190 lines
24 KiB
XML
190 lines
24 KiB
XML
<document id="E75EA44F2061C3E2A64FBDBB074EEF3F" ID-DOI="10.5281/zenodo.5377863" ID-ISSN="1638-9395" ID-Zenodo-Dep="5377863" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630283615740" checkinUser="marcus" docAuthor="Fleck, Günther, Bechly, Günter, Martínez-Delclòs, Xavier, Jarzembowski, Edmund A. & Nel, André" docDate="2004" docId="03AF87A4E1151708FD2DFAF3FE6BFAEE" docLanguage="en" docName="Geodiversitas.2004.26.1.33-60.pdf" docOrigin="Geodiversitas 26 (1)" docStyle="DocumentStyle:EA833BFD04EE0E3F39962C07E351CEA5.3:Geodiversitas.2000-2008.journal_article.type1" docStyleId="EA833BFD04EE0E3F39962C07E351CEA5" docStyleName="Geodiversitas.2000-2008.journal_article.type1" docStyleVersion="3" docTitle="Tarsophlebiidae Handlirsch 1906" docType="treatment" docVersion="5" lastPageNumber="36" masterDocId="FF96FFDCE117170BFFC5FFE1FFAAFF8D" masterDocTitle="A revision of the Upper Jurassic-Lower Cretaceous dragonfly family Tarsophlebiidae, with a discussion on the phylogenetic positions of the Tarsophlebiidae and Sieblosiidae (Insecta, Odonatoptera, Panodonata)" masterLastPageNumber="60" masterPageNumber="33" pageNumber="35" updateTime="1699213624068" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="1A733DDAA306B0F94C24B8FF6FFEBAC3">A revision of the Upper Jurassic-Lower Cretaceous dragonfly family Tarsophlebiidae, with a discussion on the phylogenetic positions of the Tarsophlebiidae and Sieblosiidae (Insecta, Odonatoptera, Panodonata)</mods:title>
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<mods:namePart id="26B101357FCFDF685671C72B0D200075">Fleck, Günther</mods:namePart>
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<mods:affiliation id="9CE6D2671A5729995474252FAE7A2F10">Département Histoire de la Terre, USM 0203 and CNRS-Muséum UMR 5143, national d’Histoire naturelle, 45 rue Buffon, F- 75231 Paris cedex 05 (France) fleck @ mnhn. fr</mods:affiliation>
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<mods:nameIdentifier id="E3D270D5FEF44E8C3039C3E2E8696A5A" type="email">fleck@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="6520A01D60F1038BD2B52B5A3887F548">Bechly, Günter</mods:namePart>
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<mods:affiliation id="611F1E1609E56BF69ACC0A930F3ED7F5">Staatliches Museum für Naturkunde Stuttgart, Rosenstein 1, D- 70191 Stuttgart (Germany) bechly. smns @ naturkundemuseum-bw. de</mods:affiliation>
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<mods:namePart id="3BDDD33C3E15DB3D521E125F73F0BA4F">Martínez-Delclòs, Xavier</mods:namePart>
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<mods:affiliation id="B7661C8B528A9B3373D37CA127914C10">Departament d’Estratigrafia, Paleontologia i Geociències Marines, Facultat de Geologia, Universitat de Barcelona, E- 08071 Barcelona (Spain) xdelclos @ natura. geo. ub. es</mods:affiliation>
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<mods:namePart id="5A6CDB5910D91D150DA886A5DD438BBC">Jarzembowski, Edmund A.</mods:namePart>
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<mods:affiliation id="9C22200A91F630AE180CF4E6918F8085">Maidstone Museum and Bentlif Art Gallery, St Faith’s St, Maidstone, Kent ME 14 1 LH (UK) and Postgraduate Research Institute for Sedimentology & PRIS, University of Reading, Reading RG 6 2 AB (UK) ed @ mbcmus 1. demon. co. uk</mods:affiliation>
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<mods:namePart id="7DF944E16EDFE952C3A30372E619E6AE">Nel, André</mods:namePart>
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<mods:affiliation id="3915CBD9E87F200D7DE9A7C4E60ECACF">Département Histoire de la Terre, USM 0203 and CNRS-Muséum UMR 5143, national d’Histoire naturelle, 45 rue Buffon, F- 75231 Paris cedex 05 (France) anel @ mnhn. fr</mods:affiliation>
