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<document id="5FE5424E25E9F8F091BFED78E43C1403" ID-DOI="10.5281/zenodo.176375" ID-GBIF-Dataset="4ae7ea7a-e406-4c68-9777-ec29e3521997" ID-ISSN="1175-5326" ID-Zenodo-Dep="176375" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459832640299" checkinUser="plazi" docAuthor="Williams, Jason D." docDate="2007" docId="03AF87BA6E3C4D7AFF7AF96721CEFB11" docLanguage="en" docName="zt01459p035.pdf" docOrigin="Zootaxa 1459" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Scolelepis alisonae Williams, 2007, sp. n." docType="treatment" docVersion="9" lastPageNumber="17" masterDocId="FF96FFC26E304D6AFFEDFF9D2452FFD3" masterDocTitle="New records and description of four new species of spionids (Annelida: Polychaeta: Spionidae) from the Philippines: the genera Dispio, Malacoceros, Polydora, and Scolelepis, with notes on palp ciliation patterns of the genus Scolelepis" masterLastPageNumber="35" masterPageNumber="1" pageNumber="13" updateTime="1698223090912" updateUser="plazi">
<mods:mods id="73BF989793C91F543B11CCE29DD38610" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="34C120FE8ADB6D5F9550E27FAB4EAE83">
<mods:title id="F2E394B233234A4F7C562C39BA943987">New records and description of four new species of spionids (Annelida: Polychaeta: Spionidae) from the Philippines: the genera Dispio, Malacoceros, Polydora, and Scolelepis, with notes on palp ciliation patterns of the genus Scolelepis</mods:title>
</mods:titleInfo>
<mods:name id="885B966D070739E8C599E930530D4DF1" type="personal">
<mods:role id="205F2BECBE694F9CA9C069904E93D300">
<mods:roleTerm id="4F940D49D862E729BA531CDF239B2F99">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F66CE485A49BA24EC7930C41C6159AD9">Williams, Jason D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5719CFC9204F3D88DF9C6A524BDE8BDA">text</mods:typeOfResource>
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<mods:title id="AEB37A3EBABF251E198EDC36727479A3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BBBDBDBE5D3E6493316DE7DDA69DF808">
<mods:date id="91B722A6718AB77033A8A4FE676A1BB1">2007</mods:date>
<mods:detail id="F6A9BC7588882939403B8C15B6D89B11" type="volume">
<mods:number id="8CDD1C076C957B8C7A0916A22F41F01B">1459</mods:number>
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<mods:classification id="CDDEECBED574F66962E8C928CA8CFA1A">journal article</mods:classification>
<mods:identifier id="A9B66336304421EBEE9ED753EB8C5993" type="DOI">10.5281/zenodo.176375</mods:identifier>
<mods:identifier id="12E95295D9290ECF364EA54065E14D97" type="GBIF-Dataset">4ae7ea7a-e406-4c68-9777-ec29e3521997</mods:identifier>
<mods:identifier id="EB9FC71537AA68591B7BFC0ABCD8DBC0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FF113D8C9C273769B460AD3AE0AA8A1E" type="Zenodo-Dep">176375</mods:identifier>
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<treatment id="03AF87BA6E3C4D7AFF7AF96721CEFB11" ID-DOI="http://doi.org/10.5281/zenodo.6242415" ID-GBIF-Taxon="119358169" ID-Zenodo-Dep="6242415" LSID="urn:lsid:plazi:treatment:03AF87BA6E3C4D7AFF7AF96721CEFB11" httpUri="http://treatment.plazi.org/id/03AF87BA6E3C4D7AFF7AF96721CEFB11" lastPageId="16" lastPageNumber="17" pageId="12" pageNumber="13">
<subSubSection id="C31C65276E3C4D66FF7AF9672574F8E4" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BB936AC6E3C4D66FF7AF9672597F8C7" blockId="12.[151,453,1786,1847]" box="[151,453,1786,1812]" pageId="12" pageNumber="13">
<heading id="D0F181C06E3C4D66FF7AF9672597F8C7" bold="true" box="[151,453,1786,1812]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="B972EABE6E3C4D66FF7AF9672597F8C7" bold="true" box="[151,453,1786,1812]" pageId="12" pageNumber="13">
<taxonomicName id="4C064D2F6E3C4D66FF7AF9672529F8C7" ID-CoL="4VRNK" box="[151,379,1786,1812]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="12" pageNumber="13" phylum="Annelida" rank="species" species="alisonae" status="sp. nov.">
<emphasis id="B972EABE6E3C4D66FF7AF9672529F8C7" bold="true" box="[151,379,1786,1812]" italics="true" pageId="12" pageNumber="13">Scolelepis alisonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24157C56E3C4D66FE6EF9672597F8C7" box="[387,453,1786,1812]" pageId="12" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB936AC6E3C4D66FF7AF8802574F8E4" blockId="12.[151,453,1786,1847]" box="[151,294,1821,1847]" pageId="12" pageNumber="13">
(
<figureCitation id="133D2A296E3C4D66FF72F880254FF8E4" box="[159,285,1821,1847]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStartId-0="13.[153,257,1798,1822]" captionStartId-1="14.[153,257,1432,1456]" captionStartId-2="15.[151,255,1818,1842]" captionTargetBox-1="[274,1323,190,1413]" captionTargetBox-2="[278,1318,1106,1801]" captionTargetId-1="figure@14.[148,1439,174,1427]" captionTargetId-2="figure@15.[148,1439,1095,1814]" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." captionText-1="FIGURE 9. Scolelepis alisonae sp. n. A – H, paratypes (USNM 1096806) SEM micrographs. A, anterior end, oblique lateral view (palps missing); B, anterior end, dorsal view; C, anterior end, lateral view; D, setiger 1 and base of palp, lateral view; E, tips of neurosetae from setiger 3; F, setiger 2, lateral view; G, setiger 19 &amp; 20 (left to right), lateral view (arrowheads indicate lateral organs); H, setigers 49 – 51 (top to bottom), lateral view. Scale bars: A – C = 250 µm; D = 100 µm; E – H = 200 µm." captionText-2="FIGURE 10. Scolelepis alisonae sp. n. A – D, paratypes (USNM 1096806) SEM micrographs. A, proximal region of palp, frontal view (arrowhead indicates faint division in transverse rows of cilia; short rows top, long rows bottom; note that at point of arrowhead there are two long rows below and only one short row above); B, distal tip of palp, frontal view; C, setiger 68, oblique dorsolateral view; D, posterior setigers and pygidium, dorsal view. Scale bars: A – B = 25 µm; C = 100 µm; D = 250 µm." httpUri-1="https://zenodo.org/record/176380/files/figure.png" httpUri-2="https://zenodo.org/record/176381/files/figure.png" pageId="12" pageNumber="13">Figs. 8–10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31C65276E3C4D67FF7AF8F02131FF09" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8BB936AC6E3C4D67FF7AF8F02131FF09" blockId="12.[151,1436,1901,2007]" lastBlockId="13.[151,1437,152,738]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="B972EABE6E3C4D66FF7AF8F027FCF854" bold="true" box="[151,942,1901,1927]" pageId="12" pageNumber="13">
Material examined.
