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<document id="D260C60FAAAEA5C4C3DF162B495B4D31" ID-CLB-Dataset="51470" ID-DOI="10.11646/zootaxa.4097.2.2" ID-GBIF-Dataset="fcd803c5-6723-4fbe-8db8-6f39b2168947" ID-ISSN="1175-5326" ID-Zenodo-Dep="262363" ID-ZooBank="732FE827-45BC-4AA6-AA6C-658C02E00FAC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459409865110" checkinUser="plazi" docAuthor="Lee, Young June, Marshall, David C., Mohagan, Alma &amp; Hill, Kathy B. R." docDate="2016" docId="03AF878BFFC8FFB5FF32FF51C68CFA08" docLanguage="en" docName="zootaxa.4097.2.2.pdf.imf" docOrigin="Zootaxa 4097 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Philipsalta Lee, Marshall &amp; Hill, gen. nov." docType="treatment" docVersion="15" lastPageNumber="197" masterDocId="FF96FFF3FFC2FFBFFFA5FFC9C421FC2E" masterDocTitle="Cicadas (Hemiptera: Cicadidae) from Camiguin of Mindanao Province and Dinagat Island in the Philippines, with a new genus and three new species" masterLastPageNumber="202" masterPageNumber="187" pageNumber="197" updateTime="1698191717392" updateUser="ExternalLinkService">
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<mods:title id="ED234442AECC6C5CCBBF68DEEFF7C1D6">Cicadas (Hemiptera: Cicadidae) from Camiguin of Mindanao Province and Dinagat Island in the Philippines, with a new genus and three new species</mods:title>
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<mods:namePart id="121B007D40D95A585D958F29D9181439">Lee, Young June</mods:namePart>
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<mods:namePart id="398C96F4826E6E395BE659B1641EE1FC">Marshall, David C.</mods:namePart>
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<mods:namePart id="CAECC6239538380F7AA4C25C88976401">Hill, Kathy B. R.</mods:namePart>
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<paragraph id="8BB9369DFFC8FFB5FF32FF51C6D0FC9C" blockId="10.[151,753,151,178]" box="[151,753,151,178]" pageId="10" pageNumber="197">
<heading id="D0F181F1FFC8FFB5FF32FF51C6D0FC9C" bold="true" box="[151,753,151,178]" fontSize="11" level="1" pageId="10" pageNumber="197" reason="1">
<emphasis id="B972EA8FFFC8FFB5FF32FF51C6D0FC9C" bold="true" box="[151,753,151,178]" pageId="10" pageNumber="197">
Genus
<taxonomicName id="4C064D1EFFC8FFB5FF4BFF5EC65AFC9C" ID-CoL="8HPDV" authority="Lee, Marshall &amp; Hill" authorityName="Lee, Marshall &amp; Hill" box="[238,635,151,178]" class="Insecta" family="Cicadidae" genus="Philipsalta" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B972EA8FFFC8FFB5FF4BFF5EC551FC9F" bold="true" box="[238,368,151,177]" italics="true" pageId="10" pageNumber="197">Philipsalta</emphasis>
Lee,
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&amp; Hill
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<taxonomicNameLabel id="A24157F4FFC8FFB5FD26FF51C6D0FC9C" box="[643,753,152,178]" pageId="10" pageNumber="197" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C31C6516FFC8FFB5FF32FF17C73AFCDA" box="[151,795,222,245]" pageId="10" pageNumber="197" type="materials_examined">
<paragraph id="8BB9369DFFC8FFB5FF32FF17C73AFCDA" blockId="10.[151,795,222,245]" box="[151,795,222,245]" pageId="10" pageNumber="197">
<emphasis id="B972EA8FFFC8FFB5FF32FF17C503FCDA" bold="true" box="[151,290,222,244]" pageId="10" pageNumber="197">
<typeStatus id="54BD883FFFC8FFB5FF32FF17C4EDFCDA" box="[151,204,222,244]" pageId="10" pageNumber="197">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C064D1EFFC8FFB5FE8CFF17C6E2FCDA" authority="Lee, Marshall &amp; Hill" authorityName="Lee, Marshall &amp; Hill" box="[297,707,222,245]" class="Insecta" family="Cicadidae" genus="Philipsalta" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="species" species="nigrina" status="sp. nov.">
<emphasis id="B972EA8FFFC8FFB5FE8CFF17C5CBFCDB" box="[297,490,222,245]" italics="true" pageId="10" pageNumber="197">Philipsalta nigrina</emphasis>
Lee,
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&amp; Hill
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<taxonomicNameLabel id="A24157F4FFC8FFB5FD6FFF17C73AFCDA" box="[714,795,222,244]" pageId="10" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C31C6516FFC8FFB5FF32FEE8C109FE50" pageId="10" pageNumber="197" type="description">
<paragraph id="8BB9369DFFC8FFB5FF32FEE8C109FE50" blockId="10.[151,1437,289,1574]" pageId="10" pageNumber="197">
<emphasis id="B972EA8FFFC8FFB5FF32FEE8C508FD14" bold="true" box="[151,297,289,314]" pageId="10" pageNumber="197">Description.</emphasis>
Body tiny, about
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long, monotonous in coloration. Head comparatively long, about as long as or longer than two-thirds the median length of pronotum. Head including eyes slightly wider than base of mesonotum. Postclypeus slightly swollen anteriad and rather flat ventrally. Pronotal collar very narrow, not dentate. Fore-femur with primary, secondary, and small subapical spine. Wings hyaline without infuscation. Forewing radial cell spacious, broader than ulnar cell 3. Forewing cubital cell spacious, about as broad as medial cell.