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<mods:title id="60BABDE15AB90B2E089BDF26F34CCA28">Geodiversitas</mods:title>
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<mods:date id="E7D0FDF5A8B7C33751851A7E32F21F98">2004</mods:date>
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<mods:number id="92587BE181F6DE30137F68E8F33EFA5A">26</mods:number>
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<mods:start id="144948D6968C10EBC6A9670359A1B3A6">33</mods:start>
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<mods:end id="F5C8DEA3083E105D5FA6B8130566BEA6">60</mods:end>
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<mods:identifier id="A6DA2674364DD054EFA33E7BCD6238B3" type="DOI">10.5281/zenodo.5377863</mods:identifier>
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<mods:identifier id="C70E42DB6D171E57E5BE0DC0FEB70E32" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="3AEDB6A996D42482015D454A31624582" type="Zenodo-Dep">5377863</mods:identifier>
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<treatment id="03AF87A4E1151708FD2DFAF3FE6BFAEE" LSID="urn:lsid:plazi:treatment:03AF87A4E1151708FD2DFAF3FE6BFAEE" httpUri="http://treatment.plazi.org/id/03AF87A4E1151708FD2DFAF3FE6BFAEE" lastPageId="3" lastPageNumber="36" pageId="2" pageNumber="35">
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<heading id="D0F181DEE1151709FD2DFAF3FB6BFAA6" box="[744,1217,1297,1324]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="35" reason="3">
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Family
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<taxonomicName id="4C064D31E1151709FCFFFAF0FB6BFAA6" authority="Handlirsch, 1906" authorityName="Handlirsch" authorityYear="1906" box="[826,1217,1297,1323]" class="Insecta" family="Tarsophlebiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">
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TARSOPHLEBIIDAE
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<bibRefCitation id="EF974B43E1151709FBC7FAF0FB6BFAA6" author="HANDLIRSCH A." box="[1026,1217,1297,1323]" pageId="2" pageNumber="35" refId="ref17865" refString="HANDLIRSCH A. 1906 - 1908. - Die fossilen Insekten und die Phylogenie der rezenten Formen. Ein Handbuch fur Palaontologen und Zoologen. Engelman V. W., Leipzig, 1430 p." type="book" year="1906">Handlirsch, 1906</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C31C6539E1151709FD0FFAB3FD5DFA0E" pageId="2" pageNumber="35" type="reference_group">
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<treatmentCitation id="0AA710A3E1151709FD0FFAB3FBC6FAE7" author="HANDLIRSCH A." box="[714,1132,1362,1386]" page="467" pageId="2" pageNumber="35" year="1906">
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<taxonomicName id="4C064D31E1151709FD0FFAB3FBC6FAE7" authority="Handlirsch, 1906: 467" authorityName="Handlirsch" authorityPageNumber="467" authorityYear="1906" box="[714,1132,1362,1386]" class="Insecta" family="Tarsophlebiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">
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Tarsophlebiidae
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<bibRefCitation id="EF974B43E1151709FCBFFAB3FBC6FAE7" author="HANDLIRSCH A." box="[890,1132,1362,1386]" pageId="2" pageNumber="35" refId="ref17865" refString="HANDLIRSCH A. 1906 - 1908. - Die fossilen Insekten und die Phylogenie der rezenten Formen. Ein Handbuch fur Palaontologen und Zoologen. Engelman V. W., Leipzig, 1430 p." type="book" year="1906">Handlirsch, 1906: 467</bibRefCitation>
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, 468, 580, 581.
