291 lines
27 KiB
XML
291 lines
27 KiB
XML
<document ID-DOI="10.5733/afin.056.0103" ID-GBIF-Dataset="30501623-9ce2-4263-8fa4-baa5ecb310de" ID-ISSN="2305-2562" ID-Zenodo-Dep="7670419" checkinTime="1676640544055" checkinUser="felipe" docAuthor="Csuzdi, Cs., Sherlock, E., Kouete, M. Talla & Doherty-Bone, T. M." docDate="2015" docId="03AF87882654A670228DFDB9FBFFD30E" docLanguage="en" docName="AfrInvertebr.56.1.25-38.pdf" docOrigin="African Invertebrates 56 (1)" docSource="http://dx.doi.org/10.5733/afin.056.0103" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.2:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="2" docTitle="Dichogaster (Diplothecodrilus) asymmetrica Csuzdi & Sherlock & Kouete & Doherty-Bone 2015, sp. n." docType="treatment" docVersion="3" lastPageNumber="35" masterDocId="FF96FFF0265DA67B2273FFBBFFF5D208" masterDocTitle="Four new earthworm species from the highlands of Cameroon with description of a new genus Okudrilus gen. n. (Oligochaeta: Eudrilidae & Acanthodrilidae)" masterLastPageNumber="38" masterPageNumber="25" pageNumber="34" updateTime="1677158441759" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Four new earthworm species from the highlands of Cameroon with description of a new genus Okudrilus gen. n. (Oligochaeta: Eudrilidae & Acanthodrilidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Csuzdi, Cs.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sherlock, E.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kouete, M. Talla</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Doherty-Bone, T. M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>African Invertebrates</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2015-01-27</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>56</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>25</mods:start>
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<mods:end>38</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.5733/afin.056.0103</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5733/afin.056.0103</mods:identifier>
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<mods:identifier type="GBIF-Dataset">30501623-9ce2-4263-8fa4-baa5ecb310de</mods:identifier>
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<mods:identifier type="ISSN">2305-2562</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7670419</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7670476" ID-Zenodo-Dep="7670476" LSID="urn:lsid:plazi:treatment:03AF87882654A670228DFDB9FBFFD30E" httpUri="http://treatment.plazi.org/id/03AF87882654A670228DFDB9FBFFD30E" lastPageId="11" lastPageNumber="35" pageId="9" pageNumber="34">
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<subSubSection box="[254,1069,514,541]" pageId="9" pageNumber="34" type="nomenclature">
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<paragraph blockId="9.[181,1142,514,955]" box="[254,1069,514,541]" pageId="9" pageNumber="34">
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<heading box="[254,1069,514,541]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="34" reason="2">
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<taxonomicName authority="Csuzdi & Sherlock" authorityName="Csuzdi & Sherlock & Kouete & Doherty-Bone" authorityYear="2015" box="[254,1003,514,540]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="34" phylum="Annelida" rank="species" species="asymmetrica" status="sp. n." subGenus="Diplothecodrilus">
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<emphasis bold="true" box="[254,782,514,540]" pageId="9" pageNumber="34">Dichogaster (Diplothecodrilus) asymmetrica</emphasis>
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Csuzdi & Sherlock
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</taxonomicName>
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<taxonomicNameLabel box="[1009,1069,515,541]" pageId="9" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection lastPageId="11" lastPageNumber="36" pageId="9" pageNumber="34" type="description">
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||
<paragraph blockId="9.[181,1142,514,955]" box="[599,724,556,582]" pageId="9" pageNumber="34">
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||
<figureCitation box="[599,724,556,582]" captionStart="Figs 11–14" captionStartId="10.[144,183,1146,1168]" captionTargetBox="[152,1060,214,1126]" captionTargetId="figure-205@10.[144,1104,204,1138]" captionTargetPageId="10" captionText="Figs 11–14. Dichogaster (Diplothecodrilus) asymmetrica sp. n.: (11) ventral view of the clitellar region; (12) ectal third of the larger penial seta; (13) ectal third of the smaller penial seta; (14) spermatheca. Abbreviations: Fp = female pore, Pr = prostate pore." figureDoi="http://doi.org/10.5281/zenodo.7670438" httpUri="https://zenodo.org/record/7670438/files/figure.png" pageId="9" pageNumber="34">Figs 11–14</figureCitation>
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</paragraph>
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<paragraph blockId="9.[181,1142,514,955]" pageId="9" pageNumber="34">EtymOlOgy:The specific epithet refers tO the asymmetrical pOsitiOn Of the spermathecal duct.</paragraph>
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<paragraph blockId="9.[181,1142,514,955]" pageId="9" pageNumber="34">
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Diagnosis: L:
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, D:
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<quantity box="[519,638,667,694]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="9" pageNumber="34" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
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. Colour slightly reddish on dorsum, pale on Ventrum. First dOrsal pOre in 5/6.