<collectingCountry id="F311760DFFC8FFB5FF75FE1CC529FDC0" box="[208,264,469,494]" name="India" pageId="10" pageNumber="197">Hind</collectingCountry>
wing with six apical cells.
<collectingCountry id="F311760DFFC8FFB5FDE9FE1CC6A4FDC0" box="[588,645,469,494]" name="Malta" pageId="10" pageNumber="197">Male</collectingCountry>
operculum small, wider than long, not reaching posterior margin of sternite II. Abdomen triangular in dorsal view, slightly longer than head and thorax together. Tergite 1 very short, less than one-fourth as long as tergite
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median length. Timbal cover absent. Timbals not extending below wing bases ventrally. Pygofer ovate with triangular dorsal beak in ventral view. Upper lobe of pygofer present. Dorsal beak triangular. Median lobe of uncus very short. A pair of clasper-like processes protruding downward.
</paragraph>
</subSubSection>
<subSubSection id="C31C6516FFC8FFB5FF62FD40C5DFFEE8" pageId="10" pageNumber="197" type="etymology">
<paragraph id="8BB9369DFFC8FFB5FF62FD40C5DFFEE8" blockId="10.[151,1437,289,1574]" pageId="10" pageNumber="197">
<emphasis id="B972EA8FFFC8FFB5FF62FD40C56FFE8C" bold="true" box="[199,334,649,674]" pageId="10" pageNumber="197">Etymology.</emphasis>
The generic name is a feminine noun from the combination of
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<collectingCountry id="F311760DFFC8FFB5FB95FD40C084FE8C" box="[1072,1189,649,674]" name="Philippines" pageId="10" pageNumber="197">Philippine</collectingCountry>
</emphasis>
and the Latin
<emphasis id="B972EA8FFFC8FFB5FAF5FD40C1B5FE8C" box="[1360,1428,649,674]" italics="true" pageId="10" pageNumber="197">psalta</emphasis>
, meaning “
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musician”.
</paragraph>
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<subSubSection id="C31C6516FFC8FFB5FF62FD18C68CFA08" pageId="10" pageNumber="197" type="discussion">
<paragraph id="8BB9369DFFC8FFB5FF62FD18C6C2FF78" blockId="10.[151,1437,289,1574]" pageId="10" pageNumber="197">
<emphasis id="B972EA8FFFC8FFB5FF62FD18C51AFEC4" bold="true" box="[199,315,721,746]" pageId="10" pageNumber="197">Remarks.</emphasis>
This genus is peculiar among the genera of
<taxonomicName id="4C064D1EFFC8FFB5FCEBFD18C7E8FEC4" box="[846,969,721,746]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="tribe" tribe="Cicadettini">Cicadettini</taxonomicName>
in having a very short tergite 1, and is distinguished from the other
<collectingCountry id="F311760DFFC8FFB5FE7EFD3CC66FFF20" box="[475,590,757,782]" name="Philippines" pageId="10" pageNumber="197">Philippine</collectingCountry>
<taxonomicName id="4C064D1EFFC8FFB5FDF0FD3CC6F1FF20" box="[597,720,757,782]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="tribe" tribe="Cicadettini">Cicadettini</taxonomicName>
genus
<taxonomicName id="4C064D1EFFC8FFB5FCBAFD3CC00CFF20" authority="Lee, 2009" authorityName="Lee" authorityYear="2009" box="[799,1069,757,782]" class="Insecta" family="Cicadidae" genus="Ggomapsalta" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFC8FFB5FCBAFD3CC797FF20" box="[799,950,757,782]" italics="true" pageId="10" pageNumber="197">Ggomapsalta</emphasis>
Lee, 2009
</taxonomicName>
by the following characteristics: body dark-colored; forewing radial and cubital cells spacious and broad; hind wing with six apical cells; upper lobe of pygofer present; median lobe of uncus very short.
</paragraph>
<paragraph id="8BB9369DFFC8FFB5FF62FCA8C69FF890" blockId="10.[151,1437,289,1574]" pageId="10" pageNumber="197">
According to a molecular phylogenetic study by
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<collectingCountry id="F311760DFFC8FFB5FD56FCA8C776FF54" box="[755,855,865,890]" name="Marshall Islands" pageId="10" pageNumber="197">Marshall</collectingCountry>
<emphasis id="B972EA8FFFC8FFB5FCFAFCABC7B0FF54" box="[863,913,865,890]" italics="true" pageId="10" pageNumber="197">et al</emphasis>
. (2015)
</bibRefCitation>
, this new genus forms a phylogenetic group together with
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<emphasis id="B972EA8FFFC8FFB5FEDEFC4CC5FAFFB0" box="[379,475,901,926]" italics="true" pageId="10" pageNumber="197">Huechys</emphasis>
</taxonomicName>
and a few other European, North American, and Micronesian genera. The new genus is morphologically distinguished from
<taxonomicName id="4C064D1EFFC8FFB5FDF4FC60C7C3FFEC" authority="Degeer, 1773" authorityName="Degeer" authorityYear="1773" box="[593,994,937,962]" class="Insecta" family="Cicadidae" genus="Huechys" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="species" species="sanguinea">
<emphasis id="B972EA8FFFC8FFB5FDF4FC60C70CFFEF" box="[593,813,937,962]" italics="true" pageId="10" pageNumber="197">Huechys sanguinea</emphasis>
(Degeer, 1773)
</taxonomicName>
, the
<typeStatus id="54BD883FFFC8FFB5FBBEFC63C06AFFEC" box="[1051,1099,938,962]" pageId="10" pageNumber="197">type</typeStatus>
species of
<taxonomicName id="4C064D1EFFC8FFB5FB68FC60C10CFFEC" box="[1229,1325,937,962]" class="Insecta" family="Cicadidae" genus="Huechys" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFC8FFB5FB68FC60C10CFFEC" box="[1229,1325,937,962]" italics="true" pageId="10" pageNumber="197">Huechys</emphasis>
</taxonomicName>
(the only Asian genus among the phylogenetic group), and its congeners by the following characteristics: head including eyes slightly wider [slightly narrower in
<taxonomicName id="4C064D1EFFC8FFB5FDFEFC38C69AF824" box="[603,699,1009,1034]" class="Insecta" family="Cicadidae" genus="Huechys" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFC8FFB5FDFEFC38C69AF824" box="[603,699,1009,1034]" italics="true" pageId="10" pageNumber="197">Huechys</emphasis>
</taxonomicName>
] than base of mesonotum; postclypeus slightly swollen [distinctly swollen in
<taxonomicName id="4C064D1EFFC8FFB5FEB3FBDCC557F800" box="[278,374,1045,1070]" class="Insecta" family="Cicadidae" genus="Huechys" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFC8FFB5FEB3FBDCC557F800" box="[278,374,1045,1070]" italics="true" pageId="10" pageNumber="197">Huechys</emphasis>
</taxonomicName>
] anteriad; wings hyaline without infuscation [forewing dark-colored in
<taxonomicName id="4C064D1EFFC8FFB5FB0BFBDCC12FF800" box="[1198,1294,1045,1070]" class="Insecta" family="Cicadidae" genus="Huechys" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFC8FFB5FB0BFBDCC12FF800" box="[1198,1294,1045,1070]" italics="true" pageId="10" pageNumber="197">Huechys</emphasis>
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]; hind wing apical cell 1 normal-sized, about half the area of apical cell 2 [minimal, less than one-fourth the area of apical cell
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<emphasis id="B972EA8FFFC8FFB5FF62FB94C506F858" box="[199,295,1117,1142]" italics="true" pageId="10" pageNumber="197">Huechys</emphasis>
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]; male tergite 1 very short, less than one-fourth as long as tergite
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median length [broadly exposed, more than one-third as long as tergite
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median length in
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<emphasis id="B972EA8FFFC8FFB5FCFDFB48C799F8B4" box="[856,952,1153,1178]" italics="true" pageId="10" pageNumber="197">Huechys</emphasis>
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].
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genitalia is very similar to that of
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<emphasis id="B972EA8FFFC8FFB5FF32FB6CC4D6F890" box="[151,247,1189,1214]" italics="true" pageId="10" pageNumber="197">Huechys</emphasis>
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, but the median lobe of uncus is shorter.
</paragraph>
<paragraph id="8BB9369DFFC8FFB5FF62FB00C68CFA08" blockId="10.[151,1437,289,1574]" pageId="10" pageNumber="197">
This new genus is morphologically similar to
<taxonomicName id="4C064D1EFFC8FFB5FCA9FB00C04FF8CC" authority="Chou &amp; Lei, 1997" authorityName="Chou &amp; Lei" authorityYear="1997" box="[780,1134,1225,1250]" class="Insecta" family="Cicadidae" genus="Scolopita" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFC8FFB5FCA9FB00C757F8CC" box="[780,886,1225,1250]" italics="true" pageId="10" pageNumber="197">Scolopita</emphasis>
Chou &amp; Lei, 1997
</taxonomicName>
[
<typeStatus id="54BD883FFFC8FFB5FB20FB03C099F8CC" box="[1157,1208,1226,1250]" pageId="10" pageNumber="197">type</typeStatus>
species:
<taxonomicName id="4C064D1EFFC8FFB5FA97FB00C5DCF928" authority="Ouchi, 1938" authorityName="Ouchi" authorityYear="1938" class="Insecta" family="Cicadidae" genus="Scolopita" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="species" species="mokanshanensis">
<emphasis id="B972EA8FFFC8FFB5FA97FB00C56EF928" italics="true" pageId="10" pageNumber="197">Scolopita mokanshanensis</emphasis>
(Ouchi, 1938)
</taxonomicName>
] but is distinguished by the following characteristics: body much smaller, with body length of
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[body longer than
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in
<taxonomicName id="4C064D1EFFC8FFB5FCB5FAD8C75BF904" box="[784,890,1297,1322]" class="Insecta" family="Cicadidae" genus="Scolopita" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="genus">
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</taxonomicName>
]; body with monotonous [colorful in
<taxonomicName id="4C064D1EFFC8FFB5FA8CFAD8C1B2F904" box="[1321,1427,1297,1322]" class="Insecta" family="Cicadidae" genus="Scolopita" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFC8FFB5FA8CFAD8C1B2F904" box="[1321,1427,1297,1322]" italics="true" pageId="10" pageNumber="197">Scolopita</emphasis>
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] coloration; head about as long as two-thirds [about half in
<taxonomicName id="4C064D1EFFC8FFB5FCF2FAFCC7E0F960" box="[855,961,1333,1358]" class="Insecta" family="Cicadidae" genus="Scolopita" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFC8FFB5FCF2FAFCC7E0F960" box="[855,961,1333,1358]" italics="true" pageId="10" pageNumber="197">Scolopita</emphasis>
</taxonomicName>
] the median length of pronotum; wings comparatively small, with ratio of forewing length to body length of less than 1.2 [more than
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<taxonomicName id="4C064D1EFFC8FFB5FA84FA90C1AAF95C" box="[1313,1419,1369,1394]" class="Insecta" family="Cicadidae" genus="Scolopita" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="genus">
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]; forewing costa slightly concave basally [never concave basally in
<taxonomicName id="4C064D1EFFC8FFB5FC27FAB4C7CDF9B8" box="[898,1004,1405,1430]" class="Insecta" family="Cicadidae" genus="Scolopita" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFC8FFB5FC27FAB4C7CDF9B8" box="[898,1004,1405,1430]" italics="true" pageId="10" pageNumber="197">Scolopita</emphasis>
</taxonomicName>
]; anal cell 2 (vannus) more than three times [less than three times in
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<emphasis id="B972EA8FFFC8FFB5FE54FA68C67AF994" box="[497,603,1441,1466]" italics="true" pageId="10" pageNumber="197">Scolopita</emphasis>
</taxonomicName>
] as long as wide; male operculum distinctly wider than long [longer than wide in
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<emphasis id="B972EA8FFFC8FFB5FF56FA0CC57CF9F0" box="[243,349,1477,1502]" italics="true" pageId="10" pageNumber="197">Scolopita</emphasis>
</taxonomicName>
]; male opercula apart from each other with gap of about one-fourth as wide as operculum [with gap of about half as wide as operculum in
<taxonomicName id="4C064D1EFFC8FFB5FDDCFA20C6C2FA2C" box="[633,739,1513,1538]" class="Insecta" family="Cicadidae" genus="Scolopita" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFC8FFB5FDDCFA20C6C2FA2C" box="[633,739,1513,1538]" italics="true" pageId="10" pageNumber="197">Scolopita</emphasis>
</taxonomicName>
]; male tergite 1 less than one-fourth [more than one-fourth in
<taxonomicName id="4C064D1EFFC8FFB5FF32F9C4C520FA08" box="[151,257,1549,1574]" class="Insecta" family="Cicadidae" genus="Scolopita" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFC8FFB5FF32F9C4C520FA08" box="[151,257,1549,1574]" italics="true" pageId="10" pageNumber="197">Scolopita</emphasis>
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] as long as tergite
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median length.
</paragraph>
</subSubSection>
</treatment>
</document>