<typeStatus id="54BD880E6E3C4D66FE47F8F0264EF854" box="[426,540,1901,1927]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
<collectingCountry id="F311763C6E3C4D66FDC2F8F026EFF854" box="[559,701,1901,1927]" name="Philippines" pageId="12" pageNumber="13">Philippines</collectingCountry>
, Morong, Bataan
</emphasis>
, sandy beach,
<date id="FFB8106C6E3C4D66FB92F8F02178F854" box="[1151,1322,1901,1927]" pageId="12" pageNumber="13" value="1999-02-28">28 Feb 1999</date>
(
<collectionCode id="ED17AE696E3C4D66FAA8F8F021CEF854" LSID="urn:lsid:biocol.org:col:34871" box="[1349,1436,1901,1927]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="13">USNM</collectionCode>
1096804).—
<typeStatus id="54BD880E6E3C4D66FEC7F80825F2F87C" box="[298,416,1941,1967]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
, same data as
<typeStatus id="54BD880E6E3C4D66FDA5F80826FCF87C" box="[584,686,1941,1967]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
(16 anterior ends, two posterior ends, one middle piece in alcohol,
<collectionCode id="ED17AE696E3C4D66FF26F8202572F804" LSID="urn:lsid:biocol.org:col:34871" box="[203,288,1981,2007]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="13">USNM</collectionCode>
1096805; one complete specimen on SEM stub, four anterior ends on SEM stubs, one set of palps on SEM stub,
<collectionCode id="ED17AE696E3D4D67FEADFF0525C7FF61" LSID="urn:lsid:biocol.org:col:34871" box="[320,405,152,178]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="14">USNM</collectionCode>
1096806); same location as
<typeStatus id="54BD880E6E3D4D67FD0AFF052702FF61" box="[743,848,152,178]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
,
<date id="FFB8106C6E3D4D67FCB0FF0527B5FF61" box="[861,999,152,178]" pageId="13" pageNumber="14" value="1999-03-01">1 Mar 1999</date>
(one anterior end,
<collectionCode id="ED17AE696E3D4D67FB2AFF05214EFF61" LSID="urn:lsid:biocol.org:col:34871" box="[1223,1308,152,178]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="14">USNM</collectionCode>
1096807); same location as
<typeStatus id="54BD880E6E3D4D67FEB2FF5D259BFF09" box="[351,457,192,218]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
,
<date id="FFB8106C6E3D4D67FE38FF5D2628FF09" box="[469,634,192,218]" pageId="13" pageNumber="14" value="1999-04-25">25 April 1999</date>
(one complete specimen, eight anterior ends,
<collectionCode id="ED17AE696E3D4D67FB79FF5D209FFF09" box="[1172,1229,192,218]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="13" pageNumber="14">ZRC</collectionCode>
2006.0222).
</paragraph>
</subSubSection>
<subSubSection id="C31C65276E3D4D67FF2AFF7A2533FEA9" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8BB936AC6E3D4D67FF2AFF7A2533FEA9" blockId="13.[151,1437,152,738]" pageId="13" pageNumber="14">
<emphasis id="B972EABE6E3D4D67FF2AFF7A2501FED2" bold="true" box="[199,339,231,257]" pageId="13" pageNumber="14">Etymology.</emphasis>
The species is named in honor of my wife, Dr. Alison S. Carson, for her help in collection of these specimens (sometimes under quite adverse conditions). Her cultural psychology studies in the
<collectingCountry id="F311763C6E3D4D67FAA1FE8D2487FE81" name="Philippines" pageId="13" pageNumber="14">Philippines</collectingCountry>
(supported by the National Science Foundation and a Fulbright Scholarship) allowed for travel to make these collections.
</paragraph>
</subSubSection>
<subSubSection id="C31C65276E3D4D7AFF2AFE1A2488FEA9" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8BB936AC6E3D4D67FF2AFE1A253BFD31" blockId="13.[151,1437,152,738]" pageId="13" pageNumber="14">
<emphasis id="B972EABE6E3D4D67FF2AFE1A2512FE72" bold="true" box="[199,320,391,417]" pageId="13" pageNumber="14">Diagnosis</emphasis>
. A species of
<taxonomicName id="4C064D2F6E3D4D67FE01FE152631FE72" box="[492,611,392,417]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="13" pageNumber="14" phylum="Annelida" rank="genus">
<emphasis id="B972EABE6E3D4D67FE01FE152631FE72" box="[492,611,392,417]" italics="true" pageId="13" pageNumber="14">Scolelepis</emphasis>
</taxonomicName>
with notosetae on setiger 1, notopodial hooded hooks and lacking an occipital tentacle. Palps short, palp sheaths absent, palps with two weakly separated transverse rows of cilia. Prostomium conical, with two pairs of eyes. Caruncle extending to middle of setiger 1, nuchal cilia in Ushaped pattern on sides of caruncle. Postsetal notopodial lamellae fused with branchiae from setiger 2, lamellae composed of up to nine conical lobes by setiger 5, number of lobes gradually decreasing posteriorly. Bidentate notopodial hooded hooks from setiger 68–99, with up to three per fascicle. Postsetal neuropodial lamellae with up to three lobes in anterior setigers. Bidentate neuropodial hooded hooks from setiger 25–33, with up to 11 hooks in middle and posterior fascicles. Pygidium broadly rounded, with short conical papillae surrounding anus.
</paragraph>
<paragraph id="8BB936AC6E3D4D64FEDEFCEF2125F901" blockId="13.[307,1137,882,915]" lastBlockId="14.[151,1436,1640,2026]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
A B C Pa C-G D E F G H I J K L M J-M
<emphasis id="B972EABE6E3E4D64FF2AF9F5250DF951" bold="true" box="[199,351,1640,1666]" pageId="14" pageNumber="15">Description.</emphasis>
Largest specimen of 136 setigers, 47.6-mm long and 0.9-mm wide by setiger 8;
<typeStatus id="54BD880E6E3E4D64FAF8F9F52129F951" box="[1301,1403,1640,1666]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
of 125 setigers, 37.5-mm long, 0.52-mm wide by setiger 8. Body widest anteriorly, gradually tapering to posterior end; body nearly rectangular in cross section. Color in alcohol opaque white, no pigmentation present.
</paragraph>
<caption id="DF7966246E3D4D67FF74F89B257DF83E" pageId="13" pageNumber="14">
<paragraph id="8BB936AC6E3D4D67FF74F89B257DF83E" blockId="13.[151,1436,1798,2029]" pageId="13" pageNumber="14">
<emphasis id="B972EABE6E3D4D67FF74F89B254EF8CD" bold="true" box="[153,284,1798,1822]" pageId="13" pageNumber="14">FIGURE 8.</emphasis>
<taxonomicName id="4C064D2F6E3D4D67FECEF89A25A2F8CD" box="[291,496,1799,1822]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="13" pageNumber="14" phylum="Annelida" rank="species" species="alisonae" status="sp. nov.">
<emphasis id="B972EABE6E3D4D67FECEF89A25A2F8CD" box="[291,496,1799,1822]" italics="true" pageId="13" pageNumber="14">Scolelepis alisonae</emphasis>
</taxonomicName>
<emphasis id="B972EABE6E3D4D67FE15F89B2666F8CD" bold="true" box="[504,564,1798,1822]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A24157C56E3D4D67FE15F89B2666F8CD" box="[504,564,1798,1822]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A–B, H–I, holotype (USNM 1096804); C–G, J–M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A–B, H–I = 250 µm; C–G = 100 µm; J–M = 25 µm.
</paragraph>
</caption>
<caption id="DF7966246E3E4D64FF74FA052522F9E9" httpUri="https://zenodo.org/record/176380/files/figure.png" pageId="14" pageNumber="15" targetBox="[274,1323,190,1413]" targetPageId="14">
<paragraph id="8BB936AC6E3E4D64FF74FA052522F9E9" blockId="14.[151,1436,1432,1594]" pageId="14" pageNumber="15">
<emphasis id="B972EABE6E3E4D64FF74FA05254FFA63" bold="true" box="[153,285,1432,1456]" pageId="14" pageNumber="15">FIGURE 9.</emphasis>
<taxonomicName id="4C064D2F6E3E4D64FECBFA0425A7FA63" box="[294,501,1433,1456]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="14" pageNumber="15" phylum="Annelida" rank="species" species="alisonae" status="sp. nov.">
<emphasis id="B972EABE6E3E4D64FECBFA0425A7FA63" box="[294,501,1433,1456]" italics="true" pageId="14" pageNumber="15">Scolelepis alisonae</emphasis>
</taxonomicName>
<emphasis id="B972EABE6E3E4D64FE10FA052669FA63" bold="true" box="[509,571,1432,1456]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A24157C56E3E4D64FE10FA052669FA63" box="[509,571,1432,1456]" pageId="14" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A–H, paratypes (USNM 1096806) SEM micrographs. A, anterior end, oblique lateral view (palps missing); B, anterior end, dorsal view; C, anterior end, lateral view; D, setiger 1 and base of palp, lateral view; E, tips of neurosetae from setiger 3; F, setiger 2, lateral view; G, setiger 19 &amp; 20 (left to right), lateral view (arrowheads indicate lateral organs); H, setigers 49–51 (top to bottom), lateral view. Scale bars: A–C = 250 µm; D = 100 µm; E–H = 200 µm.
</paragraph>
</caption>
<paragraph id="8BB936AC6E3E4D65FF2AF97D269BFED1" blockId="14.[151,1436,1640,2026]" lastBlockId="15.[151,1436,152,1058]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
Prostomium conical, extending anteriorly to long, tapering point, posteriorly extending to short caruncle with rounded posterior margin, ending in middle of setiger 1, caruncle not free on posterior margin (
<figureCitation id="133D2A296E3E4D64FAC5F8952124F8F1" box="[1320,1398,1800,1826]" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="14" pageNumber="15">Figs. 8</figureCitation>
A– B, 9A–C). Two pairs of eyes, one pair of kidney-shaped eyes laterally, one pair of round eyes medially, all eyes in nearly straight row between base of palps (
<figureCitation id="133D2A296E3E4D64FD0EF8C5277BF8A1" box="[739,809,1880,1906]" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="14" pageNumber="15">Fig. 8</figureCitation>
B); occipital tentacle absent. Palps short, extending to setigers 6–10 (
<figureCitation id="133D2A296E3E4D64FEA5F81D25CEF849" box="[328,412,1920,1946]" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="14" pageNumber="15">Figs. 8</figureCitation>
B, 9C), with two weakly separated transverse rows of cilia along the ventral surface, long rows of cilia approximately 45 µm long, short rows approximately 24 µm long (short rows on medial side), long and short rows in approximately 1:1 ratio, rows of mucus secreting cells represented by tubular necks present proximal to transverse ciliary rows; ciliary rows extending to distal ends of palps, median ciliated groove lacking (
<figureCitation id="133D2A296E3F4D65FE61FF5D25B9FF09" box="[396,491,192,218]" captionStart="FIGURE 10" captionStartId="15.[151,255,1818,1842]" captionTargetBox="[278,1318,1106,1801]" captionTargetId="figure@15.[148,1439,1095,1814]" captionTargetPageId="15" captionText="FIGURE 10. Scolelepis alisonae sp. n. A – D, paratypes (USNM 1096806) SEM micrographs. A, proximal region of palp, frontal view (arrowhead indicates faint division in transverse rows of cilia; short rows top, long rows bottom; note that at point of arrowhead there are two long rows below and only one short row above); B, distal tip of palp, frontal view; C, setiger 68, oblique dorsolateral view; D, posterior setigers and pygidium, dorsal view. Scale bars: A – B = 25 µm; C = 100 µm; D = 250 µm." httpUri="https://zenodo.org/record/176381/files/figure.png" pageId="15" pageNumber="16">Figs. 10</figureCitation>
A–B, 18C). Palpal sheaths absent. Nuchal cilia in U-shaped pattern on both sides of caruncle, posterior to base of palps (
<figureCitation id="133D2A296E3F4D65FDB2FF7526F7FED1" box="[607,677,232,258]" captionStart="FIGURE 9" captionStartId="14.[153,257,1432,1456]" captionTargetBox="[274,1323,190,1413]" captionTargetId="figure@14.[148,1439,174,1427]" captionTargetPageId="14" captionText="FIGURE 9. Scolelepis alisonae sp. n. A – H, paratypes (USNM 1096806) SEM micrographs. A, anterior end, oblique lateral view (palps missing); B, anterior end, dorsal view; C, anterior end, lateral view; D, setiger 1 and base of palp, lateral view; E, tips of neurosetae from setiger 3; F, setiger 2, lateral view; G, setiger 19 &amp; 20 (left to right), lateral view (arrowheads indicate lateral organs); H, setigers 49 – 51 (top to bottom), lateral view. Scale bars: A – C = 250 µm; D = 100 µm; E – H = 200 µm." httpUri="https://zenodo.org/record/176380/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
B).
</paragraph>
<paragraph id="8BB936AC6E3F4D65FF2AFE8D21DEFD31" blockId="15.[151,1436,152,1058]" pageId="15" pageNumber="16">
Setiger 1 well developed with bluntly pointed, triangular notopodial and neuropodial postsetal lamellae, notosetae and neurosetae present (
<figureCitation id="133D2A296E3F4D65FDCAFEA5262AFE81" box="[551,632,312,338]" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="15" pageNumber="16">Figs. 8</figureCitation>
C, 9D). Postsetal notopodial lamellae fused with branchiae from setiger 2 (
<figureCitation id="133D2A296E3F4D65FF01FEFD256FFEA9" box="[236,317,352,378]" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="15" pageNumber="16">Figs. 8</figureCitation>
D, 9F), lamella composed of 2–6 conical lobes, number of lobes increasing to 5–9 on lamellae by setiger 5 (
<figureCitation id="133D2A296E3F4D65FEDCFE1525D0FE71" box="[305,386,392,418]" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="15" pageNumber="16">Figs. 8</figureCitation>
E, 9A, C), then gradually decreasing to two conical lobes distally and rounded basal portion by setigers 20–30 (
<figureCitation id="133D2A296E3F4D65FE44FE2D25AAFE19" box="[425,504,432,458]" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="15" pageNumber="16">Figs. 8</figureCitation>
F, 9G), posteriormost setigers with one conical lobe distally, with rounded basal portion free from branchiae (
<figureCitation id="133D2A296E3F4D65FE06FE452668FE21" box="[491,570,472,498]" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="15" pageNumber="16">Figs. 8</figureCitation>
G, 9H, 10C). Neuropodial postsetal lamellae of setiger 1 rounded or conical (
<figureCitation id="133D2A296E3F4D65FF26FD9D254FFDC9" box="[203,285,512,538]" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="15" pageNumber="16">Figs. 8</figureCitation>
C, 9D), lamellae of setiger 2 with rounded base extending into short, bluntly pointed lobe (
<figureCitation id="133D2A296E3F4D65FA8CFD9D24F7FD91" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="15" pageNumber="16">Figs. 8</figureCitation>
D, 9F), lamellae of setigers 3–5 sometimes with 2–3 bluntly pointed lobes (
<figureCitation id="133D2A296E3F4D65FBCEFDB52038FD91" box="[1059,1130,552,578]" captionStart="FIGURE 9" captionStartId="14.[153,257,1432,1456]" captionTargetBox="[274,1323,190,1413]" captionTargetId="figure@14.[148,1439,174,1427]" captionTargetPageId="14" captionText="FIGURE 9. Scolelepis alisonae sp. n. A – H, paratypes (USNM 1096806) SEM micrographs. A, anterior end, oblique lateral view (palps missing); B, anterior end, dorsal view; C, anterior end, lateral view; D, setiger 1 and base of palp, lateral view; E, tips of neurosetae from setiger 3; F, setiger 2, lateral view; G, setiger 19 &amp; 20 (left to right), lateral view (arrowheads indicate lateral organs); H, setigers 49 – 51 (top to bottom), lateral view. Scale bars: A – C = 250 µm; D = 100 µm; E – H = 200 µm." httpUri="https://zenodo.org/record/176380/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
C), lamellae from setigers 6-20 with broadly rounded lobe (
<figureCitation id="133D2A296E3F4D65FDF3FDCD2630FDB9" box="[542,610,592,618]" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="15" pageNumber="16">Fig. 8</figureCitation>
F, 9G), developing notch dividing lamellae by setiger 30, on posterior setigers ventral lobe small, triangular, dorsal lobe elongate with dorsal end free and pointed, ventral end rounded (
<figureCitation id="133D2A296E3F4D65FEE6FD3D250DFD69" box="[267,351,672,698]" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="15" pageNumber="16">
Figs.
<date id="FFB8106C6E3F4D65FEBDFD3D250DFD69" box="[336,351,672,698]" pageId="15" pageNumber="16">8</date>
</figureCitation>
I, 9H), by setiger 65 ventral lobe reduced, dorsal lobe digitate (
<figureCitation id="133D2A296E3F4D65FB84FD3D2090FD69" box="[1129,1218,672,698]" captionStart="FIGURE 10" captionStartId="15.[151,255,1818,1842]" captionTargetBox="[278,1318,1106,1801]" captionTargetId="figure@15.[148,1439,1095,1814]" captionTargetPageId="15" captionText="FIGURE 10. Scolelepis alisonae sp. n. A – D, paratypes (USNM 1096806) SEM micrographs. A, proximal region of palp, frontal view (arrowhead indicates faint division in transverse rows of cilia; short rows top, long rows bottom; note that at point of arrowhead there are two long rows below and only one short row above); B, distal tip of palp, frontal view; C, setiger 68, oblique dorsolateral view; D, posterior setigers and pygidium, dorsal view. Scale bars: A – B = 25 µm; C = 100 µm; D = 250 µm." httpUri="https://zenodo.org/record/176381/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
C). Lateral organs between notopodial and neuropodial postsetal lamellae present from setiger 1 to posterior setigers (
<figureCitation id="133D2A296E3F4D65FACDFD552134FD31" box="[1312,1382,712,738]" captionStart="FIGURE 9" captionStartId="14.[153,257,1432,1456]" captionTargetBox="[274,1323,190,1413]" captionTargetId="figure@14.[148,1439,174,1427]" captionTargetPageId="14" captionText="FIGURE 9. Scolelepis alisonae sp. n. A – H, paratypes (USNM 1096806) SEM micrographs. A, anterior end, oblique lateral view (palps missing); B, anterior end, dorsal view; C, anterior end, lateral view; D, setiger 1 and base of palp, lateral view; E, tips of neurosetae from setiger 3; F, setiger 2, lateral view; G, setiger 19 &amp; 20 (left to right), lateral view (arrowheads indicate lateral organs); H, setigers 49 – 51 (top to bottom), lateral view. Scale bars: A – C = 250 µm; D = 100 µm; E – H = 200 µm." httpUri="https://zenodo.org/record/176380/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
G).
</paragraph>
<paragraph id="8BB936AC6E3F4D65FF2AFD6D2028FBF1" blockId="15.[151,1436,152,1058]" pageId="15" pageNumber="16">
Notosetae of setiger 1 and subsequent setigers arranged in two vertical rows of bilimbate capillaries, the dorsal notosetae of these rows longer than ventral ones, often loosely coiled along their distal ends (
<figureCitation id="133D2A296E3F4D65FADCFC8521D0FCE1" box="[1329,1410,792,818]" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="15" pageNumber="16">Figs. 8</figureCitation>
L, 9A, C, F), one notopodial hooded hook beginning on setiger 68–99 (
<figureCitation id="133D2A296E3F4D65FC2CFCDD2041FC89" box="[961,1043,832,858]" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="15" pageNumber="16">Figs. 8</figureCitation>
J, K, 10C), with up to three notopodial hooded hooks in posterior most setigers; notopodial hooded hooks with acute main fang and single accessory tooth. Neurosetae of setiger 1 and subsequent setigers arranged in two vertical rows of bilimbate capillaries (
<figureCitation id="133D2A296E3F4D65FECCFC25253AFC01" box="[289,360,952,978]" captionStart="FIGURE 9" captionStartId="14.[153,257,1432,1456]" captionTargetBox="[274,1323,190,1413]" captionTargetId="figure@14.[148,1439,174,1427]" captionTargetPageId="14" captionText="FIGURE 9. Scolelepis alisonae sp. n. A – H, paratypes (USNM 1096806) SEM micrographs. A, anterior end, oblique lateral view (palps missing); B, anterior end, dorsal view; C, anterior end, lateral view; D, setiger 1 and base of palp, lateral view; E, tips of neurosetae from setiger 3; F, setiger 2, lateral view; G, setiger 19 &amp; 20 (left to right), lateral view (arrowheads indicate lateral organs); H, setigers 49 – 51 (top to bottom), lateral view. Scale bars: A – C = 250 µm; D = 100 µm; E – H = 200 µm." httpUri="https://zenodo.org/record/176380/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
F–G), 1–3 neuropodial hooded hooks from setiger 25–33, up to 5–11 neuropodial hooded hooks in middle to posterior setigers (
<figureCitation id="133D2A296E3F4D65FDB2FC7D26F4FC29" box="[607,678,992,1018]" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="15" pageNumber="16">Fig. 8</figureCitation>
G); neuropodial hooded hooks with acute main fang and single accessory tooth (
<figureCitation id="133D2A296E3F4D65FEB1FB9525F3FBF1" box="[348,417,1032,1058]" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="15" pageNumber="16">Fig. 8</figureCitation>
M). Notosetae and neurosetae with minute spinelets (
<figureCitation id="133D2A296E3F4D65FBFFFB952005FBF1" box="[1042,1111,1032,1058]" captionStart="FIGURE 9" captionStartId="14.[153,257,1432,1456]" captionTargetBox="[274,1323,190,1413]" captionTargetId="figure@14.[148,1439,174,1427]" captionTargetPageId="14" captionText="FIGURE 9. Scolelepis alisonae sp. n. A – H, paratypes (USNM 1096806) SEM micrographs. A, anterior end, oblique lateral view (palps missing); B, anterior end, dorsal view; C, anterior end, lateral view; D, setiger 1 and base of palp, lateral view; E, tips of neurosetae from setiger 3; F, setiger 2, lateral view; G, setiger 19 &amp; 20 (left to right), lateral view (arrowheads indicate lateral organs); H, setigers 49 – 51 (top to bottom), lateral view. Scale bars: A – C = 250 µm; D = 100 µm; E – H = 200 µm." httpUri="https://zenodo.org/record/176380/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
E).
</paragraph>
<caption id="DF7966246E3F4D65FF7AF88725FDF86F" httpUri="https://zenodo.org/record/176381/files/figure.png" pageId="15" pageNumber="16" targetBox="[278,1318,1106,1801]" targetPageId="15">
<paragraph id="8BB936AC6E3F4D65FF7AF88725FDF86F" blockId="15.[151,1436,1818,1980]" pageId="15" pageNumber="16">
<emphasis id="B972EABE6E3F4D65FF7AF887257BF8E1" bold="true" box="[151,297,1818,1842]" pageId="15" pageNumber="16">FIGURE 10.</emphasis>
<taxonomicName id="4C064D2F6E3F4D65FEDEF8872650F8E2" box="[307,514,1818,1841]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="15" pageNumber="16" phylum="Annelida" rank="species" species="alisonae" status="sp. nov.">
<emphasis id="B972EABE6E3F4D65FEDEF8872650F8E2" box="[307,514,1818,1841]" italics="true" pageId="15" pageNumber="16">Scolelepis alisonae</emphasis>
</taxonomicName>
<emphasis id="B972EABE6E3F4D65FDE1F8872618F8E1" bold="true" box="[524,586,1818,1842]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="A24157C56E3F4D65FDE1F8872618F8E1" box="[524,586,1818,1842]" pageId="15" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A–D, paratypes (USNM 1096806) SEM micrographs. A, proximal region of palp, frontal view (arrowhead indicates faint division in transverse rows of cilia; short rows top, long rows bottom; note that at point of arrowhead there are two long rows below and only one short row above); B, distal tip of palp, frontal view; C, setiger 68, oblique dorsolateral view; D, posterior setigers and pygidium, dorsal view. Scale bars: A–B = 25 µm; C = 100 µm; D = 250 µm.
</paragraph>
</caption>
<paragraph id="8BB936AC6E204D7AFF2AFF052513FEF9" blockId="16.[151,1437,152,1218]" pageId="16" pageNumber="17">
Branchiae present from setiger 2 to end of body, fused with postsetal notopodial lamellae but with tips free in anterior setigers (
<figureCitation id="133D2A296E204D7AFE54FF5D2652FF09" box="[441,512,192,218]" captionStart="FIGURE 9" captionStartId="14.[153,257,1432,1456]" captionTargetBox="[274,1323,190,1413]" captionTargetId="figure@14.[148,1439,174,1427]" captionTargetPageId="14" captionText="FIGURE 9. Scolelepis alisonae sp. n. A – H, paratypes (USNM 1096806) SEM micrographs. A, anterior end, oblique lateral view (palps missing); B, anterior end, dorsal view; C, anterior end, lateral view; D, setiger 1 and base of palp, lateral view; E, tips of neurosetae from setiger 3; F, setiger 2, lateral view; G, setiger 19 &amp; 20 (left to right), lateral view (arrowheads indicate lateral organs); H, setigers 49 – 51 (top to bottom), lateral view. Scale bars: A – C = 250 µm; D = 100 µm; E – H = 200 µm." httpUri="https://zenodo.org/record/176380/files/figure.png" pageId="16" pageNumber="17">Fig. 9</figureCitation>
F), separated from notopodial lamellae in posterior setigers, with band of cilia along inner edge and joined to corresponding branchiae on opposite side by a single band of cilia across dorsum (
<figureCitation id="133D2A296E204D7AFF35FE8D254CFEF9" box="[216,286,272,298]" captionStart="FIGURE 9" captionStartId="14.[153,257,1432,1456]" captionTargetBox="[274,1323,190,1413]" captionTargetId="figure@14.[148,1439,174,1427]" captionTargetPageId="14" captionText="FIGURE 9. Scolelepis alisonae sp. n. A – H, paratypes (USNM 1096806) SEM micrographs. A, anterior end, oblique lateral view (palps missing); B, anterior end, dorsal view; C, anterior end, lateral view; D, setiger 1 and base of palp, lateral view; E, tips of neurosetae from setiger 3; F, setiger 2, lateral view; G, setiger 19 &amp; 20 (left to right), lateral view (arrowheads indicate lateral organs); H, setigers 49 – 51 (top to bottom), lateral view. Scale bars: A – C = 250 µm; D = 100 µm; E – H = 200 µm." httpUri="https://zenodo.org/record/176380/files/figure.png" pageId="16" pageNumber="17">Fig. 9</figureCitation>
B).
</paragraph>
<paragraph id="8BB936AC6E204D7AFF2AFEA52488FEA9" blockId="16.[151,1437,152,1218]" pageId="16" pageNumber="17">
Pygidium broadly rounded, with short conical papillae surrounding dorsal anal opening (
<figureCitation id="133D2A296E204D7AFAFCFEA52137FE81" box="[1297,1381,312,338]" captionStart="FIGURE 8" captionStartId="13.[153,257,1798,1822]" captionText="FIGURE 8. Scolelepis alisonae sp. n. A – B, H – I, holotype (USNM 1096804); C – G, J – M, paratypes (USNM 1096805). A, anterior end, lateral view; B, anterior end, dorsal view; C, setiger 1, lateral view (Pa indicates base of palp); D, setiger 2, anterior lateral view; E, setiger 4, anterior lateral view; F, setiger 10, anterior lateral view; G, setiger 89, anterior lateral view; H, posterior setigers and pygidium, dorsal view; I, posterior setigers and pygidium, lateral view; J, notopodial hooded hook from setiger 89, lateral view; K, notopodial hooded hook from setiger 89, frontal view; L, notoseta from setiger 89; M, neuropodial hooded hook from setiger 89, lateral view. Scale bars: A – B, H – I = 250 µm; C – G = 100 µm; J – M = 25 µm." pageId="16" pageNumber="17">Figs. 8</figureCitation>
H–I, 10D).
</paragraph>
</subSubSection>
<subSubSection id="C31C65276E204D7AFF2AFE1A27A6FD69" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BB936AC6E204D7AFF2AFE1A27A6FD69" blockId="16.[151,1437,152,1218]" pageId="16" pageNumber="17">
<emphasis id="B972EABE6E204D7AFF2AFE1A2512FE72" bold="true" box="[199,320,391,417]" pageId="16" pageNumber="17">Remarks.</emphasis>
The genus
<taxonomicName id="4C064D2F6E204D7AFE24FE152612FE72" box="[457,576,392,417]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="genus">
<emphasis id="B972EABE6E204D7AFE24FE152612FE72" box="[457,576,392,417]" italics="true" pageId="16" pageNumber="17">Scolelepis</emphasis>
</taxonomicName>
was reviewed by
<bibRefCitation id="EF974B5D6E204D7AFCF9FE15278BFE71" author="Maciolek" box="[788,985,392,418]" pageId="16" pageNumber="33" refString="Maciolek, N. J. (1987) New species and records of Scolelepis (Polychaeta: Spionidae) from the east coast of North America, with a revision of the subgenera. Bulletin of the Biological Society of Washington, No. 7, 15 - 40." type="book chapter" year="1987">Maciolek (1987)</bibRefCitation>
and more recently by Hutchings
<emphasis id="B972EABE6E204D7AFA8CFE1521CEFE72" box="[1377,1436,392,417]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
(1998), both studies provided summary tables for the known members of the genus. Since then, nine species of
<taxonomicName id="4C064D2F6E204D7AFF57FE452563FE22" box="[186,305,472,497]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="genus">
<emphasis id="B972EABE6E204D7AFF57FE452563FE22" box="[186,305,472,497]" italics="true" pageId="16" pageNumber="17">Scolelepis</emphasis>
</taxonomicName>
, including those treated in this paper, have been described, and there are now 58–59 species within this genus (
<tableCitation id="C68403176E204D7AFE97FD9D2584FDC9" box="[378,470,512,538]" captionStart="TABLE 1" captionStartId="17.[124,212,771,794]" captionText="TABLE 1. Taxonomic characters of species in the genus Scolelepis described since Hutchings et al. (1998)." pageId="16" pageNumber="17">Table 1</tableCitation>
). Confusion exists in the total number of species because
<bibRefCitation id="EF974B5D6E204D7AFB45FD9D24B7FD91" author="MacCord" pageId="16" pageNumber="33" refString="MacCord, F. S. &amp; Amaral, A. C. Z. (2005) Morphometric analysis of two species of Scolelepis (Polychaeta: Spionidae). Journal of the Marine Biological Association of the United Kingdom, 84, 769 - 784." type="journal article" year="2005">MacCord &amp; Amaral (2005)</bibRefCitation>
resurrected
<taxonomicName id="4C064D2F6E204D7AFE99FDB52747FD91" authority="Jones, 1962" authorityName="Jones" authorityYear="1962" box="[372,789,552,578]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="goodbodyi">
<emphasis id="B972EABE6E204D7AFE99FDB5263CFD92" box="[372,622,552,577]" italics="true" pageId="16" pageNumber="17">Scolelepis goodbodyi</emphasis>
(
<bibRefCitation id="EF974B5D6E204D7AFD90FDB5275EFD91" author="Jones" box="[637,780,552,578]" pageId="16" pageNumber="33" refString="Jones, M. L. (1962) On some polychaetous annelids from Jamaica. Bulletin of the America Museum of Natural History, 124, 173 - 212." type="journal article" year="1962">Jones, 1962</bibRefCitation>
)
</taxonomicName>
, a species that was previously considered a synonym of
<taxonomicName id="4C064D2F6E204D7AFF7AFDCD2589FDB9" authority="Muller, 1806" authorityName="Muller" authorityYear="1806" box="[151,475,592,618]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="squamata">
<emphasis id="B972EABE6E204D7AFF7AFDCD2575FDBA" box="[151,295,592,617]" italics="true" pageId="16" pageNumber="17">S. squamata</emphasis>
(
<bibRefCitation id="EF974B5D6E204D7AFEDAFDCD2580FDB9" author="Muller" box="[311,466,592,618]" pageId="16" pageNumber="34" refString="Muller, O. F. (1806) Zoologica Danica seu Animalium Daniae et Norvegiae ruriorum ac minus notorum, Descriptiones et Historia, Havniaem, 160 pp." type="book" year="1806">Müller, 1806</bibRefCitation>
)
</taxonomicName>
(
<bibRefCitation id="EF974B5D6E204D7AFE06FDCD26F3FDB9" author="Pettibone" box="[491,673,592,618]" pageId="16" pageNumber="34" refString="Pettibone, M. H. (1963) Revision of some genera of polychaete worms of the family Spionidae, including the description of a new species of Scolelepis. Proceedings of the Biological Society of Washington, 76, 89 - 104." type="journal article" year="1963">Pettibone 1963</bibRefCitation>
;
<bibRefCitation id="EF974B5D6E204D7AFD43FDCD2718FDB9" author="Foster" box="[686,842,592,618]" pageId="16" pageNumber="32" refString="Foster, N. M. (1971 a) Spionidae (Polychaeta) of the Gulf of Mexico and the Caribbean Sea. Studies on the Fauna of Curacao and other Caribbean Islands, 36, 1 - 183." type="journal article" year="1971" yearSuffix="a">Foster 1971a</bibRefCitation>
;
<bibRefCitation id="EF974B5D6E204D7AFCBAFDCD278EFDB9" author="Light" box="[855,988,592,618]" pageId="16" pageNumber="33" refString="Light, W. J. (1978) Invertebrates of the San Francisco Bay Estuary System. Family Spionidae (Annelida: Polychaeta), The Boxwood Press, Pacific Grove, California, 211 pp." type="book" year="1978">Light 1978</bibRefCitation>
;
<bibRefCitation id="EF974B5D6E204D7AFC04FDCD20CDFDB9" author="Maciolek" box="[1001,1183,592,618]" pageId="16" pageNumber="33" refString="Maciolek, N. J. (1987) New species and records of Scolelepis (Polychaeta: Spionidae) from the east coast of North America, with a revision of the subgenera. Bulletin of the Biological Society of Washington, No. 7, 15 - 40." type="book chapter" year="1987">Maciolek 1987</bibRefCitation>
). However,
<bibRefCitation id="EF974B5D6E204D7AFAC0FDCD253AFD41" author="MacCord" pageId="16" pageNumber="33" refString="MacCord, F. S. &amp; Amaral, A. C. Z. (2005) Morphometric analysis of two species of Scolelepis (Polychaeta: Spionidae). Journal of the Marine Biological Association of the United Kingdom, 84, 769 - 784." type="journal article" year="2005">MacCord &amp; Amaral (2005)</bibRefCitation>
provided no taxonomic evidence to support the conclusion that these species are distinct (their identification was based on the authority of Radashevsky; pg. 829).
</paragraph>
</subSubSection>
<subSubSection id="C31C65276E204D7AFF2AFD552071FB49" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8BB936AC6E204D7AFF2AFD5527D4FBF1" blockId="16.[151,1437,152,1218]" pageId="16" pageNumber="17">
<taxonomicName id="4C064D2F6E204D7AFF2AFD5525FAFD32" ID-CoL="4VRNK" box="[199,424,712,737]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="alisonae" status="sp. nov.">
<emphasis id="B972EABE6E204D7AFF2AFD5525FAFD32" box="[199,424,712,737]" italics="true" pageId="16" pageNumber="17">Scolelepis alisonae</emphasis>
</taxonomicName>
<emphasis id="B972EABE6E204D7AFE5DFD5A25A0FD32" bold="true" box="[432,498,711,737]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A24157C56E204D7AFE5DFD5A25A0FD32" box="[432,498,711,737]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
belongs to a group of six species including
<taxonomicName id="4C064D2F6E204D7AFC1BFD5521C5FD31" ID-CoL="4VRML" authority="Hartmann-Schroder, 1980" authorityName="Hartmann-Schroder" authorityYear="1980" box="[1014,1431,712,738]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="blakei">
<emphasis id="B972EABE6E204D7AFC1BFD55200EFD32" box="[1014,1116,712,737]" italics="true" pageId="16" pageNumber="17">S. blakei</emphasis>
<bibRefCitation id="EF974B5D6E204D7AFB8EFD5521C5FD31" author="Hartmann-Schroder" box="[1123,1431,712,738]" pageId="16" pageNumber="33" refString="Hartmann-Schroder, G. (1980) Teil 4. Die Polychaeten der tropischen Nordwestkuste Australiens (zwischen Port Samson im Norden und Exmouth im Suden). In: Hartmann-Schroder, G. &amp; Hartman, G. (Eds.), Zur Kenntnis des Eulitorals der australischen Kusten unter besonderer Berucksichtigung der Polychaeten und Ostracoden. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 77, 41 - 110." type="journal article" year="1980">Hartmann-Schröder, 1980</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C064D2F6E204D7AFF7AFD6D263AFCD9" ID-CoL="4VRNS" authority="Blake &amp; Kudenov, 1978" authorityName="Blake &amp; Kudenov" authorityYear="1978" box="[151,616,752,778]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="carunculata">
<emphasis id="B972EABE6E204D7AFF7AFD6D2510FCDA" box="[151,322,752,777]" italics="true" pageId="16" pageNumber="17">S. carunculata</emphasis>
<bibRefCitation id="EF974B5D6E204D7AFEA4FD6D263AFCD9" author="Blake" box="[329,616,752,778]" pageId="16" pageNumber="31" refString="Blake, J. A. &amp; Kudenov, J. D. (1978) The Spionidae (Polychaeta) from southeastern Australia and adjacent areas with a revision of the genera. Memoirs of the National Museum of Victoria, 39, 171 - 280." type="journal article" year="1978">Blake &amp; Kudenov, 1978</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C064D2F6E204D7AFD99FD6D2015FCD9" ID-CoL="4VRNT" authority="Hartmann-Schroder, 1962" authorityName="Hartmann-Schroder" authorityYear="1962" box="[628,1095,752,778]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="chilensis">
<emphasis id="B972EABE6E204D7AFD99FD6D26A5FCDA" box="[628,759,752,777]" italics="true" pageId="16" pageNumber="17">S. chilensis</emphasis>
(
<bibRefCitation id="EF974B5D6E204D7AFCEBFD6D206FFCD9" author="Hartmann-Schroder" box="[774,1085,752,778]" pageId="16" pageNumber="33" refString="Hartmann-Schroder, G. (1962) Die Polychaeten des Eulittorals. In: Hartmann-Schroder, G. &amp; Hartman, G. (Eds.), Zur Kenntnis des Eulitorals der chilenischen Pazifikkuste und argentinischen Kuste Sudpatagoniens unter besonderer Berucksichtigung der Polychaeten und Ostracoden. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, Supplement to 60, 57 - 167." type="journal article" year="1962">Hartmann-Schröder, 1962</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C064D2F6E204D7AFBBFFD6D21C5FCD9" ID-CoL="4VRP6" authority="Dauer, 1985" authorityName="Dauer" authorityYear="1985" box="[1106,1431,752,778]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="hutchingsae">
<emphasis id="B972EABE6E204D7AFBBFFD6D20AFFCDA" box="[1106,1277,752,777]" italics="true" pageId="16" pageNumber="17">S. hutchingsae</emphasis>
<bibRefCitation id="EF974B5D6E204D7AFAE9FD6D21C5FCD9" author="Dauer" box="[1284,1431,752,778]" pageId="16" pageNumber="32" refString="Dauer, D. M. (1985) A new species of Scolelepis (Polychaeta: Spionidae) from Lizard Island, Australia. Proceedings of the Biological Society of Washington, 98, 678 - 681." type="journal article" year="1985">Dauer, 1985</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="4C064D2F6E204D7AFF21FC8526C9FCE1" ID-CoL="4VRMY" authority="Hartmann-Schroder, 1981" authorityName="Hartmann-Schroder" authorityYear="1981" box="[204,667,792,818]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="kudenovi">
<emphasis id="B972EABE6E204D7AFF21FC85250AFCE2" box="[204,344,792,817]" italics="true" pageId="16" pageNumber="17">S. kudenovi</emphasis>
<bibRefCitation id="EF974B5D6E204D7AFE8EFC8526C9FCE1" author="Hartmann-Schroder" box="[355,667,792,818]" pageId="16" pageNumber="33" refString="Hartmann-Schroder, G. (1981) Teil 6. Die Polychaeten der tropischen-subtropischen Westkuste Australiens (zwischen Exmouth im Norden und Cervantes im Suden). In: Hartmann-Schroder, G. &amp; Hartman, G. (Eds.), Zur Kenntnis des Eulitorals der australischen Kusten unter besonderer Berucksichtigung der Polychaeten und Ostracoden. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 78, 19 - 96." type="journal article" year="1981">Hartmann-Schröder, 1981</bibRefCitation>
</taxonomicName>
, that possess notosetae on setiger 1, notopodial hooded hooks, bidentate neuropodial hooded hooks and that lack an occipital tentacle.
<taxonomicName id="4C064D2F6E204D7AFC1AFCDD2089FC8A" ID-CoL="4VRNK" box="[1015,1243,832,857]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="alisonae" status="sp. nov.">
<emphasis id="B972EABE6E204D7AFC1AFCDD2089FC8A" box="[1015,1243,832,857]" italics="true" pageId="16" pageNumber="17">Scolelepis alisonae</emphasis>
</taxonomicName>
<emphasis id="B972EABE6E204D7AFB08FCA22178FC8A" bold="true" box="[1253,1322,831,857]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A24157C56E204D7AFB08FCA22178FC8A" box="[1253,1322,831,857]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is distinguished from all these species in the possession of notopodial postsetal lamellae with up to nine lobes in anterior setigers and neuropodial postsetal lamellae with up to 3 lobes in anterior setigers.
<taxonomicName id="4C064D2F6E204D7AFB62FC0D2120FC7A" ID-CoL="4VRNK" box="[1167,1394,912,937]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="alisonae" status="sp. nov.">
<emphasis id="B972EABE6E204D7AFB62FC0D2120FC7A" box="[1167,1394,912,937]" italics="true" pageId="16" pageNumber="17">Scolelepis alisonae</emphasis>
</taxonomicName>
<emphasis id="B972EABE6E204D7AFA94FC1224FDFC02" bold="true" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A24157C56E204D7AFA94FC1224FDFC02" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C064D2F6E204D7AFF06FC25262DFC01" ID-CoL="4VRMZ" authority="Eibye-Jacobsen, 1997" authorityName="Eibye-Jacobsen" authorityYear="1997" box="[235,639,952,978]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="laciniata">
<emphasis id="B972EABE6E204D7AFF06FC252520FC02" box="[235,370,952,977]" italics="true" pageId="16" pageNumber="17">S. laciniata</emphasis>
<bibRefCitation id="EF974B5D6E204D7AFE91FC25262DFC01" author="Eibye-Jacobsen" box="[380,639,952,978]" pageId="16" pageNumber="32" refString="Eibye-Jacobsen, D. (1997) A new species of Scolelepis (Polychaeta: Spionidae), highly abundant on the sand beaches of western Phuket Island, Thailand. Bulletin of Marine Science, 60, 240 - 251." type="journal article" year="1997">Eibye-Jacobsen, 1997</bibRefCitation>
</taxonomicName>
are similar in the possession of notopodial postsetal lamellae with conical lobes; however,
<taxonomicName id="4C064D2F6E204D7AFE5EFC7D26C7FC2A" ID-CoL="4VRNK" box="[435,661,992,1017]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="alisonae" status="sp. nov.">
<emphasis id="B972EABE6E204D7AFE5EFC7D26C7FC2A" box="[435,661,992,1017]" italics="true" pageId="16" pageNumber="17">Scolelepis alisonae</emphasis>
</taxonomicName>
<emphasis id="B972EABE6E204D7AFD71FC42268CFC2A" bold="true" box="[668,734,991,1017]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A24157C56E204D7AFD71FC42268CFC2A" box="[668,734,991,1017]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is easily distinguished by the presence of notosetae on setiger 1 and notopodial hooded hooks (both absent in
<taxonomicName id="4C064D2F6E204D7AFD1DFB952724FBF2" ID-CoL="4VRMZ" box="[752,886,1032,1057]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="laciniata">
<emphasis id="B972EABE6E204D7AFD1DFB952724FBF2" box="[752,886,1032,1057]" italics="true" pageId="16" pageNumber="17">S. laciniata</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BB936AC6E204D7AFF2AFBAD2071FB49" blockId="16.[151,1437,152,1218]" pageId="16" pageNumber="17">
The palp morphology of
<taxonomicName id="4C064D2F6E204D7AFE17FBAD262CFB9A" ID-CoL="4VRNK" box="[506,638,1072,1097]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="alisonae" status="sp. nov.">
<emphasis id="B972EABE6E204D7AFE17FBAD262CFB9A" box="[506,638,1072,1097]" italics="true" pageId="16" pageNumber="17">S. alisonae</emphasis>
</taxonomicName>
<emphasis id="B972EABE6E204D7AFD64FBB2269DFB9A" bold="true" box="[649,719,1071,1097]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A24157C56E204D7AFD64FBB2269DFB9A" box="[649,719,1071,1097]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, is uniquely different from previously described species in exhibiting weakly separated long and short transverse rows of cilia in a 1:1 ratio. See Discussion for a comparative analysis of the palp morphology in members of the genus
<taxonomicName id="4C064D2F6E204D7AFC48FB1D204EFB4A" ID-CoL="84THX" box="[933,1052,1152,1177]" class="Polychaeta" family="Spionidae" genus="Scolelepis" kingdom="Animalia" order="Spionida" pageId="16" pageNumber="17" phylum="Annelida" rank="genus">
<emphasis id="B972EABE6E204D7AFC48FB1D204EFB4A" box="[933,1052,1152,1177]" italics="true" pageId="16" pageNumber="17">Scolelepis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C65276E204D7AFF2AFB3A21CEFB11" box="[199,1436,1191,1218]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8BB936AC6E204D7AFF2AFB3A21CEFB11" blockId="16.[151,1437,152,1218]" box="[199,1436,1191,1218]" pageId="16" pageNumber="17">
<emphasis id="B972EABE6E204D7AFF2AFB3A2535FB12" bold="true" box="[199,359,1191,1217]" pageId="16" pageNumber="17">Distribution.</emphasis>
Sandy beach in Morong of the Bataan province in the
<collectingCountry id="F311763C6E204D7AFC09FB35203EFB11" box="[996,1132,1192,1218]" name="Philippines" pageId="16" pageNumber="17">Philippines</collectingCountry>
; shallow subtidal (&lt;
<quantity id="4CFE9B496E204D7AFAB3FB3521DDFB11" box="[1374,1423,1192,1218]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="17" unit="m" value="5.0">5 m</quantity>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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