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<taxonomicName box="[551,657,699,725]" class="Ascidiacea" family="Didemnidae" genus="Clitellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aplousobranchia" pageId="9" pageNumber="34" phylum="Chordata" rank="genus">Clitellum</taxonomicName>
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13–20.
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<specimenCount box="[746,767,700,726]" pageId="9" pageNumber="34" type="female">♀</specimenCount>
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14 paired, between
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<emphasis box="[989,1030,700,725]" italics="true" pageId="9" pageNumber="34">a–a</emphasis>
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. Prostatic pores 17, 19. Spermathecal pores 7/8, 8/9, spermathecae with a subdivided ampulla, and an asymmetrically attached duct. The lower ampulla part bears a unilocular diverticulum. Gizzard large in 5–6, last pair of hearts in 12. Excretory system meroic with 6 meronephridia on each side. Penial setae are of
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<specimenCount box="[794,906,826,852]" pageId="9" pageNumber="34" type="generic" typeStatus="types">two types</specimenCount>
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, larger L:
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<quantity box="[1021,1107,826,852]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="34" unit="mm" value="2.3">2.3 mm</quantity>
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D:
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<quantity box="[181,295,858,884]" metricMagnitude="-6" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="34" unit="mm" value="0.006">0.006 mm</quantity>
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, tip slightly undulated, tapering. Ornamentation scattered small teeth. Smaller seta L:
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<quantity box="[264,350,889,916]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="34" unit="mm" value="2.1">2.1 mm</quantity>
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D:
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<quantity box="[391,507,889,916]" metricMagnitude="-6" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="34" unit="mm" value="0.006">0.006 mm</quantity>
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, tip spoon-shaped, ornamentation minute teeth.
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</paragraph>
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<paragraph blockId="9.[181,1142,514,955]" box="[181,321,929,955]" pageId="9" pageNumber="34">Description:</paragraph>
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<paragraph blockId="9.[181,1142,971,1677]" box="[181,414,971,997]" pageId="9" pageNumber="34">
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<emphasis box="[181,414,971,997]" italics="true" pageId="9" pageNumber="34">External characters:</emphasis>
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</paragraph>
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<paragraph blockId="9.[181,1142,971,1677]" pageId="9" pageNumber="34">
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<emphasis box="[181,279,1004,1030]" italics="true" pageId="9" pageNumber="34">
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<typeStatus box="[181,279,1004,1030]" pageId="9" pageNumber="34">Syntypes</typeStatus>
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</emphasis>
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: L:
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<quantity box="[327,451,1004,1031]" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="9" pageNumber="34" unit="mm" value="67.5" valueMax="70.0" valueMin="65.0">65–70 mm</quantity>
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, D:
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<quantity box="[499,615,1004,1031]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="9" pageNumber="34" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
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, the number of segments was uncountable due to the poor condition of the specimens.
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<emphasis box="[627,706,1036,1062]" italics="true" pageId="9" pageNumber="34">Colour</emphasis>
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: Alive unknown, preserved reddish on dorsum, pale on ventrum.
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<emphasis box="[474,610,1068,1093]" italics="true" pageId="9" pageNumber="34">Prostomium</emphasis>
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: Schizolobous.
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<emphasis box="[789,979,1067,1093]" italics="true" pageId="9" pageNumber="34">First dorsal pore</emphasis>
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: In 5/6.
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<emphasis box="[1073,1134,1068,1094]" italics="true" pageId="9" pageNumber="34">Setae</emphasis>
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: All ventral, setal arrangement on segment 14 aa:ab:bc:cd:dd=6:1:7:1:30, after clitellum not seen.
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<taxonomicName authority=": Circular" authorityName="Circular" box="[292,505,1131,1157]" class="Ascidiacea" family="Didemnidae" genus="Clitellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aplousobranchia" pageId="9" pageNumber="34" phylum="Chordata" rank="genus">
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<emphasis box="[292,396,1131,1157]" italics="true" pageId="9" pageNumber="34">Clitellum</emphasis>
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: Circular
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</taxonomicName>
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extends over segments 13–20.
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<emphasis box="[867,981,1131,1157]" italics="true" pageId="9" pageNumber="34">Male field</emphasis>
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: Oblong with prostatic pores paired on 17,
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<quantity box="[511,568,1162,1189]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="9" pageNumber="34" unit="in" value="19.0">19 in</quantity>
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the setal line
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<emphasis box="[727,741,1163,1188]" italics="true" pageId="9" pageNumber="34">a</emphasis>
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.
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<emphasis box="[756,882,1162,1188]" italics="true" pageId="9" pageNumber="34">Male pores</emphasis>
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: On 18, in the seminal groves.
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<emphasis box="[268,418,1194,1220]" italics="true" pageId="9" pageNumber="34">Female pores</emphasis>
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: Paired on
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<quantity box="[541,596,1194,1220]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="9" pageNumber="34" unit="in" value="14.0">14 in</quantity>
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between setae
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<emphasis box="[760,801,1195,1220]" italics="true" pageId="9" pageNumber="34">a–a</emphasis>
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(
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<figureCitation box="[815,893,1194,1220]" captionStart="Figs 11–14" captionStartId="10.[144,183,1146,1168]" captionTargetBox="[152,1060,214,1126]" captionTargetId="figure-205@10.[144,1104,204,1138]" captionTargetPageId="10" captionText="Figs 11–14. Dichogaster (Diplothecodrilus) asymmetrica sp. n.: (11) ventral view of the clitellar region; (12) ectal third of the larger penial seta; (13) ectal third of the smaller penial seta; (14) spermatheca. Abbreviations: Fp = female pore, Pr = prostate pore." figureDoi="http://doi.org/10.5281/zenodo.7670438" httpUri="https://zenodo.org/record/7670438/files/figure.png" pageId="9" pageNumber="34">Fig. 11</figureCitation>
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).
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<emphasis box="[914,1134,1194,1220]" italics="true" pageId="9" pageNumber="34">Spermathecal pores</emphasis>
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: Paired in 7/8, 8/
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<quantity box="[361,403,1226,1252]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="9" pageNumber="34" unit="in" value="9.0">9 in</quantity>
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the setal line
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<emphasis box="[559,573,1227,1252]" italics="true" pageId="9" pageNumber="34">a</emphasis>
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.
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</paragraph>
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<paragraph blockId="9.[181,1142,971,1677]" box="[181,407,1267,1293]" pageId="9" pageNumber="34">
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<emphasis box="[181,407,1267,1293]" italics="true" pageId="9" pageNumber="34">Internal characters:</emphasis>
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</paragraph>
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<paragraph blockId="9.[181,1142,971,1677]" lastBlockId="10.[144,1104,1267,1677]" lastPageId="10" lastPageNumber="35" pageId="9" pageNumber="34">
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<taxonomicName authorityName="Perry" authorityYear="1810" box="[181,243,1301,1327]" class="Gastropoda" family="Cymatiidae" genus="Septa" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="34" phylum="Mollusca" rank="genus">
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<emphasis box="[181,243,1301,1327]" italics="true" pageId="9" pageNumber="34">Septa</emphasis>
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</taxonomicName>
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: 9/10–12/13 slightly thickened.
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<emphasis box="[609,708,1300,1326]" italics="true" pageId="9" pageNumber="34">Gizzards</emphasis>
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: Large in 5 and 6.
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<emphasis box="[922,1134,1300,1326]" italics="true" pageId="9" pageNumber="34">Calciferous glands</emphasis>
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: Three pairs in segment 15–17, their size slightly increasing backwards.
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<emphasis italics="true" pageId="9" pageNumber="34">Excretory system</emphasis>
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: Meroic, with 6 meronephridia on each side.
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<emphasis box="[759,835,1365,1390]" italics="true" pageId="9" pageNumber="34">Hearts</emphasis>
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: Paired in segments 10–12.
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<emphasis box="[181,302,1396,1422]" italics="true" pageId="9" pageNumber="34">Typhlosole</emphasis>
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: Lamellar, but due to the poor preservation of the intestine its beginning can not be determined.
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<emphasis box="[398,558,1427,1454]" italics="true" pageId="9" pageNumber="34">Sperm funnels</emphasis>
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: 10, 11.
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<emphasis box="[656,840,1427,1453]" italics="true" pageId="9" pageNumber="34">Seminal vesicles</emphasis>
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: Small in 11, 12.
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<emphasis italics="true" pageId="9" pageNumber="34">Prostatic glands</emphasis>
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: Two pairs in 17 and 19; those in 17 are larger, in 19 smaller.
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<emphasis box="[958,1097,1459,1485]" italics="true" pageId="9" pageNumber="34">Penial setae</emphasis>
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: Of
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<specimenCount box="[181,292,1491,1517]" pageId="9" pageNumber="34" type="generic" typeStatus="types">two types</specimenCount>
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, the longer about
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||
<quantity box="[491,577,1491,1517]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="34" unit="mm" value="2.3">2.3 mm</quantity>
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||
long and
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<quantity box="[687,801,1491,1517]" metricMagnitude="-6" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="34" unit="mm" value="0.006">0.006 mm</quantity>
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wide in the middle, ectal third undulated, tip acute, ornamentation consists of small teeth (
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<figureCitation box="[852,936,1523,1549]" captionStart="Figs 11–14" captionStartId="10.[144,183,1146,1168]" captionTargetBox="[152,1060,214,1126]" captionTargetId="figure-205@10.[144,1104,204,1138]" captionTargetPageId="10" captionText="Figs 11–14. Dichogaster (Diplothecodrilus) asymmetrica sp. n.: (11) ventral view of the clitellar region; (12) ectal third of the larger penial seta; (13) ectal third of the smaller penial seta; (14) spermatheca. Abbreviations: Fp = female pore, Pr = prostate pore." figureDoi="http://doi.org/10.5281/zenodo.7670438" httpUri="https://zenodo.org/record/7670438/files/figure.png" pageId="9" pageNumber="34">Fig. 12</figureCitation>
|
||
). The smaller seta
|
||
<quantity box="[181,264,1554,1581]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="34" unit="mm" value="2.1">2.1 mm</quantity>
|
||
long, diameter
|
||
<quantity box="[434,545,1554,1581]" metricMagnitude="-6" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="34" unit="mm" value="0.006">0.006 mm</quantity>
|
||
in the middle, ectal third slightly undulated, tip spoonshaped, ornamentation minute teeth (
|
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<figureCitation box="[599,683,1586,1612]" captionStart="Figs 11–14" captionStartId="10.[144,183,1146,1168]" captionTargetBox="[152,1060,214,1126]" captionTargetId="figure-205@10.[144,1104,204,1138]" captionTargetPageId="10" captionText="Figs 11–14. Dichogaster (Diplothecodrilus) asymmetrica sp. n.: (11) ventral view of the clitellar region; (12) ectal third of the larger penial seta; (13) ectal third of the smaller penial seta; (14) spermatheca. Abbreviations: Fp = female pore, Pr = prostate pore." figureDoi="http://doi.org/10.5281/zenodo.7670438" httpUri="https://zenodo.org/record/7670438/files/figure.png" pageId="9" pageNumber="34">Fig. 13</figureCitation>
|
||
).
|
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<emphasis box="[704,863,1586,1612]" italics="true" pageId="9" pageNumber="34">Spermathecae</emphasis>
|
||
: Two pairs in segment 8 and 9. The ampulla subdivided with smaller ectal and larger ental part. The duct as long as the ental ampulla part joins asymmetrically at the junction of the two ampulla parts. The ectal ampulla part bears a small unilocular diverticulum (
|
||
<figureCitation box="[835,914,1267,1294]" captionStart="Figs 11–14" captionStartId="10.[144,183,1146,1168]" captionTargetBox="[152,1060,214,1126]" captionTargetId="figure-205@10.[144,1104,204,1138]" captionTargetPageId="10" captionText="Figs 11–14. Dichogaster (Diplothecodrilus) asymmetrica sp. n.: (11) ventral view of the clitellar region; (12) ectal third of the larger penial seta; (13) ectal third of the smaller penial seta; (14) spermatheca. Abbreviations: Fp = female pore, Pr = prostate pore." figureDoi="http://doi.org/10.5281/zenodo.7670438" httpUri="https://zenodo.org/record/7670438/files/figure.png" pageId="10" pageNumber="35">Fig.14</figureCitation>
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||
.).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7670438" ID-Zenodo-Dep="7670438" httpUri="https://zenodo.org/record/7670438/files/figure.png" pageId="10" pageNumber="35" startId="10.[144,183,1146,1168]" targetBox="[152,1060,214,1126]" targetPageId="10">
|
||
<paragraph blockId="10.[144,1105,1146,1216]" pageId="10" pageNumber="35">
|
||
Figs 11–14.
|
||
<taxonomicName authority="Csuzdi & Sherlock & Kouete & Doherty-Bone, 2015" authorityName="Csuzdi & Sherlock & Kouete & Doherty-Bone" authorityYear="2015" box="[259,669,1146,1168]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="35" phylum="Annelida" rank="species" species="asymmetrica" status="sp. n." subGenus="Diplothecodrilus">
|
||
<emphasis box="[259,371,1146,1167]" italics="true" pageId="10" pageNumber="35">Dichogaster</emphasis>
|
||
(
|
||
<emphasis box="[385,538,1146,1167]" italics="true" pageId="10" pageNumber="35">Diplothecodrilus</emphasis>
|
||
)
|
||
<emphasis box="[553,669,1147,1168]" italics="true" pageId="10" pageNumber="35">asymmetrica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[675,728,1147,1168]" pageId="10" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
|
||
: (11) ventral view of the clitellar region; (12) ectal third of the larger penial seta; (13) ectal third of the smaller penial seta; (14) spermatheca. Abbreviations: Fp = female pore, Pr = prostate pore.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="10.[144,1104,1267,1677]" pageId="10" pageNumber="35">
|
||
<materialsCitation collectingDate="2012-06-15" collectionCode="T, HNHM" country="Cameroon" elevation="2895" location="Bone" municipality="Mt. Oku" pageId="10" pageNumber="35" specimenCode="NHM 2013.448, 450" specimenCount="4" stateProvince="North-West" typeStatus="syntype">
|
||
<typeStatus box="[144,231,1303,1324]" pageId="10" pageNumber="35">Syntypes</typeStatus>
|
||
:
|
||
<collectingCountry box="[239,372,1303,1324]" name="Cameroon" pageId="10" pageNumber="35">CAMEROON</collectingCountry>
|
||
:
|
||
<collectingRegion box="[380,559,1303,1325]" country="Cameroon" name="North-West" pageId="10" pageNumber="35">North-West Region</collectingRegion>
|
||
, summit of
|
||
<collectingMunicipality box="[666,745,1303,1325]" pageId="10" pageNumber="35">Mt. Oku</collectingMunicipality>
|
||
,
|
||
<quantity box="[754,882,1303,1325]" metricMagnitude="3" metricUnit="m" metricValue="2.8945" metricValueMax="2.9219999999999997" metricValueMin="2.867" pageId="10" pageNumber="35" unit="m" value="2894.5" valueMax="2922.0" valueMin="2867.0">
|
||
<elevation box="[754,882,1303,1325]" metricMagnitude="3" metricUnit="m" metricValue="2.8945" metricValueMax="2.9219999999999997" metricValueMin="2.867" pageId="10" pageNumber="35" unit="m" value="2894.5" valueMax="2922.0" valueMin="2867.0">2867–2922 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date box="[891,990,1303,1325]" pageId="10" pageNumber="35" value="2012-06-15">
|
||
<collectingDate box="[891,990,1303,1325]" pageId="10" pageNumber="35" value="2012-06-15">15.vi.2012</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode box="[998,1013,1303,1324]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="35">T</collectionCode>
|
||
.