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</paragraph>
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<subSubSection id="C31C6539E1151708FD0FFA72FD21FF78" lastPageId="3" lastPageNumber="36" pageId="2" pageNumber="35" type="materials_examined">
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<paragraph id="8BB936B2E1151709FD0FFA72FB08FA49" blockId="2.[714,1247,1362,1669]" pageId="2" pageNumber="35">
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<typeStatus id="54BD8810E1151709FD0FFA72FD56FA25" box="[714,764,1427,1450]" pageId="2" pageNumber="35">TYPE</typeStatus>
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GENUS. —
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<taxonomicName id="4C064D31E1151709FCB2FA72FB2FFA26" authority="Hagen, 1866" authorityName="Hagen" authorityYear="1866" box="[887,1157,1427,1451]" class="Insecta" family="Tarsophlebiidae" genus="Tarsophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="B972EAA0E1151709FCB2FA72FC5EFA26" box="[887,1012,1427,1451]" italics="true" pageId="2" pageNumber="35">Tarsophlebia</emphasis>
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<bibRefCitation id="EF974B43E1151709FC38FA72FB2FFA26" author="HAGEN H. A." box="[1021,1157,1427,1451]" pageId="2" pageNumber="35" pagination="57 - 96" refId="ref17842" refString="HAGEN H. A. 1866. - Die Neuroptera des lithographischen Schiefers in Bayern. Palaeontographica 15: 57 - 96." type="journal article" year="1866">Hagen, 1866</bibRefCitation>
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</taxonomicName>
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||
, original designation of
|
||
<bibRefCitation id="EF974B43E1151709FC9EFA4CFBBBFA48" author="HANDLIRSCH A." box="[859,1041,1453,1477]" pageId="2" pageNumber="35" refId="ref17865" refString="HANDLIRSCH A. 1906 - 1908. - Die fossilen Insekten und die Phylogenie der rezenten Formen. Ein Handbuch fur Palaontologen und Zoologen. Engelman V. W., Leipzig, 1430 p." type="book" year="1906">Handlirsch (1906)</bibRefCitation>
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by monotypy.
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</paragraph>
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<paragraph id="8BB936B2E1151708FD0FFA35FD21FF78" blockId="2.[714,1247,1362,1669]" lastBlockId="3.[151,684,169,245]" lastPageId="3" lastPageNumber="36" pageId="2" pageNumber="35">
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INCLUDED GENERA. —
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<taxonomicName id="4C064D31E1151709FC7DFA34FB6BFA60" authority="Hagen, 1866" authorityName="Hagen" authorityYear="1866" box="[952,1217,1492,1517]" class="Insecta" family="Tarsophlebiidae" genus="Tarsophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="B972EAA0E1151709FC7DFA34FB9EFA60" box="[952,1076,1493,1517]" italics="true" pageId="2" pageNumber="35">Tarsophlebia</emphasis>
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<bibRefCitation id="EF974B43E1151709FBF8FA35FB6BFA60" author="HAGEN H. A." box="[1085,1217,1492,1517]" pageId="2" pageNumber="35" pagination="57 - 96" refId="ref17842" refString="HAGEN H. A. 1866. - Die Neuroptera des lithographischen Schiefers in Bayern. Palaeontographica 15: 57 - 96." type="journal article" year="1866">Hagen, 1866</bibRefCitation>
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</taxonomicName>
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(=
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<taxonomicName id="4C064D31E1151709FD0FFA0EFBA7F98A" authority="Tillyard, 1923" authorityName="Tillyard" authorityYear="1923" box="[714,1037,1519,1543]" class="Insecta" genus="Tarsophlebiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="B972EAA0E1151709FD0FFA0EFCC1F98A" box="[714,875,1519,1543]" italics="true" pageId="2" pageNumber="35">Tarsophlebiopsis</emphasis>
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<bibRefCitation id="EF974B43E1151709FCB1FA0EFBA7F98A" author="TILLYARD R. J." box="[884,1037,1519,1543]" pageId="2" pageNumber="35" pagination="146 - 152" refId="ref18836" refString="TILLYARD R. J. 1923. - Tarsophlebiopsis mayi n. gen. et n. sp., a dragon-fly, found in the body-chamber of a Corallian ammonite. Geological Magazine 60: 146 - 152." type="journal article" year="1923">Tillyard, 1923</bibRefCitation>
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</taxonomicName>
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) and
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<taxonomicName id="4C064D31E1151709FB89FA0EFCC6F9AD" authority="Pritykina, 1968" authorityName="Pritykina" authorityYear="1968" class="Insecta" family="Tarsophlebiidae" genus="Turanophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="B972EAA0E1151709FB89FA0EFB75F98A" box="[1100,1247,1519,1543]" italics="true" pageId="2" pageNumber="35">Turanophlebia</emphasis>
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<bibRefCitation id="EF974B43E1151709FD0FF9E9FCC6F9AD" author="PRITYKINA L. N." box="[714,876,1544,1568]" pageId="2" pageNumber="35" pagination="26 - 55" refId="ref18579" refString="PRITYKINA L. N. 1968. - Strekozy Karatau (Odonata) [= Dragonflies from Karatau (Odonata)], in PANFILOV D. V. (ed.), Yurskie nasekomye Karatau. [= Fossil Insects from Karatau]. Academy of Sciences of the SSSR, Nauka, Moscow: 26 - 55 (in Russian)." type="book chapter" year="1968">Pritykina, 1968</bibRefCitation>
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</taxonomicName>
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||
. Since
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||
<bibRefCitation id="EF974B43E1151709FC73F9E9FBEAF9AD" author="BECHLY G." box="[950,1088,1544,1568]" pageId="2" pageNumber="35" refId="ref17284" refString="BECHLY G. 1996. - Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata), unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der * Odonata. Petalura Boblingen, Special Volume 2, 402 p." type="book" year="1996">Bechly (1996</bibRefCitation>
|
||
,
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||
<bibRefCitation id="EF974B43E1151709FB88F9E9FB28F9AD" author="BECHLY G." box="[1101,1154,1544,1568]" pageId="2" pageNumber="35" pagination="31 - 70" refId="ref17331" refString="BECHLY G. 1997. - New fossil Odonata from the Upper Triassic of Italy, with a redescription of Italophlebia gervasutti, and a reclassification of Triassic dragonflies. Rivista del Museo Civico di Scienze Naturale E. Caffi Bergamo 19: 31 - 70." type="journal article" year="1997">1997</bibRefCitation>
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) demonstrated that the genera
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<taxonomicName id="4C064D31E1151709FC0FF9C0FB75F9B4" authority="Pritykina, 1968" authorityName="Pritykina" authorityYear="1968" box="[970,1247,1569,1593]" class="Magnoliopsida" family="Ochnaceae" genus="Euthemis" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="genus">
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||
<emphasis id="B972EAA0E1151709FC0FF9C0FB80F9B4" box="[970,1066,1569,1593]" italics="true" pageId="2" pageNumber="35">Euthemis</emphasis>
|
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<bibRefCitation id="EF974B43E1151709FBF1F9C0FB75F9B4" author="PRITYKINA L. N." box="[1076,1247,1569,1593]" pageId="2" pageNumber="35" pagination="26 - 55" refId="ref18579" refString="PRITYKINA L. N. 1968. - Strekozy Karatau (Odonata) [= Dragonflies from Karatau (Odonata)], in PANFILOV D. V. (ed.), Yurskie nasekomye Karatau. [= Fossil Insects from Karatau]. Academy of Sciences of the SSSR, Nauka, Moscow: 26 - 55 (in Russian)." type="book chapter" year="1968">Pritykina, 1968</bibRefCitation>
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</taxonomicName>
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||
(Upper Jurassic) and
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<taxonomicName id="4C064D31E1151709FC7EF9DAFB75F9DE" authority="Bode, 1953" authorityName="Bode" authorityYear="1953" box="[955,1247,1595,1619]" class="Insecta" family="Sphenophlebiidae" genus="Sphenophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="35" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B972EAA0E1151709FC7EF9DAFBFFF9DE" box="[955,1109,1595,1619]" italics="true" pageId="2" pageNumber="35">Sphenophlebia</emphasis>
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Bode, 1953
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</taxonomicName>
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(Upper Liassic) are isophlebioid “anisozygoptères” and thus not related to
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<taxonomicName id="4C064D31E1151709FC48F98CFB84F908" authorityName="Handlirsch" authorityYear="1906" box="[909,1070,1645,1669]" class="Insecta" family="Tarsophlebiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="35" phylum="Arthropoda" rank="family">Tarsophlebiidae</taxonomicName>
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, there is no more evidence for a Lower Jurassic occurrence of
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<taxonomicName id="4C064D31E1141708FF52FF22FE9EFF56" authorityName="Handlirsch" authorityYear="1906" box="[151,308,195,219]" class="Insecta" family="Tarsophlebiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="36" phylum="Arthropoda" rank="family">Tarsophlebiidae</taxonomicName>
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and also no need for a further redundant taxon Tarsophlebioidea or Tarsophlebioptera.
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31C6539E1141708FF52FEEAFE6BFAEE" pageId="3" pageNumber="36" type="description">
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||
<paragraph id="8BB936B2E1141708FF52FEEAFEAAFEDB" blockId="3.[151,684,267,342]" pageId="3" pageNumber="36">STRATIGRAPHIC AND GEOGRAPHIC RANGE. — Only known from the Upper Jurassic and Lower Cretaceous of Eurasia.</paragraph>
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<paragraph id="8BB936B2E1141708FF52FE8DFE3BFCE9" blockId="3.[151,685,364,1379]" pageId="3" pageNumber="36">
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||
EMENDED DIAGNOSIS. — Nel
|
||
<emphasis id="B972EAA0E1141708FE05FE8DFE5AFE08" box="[448,496,364,389]" italics="true" pageId="3" pageNumber="36">et al.</emphasis>
|
||
(1993) proposed a diagnosis of the
|
||
<taxonomicName id="4C064D31E1141708FEFCFE67FE73FE13" authorityName="Handlirsch" authorityYear="1906" box="[313,473,390,414]" class="Insecta" family="Tarsophlebiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="36" phylum="Arthropoda" rank="family">Tarsophlebiidae</taxonomicName>
|
||
. We here emend this diagnosis with the following characters: 1) primary antenodal braces Ax1 and Ax2 stronger than the secondary antenodal cross-veins; 2) in all fore and hind wings, there are pairs of secondary longitudinal concave veins “above” and “below” the convex veins CuA, MA, and IR2, and closely parallel to them. We propose to name them respectively antero-CuA, postero- CuA, antero-MA, postero-MA, antero-IR2 and postero-IR
|
||
<specimenCount id="9D00FD3BE1141708FEC4FD8BFED3FD0F" box="[257,377,618,642]" pageId="3" pageNumber="36" type="intercalary">2 intercalary</specimenCount>
|
||
veins. These veins are more or less long and defined in the different taxa. The posterointercalaries are always longer than the associated antero-intercalaries; and 3) in male, one pair of well separated anal appendages visible, of very particular shape, basally strongly sclerotized, with high humps and a non-sclerotized paddle-like distal part. No median anal appendage visible. These body characters are only known for
|
||
<taxonomicName id="4C064D31E1141708FE95FCD4FD85FCC6" baseAuthorityName="Hagen" baseAuthorityYear="1862" box="[336,559,820,845]" class="Insecta" family="Tarsophlebiidae" genus="Tarsophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="eximia">
|
||
<emphasis id="B972EAA0E1141708FE95FCD4FD85FCC6" box="[336,559,820,845]" italics="true" pageId="3" pageNumber="36">Tarsophlebia eximia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C064D31E1141708FDA9FCD5FEFEFCE9" authority="Fleck & Bechly & Martínez-Delclòs & Jarzembowski & Nel, 2004" authorityName="Fleck & Bechly & Martínez-Delclòs & Jarzembowski & Nel" authorityYear="2004" class="Insecta" family="Tarsophlebiidae" genus="Turanophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="vitimensis" status="sp. nov.">
|
||
<emphasis id="B972EAA0E1141708FDA9FCD5FEFEFCE9" italics="true" pageId="3" pageNumber="36">Turanophlebia vitimensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24157DBE1141708FE9EFCACFE3BFCE9" box="[347,401,845,868]" pageId="3" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BB936B2E1141708FF52FC8BFE6BFAEE" blockId="3.[151,685,364,1379]" pageId="3" pageNumber="36">
|
||
The monophyly of
|
||
<taxonomicName id="4C064D31E1141708FEA4FC8BFDAEFC0F" authorityName="Handlirsch" authorityYear="1906" box="[353,516,874,898]" class="Insecta" family="Tarsophlebiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="36" phylum="Arthropoda" rank="family">Tarsophlebiidae</taxonomicName>
|
||
is supported by several strong autapomorphies (Nel
|
||
<emphasis id="B972EAA0E1141708FE3BFC63FD85FC16" box="[510,559,898,923]" italics="true" pageId="3" pageNumber="36">et al.</emphasis>
|
||
1993), such as: hypertrophied hind wing subdiscoidal cell, developed as “pseudo discoidal cell”; in hind wing, fusion of veins [MAb + MP + CuA] for a considerable distance before separation of MP and CuA; vein AA strongly bent at insertion of CuP-crossing; extremely acute distal angles of fore wing discoidal and subdiscoidal cell. The characters “distinctly prolonged legs, with very long tarsi” and “male cerci with strange distal expansions” are present in both
|
||
<taxonomicName id="4C064D31E1141708FE5AFB87FE56FBF0" baseAuthorityName="Hagen" baseAuthorityYear="1862" box="[415,508,1126,1149]" class="Insecta" family="Tarsophlebiidae" genus="Tarsophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="eximia">
|
||
<emphasis id="B972EAA0E1141708FE5AFB87FE56FBF0" box="[415,508,1126,1149]" italics="true" pageId="3" pageNumber="36">T. eximia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C064D31E1141708FDF4FB87FD01FBF0" authority="Fleck & Bechly & Martínez-Delclòs & Jarzembowski & Nel, 2004" authorityName="Fleck & Bechly & Martínez-Delclòs & Jarzembowski & Nel" authorityYear="2004" box="[561,683,1126,1149]" class="Insecta" family="Tarsophlebiidae" genus="Turanophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="vitimensis" status="sp. nov.">
|
||
<emphasis id="B972EAA0E1141708FDF4FB87FD01FBF0" box="[561,683,1126,1149]" italics="true" pageId="3" pageNumber="36">T. vitimensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24157DBE1141708FF52FB61FF79FB1A" box="[151,211,1152,1175]" pageId="3" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
|
||
, and probably also in other
|
||
<taxonomicName id="4C064D31E1141708FE3AFB61FD03FB15" authorityName="Handlirsch" authorityYear="1906" box="[511,681,1152,1176]" class="Insecta" family="Tarsophlebiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="36" phylum="Arthropoda" rank="family">Tarsophlebiidae</taxonomicName>
|
||
. The “extremely prolonged female ovipositor” could be an autapomorphy of the family, too, but it is only known from
|
||
<taxonomicName id="4C064D31E1141708FEDEFB2DFED3FB6E" baseAuthorityName="Hagen" baseAuthorityYear="1862" box="[283,377,1228,1251]" class="Insecta" family="Tarsophlebiidae" genus="Tarsophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="eximia">
|
||
<emphasis id="B972EAA0E1141708FEDEFB2DFED3FB6E" box="[283,377,1228,1251]" italics="true" pageId="3" pageNumber="36">T. eximia</emphasis>
|
||
</taxonomicName>
|
||
. A further alleged autapomorphy mentioned by Nel
|
||
<emphasis id="B972EAA0E1141708FEB9FB04FE06FB73" box="[380,428,1253,1278]" italics="true" pageId="3" pageNumber="36">et al.</emphasis>
|
||
(1993) rather seems to be a symplesiomorphy (
|
||
<bibRefCitation id="EF974B43E1141708FE43FB1EFDB7FA9A" author="BECHLY G." box="[390,541,1279,1303]" pageId="3" pageNumber="36" refId="ref17284" refString="BECHLY G. 1996. - Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata), unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der * Odonata. Petalura Boblingen, Special Volume 2, 402 p." type="book" year="1996">Bechly 1996</bibRefCitation>
|
||
), viz. “veins [RP + MA] - MAb - CuA aligned”. It is also present in the
|
||
<taxonomicName id="4C064D31E1141708FF05FAD3FD83FAC7" authority="Muttkowski, 1910" authorityName="Muttkowski" authorityYear="1910" box="[192,553,1330,1354]" class="Insecta" family="Epiophlebiidae" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="36" phylum="Arthropoda" rank="family">Epiophlebiidae Muttkowski, 1910</taxonomicName>
|
||
, Isophlebioptera and Heterophlebioptera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |