName>
Marinković- Gospodnetić, 1976 (left), and 
<taxonomicName id="4C066E97FFF3FFCB8CA2502C7B152D60" box="[951,1288,1330,1353]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="septentrionis">
<emphasis id="B972C906FFF3FFCB8CA2502C7B152D60" box="[951,1288,1330,1353]" italics="true" pageId="13" pageNumber="14">D. radovanovici septentrionis</emphasis>
</taxonomicName>
Marinković- Gospodnetić, 1976 (right): parietal part of the head at eye level showing spinules, right lateral view.
</paragraph>
</caption>
<paragraph id="8BB91514FFF3FFCB8FD3508C7A9B2EE8" blockId="13.[151,1437,1426,2025]" pageId="13" pageNumber="14">
<taxonomicName id="4C066E97FFF3FFCB8FD3508C7C3E2D82" box="[198,547,1426,1451]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="septentrionis">
<emphasis id="B972C906FFF3FFCB8FD3508C7C3E2D82" box="[198,547,1426,1451]" italics="true" pageId="13" pageNumber="14">D. radovanovici septentrionis</emphasis>
</taxonomicName>
is morphologically quite similar to 
<taxonomicName id="4C066E97FFF3FFCB8CDC508C7A432D82" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[969,1118,1426,1451]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF3FFCB8CDC508C7A432D82" box="[969,1118,1426,1451]" italics="true" pageId="13" pageNumber="14">D. medianus</emphasis>
</taxonomicName>
but some subtle characters distinguish larvae of these two species: 
<taxonomicName id="4C066E97FFF3FFCB8D6C50AA7D0D2DE4" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[633,784,1460,1485]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF3FFCB8D6C50AA7D0D2DE4" box="[633,784,1460,1485]" italics="true" pageId="13" pageNumber="14">D. medianus</emphasis>
</taxonomicName>
bears white recumbent setae on the whole area of the pronotum (
<figureCitation id="133D0991FFF3FFCB8E0C50C97F6B2DD8" box="[281,374,1495,1521]" captionStart="FIGURES 33 – 36. 33" captionStartId="11.[151,269,1363,1387]" captionTargetBox="[151,1436,370,1342]" captionTargetId="figure@11.[151,1436,370,1342]" captionTargetPageId="11" captionText="FIGURES 33 – 36. 33, Drusus croaticus Marinković- Gospodnetić, 1971: head and pronotum protruding from case, right lateral view; 34, Drusus ramae Marinković- Gospodnetić, 1971: head and pronotum, right lateral view; 35, Drusus medianus Marinković- Gospodnetić, 1976: head and pronotum, right lateral view; 36, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: head and pronotum, right lateral view." httpUri="https://zenodo.org/record/1051922/files/figure.png" pageId="13" pageNumber="14">Figs 35</figureCitation>
, 
<figureCitation id="133D0991FFF3FFCB8E9750C97FBE2DD8" box="[386,419,1495,1521]" captionStart="FIGURES 41 – 45. 41" captionStartId="13.[151,269,1233,1257]" captionTargetBox="[151,1436,197,1206]" captionTargetId="figure@13.[151,1436,197,1209]" captionTargetPageId="13" captionText="FIGURES 41 – 45. 41, Drusus croaticus Marinković- Gospodnetić, 1971: pronotum, right lateral view; 42, Drusus ramae Marinković- Gospodnetić, 1971: pronotum, right lateral view; 43, Drusus medianus Marinković- Gospodnetić, 1976: pronotum, right lateral view; 44, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: pronotum, right lateral view; 45, Drusus medianus Marinković- Gospodnetić, 1976 (left), and D. radovanovici septentrionis Marinković- Gospodnetić, 1976 (right): parietal part of the head at eye level showing spinules, right lateral view." httpUri="https://zenodo.org/record/1051927/files/figure.png" pageId="13" pageNumber="14">43</figureCitation>
), but 
<taxonomicName id="4C066E97FFF3FFCB8EF250C97D5A2DD9" box="[487,839,1495,1520]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="septentrionis">
<emphasis id="B972C906FFF3FFCB8EF250C97D5A2DD9" box="[487,839,1495,1520]" italics="true" pageId="13" pageNumber="14">D. radovanovici septentrionis</emphasis>
</taxonomicName>
bears white recumbent setae only on the posterior and lateral parts of its pronotum (
<figureCitation id="133D0991FFF3FFCB8D3250E47C992E3D" box="[551,644,1530,1556]" captionStart="FIGURES 33 – 36. 33" captionStartId="11.[151,269,1363,1387]" captionTargetBox="[151,1436,370,1342]" captionTargetId="figure@11.[151,1436,370,1342]" captionTargetPageId="11" captionText="FIGURES 33 – 36. 33, Drusus croaticus Marinković- Gospodnetić, 1971: head and pronotum protruding from case, right lateral view; 34, Drusus ramae Marinković- Gospodnetić, 1971: head and pronotum, right lateral view; 35, Drusus medianus Marinković- Gospodnetić, 1976: head and pronotum, right lateral view; 36, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: head and pronotum, right lateral view." httpUri="https://zenodo.org/record/1051922/files/figure.png" pageId="13" pageNumber="14">Figs 36</figureCitation>
, 
<figureCitation id="133D0991FFF3FFCB8D8450E47CAE2E3D" box="[657,691,1530,1556]" captionStart="FIGURES 41 – 45. 41" captionStartId="13.[151,269,1233,1257]" captionTargetBox="[151,1436,197,1206]" captionTargetId="figure@13.[151,1436,197,1209]" captionTargetPageId="13" captionText="FIGURES 41 – 45. 41, Drusus croaticus Marinković- Gospodnetić, 1971: pronotum, right lateral view; 42, Drusus ramae Marinković- Gospodnetić, 1971: pronotum, right lateral view; 43, Drusus medianus Marinković- Gospodnetić, 1976: pronotum, right lateral view; 44, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: pronotum, right lateral view; 45, Drusus medianus Marinković- Gospodnetić, 1976 (left), and D. radovanovici septentrionis Marinković- Gospodnetić, 1976 (right): parietal part of the head at eye level showing spinules, right lateral view." httpUri="https://zenodo.org/record/1051927/files/figure.png" pageId="13" pageNumber="14">44</figureCitation>
). About 10 % of 
<taxonomicName id="4C066E97FFF3FFCB8C9750E47AF82E3A" box="[898,1253,1530,1555]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="septentrionis">
<emphasis id="B972C906FFF3FFCB8C9750E47AF82E3A" box="[898,1253,1530,1555]" italics="true" pageId="13" pageNumber="14">D. radovanovici septentrionis</emphasis>
</taxonomicName>
specimens had only a few, small, white recumbent setae anteromesally on the pronotum as opposed to 
<taxonomicName id="4C066E97FFF3FFCB8BA153027B562E1C" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[1204,1355,1564,1589]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF3FFCB8BA153027B562E1C" box="[1204,1355,1564,1589]" italics="true" pageId="13" pageNumber="14">D. medianus</emphasis>
</taxonomicName>
larvae that had numerous and longer white setae. In some cases (10–15% specimens), areas with spinules are larger in 
<taxonomicName id="4C066E97FFF3FFCB8FA0537C7F572E52" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[181,330,1634,1659]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF3FFCB8FA0537C7F572E52" box="[181,330,1634,1659]" italics="true" pageId="13" pageNumber="14">D. medianus</emphasis>
</taxonomicName>
(
<figureCitation id="133D0991FFF3FFCB8E4C537C7FFF2E55" box="[345,482,1634,1660]" captionStart="FIGURES 16 – 21" captionStartId="8.[151,270,1652,1676]" captionTargetBox="[151,1436,197,1629]" captionTargetId="figure@8.[151,1436,197,1629]" captionTargetPageId="8" captionText="FIGURES 16 – 21. Drusus medianus Marinković- Gospodnetić, 1976. 16, larva in its case, right lateral view; 17, larva, right lateral view; 18, head, pronotum and mesonotum, right lateral view; 19, head with spinules above eye, right lateral view; 20, area with spinules, right lateral view; 21, spinules, right lateral view." httpUri="https://zenodo.org/record/1051916/files/figure.png" pageId="13" pageNumber="14">Figs 19, 20</figureCitation>
, 
<figureCitation id="133D0991FFF3FFCB8EFB537C7C132E55" box="[494,526,1634,1660]" captionStart="FIGURES 37 – 40. 37" captionStartId="12.[151,269,1191,1215]" captionTargetBox="[151,1436,197,1167]" captionTargetId="figure@12.[151,1436,197,1167]" captionTargetPageId="12" captionText="FIGURES 37 – 40. 37, Drusus croaticus Marinković- Gospodnetić, 1971: posterior part of the head, right lateral view; 38, Drusus ramae Marinković- Gospodnetić, 1971: posterior part of the head, right lateral view; 39, Drusus medianus Marinković- Gospodnetić, 1976: parietal part of the head at eye level, right lateral view; 40, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: parietal part of the head at eye level, right lateral view." httpUri="https://zenodo.org/record/1051924/files/figure.png" pageId="13" pageNumber="14">39</figureCitation>
, 
<figureCitation id="133D0991FFF3FFCB8D0F537C7C242E55" box="[538,569,1634,1660]" captionStart="FIGURES 41 – 45. 41" captionStartId="13.[151,269,1233,1257]" captionTargetBox="[151,1436,197,1206]" captionTargetId="figure@13.[151,1436,197,1209]" captionTargetPageId="13" captionText="FIGURES 41 – 45. 41, Drusus croaticus Marinković- Gospodnetić, 1971: pronotum, right lateral view; 42, Drusus ramae Marinković- Gospodnetić, 1971: pronotum, right lateral view; 43, Drusus medianus Marinković- Gospodnetić, 1976: pronotum, right lateral view; 44, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: pronotum, right lateral view; 45, Drusus medianus Marinković- Gospodnetić, 1976 (left), and D. radovanovici septentrionis Marinković- Gospodnetić, 1976 (right): parietal part of the head at eye level showing spinules, right lateral view." httpUri="https://zenodo.org/record/1051927/files/figure.png" pageId="13" pageNumber="14">45</figureCitation>
) compared to 
<taxonomicName id="4C066E97FFF3FFCB8DF6537C7A5D2E52" box="[739,1088,1634,1659]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="septentrionis">
<emphasis id="B972C906FFF3FFCB8DF6537C7A5D2E52" box="[739,1088,1634,1659]" italics="true" pageId="13" pageNumber="14">D. radovanovici septentrionis</emphasis>
</taxonomicName>
(
<figureCitation id="133D0991FFF3FFCB8B44537C7AB42E55" box="[1105,1193,1634,1660]" captionStart="FIGURES 41 – 45. 41" captionStartId="13.[151,269,1233,1257]" captionTargetBox="[151,1436,197,1206]" captionTargetId="figure@13.[151,1436,197,1209]" captionTargetPageId="13" captionText="FIGURES 41 – 45. 41, Drusus croaticus Marinković- Gospodnetić, 1971: pronotum, right lateral view; 42, Drusus ramae Marinković- Gospodnetić, 1971: pronotum, right lateral view; 43, Drusus medianus Marinković- Gospodnetić, 1976: pronotum, right lateral view; 44, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: pronotum, right lateral view; 45, Drusus medianus Marinković- Gospodnetić, 1976 (left), and D. radovanovici septentrionis Marinković- Gospodnetić, 1976 (right): parietal part of the head at eye level showing spinules, right lateral view." httpUri="https://zenodo.org/record/1051927/files/figure.png" pageId="13" pageNumber="14">Fig. 45</figureCitation>
). Moreover, the area covered with spinules is always compact in 
<taxonomicName id="4C066E97FFF3FFCB8DCD539A7D612EB4" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[728,892,1668,1693]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF3FFCB8DCD539A7D612EB4" box="[728,892,1668,1693]" italics="true" pageId="13" pageNumber="14">D. medianus</emphasis>
</taxonomicName>
, whereas some larvae of 
<taxonomicName id="4C066E97FFF3FFCB8BDF539A7F322EE9" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="septentrionis">
<emphasis id="B972C906FFF3FFCB8BDF539A7F322EE9" italics="true" pageId="13" pageNumber="14">D. radovanovici septentrionis</emphasis>
</taxonomicName>
had only a small number of sparsely distributed spinules (
<figureCitation id="133D0991FFF3FFCB8CCF53B97A2C2EE8" box="[986,1073,1703,1729]" captionStart="FIGURES 41 – 45. 41" captionStartId="13.[151,269,1233,1257]" captionTargetBox="[151,1436,197,1206]" captionTargetId="figure@13.[151,1436,197,1209]" captionTargetPageId="13" captionText="FIGURES 41 – 45. 41, Drusus croaticus Marinković- Gospodnetić, 1971: pronotum, right lateral view; 42, Drusus ramae Marinković- Gospodnetić, 1971: pronotum, right lateral view; 43, Drusus medianus Marinković- Gospodnetić, 1976: pronotum, right lateral view; 44, D. radovanovici septentrionis Marinković- Gospodnetić, 1976: pronotum, right lateral view; 45, Drusus medianus Marinković- Gospodnetić, 1976 (left), and D. radovanovici septentrionis Marinković- Gospodnetić, 1976 (right): parietal part of the head at eye level showing spinules, right lateral view." httpUri="https://zenodo.org/record/1051927/files/figure.png" pageId="13" pageNumber="14">Fig. 45</figureCitation>
, right).
</paragraph>
<paragraph id="8BB91514FFF3FFCB8FD353D47B812FC0" blockId="13.[151,1437,1426,2025]" pageId="13" pageNumber="14">
Besides the above mentioned characters, we have found some differences in the shape and size of the anterior-median sclerites of the metanotum that separate those 4 species into 2 distinct groups (
<figureCitation id="133D0991FFF3FFCB8A1153F17B922F20" box="[1284,1423,1775,1801]" captionStart="FIGURES 46 – 53" captionStartId="15.[151,272,1869,1893]" captionTargetBox="[151,1424,197,1846]" captionTargetId="figure@15.[151,1436,197,1846]" captionTargetPageId="15" captionText="FIGURES 46 – 53. Metanota of Drusus spp., showing anteromedian (sa 1) and posteromedian (sa 2) sclerites. 46 – 47, Drusus ramae Marinković- Gospodnetić, 1976; 48 – 49, Drusus medianus Marinković- Gospodnetić, 1971; 50 – 51, D. radovanovici septentrionis Marinković- Gospodnetić, 1976; 52 – 53, Drusus croaticus Marinković- Gospodnetić, 1971." httpUri="https://zenodo.org/record/1051929/files/figure.png" pageId="13" pageNumber="14">Figs 46–53</figureCitation>
). All 3 species from the 
<taxonomicName id="4C066E97FFF3FFCB8EB1520A7C322F04" authorityName="Klapalek" authorityYear="1899" box="[420,559,1812,1837]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bosnicus">
<emphasis id="B972C906FFF3FFCB8EB1520A7C322F04" box="[420,559,1812,1837]" italics="true" pageId="13" pageNumber="14">D. bosnicus</emphasis>
</taxonomicName>
Group have similarly shaped triangular sclerites (
<figureCitation id="133D0991FFF3FFCB8B69520A7B182F07" box="[1148,1285,1812,1838]" captionStart="FIGURES 46 – 53" captionStartId="15.[151,272,1869,1893]" captionTargetBox="[151,1424,197,1846]" captionTargetId="figure@15.[151,1436,197,1846]" captionTargetPageId="15" captionText="FIGURES 46 – 53. Metanota of Drusus spp., showing anteromedian (sa 1) and posteromedian (sa 2) sclerites. 46 – 47, Drusus ramae Marinković- Gospodnetić, 1976; 48 – 49, Drusus medianus Marinković- Gospodnetić, 1971; 50 – 51, D. radovanovici septentrionis Marinković- Gospodnetić, 1976; 52 – 53, Drusus croaticus Marinković- Gospodnetić, 1971." httpUri="https://zenodo.org/record/1051929/files/figure.png" pageId="13" pageNumber="14">Figs 46–51</figureCitation>
), as opposed to 
<taxonomicName id="4C066E97FFF3FFCB8FA152247F592F7A" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[180,324,1850,1875]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="croaticus">
<emphasis id="B972C906FFF3FFCB8FA152247F592F7A" box="[180,324,1850,1875]" italics="true" pageId="13" pageNumber="14">D. croaticus</emphasis>
</taxonomicName>
on which they are ellipsoid to quadratic (
<figureCitation id="133D0991FFF3FFCB8C3152247DB02F7D" box="[804,941,1850,1876]" captionStart="FIGURES 46 – 53" captionStartId="15.[151,272,1869,1893]" captionTargetBox="[151,1424,197,1846]" captionTargetId="figure@15.[151,1436,197,1846]" captionTargetPageId="15" captionText="FIGURES 46 – 53. Metanota of Drusus spp., showing anteromedian (sa 1) and posteromedian (sa 2) sclerites. 46 – 47, Drusus ramae Marinković- Gospodnetić, 1976; 48 – 49, Drusus medianus Marinković- Gospodnetić, 1971; 50 – 51, D. radovanovici septentrionis Marinković- Gospodnetić, 1976; 52 – 53, Drusus croaticus Marinković- Gospodnetić, 1971." httpUri="https://zenodo.org/record/1051929/files/figure.png" pageId="13" pageNumber="14">Figs 52–53</figureCitation>
). 
<taxonomicName id="4C066E97FFF3FFCB8CD552247A912F7A" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[960,1164,1850,1875]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF3FFCB8CD552247A912F7A" box="[960,1164,1850,1875]" italics="true" pageId="13" pageNumber="14">Drusus medianus</emphasis>
</taxonomicName>
has the most triangular shape of sclerites (
<figureCitation id="133D0991FFF3FFCB8E6352417C1E2F50" box="[374,515,1887,1913]" captionStart="FIGURES 46 – 53" captionStartId="15.[151,272,1869,1893]" captionTargetBox="[151,1424,197,1846]" captionTargetId="figure@15.[151,1436,197,1846]" captionTargetPageId="15" captionText="FIGURES 46 – 53. Metanota of Drusus spp., showing anteromedian (sa 1) and posteromedian (sa 2) sclerites. 46 – 47, Drusus ramae Marinković- Gospodnetić, 1976; 48 – 49, Drusus medianus Marinković- Gospodnetić, 1971; 50 – 51, D. radovanovici septentrionis Marinković- Gospodnetić, 1976; 52 – 53, Drusus croaticus Marinković- Gospodnetić, 1971." httpUri="https://zenodo.org/record/1051929/files/figure.png" pageId="13" pageNumber="14">Figs 48–49</figureCitation>
); those of 
<taxonomicName id="4C066E97FFF3FFCB8D9552417D0F2F51" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[640,786,1887,1912]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="croaticus">
<emphasis id="B972C906FFF3FFCB8D9552417D0F2F51" box="[640,786,1887,1912]" italics="true" pageId="13" pageNumber="14">D. croaticus</emphasis>
</taxonomicName>
are more ellipsoid to quadratic, with rounded anterior and posterior margins (
<figureCitation id="133D0991FFF3FFCB8EB3529A7C2D2FB7" box="[422,560,1924,1950]" captionStart="FIGURES 46 – 53" captionStartId="15.[151,272,1869,1893]" captionTargetBox="[151,1424,197,1846]" captionTargetId="figure@15.[151,1436,197,1846]" captionTargetPageId="15" captionText="FIGURES 46 – 53. Metanota of Drusus spp., showing anteromedian (sa 1) and posteromedian (sa 2) sclerites. 46 – 47, Drusus ramae Marinković- Gospodnetić, 1976; 48 – 49, Drusus medianus Marinković- Gospodnetić, 1971; 50 – 51, D. radovanovici septentrionis Marinković- Gospodnetić, 1976; 52 – 53, Drusus croaticus Marinković- Gospodnetić, 1971." httpUri="https://zenodo.org/record/1051929/files/figure.png" pageId="13" pageNumber="14">Figs 52–53</figureCitation>
). The shortest distance between 
<emphasis id="B972C906FFF3FFCB8CB8529A7DDA2FB4" box="[941,967,1924,1949]" italics="true" pageId="13" pageNumber="14">sa</emphasis>
1 sclerites was recorded in 
<taxonomicName id="4C066E97FFF3FFCB8A12529A7B812FB4" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[1287,1436,1924,1949]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF3FFCB8A12529A7B812FB4" box="[1287,1436,1924,1949]" italics="true" pageId="13" pageNumber="14">D. medianus</emphasis>
</taxonomicName>
(average = 0.08 mm) and the longest for 
<taxonomicName id="4C066E97FFF3FFCB8D9752B47CEE2FEA" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[642,755,1962,1987]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF3FFCB8D9752B47CEE2FEA" box="[642,755,1962,1987]" italics="true" pageId="13" pageNumber="14">D. ramae</emphasis>
</taxonomicName>
(average = 0.19 mm) (
<tableCitation id="C68420AFFFF3FFCB8B1352B47A482FED" box="[1030,1109,1962,1988]" captionStart="TABLE 2" captionStartId="18.[151,241,1122,1146]" captionText="TABLE 2. Measures of length (1) and width (2) of the metathoracic anteromedian sclerites (metanotal sa 1 sclerites), distance between them (3) and length of the metanotal posteromedian sclerites (metanotal sa 2 sclerites) (4) for D. ramae Marinković- Gospodnetić, 1971, D. medianus Marinković- Gospodnetić, 1976, D. radovanovici septentrionis Marinković- Gospodnetić, 1976 and D. croaticus Marinković- Gospodnetić, 1971 (all values in mm)." pageId="13" pageNumber="14">Tab. 2</tableCitation>
). The shortest width of 
<emphasis id="B972C906FFF3FFCB8A6652B47B902FEA" box="[1395,1421,1962,1987]" italics="true" pageId="13" pageNumber="14">sa</emphasis>
1 sclerites was recorded in 
<taxonomicName id="4C066E97FFF3FFCB8ED552D17C4C2FC1" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[448,593,1999,2024]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="croaticus">
<emphasis id="B972C906FFF3FFCB8ED552D17C4C2FC1" box="[448,593,1999,2024]" italics="true" pageId="13" pageNumber="14">D. croaticus</emphasis>
</taxonomicName>
(average = 0.23) and the longest in 
<taxonomicName id="4C066E97FFF3FFCB8CEF52D17A932FC1" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[1018,1166,1999,2024]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF3FFCB8CEF52D17A932FC1" box="[1018,1166,1999,2024]" italics="true" pageId="13" pageNumber="14">D. medianus</emphasis>
</taxonomicName>
(average = 0.33) (
<tableCitation id="C68420AFFFF3FFCB8A7D52D17B812FC0" box="[1384,1436,1999,2025]" captionStart="TABLE 2" captionStartId="18.[151,241,1122,1146]" captionText="TABLE 2. Measures of length (1) and width (2) of the metathoracic anteromedian sclerites (metanotal sa 1 sclerites), distance between them (3) and length of the metanotal posteromedian sclerites (metanotal sa 2 sclerites) (4) for D. ramae Marinković- Gospodnetić, 1971, D. medianus Marinković- Gospodnetić, 1976, D. radovanovici septentrionis Marinković- Gospodnetić, 1976 and D. croaticus Marinković- Gospodnetić, 1971 (all values in mm)." pageId="13" pageNumber="14">Tab.</tableCitation>
</paragraph>
<paragraph id="8BB91514FFF0FFC88F8255867A57299E" blockId="14.[151,1437,152,1895]" pageId="14" pageNumber="15">
2). 
<taxonomicName id="4C066E97FFF0FFC88FD555867F702898" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[192,365,152,177]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF0FFC88FD555867F702898" box="[192,365,152,177]" italics="true" pageId="14" pageNumber="15">Drusus ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF0FFC88EBF55867C222898" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[426,575,152,177]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="croaticus">
<emphasis id="B972C906FFF0FFC88EBF55867C222898" box="[426,575,152,177]" italics="true" pageId="14" pageNumber="15">D. croaticus</emphasis>
</taxonomicName>
showed more similarity in average width of 
<emphasis id="B972C906FFF0FFC88B7355867A9C2898" box="[1126,1153,152,177]" italics="true" pageId="14" pageNumber="15">sa</emphasis>
1 sclerites and distance between them, in contrast to 
<taxonomicName id="4C066E97FFF0FFC88EEB55A37C8728FF" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[510,666,189,214]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF0FFC88EEB55A37C8728FF" box="[510,666,189,214]" italics="true" pageId="14" pageNumber="15">D. medianus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF0FFC88DC255A37A5A28FF" box="[727,1095,189,214]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="septentrionis">
<emphasis id="B972C906FFF0FFC88DC255A37A5A28FF" box="[727,1095,189,214]" italics="true" pageId="14" pageNumber="15">D. radovanovici septentrionis</emphasis>
</taxonomicName>
(
<tableCitation id="C68420AFFFF0FFC88B4D55A37AB728FE" box="[1112,1194,189,215]" captionStart="TABLE 2" captionStartId="18.[151,241,1122,1146]" captionText="TABLE 2. Measures of length (1) and width (2) of the metathoracic anteromedian sclerites (metanotal sa 1 sclerites), distance between them (3) and length of the metanotal posteromedian sclerites (metanotal sa 2 sclerites) (4) for D. ramae Marinković- Gospodnetić, 1971, D. medianus Marinković- Gospodnetić, 1976, D. radovanovici septentrionis Marinković- Gospodnetić, 1976 and D. croaticus Marinković- Gospodnetić, 1971 (all values in mm)." pageId="14" pageNumber="15">Tab. 2</tableCitation>
). Length of the 
<emphasis id="B972C906FFF0FFC88A6755A37B9028FF" box="[1394,1421,189,214]" italics="true" pageId="14" pageNumber="15">sa</emphasis>
1 sclerites is similar in all 4 species (
<tableCitation id="C68420AFFFF0FFC88D3E55FC7C6A28D5" box="[555,631,226,252]" captionStart="TABLE 2" captionStartId="18.[151,241,1122,1146]" captionText="TABLE 2. Measures of length (1) and width (2) of the metathoracic anteromedian sclerites (metanotal sa 1 sclerites), distance between them (3) and length of the metanotal posteromedian sclerites (metanotal sa 2 sclerites) (4) for D. ramae Marinković- Gospodnetić, 1971, D. medianus Marinković- Gospodnetić, 1976, D. radovanovici septentrionis Marinković- Gospodnetić, 1976 and D. croaticus Marinković- Gospodnetić, 1971 (all values in mm)." pageId="14" pageNumber="15">Tab. 2</tableCitation>
). Posteromedian (
<emphasis id="B972C906FFF0FFC88C5955FC7D7B28D2" box="[844,870,226,251]" italics="true" pageId="14" pageNumber="15">sa</emphasis>
2) sclerites in the 4 species vary in form and size (
<figureCitation id="133D0991FFF0FFC88F8A54167F37290B" box="[159,298,264,290]" captionStart="FIGURES 46 – 53" captionStartId="15.[151,272,1869,1893]" captionTargetBox="[151,1424,197,1846]" captionTargetId="figure@15.[151,1436,197,1846]" captionTargetPageId="15" captionText="FIGURES 46 – 53. Metanota of Drusus spp., showing anteromedian (sa 1) and posteromedian (sa 2) sclerites. 46 – 47, Drusus ramae Marinković- Gospodnetić, 1976; 48 – 49, Drusus medianus Marinković- Gospodnetić, 1971; 50 – 51, D. radovanovici septentrionis Marinković- Gospodnetić, 1976; 52 – 53, Drusus croaticus Marinković- Gospodnetić, 1971." httpUri="https://zenodo.org/record/1051929/files/figure.png" pageId="14" pageNumber="15">Figs 46–53</figureCitation>
; 
<tableCitation id="C68420AFFFF0FFC88E2354167F99290B" box="[310,388,264,290]" captionStart="TABLE 2" captionStartId="18.[151,241,1122,1146]" captionText="TABLE 2. Measures of length (1) and width (2) of the metathoracic anteromedian sclerites (metanotal sa 1 sclerites), distance between them (3) and length of the metanotal posteromedian sclerites (metanotal sa 2 sclerites) (4) for D. ramae Marinković- Gospodnetić, 1971, D. medianus Marinković- Gospodnetić, 1976, D. radovanovici septentrionis Marinković- Gospodnetić, 1976 and D. croaticus Marinković- Gospodnetić, 1971 (all values in mm)." pageId="14" pageNumber="15">Tab. 2</tableCitation>
). The shape and size of 
<emphasis id="B972C906FFF0FFC88DB054167CA22908" box="[677,703,264,289]" italics="true" pageId="14" pageNumber="15">sa</emphasis>
1 sclerites are only relatively useful for diagnosis because we found variability among some specimens, especially in species from the 
<taxonomicName id="4C066E97FFF0FFC88B1954337A86296F" authorityName="Klapalek" authorityYear="1899" box="[1036,1179,301,326]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bosnicus">
<emphasis id="B972C906FFF0FFC88B1954337A86296F" box="[1036,1179,301,326]" italics="true" pageId="14" pageNumber="15">D. bosnicus</emphasis>
</taxonomicName>
Group (
<figureCitation id="133D0991FFF0FFC88A1454337B8D296E" box="[1281,1424,301,327]" captionStart="FIGURES 46 – 53" captionStartId="15.[151,272,1869,1893]" captionTargetBox="[151,1424,197,1846]" captionTargetId="figure@15.[151,1436,197,1846]" captionTargetPageId="15" captionText="FIGURES 46 – 53. Metanota of Drusus spp., showing anteromedian (sa 1) and posteromedian (sa 2) sclerites. 46 – 47, Drusus ramae Marinković- Gospodnetić, 1976; 48 – 49, Drusus medianus Marinković- Gospodnetić, 1971; 50 – 51, D. radovanovici septentrionis Marinković- Gospodnetić, 1976; 52 – 53, Drusus croaticus Marinković- Gospodnetić, 1971." httpUri="https://zenodo.org/record/1051929/files/figure.png" pageId="14" pageNumber="15">Figs 46–51</figureCitation>
). Therefore, we recommend stronger reliance on other morphological features: shape of pronotum, area covered with spinules, position of white recumbent setae on the pronotum. Aside from morphological differences, molecular data clearly distinguish larvae of these 4 species (
<bibRefCitation id="EF9768E5FFF0FFC88C4354837A23299E" author="Previsic" box="[854,1086,412,439]" pageId="14" pageNumber="15" refString="Previsic, A., Walton, C., Kucinic, M., Mitrikeski, P. T. &amp; Kerovec, M. (2009) Pleistocene divergence of Dinaric Drusus endemics (Trichoptera, Limnephilidae) in multiple microrefugia within the Balkan Peninsula. Molecular Ecology, 18 (4), 634 - 647." type="journal article" year="2009">Previšić et al. 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31C469FFFF0FFD68FD354DC7A0D2BEB" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BB91514FFF0FFC88FD354DC7A482ACB" blockId="14.[151,1437,152,1895]" pageId="14" pageNumber="15">
Subfamily 
<taxonomicName id="4C066E97FFF0FFC88E5854DC7FAA29F5" authorityName="Banks" authorityYear="1916" box="[333,439,450,476]" class="Insecta" family="Limnephilidae" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subFamily" subFamily="Drusinae">Drusinae</taxonomicName>
is divided into 8 genera: 
<taxonomicName id="4C066E97FFF0FFC88DE854DC7DC029F2" authorityName="Fischer" authorityYear="1966" box="[765,989,450,475]" class="Insecta" family="Limnephilidae" genus="Anomalopterygella" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B972C906FFF0FFC88DE854DC7DC029F2" box="[765,989,450,475]" italics="true" pageId="14" pageNumber="15">Anomalopterygella</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C066E97FFF0FFC88CE454DC7A6429F2" authorityName="McLachlan" authorityYear="1867" box="[1009,1145,450,475]" class="Insecta" family="Limnephilidae" genus="Cryptothrix" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B972C906FFF0FFC88CE454DC7A6429F2" box="[1009,1145,450,475]" italics="true" pageId="14" pageNumber="15">Cryptothrix</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C066E97FFF0FFC88B9854DC7AFF29F2" box="[1165,1250,450,475]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B972C906FFF0FFC88B9854DC7AFF29F2" box="[1165,1250,450,475]" italics="true" pageId="14" pageNumber="15">Drusus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C066E97FFF0FFC88BE054DC7B8929F2" authorityName="Kolenati" authorityYear="1848" box="[1269,1428,450,475]" class="Insecta" family="Limnephilidae" genus="Ecclisopteryx" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B972C906FFF0FFC88BE054DC7B8929F2" box="[1269,1428,450,475]" italics="true" pageId="14" pageNumber="15">Ecclisopteryx</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C066E97FFF0FFC88F8254F67F112A28" authorityName="Sipahiler" authorityYear="2002" box="[151,268,488,513]" class="Insecta" family="Limnephilidae" genus="Hadimina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B972C906FFF0FFC88F8254F67F112A28" box="[151,268,488,513]" italics="true" pageId="14" pageNumber="15">Hadimina</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C066E97FFF0FFC88E0E54F67FB12A28" box="[283,428,488,513]" class="Insecta" family="Limnephilidae" genus="Leptodrusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B972C906FFF0FFC88E0E54F67FB12A28" box="[283,428,488,513]" italics="true" pageId="14" pageNumber="15">Leptodrusus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C066E97FFF0FFC88EAF54F67C322A28" authorityName="McLachlan" authorityYear="1880" box="[442,559,488,513]" class="Insecta" family="Limnephilidae" genus="Metanoea" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B972C906FFF0FFC88EAF54F67C322A28" box="[442,559,488,513]" italics="true" pageId="14" pageNumber="15">Metanoea</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF0FFC88D7D54F67CEA2A28" authorityName="Rambur" authorityYear="1842" box="[616,759,488,513]" class="Insecta" family="Limnephilidae" genus="Monocentra" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B972C906FFF0FFC88D7D54F67CEA2A28" box="[616,759,488,513]" italics="true" pageId="14" pageNumber="15">Monocentra</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9768E5FFF0FFC88C1254F67DBA2A2B" author="Schmid" box="[775,935,488,514]" pageId="14" pageNumber="15" refString="Schmid, F. (1956) La sous-famille des Drusinae (Trichoptera, Limnephilidae). Memoires Institut Royal des Sciences Naturelles de Belgique, 2 eme Series 55, 1 - 92." type="book chapter" year="1956">Schmid 1956</bibRefCitation>
, 
<bibRefCitation id="EF9768E5FFF0FFC88CA654F67A7F2A2B" author="Sipahiler" box="[947,1122,488,514]" pageId="14" pageNumber="15" refString="Sipahiler, F. (2002) Hadimina torosensis, new genus and new species of Drusinae from southern Turkey (Trichoptera: Limnephilidae), Nova Supplementa Entomologica, 15, 239 - 248." type="journal article" year="2002">Sipahiler 2002</bibRefCitation>
; in tribe 
<taxonomicName id="4C066E97FFF0FFC88BDC54F67B3C2A2B" authorityName="N Banks" authorityYear="1916" box="[1225,1313,488,514]" class="Insecta" family="Limnephilidae" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="tribe" tribe="Drusini">Drusini</taxonomicName>
according to 
<bibRefCitation id="EF9768E5FFF0FFC88FA357137F572A0E" author="Morse" box="[182,330,525,551]" pageId="14" pageNumber="15" refString="Morse, J. C. (Ed.) (2010) Trichoptera World Checklist. http: // entweb. clemson. edu / database / trichopt / index. htm (accessed 21 February 2010)" type="journal article" year="2010">Morse 2010</bibRefCitation>
, 
<bibRefCitation id="EF9768E5FFF0FFC88E4257137C782A0E" author="Vshivkova" box="[343,613,525,551]" pageId="14" pageNumber="15" refString="Vshivkova, T. S., Morse, J. C. &amp; Ruiter, D. (2007) Phylogeny of Limnephilidae and composition of the genus Limnephilus (Limnephilidae: Limnephilinae, Limnephilini). In: Bueno-Soria, J., Barba-Alvarez, J. &amp; Armitage, B. J. (Eds.) Proceedings of the XIIth International Symposium on Trichoptera, June 18 - 22, 2006. The Caddis Press, Columbus, Ohio, pp. 309 - 319." type="book chapter" year="2007">Vshivkova et al. 2007</bibRefCitation>
). Larval morphological features typical for species belonging to this subfamily were well-described by 
<bibRefCitation id="EF9768E5FFF0FFC88D28572C7D0F2A65" box="[573,786,562,588]" pageId="14" pageNumber="15" refString="Graf, W., Lubini, V. &amp; Pauls, S. U. (2005) Larval description of Drusus muelleri McLachlan, 1868 (Trichoptera: Limnephilidae) with some notes on its ecology and systematic position within the genus Drusus. Annales de Limnologie, 41 (2), 93 - 98." type="journal article">Graf et al. (2005)</bibRefCitation>
and 
<bibRefCitation id="EF9768E5FFF0FFC88C5B572C7A672A65" author="Waringer" box="[846,1146,562,588]" pageId="14" pageNumber="15" refString="Waringer, J. &amp; Graf, W. (1997) Atlas der osterreichischen Kocherfliegenlarven. Facultas Universitatsverlag, Wien, 286 pp." type="book" year="1997">Waringer &amp; Graf (1997)</bibRefCitation>
. The following features clearly distinguish larvae of 
<taxonomicName id="4C066E97FFF0FFC88EE457467C7F2A58" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[497,610,600,625]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF0FFC88EE457467C7F2A58" box="[497,610,600,625]" italics="true" pageId="14" pageNumber="15">D. ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF0FFC88D8A57467D252A58" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[671,824,600,625]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF0FFC88D8A57467D252A58" box="[671,824,600,625]" italics="true" pageId="14" pageNumber="15">D. medianus</emphasis>
</taxonomicName>
from other described 
<taxonomicName id="4C066E97FFF0FFC88B5057467AAF2A5B" authorityName="Banks" authorityYear="1916" box="[1093,1202,600,626]" class="Insecta" family="Limnephilidae" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subFamily" subFamily="Drusinae">Drusinae</taxonomicName>
genera and species (
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, 
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, 
<bibRefCitation id="EF9768E5FFF0FFC88D9157637CDC2ABE" author="Graf" box="[644,705,637,663]" pageId="14" pageNumber="15" refString="Graf, W., Waringer, J. &amp; Pauls, S. U. (2009) A new feeding group within larval Drusinae (Trichoptera: Limnephilidae): the Drusus alpinus Group sensu Schmid, 1956, including larval descriptions of Drusus franzi Schmid, 1956, and Drusus alpinus (Meyer-Dur, 1875). Zootaxa, 2031, 53 - 62." type="journal article" year="2009">2009</bibRefCitation>
, 
<bibRefCitation id="EF9768E5FFF0FFC88DD957637D7D2ABE" author="Hickin" box="[716,864,637,663]" pageId="14" pageNumber="15" refString="Hickin, N. E. (1967) Caddis Larvae, Larvae of British Trichoptera. Hutchinson &amp; Co., London, 476 pp." type="book" year="1967">Hickin 1967</bibRefCitation>
, 
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, 
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, 
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, 
<bibRefCitation id="EF9768E5FFF0FFC88F8257BC7F2D2A95" author="Sedlak" box="[151,304,674,700]" pageId="14" pageNumber="15" refString="Sedlak, E. (1980) R. d Chrostici-Trichoptera. In: Razkosny, R. (Ed.), Kli c vodnich larev hmyzu, C eskoslovenska Akademie Ved, Praha, 163 - 225." type="book chapter" year="1980">Sedlák 1980</bibRefCitation>
, 
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, 
<bibRefCitation id="EF9768E5FFF0FFC88EEA57BC7CF62A95" author="Urbanic" box="[511,747,674,700]" pageId="14" pageNumber="15" refString="Urbanic, G., Waringer, J. &amp; Graf, W. (2003) The Larva of Ecclisopteryx asterix Malicky, 1979 (Trichoptera: Limnephilidae: Drusinae). Lauterbornia, 46, 125 - 134." type="journal article" year="2003">Urbanič et al. 2003</bibRefCitation>
, 
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, 
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, 
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, 
<bibRefCitation id="EF9768E5FFF0FFC88FF557D67FF12ACB" author="Waringer" box="[224,492,712,738]" pageId="14" pageNumber="15" refString="Waringer, J. &amp; Graf, W. (1997) Atlas der osterreichischen Kocherfliegenlarven. Facultas Universitatsverlag, Wien, 286 pp." type="book" year="1997">Waringer &amp; Graf 1997</bibRefCitation>
, 
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, 
<bibRefCitation id="EF9768E5FFF0FFC88DE057D67D5D2ACB" author="Waringer" box="[757,832,712,738]" pageId="14" pageNumber="15" refString="Waringer, J., Graf, W. &amp; Pauls, S. U. (2007 a) Functional feeding ecology in Central European species of subfamily Drusinae (Insecta: Trichoptera). Lauterbornia, 61, 3 - 8." type="journal article" year="2007" yearSuffix="a">2007a</bibRefCitation>
, 
<bibRefCitation id="EF9768E5FFF0FFC88C5E57D67D852ACB" author="Waringer" box="[843,920,712,738]" pageId="14" pageNumber="15" refString="Waringer, J., Graf, W., Pauls, S. &amp; Lubini, V. (2007 b) The Larva of Drusus nigrescens Meyer-Dur, 1875 (Trichoptera: Limnephilidae: Drusinae) with notes on its ecology, genetic differentiation and systematic position. Annales de Limnologie, 43 (3), 161 - 166." type="journal article" year="2007" yearSuffix="b">2007b</bibRefCitation>
, 
<bibRefCitation id="EF9768E5FFF0FFC88CB657D67DF32ACB" author="Waringer" box="[931,1006,712,738]" pageId="14" pageNumber="15" refString="Waringer, J., Graf, W., Pauls, S. U. &amp; Cianficconi, F. (2008 a) The larvae of Drusus improvisus McLachlan, 1884, Drusus camerinus Moretti, 1981 and Drusus aprutiensis Moretti, 1981 (Trichoptera: Limnephilidae: Drusinae). Aquatic Insects, 30 (4), 269 - 279." type="journal article" year="2008" yearSuffix="a">2008a</bibRefCitation>
, 
<bibRefCitation id="EF9768E5FFF0FFC88CEF57D67A552ACB" author="Waringer" box="[1018,1096,712,738]" pageId="14" pageNumber="15" refString="Waringer, J., Graf, W., Pauls, S. U., Vicentini, H. &amp; Lubini, V. (2008 b) DNA based association and description of the larval stage of Drusus melanchaetes McLachlan, 1876 (Trichoptera: Limnephilidae: Drusinae) with notes on ecology and zoogeography. Limnologica, 38 (1), 34 - 42." type="journal article" year="2008" yearSuffix="b">2008b</bibRefCitation>
):
</paragraph>
<paragraph id="8BB91514FFF0FFC88FD357F27AC92BB5" blockId="14.[151,1437,152,1895]" pageId="14" pageNumber="15">
<emphasis id="B972C906FFF0FFC88FD357F27ECE2B2F" bold="true" box="[198,211,748,774]" pageId="14" pageNumber="15">•</emphasis>
<taxonomicName id="4C066E97FFF0FFC88FC157F37CA62B2E" authority="McLachlan, 1867" authorityName="McLachlan" authorityYear="1867" box="[212,699,749,775]" class="Insecta" family="Limnephilidae" genus="Cryptothrix" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nebulicola">
<emphasis id="B972C906FFF0FFC88FC157F37FFC2B2F" box="[212,481,749,774]" italics="true" pageId="14" pageNumber="15">Cryptothrix nebulicola</emphasis>
McLachlan, 1867
</taxonomicName>
(distributed in central Europe, Alps); 
<taxonomicName id="4C066E97FFF0FFC88B9157F37EC02B05" authority="(Meyer-Dur, 1875)" authorityName="Meyer-Dur" authorityYear="1875" baseAuthorityName="Meyer-Dur" baseAuthorityYear="1875" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis id="B972C906FFF0FFC88B9157F37AE22B2F" box="[1156,1279,749,774]" italics="true" pageId="14" pageNumber="15">D. alpinus</emphasis>
(Meyer-Dür, 1875)
</taxonomicName>
(distributed in western Alps), 
<taxonomicName id="4C066E97FFF0FFC88D45560C7DAD2B05" authority="(Rambur, 1842)" authorityName="Rambur" authorityYear="1842" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[592,944,786,812]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="chrysotus">
<emphasis id="B972C906FFF0FFC88D45560C7C782B02" box="[592,613,786,811]" italics="true" pageId="14" pageNumber="15">D</emphasis>
. 
<emphasis id="B972C906FFF0FFC88D63560C7CF42B02" box="[630,745,786,811]" italics="true" pageId="14" pageNumber="15">chrysotus</emphasis>
(Rambur, 1842)
</taxonomicName>
(distributed in central Europe, Alps), 
<taxonomicName id="4C066E97FFF0FFC88A6A560C7EEA2B78" baseAuthorityName="Rambur" baseAuthorityYear="1842" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="B972C906FFF0FFC88A6A560C7EEA2B78" italics="true" pageId="14" pageNumber="15">D. discolor</emphasis>
</taxonomicName>
(distributed in Europe), 
<taxonomicName id="4C066E97FFF0FFC88D0356267D3A2B7B" authority="Schmid, 1956" authorityName="Schmid" authorityYear="1956" box="[534,807,824,850]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="franzi">
<emphasis id="B972C906FFF0FFC88D0356267C602B78" box="[534,637,824,849]" italics="true" pageId="14" pageNumber="15">D. franzi</emphasis>
<bibRefCitation id="EF9768E5FFF0FFC88D9156267D3A2B7B" author="Schmid" box="[644,807,824,850]" pageId="14" pageNumber="15" refString="Schmid, F. (1956) La sous-famille des Drusinae (Trichoptera, Limnephilidae). Memoires Institut Royal des Sciences Naturelles de Belgique, 2 eme Series 55, 1 - 92." type="book chapter" year="1956">Schmid, 1956</bibRefCitation>
</taxonomicName>
(distributed in eastern Alps); 
<taxonomicName id="4C066E97FFF0FFC88B9656267ECF2B5E" authority="McLachlan, 1868" authorityName="McLachlan" authorityYear="1868" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="muelleri">
<emphasis id="B972C906FFF0FFC88B9656267A852B78" box="[1155,1176,824,849]" italics="true" pageId="14" pageNumber="15">D</emphasis>
. 
<emphasis id="B972C906FFF0FFC88BB256267B142B78" box="[1191,1289,824,849]" italics="true" pageId="14" pageNumber="15">muelleri</emphasis>
McLachlan, 1868
</taxonomicName>
(distributed in Alps) and 
<taxonomicName id="4C066E97FFF0FFC88D1556437C832B5F" authorityName="Murgoci &amp; Botosaneanu" authorityYear="1953" box="[512,670,861,886]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="romanicus">
<emphasis id="B972C906FFF0FFC88D1556437C832B5F" box="[512,670,861,886]" italics="true" pageId="14" pageNumber="15">D. romanicus</emphasis>
</taxonomicName>
. (distributed in Balkan Peninsula; 
<collectingCountry id="F3115584FFF0FFC88B2056437ABF2B5E" box="[1077,1186,861,887]" name="Romania" pageId="14" pageNumber="15">Romania</collectingCountry>
, 
<collectingCountry id="F3115584FFF0FFC88BBB56437B082B5E" box="[1198,1301,861,887]" name="Bulgaria" pageId="14" pageNumber="15">Bulgaria</collectingCountry>
) have teeth along the mesal margin of each mandible; 
<taxonomicName id="4C066E97FFF0FFC88D9C569C7CE52BB2" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[649,760,898,924]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF0FFC88D9C569C7C832BB2" box="[649,670,898,923]" italics="true" pageId="14" pageNumber="15">D</emphasis>
. 
<emphasis id="B972C906FFF0FFC88DB8569C7CE52BB2" box="[685,760,898,923]" italics="true" pageId="14" pageNumber="15">ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF0FFC88C24569C7DD82BB2" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[817,965,898,923]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF0FFC88C24569C7DD82BB2" box="[817,965,898,923]" italics="true" pageId="14" pageNumber="15">D. medianus</emphasis>
</taxonomicName>
don’t have these teeth.
</paragraph>
<paragraph id="8BB91514FFF0FFC88FD056B97C572BCE" blockId="14.[151,1437,152,1895]" pageId="14" pageNumber="15">
<emphasis id="B972C906FFF0FFC88FD056B97ECF2BE8" bold="true" box="[197,210,935,961]" pageId="14" pageNumber="15">•</emphasis>
<taxonomicName id="4C066E97FFF0FFC88FC756B67F932BE8" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[210,398,936,961]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="B972C906FFF0FFC88FC756B67F932BE8" box="[210,398,936,961]" italics="true" pageId="14" pageNumber="15">Drusus discolor</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF0FFC88EDD56B67C7A2BE8" authorityName="Murgoci &amp; Botosaneanu" authorityYear="1953" box="[456,615,936,961]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="romanicus">
<emphasis id="B972C906FFF0FFC88EDD56B67C7A2BE8" box="[456,615,936,961]" italics="true" pageId="14" pageNumber="15">D. romanicus</emphasis>
</taxonomicName>
have hair-like structure on the head and pronotum; these structure are absent in 
<taxonomicName id="4C066E97FFF0FFC88E1256D37F6B2BCF" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[263,374,973,998]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF0FFC88E1256D37F6B2BCF" box="[263,374,973,998]" italics="true" pageId="14" pageNumber="15">D. ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF0FFC88EBA56D37C5E2BCF" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[431,579,973,998]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF0FFC88EBA56D37C5E2BCF" box="[431,579,973,998]" italics="true" pageId="14" pageNumber="15">D. medianus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB91514FFF0FFC88FD056EC7F8D2C55" blockId="14.[151,1437,152,1895]" pageId="14" pageNumber="15">
<emphasis id="B972C906FFF0FFC88FD056EC7ECF2C25" bold="true" box="[197,210,1010,1036]" pageId="14" pageNumber="15">•</emphasis>
<taxonomicName id="4C066E97FFF0FFC88FC756EC7C622C25" authority="McLachlan, 1876" authorityName="McLachlan" authorityYear="1876" box="[210,639,1010,1036]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B972C906FFF0FFC88FC756EC7FB82C22" box="[210,421,1010,1035]" italics="true" pageId="14" pageNumber="15">Drusus monticola</emphasis>
McLachlan, 1876
</taxonomicName>
(distributed in eastern Alps), and 
<taxonomicName id="4C066E97FFF0FFC88B0256EC7B812C25" authority="Meyer-Duer, 1875" authorityName="Meyer-Duer" authorityYear="1875" box="[1047,1436,1010,1036]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nigrescens">
<emphasis id="B972C906FFF0FFC88B0256EC7AA42C22" box="[1047,1209,1010,1035]" italics="true" pageId="14" pageNumber="15">D. nigrescens</emphasis>
Meyer-Duer, 1875
</taxonomicName>
(distributed in western Alps), have different shape of the pronotum in lateral view; these species have a ridge of dorsal humps in the last 1/3rd of the pronotum, whereas 
<taxonomicName id="4C066E97FFF0FFC88C9E51237A1F2C7F" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[907,1026,1085,1110]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF0FFC88C9E51237A1F2C7F" box="[907,1026,1085,1110]" italics="true" pageId="14" pageNumber="15">D. ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF0FFC88B5551237AFD2C7F" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[1088,1248,1085,1110]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF0FFC88B5551237AFD2C7F" box="[1088,1248,1085,1110]" italics="true" pageId="14" pageNumber="15">D. medianus</emphasis>
</taxonomicName>
have the ridge positioned anteriorly.
</paragraph>
<paragraph id="8BB91514FFF0FFC88FD051997AE02CC5" blockId="14.[151,1437,152,1895]" pageId="14" pageNumber="15">
<emphasis id="B972C906FFF0FFC88FD051997ECF2C88" bold="true" box="[197,210,1159,1185]" pageId="14" pageNumber="15">•</emphasis>
<taxonomicName id="4C066E97FFF0FFC88FC751967FB12C88" authorityName="Meyer-Duer" authorityYear="1875" box="[210,428,1160,1185]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nigrescens">
<emphasis id="B972C906FFF0FFC88FC751967FB12C88" box="[210,428,1160,1185]" italics="true" pageId="14" pageNumber="15">Drusus nigrescens</emphasis>
</taxonomicName>
has a large, median sclerotized patch on abdominal sternum I (setal bases creating a sclerotized pattern) and lateral fringe started from posterior third of III abdominal segment; 
<taxonomicName id="4C066E97FFF0FFC88BC051B37B582CEF" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[1237,1349,1197,1223]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF0FFC88BC051B37AF72CEF" box="[1237,1258,1197,1222]" italics="true" pageId="14" pageNumber="15">D</emphasis>
. 
<emphasis id="B972C906FFF0FFC88BEF51B37B582CEF" box="[1274,1349,1197,1222]" italics="true" pageId="14" pageNumber="15">ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF0FFC88A6A51B37F152CC5" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF0FFC88A6A51B37B892CEF" box="[1407,1428,1197,1222]" italics="true" pageId="14" pageNumber="15">D</emphasis>
. 
<emphasis id="B972C906FFF0FFC88F8251CD7F152CC5" box="[151,264,1235,1260]" italics="true" pageId="14" pageNumber="15">medianus</emphasis>
</taxonomicName>
don’t have this sclerotized patch and lateral fringe started from II abdominal segment.
</paragraph>
<paragraph id="8BB91514FFF0FFC88FD051E97DCF2E3E" blockId="14.[151,1437,152,1895]" pageId="14" pageNumber="15">
<emphasis id="B972C906FFF0FFC88FD051E97ECF2D38" bold="true" box="[197,210,1271,1297]" pageId="14" pageNumber="15">•</emphasis>
<taxonomicName id="4C066E97FFF0FFC88FC751E67F902D38" baseAuthorityName="Meyer-Dur" baseAuthorityYear="1875" box="[210,397,1272,1297]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis id="B972C906FFF0FFC88FC751E67F902D38" box="[210,397,1272,1297]" italics="true" pageId="14" pageNumber="15">Drusus alpinus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C066E97FFF0FFC88EB051E67D542D3B" authority="Stephens, 1837" authorityName="Stephens" authorityYear="1837" box="[421,841,1272,1298]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="annulatus">
<emphasis id="B972C906FFF0FFC88EB051E67C632D38" box="[421,638,1272,1297]" italics="true" pageId="14" pageNumber="15">Drusus annulatus</emphasis>
Stephens, 1837
</taxonomicName>
(distributed in central &amp; western Europe), 
<taxonomicName id="4C066E97FFF0FFC88A6A51E67F092D1F" baseAuthorityName="Pictet" baseAuthorityYear="1834" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="biguttatus">
<emphasis id="B972C906FFF0FFC88A6A51E67F092D1F" italics="true" pageId="14" pageNumber="15">D. biguttatus</emphasis>
</taxonomicName>
(distributed in Europe), 
<taxonomicName id="4C066E97FFF0FFC88D5E50037DBA2D1E" authority="Moretti, 1981" authorityName="Moretti" authorityYear="1981" box="[587,935,1309,1335]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="camerinus">
<emphasis id="B972C906FFF0FFC88D5E50037CEE2D1F" box="[587,755,1309,1334]" italics="true" pageId="14" pageNumber="15">D. camerinus</emphasis>
Moretti, 1981
</taxonomicName>
(distributed in Apennine Peninsula), 
<taxonomicName id="4C066E97FFF0FFC88A6A50037F1B2D75" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="croaticus">
<emphasis id="B972C906FFF0FFC88A6A50037F1B2D75" italics="true" pageId="14" pageNumber="15">D. croaticus</emphasis>
</taxonomicName>
(distributed in Dinaric Mountains), 
<taxonomicName id="4C066E97FFF0FFC88DAB505D7A352D75" authority="(Kolenati, 1848)" authorityName="Kolenati" authorityYear="1848" baseAuthorityName="Kolenati" baseAuthorityYear="1848" box="[702,1064,1346,1372]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="destitutus">
<emphasis id="B972C906FFF0FFC88DAB505D7D4A2D75" box="[702,855,1347,1372]" italics="true" pageId="14" pageNumber="15">D. destitutus</emphasis>
(Kolenati, 1848)
</taxonomicName>
(distributed in central Europe, Alps), 
<taxonomicName id="4C066E97FFF0FFC88FF250767F952DA8" authorityName="Schmid" authorityYear="1956" box="[231,392,1384,1409]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="franzi">
<emphasis id="B972C906FFF0FFC88FF250767F952DA8" box="[231,392,1384,1409]" italics="true" pageId="14" pageNumber="15">Drusus franzi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C066E97FFF0FFC88E8D50767D092DAB" authority="McLachlan, 1884" authorityName="McLachlan" authorityYear="1884" box="[408,788,1384,1410]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="improvisus">
<emphasis id="B972C906FFF0FFC88E8D50767C212DA8" box="[408,572,1384,1409]" italics="true" pageId="14" pageNumber="15">D. improvisus</emphasis>
McLachlan, 1884
</taxonomicName>
(distributed in Apennine Peninsula), 
<taxonomicName id="4C066E97FFF0FFC88BDA50767EC62D8E" authority="(Pictet, 1834)" authorityName="Pictet" authorityYear="1834" baseAuthorityName="Pictet" baseAuthorityYear="1834" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="B972C906FFF0FFC88BDA50767B5C2DA8" box="[1231,1345,1384,1409]" italics="true" pageId="14" pageNumber="15">D. mixtus</emphasis>
(Pictet, 1834)
</taxonomicName>
(distributed in Alps), 
<taxonomicName id="4C066E97FFF0FFC88EF450937D392D8E" authority="McLachlan, 1868" authorityName="McLachlan" authorityYear="1868" box="[481,804,1421,1447]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="rectus">
<emphasis id="B972C906FFF0FFC88EF450937C502D8F" box="[481,589,1421,1446]" italics="true" pageId="14" pageNumber="15">D. rectus</emphasis>
McLachlan, 1868
</taxonomicName>
(distributed in Iberian Peninsula) and 
<taxonomicName id="4C066E97FFF0FFC88BFE50937F752DE5" authority="McLachlan, 1868" authorityName="McLachlan" authorityYear="1868" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="trifidus">
<emphasis id="B972C906FFF0FFC88BFE50937B812D8F" box="[1259,1436,1421,1446]" italics="true" pageId="14" pageNumber="15">Drusus trifidus</emphasis>
McLachlan, 1868
</taxonomicName>
(distributed in central Europe) have different shape of the pronotum in lateral view; these species have more rounded posterior part of pronotum, without pronuced hump (humps); 
<taxonomicName id="4C066E97FFF0FFC88BC750C67B5E2DD8" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[1234,1347,1496,1521]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF0FFC88BC750C67B5E2DD8" box="[1234,1347,1496,1521]" italics="true" pageId="14" pageNumber="15">D. ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF0FFC88A6A50C67F152E3F" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF0FFC88A6A50C67F152E3F" italics="true" pageId="14" pageNumber="15">D. medianus</emphasis>
</taxonomicName>
have a pronuced hump (humps) in this part of the pronotum.
</paragraph>
<paragraph id="8BB91514FFF0FFC88FD3533C7C1D2E4B" blockId="14.[151,1437,152,1895]" pageId="14" pageNumber="15">
<emphasis id="B972C906FFF0FFC88FD3533C7ECE2E15" bold="true" box="[198,211,1570,1596]" pageId="14" pageNumber="15">•</emphasis>
<taxonomicName id="4C066E97FFF0FFC88FC1533D7F952E15" authorityName="McLachlan" authorityYear="1868" box="[212,392,1571,1596]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="trifidus">
<emphasis id="B972C906FFF0FFC88FC1533D7F952E15" box="[212,392,1571,1596]" italics="true" pageId="14" pageNumber="15">Drusus trifidus</emphasis>
</taxonomicName>
has numerous light spines on pronotum; 
<taxonomicName id="4C066E97FFF0FFC88C96533D7DE82E15" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[899,1013,1571,1596]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF0FFC88C96533D7DE82E15" box="[899,1013,1571,1596]" italics="true" pageId="14" pageNumber="15">D. ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF0FFC88B21533D7AD12E15" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[1076,1228,1571,1596]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF0FFC88B21533D7AD12E15" box="[1076,1228,1571,1596]" italics="true" pageId="14" pageNumber="15">D. medianus</emphasis>
</taxonomicName>
don’t have these numerous spines on pronotum.
</paragraph>
<paragraph id="8BB91514FFF0FFC88FD053727B8C2E85" blockId="14.[151,1437,152,1895]" pageId="14" pageNumber="15">
<emphasis id="B972C906FFF0FFC88FD053727ECF2EAF" bold="true" box="[197,210,1644,1670]" pageId="14" pageNumber="15">•</emphasis>
<taxonomicName id="4C066E97FFF0FFC88FC753737C462EAE" authority="Moretti, 1981" authorityName="Moretti" authorityYear="1981" box="[210,603,1645,1671]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aprutiensis">
<emphasis id="B972C906FFF0FFC88FC753737FAC2EAF" box="[210,433,1645,1670]" italics="true" pageId="14" pageNumber="15">Drusus aprutiensis</emphasis>
Moretti, 1981
</taxonomicName>
(distributed in Apennine Peninsula), has a more flattened pronotum in lateral view; 
<taxonomicName id="4C066E97FFF0FFC88E25538D7F822E85" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[304,415,1683,1708]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF0FFC88E25538D7F822E85" box="[304,415,1683,1708]" italics="true" pageId="14" pageNumber="15">D. ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF0FFC88EC2538D7C762E85" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[471,619,1683,1708]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF0FFC88EC2538D7C762E85" box="[471,619,1683,1708]" italics="true" pageId="14" pageNumber="15">D. medianus</emphasis>
</taxonomicName>
have a pronounced hump (humps) in posterior part of the pronotum.
</paragraph>
<paragraph id="8BB91514FFF0FFC88FD053A97F0C2F6B" blockId="14.[151,1437,152,1895]" pageId="14" pageNumber="15">
<emphasis id="B972C906FFF0FFC88FD053A97ECF2EF8" bold="true" box="[197,210,1719,1745]" pageId="14" pageNumber="15">•</emphasis>
<taxonomicName id="4C066E97FFF0FFC88FC753A67F9B2EF8" baseAuthorityName="Meyer-Dur" baseAuthorityYear="1875" box="[210,390,1720,1745]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis id="B972C906FFF0FFC88FC753A67F9B2EF8" box="[210,390,1720,1745]" italics="true" pageId="14" pageNumber="15">Drusus alpinus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C066E97FFF0FFC88E8353A67C232EF8" authorityName="Moretti" authorityYear="1981" box="[406,574,1720,1745]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aprutiensis">
<emphasis id="B972C906FFF0FFC88E8353A67C232EF8" box="[406,574,1720,1745]" italics="true" pageId="14" pageNumber="15">D. aprutiensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C066E97FFF0FFC88D5B53A67CF02EF8" authorityName="Moretti" authorityYear="1981" box="[590,749,1720,1745]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="camerinus">
<emphasis id="B972C906FFF0FFC88D5B53A67CF02EF8" box="[590,749,1720,1745]" italics="true" pageId="14" pageNumber="15">D. camerinus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C066E97FFF0FFC88DE853A67AF02EFB" authority="Dziedzielewicz, 1911" authorityName="Dziedzielewicz" authorityYear="1911" box="[765,1261,1720,1746]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carpathicus">
<emphasis id="B972C906FFF0FFC88DE853A67DF82EF8" box="[765,997,1720,1745]" italics="true" pageId="14" pageNumber="15">Drusus carpathicus</emphasis>
Dziedzielewicz, 1911
</taxonomicName>
(distributed in Carpathian Mountains), 
<taxonomicName id="4C066E97FFF0FFC88EA953C37C522EDF" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[444,591,1757,1782]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="croaticus">
<emphasis id="B972C906FFF0FFC88EA953C37C522EDF" box="[444,591,1757,1782]" italics="true" pageId="14" pageNumber="15">D. croaticus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C066E97FFF0FFC88D4A53C37CD72EDF" authorityName="Schmid" authorityYear="1956" box="[607,714,1757,1782]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="franzi">
<emphasis id="B972C906FFF0FFC88D4A53C37CD72EDF" box="[607,714,1757,1782]" italics="true" pageId="14" pageNumber="15">D. franzi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C066E97FFF0FFC88DCF53C37D502EDF" baseAuthorityName="Pictet" baseAuthorityYear="1834" box="[730,845,1757,1782]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="B972C906FFF0FFC88DCF53C37D502EDF" box="[730,845,1757,1782]" italics="true" pageId="14" pageNumber="15">D. mixtus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF0FFC88C9C53C37DEB2EDF" authorityName="McLachlan" authorityYear="1868" box="[905,1014,1757,1782]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="rectus">
<emphasis id="B972C906FFF0FFC88C9C53C37DEB2EDF" box="[905,1014,1757,1782]" italics="true" pageId="14" pageNumber="15">D. rectus</emphasis>
</taxonomicName>
lack prominent, long median setae dorsally on anterior border of the pronotum; 
<taxonomicName id="4C066E97FFF0FFC88DB5521D7D122F35" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[672,783,1795,1820]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF0FFC88DB5521D7D122F35" box="[672,783,1795,1820]" italics="true" pageId="14" pageNumber="15">D. ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF0FFC88C52521D7DC62F35" box="[839,987,1795,1820]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="medinaus">
<emphasis id="B972C906FFF0FFC88C52521D7DC62F35" box="[839,987,1795,1820]" italics="true" pageId="14" pageNumber="15">D. medinaus</emphasis>
</taxonomicName>
have long, median setae in this part of pronotum.
</paragraph>
<paragraph id="8BB91514FFF0FFC88FD052527B362F4E" blockId="14.[151,1437,152,1895]" box="[197,1323,1868,1895]" pageId="14" pageNumber="15">
<emphasis id="B972C906FFF0FFC88FD052527ECF2F4F" bold="true" box="[197,210,1868,1894]" pageId="14" pageNumber="15">•</emphasis>
<taxonomicName id="4C066E97FFF0FFC88FC752537FA52F4F" authorityName="Dziedzielewicz" authorityYear="1911" box="[210,440,1869,1894]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carpathicus">
<emphasis id="B972C906FFF0FFC88FC752537FA52F4F" box="[210,440,1869,1894]" italics="true" pageId="14" pageNumber="15">Drusus carpathicus</emphasis>
</taxonomicName>
lacks dorsal gills on abdomen; 
<taxonomicName id="4C066E97FFF0FFC88C3952537D872F4F" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[812,922,1869,1894]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF0FFC88C3952537D872F4F" box="[812,922,1869,1894]" italics="true" pageId="14" pageNumber="15">D. ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFF0FFC88CC652537A7A2F4F" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[979,1127,1869,1894]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF0FFC88CC652537A7A2F4F" box="[979,1127,1869,1894]" italics="true" pageId="14" pageNumber="15">D. medianus</emphasis>
</taxonomicName>
have these gills.
</paragraph>
<caption id="DF79459CFFF1FFC98F8252537B642F8D" httpUri="https://zenodo.org/record/1051929/files/figure.png" pageId="15" pageNumber="16" targetBox="[151,1424,197,1846]" targetPageId="15">
<paragraph id="8BB91514FFF1FFC98F8252537B642F8D" blockId="15.[151,1436,1869,1957]" pageId="15" pageNumber="16">
<emphasis id="B972C906FFF1FFC98F8252537F412F4C" bold="true" box="[151,348,1869,1893]" pageId="15" pageNumber="16">FIGURES 46–53</emphasis>
. Metanota of 
<taxonomicName id="4C066E97FFF1FFC98EEC52507C552F4C" box="[505,584,1870,1893]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B972C906FFF1FFC98EEC52507C552F4C" box="[505,584,1870,1893]" italics="true" pageId="15" pageNumber="16">Drusus</emphasis>
</taxonomicName>
spp., showing anteromedian (
<emphasis id="B972C906FFF1FFC98C8252507DB22F4C" box="[919,943,1870,1893]" italics="true" pageId="15" pageNumber="16">sa</emphasis>
1) and posteromedian (
<emphasis id="B972C906FFF1FFC98BA552507AD42F4C" box="[1200,1225,1870,1893]" italics="true" pageId="15" pageNumber="16">sa</emphasis>
2) sclerites. 46–47, 
<taxonomicName id="4C066E97FFF1FFC98F8252707F292FAC" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[151,308,1902,1925]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFF1FFC98F8252707F292FAC" box="[151,308,1902,1925]" italics="true" pageId="15" pageNumber="16">Drusus ramae</emphasis>
</taxonomicName>
Marinković- Gospodnetić, 1976; 48–49, 
<taxonomicName id="4C066E97FFF1FFC98DEE52707DA12FAC" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[763,956,1902,1925]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFF1FFC98DEE52707DA12FAC" box="[763,956,1902,1925]" italics="true" pageId="15" pageNumber="16">Drusus medianus</emphasis>
</taxonomicName>
Marinković- Gospodnetić, 1971; 50–51, 
<taxonomicName id="4C066E97FFF1FFC98A9752707FAE2F8C" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="septentrionis">
<emphasis id="B972C906FFF1FFC98A9752707FAE2F8C" italics="true" pageId="15" pageNumber="16">D. radovanovici septentrionis</emphasis>
</taxonomicName>
Marinković- Gospodnetić, 1976; 52–53, 
<taxonomicName id="4C066E97FFF1FFC98C7252907A002F8C" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[871,1053,1934,1957]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="croaticus">
<emphasis id="B972C906FFF1FFC98C7252907A002F8C" box="[871,1053,1934,1957]" italics="true" pageId="15" pageNumber="16">Drusus croaticus</emphasis>
</taxonomicName>
Marinković- Gospodnetić, 1971.
</paragraph>
</caption>
<paragraph id="8BB91514FFEEFFD68FD055897F28290B" blockId="16.[151,1437,151,962]" pageId="16" pageNumber="17">
<emphasis id="B972C906FFEEFFD68FD055897ECF2898" bold="true" box="[197,210,151,177]" pageId="16" pageNumber="17">•</emphasis>
<taxonomicName id="4C066E97FFEEFFD68FC755867C6A2898" box="[210,631,152,177]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="septentrionis">
<emphasis id="B972C906FFEEFFD68FC755867C6A2898" box="[210,631,152,177]" italics="true" pageId="16" pageNumber="17">Drusus radovanovici septentrionis</emphasis>
</taxonomicName>
(distributed in Balkan Peninsula) has only a few, small, white recumbent setae anteromesally on the pronotum; 
<taxonomicName id="4C066E97FFEEFFD68DFE55A37D9F28FF" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[747,898,189,214]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFEEFFD68DFE55A37D9F28FF" box="[747,898,189,214]" italics="true" pageId="16" pageNumber="17">D. medianus</emphasis>
</taxonomicName>
has numerous and longer white setae in this part of pronotum, 
<taxonomicName id="4C066E97FFEEFFD68E6455FC7FFF28D2" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[369,482,226,251]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFEEFFD68E6455FC7FFF28D2" box="[369,482,226,251]" italics="true" pageId="16" pageNumber="17">D. ramae</emphasis>
</taxonomicName>
have more produced humps in posterior part of pronotum than 
<taxonomicName id="4C066E97FFEEFFD68BCE55FC7F332908" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="septentrionis">
<emphasis id="B972C906FFEEFFD68BCE55FC7F332908" italics="true" pageId="16" pageNumber="17">D. radovanovici septentrionis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB91514FFEEFFD68FD054327D1029F5" blockId="16.[151,1437,151,962]" pageId="16" pageNumber="17">
<emphasis id="B972C906FFEEFFD68FD054327ECF296F" bold="true" box="[197,210,300,326]" pageId="16" pageNumber="17">•</emphasis>
<taxonomicName id="4C066E97FFEEFFD68FC754337CF2296E" authority="(Stein, 1874)" authorityName="Stein" authorityYear="1874" baseAuthorityName="Stein" baseAuthorityYear="1874" box="[210,751,301,327]" class="Insecta" family="Limnephilidae" genus="Anomalopterygella" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="chauviniana">
<emphasis id="B972C906FFEEFFD68FC754337C50296F" box="[210,589,301,326]" italics="true" pageId="16" pageNumber="17">Anomalopterygella chauviniana</emphasis>
(Stein, 1874)
</taxonomicName>
(distributed in western Europe), 
<taxonomicName id="4C066E97FFEEFFD68B6F54337EC22945" authority="(McLachlan, 1880)" authorityName="McLachlan" authorityYear="1880" baseAuthorityName="McLachlan" baseAuthorityYear="1880" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B972C906FFEEFFD68B6F54337AE0296F" box="[1146,1277,301,326]" italics="true" pageId="16" pageNumber="17">D. bolivari</emphasis>
(McLachlan, 1880)
</taxonomicName>
(distributed in Iberian Peninsula) and 
<taxonomicName id="4C066E97FFEEFFD68DD6544C7A392945" authority="Klapalek, 1898" authorityName="Klapalek" authorityYear="1898" box="[707,1060,338,364]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B972C906FFEEFFD68DD6544C7D402942" box="[707,861,338,363]" italics="true" pageId="16" pageNumber="17">D. brunneus</emphasis>
Klapálek, 1898
</taxonomicName>
(distributed in the Carpathian Mountains), have different morphology of posterior part of pronotum; these species have a very pronounced mid-dorsal concavity in the dorsal hump; 
<taxonomicName id="4C066E97FFEEFFD68D9654837CEF299F" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[643,754,413,438]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFEEFFD68D9654837CEF299F" box="[643,754,413,438]" italics="true" pageId="16" pageNumber="17">D. ramae</emphasis>
</taxonomicName>
has a more-pointed ridge on each hemipronotum; and the mid-dorsal concavity is nearly absent in 
<taxonomicName id="4C066E97FFEEFFD68D6754DC7D1B29F2" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[626,774,450,475]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFEEFFD68D6754DC7D1B29F2" box="[626,774,450,475]" italics="true" pageId="16" pageNumber="17">D. medianus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB91514FFEEFFD68FD354F97FE42A65" blockId="16.[151,1437,151,962]" pageId="16" pageNumber="17">
<emphasis id="B972C906FFEEFFD68FD354F97ECE2A28" bold="true" box="[198,211,487,513]" pageId="16" pageNumber="17">•</emphasis>
<taxonomicName id="4C066E97FFEEFFD68FC154F67FB82A28" baseAuthorityName="Kolenati" baseAuthorityYear="1848" box="[212,421,488,513]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="destitutus">
<emphasis id="B972C906FFEEFFD68FC154F67FB82A28" box="[212,421,488,513]" italics="true" pageId="16" pageNumber="17">Drusus destitutus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFEEFFD68EF954F67D852A2B" authority="McLachlan, 1876" authorityName="McLachlan" authorityYear="1876" box="[492,920,488,514]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="melanchaetes">
<emphasis id="B972C906FFEEFFD68EF954F67CAB2A28" box="[492,694,488,513]" italics="true" pageId="16" pageNumber="17">D. melanchaetes</emphasis>
McLachlan, 1876
</taxonomicName>
(distributed in western Alps), have setae through the whole length on dorsal side of mid- and hind-legs tibia; 
<taxonomicName id="4C066E97FFEEFFD68CC557137A5C2A0F" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[976,1089,525,550]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFEEFFD68CC557137A5C2A0F" box="[976,1089,525,550]" italics="true" pageId="16" pageNumber="17">D. ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFEEFFD68B6857137B092A0F" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[1149,1300,525,550]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFEEFFD68B6857137B092A0F" box="[1149,1300,525,550]" italics="true" pageId="16" pageNumber="17">D. medianus</emphasis>
</taxonomicName>
don’t have these continuous row of setae.
</paragraph>
<paragraph id="8BB91514FFEEFFD68FD357497FFF2ACB" blockId="16.[151,1437,151,962]" pageId="16" pageNumber="17">
<emphasis id="B972C906FFEEFFD68FD357497ECE2A58" bold="true" box="[198,211,599,625]" pageId="16" pageNumber="17">•</emphasis>
<taxonomicName id="4C066E97FFEEFFD68FC157467CAA2A5B" authority="Kolenati, 1848" authorityName="Kolenati" authorityYear="1848" box="[212,695,600,626]" class="Insecta" family="Limnephilidae" genus="Ecclisopteryx" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="dalecarlica">
<emphasis id="B972C906FFEEFFD68FC157467C1C2A58" box="[212,513,600,625]" italics="true" pageId="16" pageNumber="17">Ecclisopteryx dalecarlica</emphasis>
Kolenati, 1848
</taxonomicName>
(distributed in Europe), 
<taxonomicName id="4C066E97FFEEFFD68CCF57467B112A5B" authority="(Pictet, 1834)" authorityName="Pictet" authorityYear="1834" baseAuthorityName="Pictet" baseAuthorityYear="1834" box="[986,1292,600,626]" class="Insecta" family="Limnephilidae" genus="Ecclisopteryx" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="guttulata">
<emphasis id="B972C906FFEEFFD68CCF57467A782A58" box="[986,1125,600,625]" italics="true" pageId="16" pageNumber="17">E. guttulata</emphasis>
(Pictet, 1834)
</taxonomicName>
(distributed in Balkan Peninsula, central and western Europe) and 
<taxonomicName id="4C066E97FFEEFFD68C3057637A922ABE" authority="(McLachlan, 1867)" authorityName="McLachlan" authorityYear="1867" baseAuthorityName="McLachlan" baseAuthorityYear="1867" box="[805,1167,637,663]" class="Insecta" family="Limnephilidae" genus="Ecclisopteryx" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="madida">
<emphasis id="B972C906FFEEFFD68C3057637DBD2ABF" box="[805,928,637,662]" italics="true" pageId="16" pageNumber="17">E. madida</emphasis>
(McLachlan, 1867)
</taxonomicName>
(distributed in central Europe) have prominent spines on head and/or pronotum; 
<taxonomicName id="4C066E97FFEEFFD68C4057BC7DDA2A92" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[853,967,674,699]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFEEFFD68C4057BC7DDA2A92" box="[853,967,674,699]" italics="true" pageId="16" pageNumber="17">D. ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFEEFFD68B1757BC7A842A92" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[1026,1177,674,699]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFEEFFD68B1757BC7A842A92" box="[1026,1177,674,699]" italics="true" pageId="16" pageNumber="17">D. medianus</emphasis>
</taxonomicName>
lack these prominent head and/or pronotal spines.
</paragraph>
<paragraph id="8BB91514FFEEFFD68FD057F27A0C2B5E" blockId="16.[151,1437,151,962]" pageId="16" pageNumber="17">
<emphasis id="B972C906FFEEFFD68FD057F27ECF2B2F" bold="true" box="[197,210,748,774]" pageId="16" pageNumber="17">•</emphasis>
<taxonomicName id="4C066E97FFEEFFD68FC757F37C602B2E" authority="Malicky, 1979" authorityName="Malicky" authorityYear="1979" box="[210,637,749,775]" class="Insecta" family="Limnephilidae" genus="Ecclisopteryx" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="asterix">
<emphasis id="B972C906FFEEFFD68FC757F37FD72B2F" box="[210,458,749,774]" italics="true" pageId="16" pageNumber="17">Ecclisopteryx asterix</emphasis>
Malicky, 1979
</taxonomicName>
(distributed in southern Alps), has ovoidal metanotal anteromedian sclerites (metanotal 
<emphasis id="B972C906FFEEFFD68E9F560C7FB92B02" box="[394,420,786,811]" italics="true" pageId="16" pageNumber="17">sa</emphasis>
1 sclerites), different shape of the pronotum in lateral view and lateral fringe started from last third of III abdominal segment; 
<taxonomicName id="4C066E97FFEEFFD68DB856267D382B78" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[685,805,824,849]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFEEFFD68DB856267D382B78" box="[685,805,824,849]" italics="true" pageId="16" pageNumber="17">D. ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFEEFFD68C7156267A192B78" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[868,1028,824,849]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFEEFFD68C7156267A192B78" box="[868,1028,824,849]" italics="true" pageId="16" pageNumber="17">D. medianus</emphasis>
</taxonomicName>
have more triangular metanotal anteromedian sclerites and lateral fringe started from II abdominal segment.
</paragraph>
<paragraph id="8BB91514FFEEFFD68FD0569C7A0D2BEB" blockId="16.[151,1437,151,962]" pageId="16" pageNumber="17">
<emphasis id="B972C906FFEEFFD68FD0569C7ECF2BB5" bold="true" box="[197,210,898,924]" pageId="16" pageNumber="17">•</emphasis>
<taxonomicName id="4C066E97FFEEFFD68FC7569C7C752BB5" authority="Schmid, 1956" authorityName="Schmid" authorityYear="1956" box="[210,616,898,924]" class="Insecta" family="Limnephilidae" genus="Metanoea" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rhaetica">
<emphasis id="B972C906FFEEFFD68FC7569C7FA92BB2" box="[210,436,898,923]" italics="true" pageId="16" pageNumber="17">Metanoea rhaetica</emphasis>
<bibRefCitation id="EF9768E5FFEEFFD68EAA569C7C752BB5" author="Schmid" box="[447,616,898,924]" pageId="16" pageNumber="17" refString="Schmid, F. (1956) La sous-famille des Drusinae (Trichoptera, Limnephilidae). Memoires Institut Royal des Sciences Naturelles de Belgique, 2 eme Series 55, 1 - 92." type="book chapter" year="1956">Schmid, 1956</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C066E97FFEEFFD68DBB569C7A152BB5" authority="(Pictet, 1834)" authorityName="Pictet" authorityYear="1834" baseAuthorityName="Pictet" baseAuthorityYear="1834" box="[686,1032,898,924]" class="Insecta" family="Limnephilidae" genus="Metanoea" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="flavipennis">
<emphasis id="B972C906FFEEFFD68DBB569C7CDB2BB2" box="[686,710,898,923]" italics="true" pageId="16" pageNumber="17">M</emphasis>
. 
<emphasis id="B972C906FFEEFFD68DCC569C7D472BB2" box="[729,858,898,923]" italics="true" pageId="16" pageNumber="17">flavipennis</emphasis>
(Pictet, 1834)
</taxonomicName>
have a large, median sclerotized patch on abdominal sternum I, that is lacking in 
<taxonomicName id="4C066E97FFEEFFD68DD856B67D212BE8" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[717,828,936,962]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFEEFFD68DD856B67CFF2BE8" box="[717,738,936,961]" italics="true" pageId="16" pageNumber="17">D</emphasis>
. 
<emphasis id="B972C906FFEEFFD68DE456B67D212BE8" box="[753,828,936,961]" italics="true" pageId="16" pageNumber="17">ramae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C066E97FFEEFFD68C6056B67A142BE8" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[885,1033,936,962]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFEEFFD68C6056B67D972BE8" box="[885,906,936,961]" italics="true" pageId="16" pageNumber="17">D</emphasis>
. 
<emphasis id="B972C906FFEEFFD68C8C56B67A142BE8" box="[921,1033,936,961]" italics="true" pageId="16" pageNumber="17">medianus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</subSection>
<subSection id="E2890CFFFFEEFFD68F82510C7B862D85" pageId="16" pageNumber="17" type="multiple">
<subSubSection id="C31C469FFFEEFFD68F82510C7B862D85" pageId="16" pageNumber="17" type="key">
<paragraph id="8BB91514FFEEFFD68F82510C7DD72C05" blockId="16.[151,970,1042,1068]" box="[151,970,1042,1068]" pageId="16" pageNumber="17">
<heading id="D0F1A278FFEEFFD68F82510C7DD72C05" bold="true" box="[151,970,1042,1068]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B972C906FFEEFFD68F82510C7DD72C05" bold="true" box="[151,970,1042,1068]" pageId="16" pageNumber="17">
Identification key of known 
<taxonomicName id="4C066E97FFEEFFD68EE3510C7C532C05" ID-CoL="62R9B" box="[502,590,1042,1068]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B972C906FFEEFFD68EE3510C7C532C05" bold="true" box="[502,590,1042,1068]" italics="true" pageId="16" pageNumber="17">Drusus</emphasis>
</taxonomicName>
larvae from Balkan Peninsula
</emphasis>
</heading>
</paragraph>
<key id="FF974225FFEEFFD68F82514B7B862D85" pageId="16" pageNumber="17">
<keyStep id="30F20DB1FFEEFFD68F82514B7B862CA5" pageId="16" pageNumber="17">
<paragraph id="8BB91514FFEEFFD68F82514B7B812C45" blockId="16.[151,1436,1109,1452]" box="[151,1436,1109,1132]" pageId="16" pageNumber="17">
<keyLead id="30F7B621FFEEFFD68F82514B7B812C45" box="[151,1436,1109,1132]" pageId="16" pageNumber="17">
1 Mandibles each with subapicomesal tooth; head with felt-like hair...................................... 
<taxonomicName id="4C066E97FFEEFFD68B92514B7AE22C45" ID-CoL="37TL6" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[1159,1279,1109,1132]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="B972C906FFEEFFD68B92514B7AE22C45" box="[1159,1279,1109,1132]" italics="true" pageId="16" pageNumber="17">D. discolor</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C066E97FFEEFFD68A19514B7B812C45" ID-CoL="37TM9" authorityName="Murgoci &amp; Botosaneanu" authorityYear="1953" box="[1292,1436,1109,1132]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="romanicus">
<emphasis id="B972C906FFEEFFD68A19514B7B812C45" box="[1292,1436,1109,1132]" italics="true" pageId="16" pageNumber="17">D. romanicus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB91514FFEEFFD68F82516B7B862CA5" blockId="16.[151,1436,1109,1452]" box="[151,1435,1141,1164]" pageId="16" pageNumber="17">
<keyLead id="30F7B621FFEEFFD68F82516B7B862CA5" box="[151,1435,1141,1164]" pageId="16" pageNumber="17">- Mandibles without teeth along mesal margin ............................................................................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F20DB1FFEEFFD68F82518B7B882CC5" pageId="16" pageNumber="17">
<paragraph id="8BB91514FFEEFFD68F82518B7B882CE5" blockId="16.[151,1436,1109,1452]" pageId="16" pageNumber="17">
<keyLead id="30F7B621FFEEFFD68F82518B7B882CE5" pageId="16" pageNumber="17">
2 Anterior margin of pronotum without prominent, black median setae; anterior part of pronotum flat, posterior part curved........................................................................................................................................................ 
<taxonomicName id="4C066E97FFEEFFD68A0751AB7B882CE5" ID-CoL="37TL3" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[1298,1429,1205,1228]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="croaticus">
<emphasis id="B972C906FFEEFFD68A0751AB7B882CE5" box="[1298,1429,1205,1228]" italics="true" pageId="16" pageNumber="17">D. croaticus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB91514FFEEFFD68F8251CB7B882CC5" blockId="16.[151,1436,1109,1452]" box="[151,1429,1237,1260]" pageId="16" pageNumber="17">
<keyLead id="30F7B621FFEEFFD68F8251CB7B882CC5" box="[151,1429,1237,1260]" pageId="16" pageNumber="17">- Anterior margin of pronotum with prominent, black median setae............................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F20DB1FFEEFFD68F8251EB7B862D05" pageId="16" pageNumber="17">
<paragraph id="8BB91514FFEEFFD68F8251EB7B862D25" blockId="16.[151,1436,1109,1452]" box="[151,1435,1269,1292]" pageId="16" pageNumber="17">
<keyLead id="30F7B621FFEEFFD68F8251EB7B862D25" box="[151,1435,1269,1292]" pageId="16" pageNumber="17">3 Head with spinules........................................................................................................................................................ 4</keyLead>
</paragraph>
<paragraph id="8BB91514FFEEFFD68F82500B7B862D05" blockId="16.[151,1436,1109,1452]" box="[151,1435,1301,1324]" pageId="16" pageNumber="17">
<keyLead id="30F7B621FFEEFFD68F82500B7B862D05" box="[151,1435,1301,1324]" pageId="16" pageNumber="17">- Head without spinules................................................................................................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F20DB1FFEEFFD68F82502B7B812D45" pageId="16" pageNumber="17">
<paragraph id="8BB91514FFEEFFD68F82502B7B8B2D65" blockId="16.[151,1436,1109,1452]" box="[151,1430,1333,1356]" pageId="16" pageNumber="17">
<keyLead id="30F7B621FFEEFFD68F82502B7B8B2D65" box="[151,1430,1333,1356]" pageId="16" pageNumber="17">
4 Anterodorsal part of pronotum without numerous white, recumbent setae...................................... 
<taxonomicName id="4C066E97FFEEFFD68BC3502B7B8B2D65" ID-CoL="5GZ8L" box="[1238,1430,1333,1356]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="radovanovici" subSpecies="septentrionis">
<emphasis id="B972C906FFEEFFD68BC3502B7B8B2D65" box="[1238,1430,1333,1356]" italics="true" pageId="16" pageNumber="17">D. r. septentrionis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB91514FFEEFFD68F82504B7B812D45" blockId="16.[151,1436,1109,1452]" box="[151,1436,1365,1388]" pageId="16" pageNumber="17">
<keyLead id="30F7B621FFEEFFD68F82504B7B812D45" box="[151,1436,1365,1388]" pageId="16" pageNumber="17">
- Anterodorsal part of pronotum with numerous white, recumbent setae..................................................... 
<taxonomicName id="4C066E97FFEEFFD68A00504B7B812D45" ID-CoL="37TLT" authorityName="Marinkovic-Gospodnetic" authorityYear="1976" box="[1301,1436,1365,1388]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B972C906FFEEFFD68A00504B7B812D45" box="[1301,1436,1365,1388]" italics="true" pageId="16" pageNumber="17">D. medianus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F20DB1FFEEFFD68F82506B7B862D85" pageId="16" pageNumber="17">
<paragraph id="8BB91514FFEEFFD68F82506B7B882DA5" blockId="16.[151,1436,1109,1452]" box="[151,1429,1397,1420]" pageId="16" pageNumber="17">
<keyLead id="30F7B621FFEEFFD68F82506B7B882DA5" box="[151,1429,1397,1420]" pageId="16" pageNumber="17">
5 Posterior part of pronotum curved ............................................................................................................ 
<taxonomicName id="4C066E97FFEEFFD68A1F506B7B882DA5" ID-CoL="37TKL" baseAuthorityName="Pictet" baseAuthorityYear="1834" box="[1290,1429,1397,1420]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="biguttatus">
<emphasis id="B972C906FFEEFFD68A1F506B7B882DA5" box="[1290,1429,1397,1420]" italics="true" pageId="16" pageNumber="17">D. biguttatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB91514FFEEFFD68F82508B7B862D85" blockId="16.[151,1436,1109,1452]" box="[151,1435,1429,1452]" pageId="16" pageNumber="17">
<keyLead id="30F7B621FFEEFFD68F82508B7B862D85" box="[151,1435,1429,1452]" pageId="16" pageNumber="17">
- Posterior part of pronotum with 2 prominent acute humps ............................................................................ 
<taxonomicName id="4C066E97FFEEFFD68A22508B7B862D85" ID-CoL="37TM6" authorityName="Marinkovic-Gospodnetic" authorityYear="1971" box="[1335,1435,1429,1452]" class="Insecta" family="Limnephilidae" genus="Drusus" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ramae">
<emphasis id="B972C906FFEEFFD68A22508B7B862D85" box="[1335,1435,1429,1452]" italics="true" pageId="16" pageNumber="17">D. ramae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</subSection>
</treatment>
</document>