|
||
<location LSID="urn:lsid:plazi:treatment:03AF87882654A670228DFDB9FBFFD30E:8ED960452657A67122E3FA94FF35D74C" box="[144,192,1327,1348]" country="Cameroon" municipality="Mt. Oku" name="Bone" pageId="10" pageNumber="35" stateProvince="North-West">DohertyBone</location>
|
||
(
|
||
<specimenCode box="[203,349,1327,1349]" collectionCode="NHM" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="10" pageNumber="35" type="Herbarium">NHM 2013.448</specimenCode>
|
||
-
|
||
<specimenCode box="[355,389,1327,1349]" collectionCode="NHM" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="10" pageNumber="35" type="Herbarium">450</specimenCode>
|
||
<specimenCount box="[395,442,1327,1348]" pageId="10" pageNumber="35" type="generic">3 ex.</specimenCount>
|
||
,
|
||
<collectionCode box="[451,520,1327,1348]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="35">HNHM</collectionCode>
|
||
AF/5574
|
||
<specimenCount box="[611,658,1327,1348]" pageId="10" pageNumber="35" type="generic">1 ex.</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="10.[144,1104,1267,1677]" pageId="10" pageNumber="35">
|
||
Remarks:
|
||
<taxonomicName authority="Csuzdi & Sherlock & Kouete & Doherty-Bone, 2015" authorityName="Csuzdi & Sherlock & Kouete & Doherty-Bone" authorityYear="2015" box="[261,773,1363,1390]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="35" phylum="Annelida" rank="species" species="asymmetrica" status="sp. n." subGenus="Diplothecodrilus">
|
||
<emphasis box="[261,399,1363,1389]" italics="true" pageId="10" pageNumber="35">Dichogaster</emphasis>
|
||
(
|
||
<emphasis box="[420,610,1363,1389]" italics="true" pageId="10" pageNumber="35">Diplothecodrilus</emphasis>
|
||
)
|
||
<emphasis box="[630,773,1364,1389]" italics="true" pageId="10" pageNumber="35">asymmetrica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[783,847,1364,1390]" pageId="10" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
|
||
belongs to the
|
||
<emphasis box="[1032,1104,1363,1389]" italics="true" pageId="10" pageNumber="35">bolaui</emphasis>
|
||
species group (Csuzdi 2010) characterized by the presence of dimorph penial setae; however, it differs from all other species in this group by the asymmetric shape of the spermathecae.
|
||
</paragraph>
|
||
<paragraph blockId="10.[144,1104,1267,1677]" pageId="10" pageNumber="35">
|
||
Habitat as for
|
||
<taxonomicName authorityName="Csuzdi & Sherlock & Kouete & Doherty-Bone" authorityYear="2015" box="[329,570,1491,1517]" class="Clitellata" family="Eudrilidae" genus="Okudrilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="10" pageNumber="35" phylum="Annelida" rank="species" species="monticolus">
|
||
<emphasis box="[329,570,1491,1517]" italics="true" pageId="10" pageNumber="35">Okudrilus monticolus</emphasis>
|
||
</taxonomicName>
|
||
. As this species is smaller and less conspicuous than
|
||
<taxonomicName authorityName="Csuzdi & Sherlock & Kouete & Doherty-Bone" authorityYear="2015" box="[197,353,1523,1550]" class="Clitellata" family="Eudrilidae" genus="Okudrilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="10" pageNumber="35" phylum="Annelida" rank="species" species="monticolus">
|
||
<emphasis box="[197,353,1523,1550]" italics="true" pageId="10" pageNumber="35">O. monticolus</emphasis>
|
||
</taxonomicName>
|
||
(hence, it has a lower likelihood of collection), the authors are more uncertain as to its occurrence in other habitats on Mount Oku.
|
||
</paragraph>
|
||
<paragraph blockId="10.[144,1104,1267,1677]" lastBlockId="11.[181,1141,204,262]" lastPageId="11" lastPageNumber="36" pageId="10" pageNumber="35">
|
||
The subfamily
|
||
<taxonomicName authorityName="Michaelsen" authorityYear="1897" box="[341,489,1588,1614]" class="Clitellata" family="Acanthodrilidae" kingdom="Animalia" order="Crassiclitellata" pageId="10" pageNumber="35" phylum="Annelida" rank="subFamily" subFamily="Benhamiinae">Benhamiinae</taxonomicName>
|
||
is regarded by some authors (e.g.
|
||
<bibRefCitation author="BLAKEMORE, R. J." box="[877,1065,1588,1614]" pageId="10" pageNumber="35" pagination="107 - 127" refId="ref5914" refString="BLAKEMORE, R. J. 2013. The major megadrile families of the world reviewed again on their taxonomic types (Annelida: Oligochaeta: Megadrilacea). Opuscula Zoologica, Budapest 44 (2): 107 - 127." type="journal article" year="2013">Blakemore 2013</bibRefCitation>
|
||
) as belonging to the family
|
||
<taxonomicName authority=", Michaelsen 1900" authorityName=", Michaelsen" authorityYear="1900" box="[403,766,1619,1645]" class="Clitellata" family="Octochaetidae" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="35" phylum="Annelida" rank="family">Octochaetidae, Michaelsen 1900</taxonomicName>
|
||
. However, both morphological and molecular data show that
|
||
<taxonomicName authorityName=", Michaelsen" authorityYear="1900" box="[484,644,1651,1677]" class="Clitellata" family="Octochaetidae" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="35" phylum="Annelida" rank="family">Octochaetidae</taxonomicName>
|
||
in its classical sense is polyphyletic and
|
||
<taxonomicName authorityName="Michaelsen" authorityYear="1897" box="[181,328,204,230]" class="Clitellata" family="Acanthodrilidae" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="35" phylum="Annelida" rank="subFamily" subFamily="Benhamiinae">Benhamiinae</taxonomicName>
|
||
is related to the classical
|
||
<taxonomicName authorityName="Claus" authorityYear="1880" box="[602,781,204,230]" class="Clitellata" family="Acanthodrilidae" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="36" phylum="Annelida" rank="family">Acanthodrilidae</taxonomicName>
|
||
species (see Csuzdi 2010;
|
||
<bibRefCitation author="JAMES, S. W. & DAVIDSON, S. K." pageId="11" pageNumber="36" pagination="213 - 229" refId="ref6287" refString="JAMES, S. W. & DAVIDSON, S. K. 2012. Molecular phylogeny of earthworms (Annelida: Crassiclitellata) based on 28 S, 18 S and 16 S gene sequences. Invertebrate Systematics 26: 213 - 229." type="journal article" year="2012">James & Davidson 2012</bibRefCitation>
|
||
); therefore, we maintain
|
||
<taxonomicName authorityName="Michaelsen" authorityYear="1897" box="[664,812,236,262]" class="Clitellata" family="Acanthodrilidae" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="35" phylum="Annelida" rank="subFamily" subFamily="Benhamiinae">Benhamiinae</taxonomicName>
|
||
in
|
||
<taxonomicName authorityName="Claus" authorityYear="1880" box="[846,1030,236,262]" class="Clitellata" family="Acanthodrilidae" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="36" phylum="Annelida" rank="family">Acanthodrilidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |