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<document id="C3C714F15B2A255F909B0520539230E0" ID-DOI="10.5281/zenodo.5351598" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351598" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630206282112" checkinUser="tatiana" docAuthor="Jäch, Manfred A., Díaz, Juan A. &amp; Skale, Andre" docDate="2013" docId="03AF8781EA3C293ECF3ABF641E5ADCFB" docLanguage="en" docName="61rbz053-071.pdf" docOrigin="Raffles Bulletin of Zoology 61 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Hydraena (Hydraenopsis) formula Orchymont 1932" docType="treatment" docVersion="4" lastPageNumber="55" masterDocId="FF96FFF9EA3D293CCC79BD781C2BDF4E" masterDocTitle="The Hydraenidae (Coleoptera) Of The Republic Of Singapore" masterLastPageNumber="71" masterPageNumber="53" pageNumber="54" updateTime="1699184966419" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="8CDEA7432E03D67C249EB4F3F20BAFDA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6D3161BB1C139D0795E242DDA1C33430">
<mods:title id="97F8E5BF8CD9A82D1EF7F4BC3D577515">The Hydraenidae (Coleoptera) Of The Republic Of Singapore</mods:title>
</mods:titleInfo>
<mods:name id="5D73591582543C1F6976A7B349DD17A4" type="personal">
<mods:role id="EA4F48C02C5402930A4228CFC1180752">
<mods:roleTerm id="45D098FAA0CFFE0C71AE12B698520D03">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="15606040A54C9C3383A9BB7E6688631A">Jäch, Manfred A.</mods:namePart>
</mods:name>
<mods:name id="411597AA2CA2E9AEB54F4DF777C4AF32" type="personal">
<mods:role id="9245DEEB8BA2F84B1E6F81FEAEE27C5C">
<mods:roleTerm id="CA5D2EC76E6A1E7C66E287A7D21A0B01">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7D6038DB4777D55CD8D5A15342080DD0">Díaz, Juan A.</mods:namePart>
</mods:name>
<mods:name id="30ED5FF148B7A0CA2A579901A60BB55B" type="personal">
<mods:role id="DDA1A857F6249B5CC1C8B573CC323E39">
<mods:roleTerm id="F624BBE408BCF624520413EFD61E0FDF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="845C2199A7D8FA8EAC6453B6CF815C89">Skale, Andre</mods:namePart>
</mods:name>
<mods:typeOfResource id="C2751523F628E2DD8FEF507EF6201CD4">text</mods:typeOfResource>
<mods:relatedItem id="D2BAFE938EF12BFD0E5041864E9033F0" type="host">
<mods:titleInfo id="DE85671DD73AD9BAD875F30E9C06838A">
<mods:title id="F1B16BBAF60D849565E8F85074F3B8CA">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="5D511AE0DE08EEBE733FA445F94CA617">
<mods:date id="49A0DED811C64BEBC5404FC3A2AE0F6F">2013</mods:date>
<mods:detail id="646C1243B3F75E22778CF3D3C1C5E982" type="pubDate">
<mods:number id="467020DF032507A05EF767467F9E2DEF">2013-02-28</mods:number>
</mods:detail>
<mods:detail id="3F5222A4986FF86E30D29EE5A4DA9146" type="volume">
<mods:number id="B552B3C09FD0AC6A31E027BEFBD599AB">61</mods:number>
</mods:detail>
<mods:detail id="2ACBE819C7B404B128AFB372DE46CF43" type="issue">
<mods:number id="49129D45D4E165C53083F68E61ED01D1">1</mods:number>
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<mods:start id="11F7DFF5B97AE544816572E3E04BB7C4">53</mods:start>
<mods:end id="5212CB4532E730691A6AC996711793A5">71</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="F63C2AF1A6066A86B537563F2E5B4661">journal article</mods:classification>
<mods:identifier id="39A53B194C96ACBF4672FF01824BE688" type="DOI">10.5281/zenodo.5351598</mods:identifier>
<mods:identifier id="781B89E4E533F4D10FC7D14A32BDA928" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="78AC264903C6E442FE91656FEF0F873A" type="Zenodo-Dep">5351598</mods:identifier>
</mods:mods>
<treatment id="03AF8781EA3C293ECF3ABF641E5ADCFB" ID-DOI="http://doi.org/10.5281/zenodo.5449811" ID-Zenodo-Dep="5449811" LSID="urn:lsid:plazi:treatment:03AF8781EA3C293ECF3ABF641E5ADCFB" httpUri="http://treatment.plazi.org/id/03AF8781EA3C293ECF3ABF641E5ADCFB" lastPageId="2" lastPageNumber="55" pageId="1" pageNumber="54">
<subSubSection id="C31C651CEA3C293DCF3ABF6419BFDD7B" box="[835,1428,540,565]" pageId="1" pageNumber="54" type="nomenclature">
<paragraph id="8BB93697EA3C293DCF3ABF6419BFDD7B" blockId="1.[835,1428,540,597]" box="[835,1428,540,565]" pageId="1" pageNumber="54">
<heading id="D0F181FBEA3C293DCF3ABF6419BFDD7B" bold="true" box="[835,1428,540,565]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="54" reason="2">
<taxonomicName id="4C064D14EA3C293DCF3ABF6419BFDD7B" ID-CoL="6MHGM" authority="Orchymont, 1932" authorityName="Orchymont" authorityYear="1932" box="[835,1428,540,565]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="formula" subGenus="Hydraenopsis">
<emphasis id="B972EA85EA3C293DCF3ABF6419BFDD7B" bold="true" box="[835,1428,540,565]" pageId="1" pageNumber="54">
<emphasis id="B972EA85EA3C293DCF3ABF641F9ADD7B" bold="true" box="[835,945,540,565]" italics="true" pageId="1" pageNumber="54">Hydraena</emphasis>
(
<emphasis id="B972EA85EA3C293DCFBBBF641872DD7B" bold="true" box="[962,1113,540,565]" italics="true" pageId="1" pageNumber="54">Hydraenopsis</emphasis>
)
<emphasis id="B972EA85EA3C293DC812BF6418EFDD7B" bold="true" box="[1131,1220,540,565]" italics="true" pageId="1" pageNumber="54">formula</emphasis>
<bibRefCitation id="EF974B66EA3C293DC8B4BF6519BFDD7B" author="Orchymont, A" box="[1229,1428,541,565]" pageId="1" pageNumber="54" pagination="623 - 714" refId="ref11600" refString="Orchymont, A. d', 1932. LIII. Zur Kenntnis der Kolbenwasserkafer (Palpicornia) von Sumatra, Java und Bali. Archiv fur Hydrobiologie, Supplement, 9: 623 - 714, pls. XIV - XIX." type="journal article" year="1932">Orchymont, 1932</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA3C293DCF86BF4518F2DD1B" box="[1023,1241,573,597]" pageId="1" pageNumber="54" type="description">
<paragraph id="8BB93697EA3C293DCF86BF4518F2DD1B" blockId="1.[835,1428,540,597]" box="[1023,1241,573,597]" pageId="1" pageNumber="54">
(
<figureCitation id="133D2A12EA3C293DC87EBF45187FDD1B" box="[1031,1108,573,597]" captionStart="Fig" captionStartId="8.[136,171,2037,2059]" captionTargetBox="[186,1444,192,1986]" captionTargetId="figure-8@8.[125,1459,174,2004]" captionTargetPageId="8" captionText="Fig. 9. Hydraena formula: a, aedeagus in lateral view (setae of both parameres omitted); b, aedeagus, dorsal view; c, aedeagus, ventral view (setae of left paramere omitted in b, c); d, left paramere; e, male sternite X and spiculum; f, gonocoxite; g, female tergite X; h, i, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351610" httpUri="https://zenodo.org/record/5351610/files/figure.png" pageId="1" pageNumber="54">Figs. 9</figureCitation>
,
<figureCitation id="133D2A12EA3C293DC81BBF451854DD1B" box="[1122,1151,573,597]" captionStart="Figs" captionStartId="12.[137,181,2091,2113]" captionTargetBox="[138,1476,1428,2061]" captionTargetId="figure-8@12.[134,1479,1425,2064]" captionTargetPageId="12" captionText="Figs. 12–14. Meso- and metaventrite of: 12, Hydraena hendrichi; 13, H. singaporensis; and 14, H. formula." figureDoi="http://doi.org/10.5281/zenodo.5351616" httpUri="https://zenodo.org/record/5351616/files/figure.png" pageId="1" pageNumber="54">14</figureCitation>
,
<figureCitation id="133D2A12EA3C293DC8F5BF451882DD1B" box="[1164,1193,573,597]" captionStart="Fig" captionStartId="14.[137,172,2090,2112]" captionTargetBox="[137,1470,1208,2063]" captionTargetId="figure-8@14.[137,1478,1202,2067]" captionTargetPageId="14" captionText="Fig. 15. Geographical map of Singapore, showing distribution of Hydraenidae." figureDoi="http://doi.org/10.5281/zenodo.5351618" httpUri="https://zenodo.org/record/5351618/files/figure.png" pageId="1" pageNumber="54">15</figureCitation>
,
<figureCitation id="133D2A12EA3C293DC8CFBF4518FADD1B" box="[1206,1233,573,597]" captionStart="Fig" captionStartId="16.[136,172,2061,2083]" captionTargetBox="[279,1324,1376,2039]" captionTargetId="figure-537@16.[276,1328,1373,2041]" captionTargetPageId="16" captionText="Fig. 17. Nee Soon Swamp Forest, Singapore; C.M. Yang on right. Four species, H. formula, H. jacobsoni, H. paulmoritz, and H. singaporensis were collected from this site. (Photograph by: L. Hendrich)." figureDoi="http://doi.org/10.5281/zenodo.5351622" httpUri="https://zenodo.org/record/5351622/files/figure.png" pageId="1" pageNumber="54">17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA3C293DCF54BF051FE3DDE0" pageId="1" pageNumber="54" type="reference_group">
<paragraph id="8BB93697EA3C293DCF54BF051FE3DDE0" blockId="1.[813,1451,636,687]" pageId="1" pageNumber="54">
<treatmentCitationGroup id="AB1611B9EA3C293DCF54BF051FE3DDE0" pageId="1" pageNumber="54">
<treatmentCitation id="0AA71086EA3C293DCF54BF0518F3DDDC" author="Orchymont, A" box="[813,1240,636,658]" page="655" pageId="1" pageNumber="54" year="1932">
<taxonomicName id="4C064D14EA3C293DCF54BF0518F3DDDC" ID-CoL="6MHGM" authority="Orchymont, 1932: 655" authorityName="Orchymont" authorityPageNumber="655" authorityYear="1932" box="[813,1240,636,658]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="formula">
<emphasis id="B972EA85EA3C293DCF54BF051FCDDDDF" box="[813,998,637,657]" italics="true" pageId="1" pageNumber="54">Hydraena formula</emphasis>
<bibRefCitation id="EF974B66EA3C293DCF96BF0418F3DDDC" author="Orchymont, A" box="[1007,1240,636,658]" pageId="1" pageNumber="54" pagination="623 - 714" refId="ref11600" refString="Orchymont, A. d', 1932. LIII. Zur Kenntnis der Kolbenwasserkafer (Palpicornia) von Sumatra, Java und Bali. Archiv fur Hydrobiologie, Supplement, 9: 623 - 714, pls. XIV - XIX." type="journal article" year="1932">Orchymont, 1932: 655</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<bibRefCitation id="EF974B66EA3C293DC89CBF041958DDDF" author="Hansen, M" box="[1253,1395,636,657]" pageId="1" pageNumber="54" refId="ref11156" refString="Hansen, M., 1998. Hydraenidae. In: Hansen, M. (ed.), World Catalogue of Insects 1. Apollo Books, Stenstrup. 168 pp." type="book" year="1998">Hansen, 1998</bibRefCitation>
;
<bibRefCitation id="EF974B66EA3C293DC9F9BF041FEFDDE0" author="Jach, M &amp; Beutel, J" pageId="1" pageNumber="54" pagination="177 - 258" refId="ref11287" refString="Jach, M. A., R. G. Beutel, J. A. Diaz &amp; J. Kodada, 2000. Subgeneric classification, description of head structures, and world check list of Hydraena Kugelann (Insecta: Coleoptera: Hydraenidae). Annalen des Naturhistorischen Museums in Wien, B, 102: 177 - 258." type="journal article" year="2000">Jäch et al., 2000</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA3C293DCF54BFAB19A5DB9F" pageId="1" pageNumber="54" type="materials_examined">
<paragraph id="8BB93697EA3C293DCF54BFAB1FAEDC4B" blockId="1.[813,1449,721,773]" pageId="1" pageNumber="54">
<materialsCitation id="3B6E3CCAEA3C293DCF54BFAB1FAADC4B" country="Vietnam" location="Lac Tho" pageId="1" pageNumber="54" specimenCount="1" stateProvince="Hoa Binh" typeStatus="holotype">
<emphasis id="B972EA85EA3C293DCF54BFAB1FEEDDA9" bold="true" box="[813,965,721,744]" italics="true" pageId="1" pageNumber="54">
<typeStatus id="54BD8835EA3C293DCF54BFAB1F4ADDA6" box="[813,865,723,744]" pageId="1" pageNumber="54">Type</typeStatus>
locality.
</emphasis>
—
<location id="8ED9604CEA3C293DCF8FBFAB187EDDA6" LSID="urn:lsid:plazi:treatment:03AF8781EA3C293ECF3ABF641E5ADCFB:8ED9604CEA3C293DCF8FBFAB187EDDA6" box="[1014,1109,722,744]" country="Vietnam" name="Lac Tho" pageId="1" pageNumber="54" stateProvince="Hoa Binh">Lac Thô</location>
,
<collectingRegion id="49C2F875EA3C293DC81FBFAA196BDDA6" box="[1126,1344,722,744]" country="Vietnam" name="Hoa Binh" pageId="1" pageNumber="54">Hòa Bình Province</collectingRegion>
, northern
<collectingCountry id="F3117607EA3C293DCF54BF971FAADC4B" box="[813,897,751,773]" name="Vietnam" pageId="1" pageNumber="54">Vietnam</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB93697EA3C293DCF54BE5119A6DC81" blockId="1.[813,1451,808,975]" pageId="1" pageNumber="54">
<emphasis id="B972EA85EA3C293DCF54BE511FF6DC70" bold="true" box="[813,989,808,830]" pageId="1" pageNumber="54">
<emphasis id="B972EA85EA3C293DCF54BE511F96DC70" bold="true" box="[813,957,808,830]" italics="true" pageId="1" pageNumber="54">Type material.</emphasis>
—
</emphasis>
<materialsCitation id="3B6E3CCAEA3C293DCF9FBE51180DDCFF" collectionCode="ISNB" country="Vietnam" determinerName="A. d'Orchymont" location="Tonkin" pageId="1" pageNumber="54" specimenCount="1" typeStatus="holotype">
<emphasis id="B972EA85EA3C293DCF9FBE511868DC70" bold="true" box="[998,1091,809,830]" pageId="1" pageNumber="54">
<typeStatus id="54BD8835EA3C293DCF9FBE511868DC70" box="[998,1091,809,830]" pageId="1" pageNumber="54" type="holotype">Holotype</typeStatus>
</emphasis>
male (
<collectionCode id="ED17AE52EA3C293DC8F0BE5118E8DC70" box="[1161,1219,809,830]" pageId="1" pageNumber="54">ISNB</collectionCode>
, examined): “ ”, “Lac Tho [handwritten]
<collectingCountry id="F3117607EA3C293DCFA7BE3D1809DC15" box="[990,1058,837,859]" name="Vietnam" pageId="1" pageNumber="54">Tonkin</collectingCountry>
De Cooman [printed]”, “
<determinerName id="7E65DA53EA3C293DC976BE3E1980DC15" box="[1295,1451,837,859]" pageId="1" pageNumber="54">A. d’Orchymont</determinerName>
det [printed]
<taxonomicName id="4C064D14EA3C293DCFCCBE1B180EDC36" authorityName="Orchymont" authorityYear="1932" box="[949,1061,866,888]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="formula">H. formula</taxonomicName>
TYPE [handwritten]”, “TYPE” [red], “Prép. micr. N° 710709.13” [green], “AEDEAGUS DRAWN BY P.D.PERKINS” [printed]
</materialsCitation>
.
<materialsCitation id="3B6E3CCAEA3C293DC84BBEE419A2DC81" collectionCode="ISNB" country="Vietnam" determinerName="A. d'Orchymont" location="Tonkin" pageId="1" pageNumber="54" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B972EA85EA3C293DC84BBEE418B1DCFF" bold="true" box="[1074,1178,924,945]" pageId="1" pageNumber="54">
<typeStatus id="54BD8835EA3C293DC84BBEE418B1DCFF" box="[1074,1178,924,945]" pageId="1" pageNumber="54" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<collectionCode id="ED17AE52EA3C293DC8D3BEE418CFDCFF" box="[1194,1252,924,945]" pageId="1" pageNumber="54">ISNB</collectionCode>
, examined):
<specimenCount id="9D00FD1EEA3C293DC91EBEE41980DCFC" box="[1383,1451,924,946]" pageId="1" pageNumber="54" type="male">1 male</specimenCount>
(aedeagus lacking),
<specimenCount id="9D00FD1EEA3C293DCF88BEC11879DC81" box="[1009,1106,953,975]" pageId="1" pageNumber="54" type="female">2 females</specimenCount>
, same locality data as
<typeStatus id="54BD8835EA3C293DC94BBEC119A2DC81" box="[1330,1417,953,975]" pageId="1" pageNumber="54" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB93697EA3C293DCF54BE8919A5DB9F" blockId="1.[813,1452,1009,1234]" pageId="1" pageNumber="54">
<emphasis id="B972EA85EA3C293DCF54BE891872DB49" bold="true" box="[813,1113,1009,1031]" italics="true" pageId="1" pageNumber="54">Additional material examined.</emphasis>
—
<materialsCitation id="3B6E3CCAEA3C293DC802BE8A1913DB6A" collectingDate="1997-04-02" collectionCode="ZRC" collectorName="Lua" country="Singapore" location="Bukit Timah Nature Reserve" pageId="1" pageNumber="54" specimenCount="1" specimenCount-female="1">
<collectingCountry id="F3117607EA3C293DC802BE8A192DDB49" box="[1147,1286,1010,1031]" name="Singapore" pageId="1" pageNumber="54">SINGAPORE</collectingCountry>
:
<specimenCount id="9D00FD1EEA3C293DC977BE8A194BDB46" box="[1294,1376,1010,1032]" pageId="1" pageNumber="54" type="female">1 female</specimenCount>
(
<collectionCode id="ED17AE52EA3C293DC914BE8A198ADB46" box="[1389,1441,1010,1032]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="54">ZRC</collectionCode>
):
<location id="8ED9604CEA3C293DCF54B977187ADB6B" LSID="urn:lsid:plazi:treatment:03AF8781EA3C293ECF3ABF641E5ADCFB:8ED9604CEA3C293DCF54B977187ADB6B" box="[813,1105,1039,1061]" country="Singapore" name="Bukit Timah Nature Reserve" pageId="1" pageNumber="54">Bukit Timah Nature Reserve</location>
, coll.
<collectorName id="26F35341EA3C293DC8EAB9681890DB6B" box="[1171,1211,1040,1061]" pageId="1" pageNumber="54">Lua</collectorName>
,
<date id="FFB81057EA3C293DC8B0B9771913DB6A" box="[1225,1336,1039,1060]" pageId="1" pageNumber="54" value="1997-04-02">
<collectingDate id="EFFCE9BFEA3C293DC8B0B9771913DB6A" box="[1225,1336,1039,1060]" pageId="1" pageNumber="54" value="1997-04-02">2 Apr.1997</collectingDate>
</date>
</materialsCitation>
“NS 224”;
<materialsCitation id="3B6E3CCAEA3C293DCF54B9541980DB0C" box="[813,1451,1068,1090]" collectingDate="1995-12-13" collectionCode="ZRC" collectorName="Lua" country="Singapore" location="Bukit Timah Nature Reserve" pageId="1" pageNumber="54" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D00FD1EEA3C293DCF54B9541FABDB0C" box="[813,896,1068,1090]" pageId="1" pageNumber="54" type="female">1 female</specimenCount>
(
<collectionCode id="ED17AE52EA3C293DCFE9B9541FEFDB0C" box="[912,964,1068,1090]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="54">ZRC</collectionCode>
): same locality, but coll.
<collectorName id="26F35341EA3C293DC8B9B95418CEDB0F" box="[1216,1253,1068,1089]" pageId="1" pageNumber="54">Lua</collectorName>
et al.,
<date id="FFB81057EA3C293DC957B9541980DB0C" box="[1326,1451,1068,1090]" pageId="1" pageNumber="54" value="1995-12-13">
<collectingDate id="EFFCE9BFEA3C293DC957B9541980DB0C" box="[1326,1451,1068,1090]" pageId="1" pageNumber="54" value="1995-12-13">13 Dec.1995</collectingDate>
</date>
</materialsCitation>
“NS 214”;
<materialsCitation id="3B6E3CCAEA3C293DCFECB93118ADDB35" collectingDate="1993-05-27" collectionCode="ZRC" collectorName="C. M. Yang" country="Singapore" location="Rifle Range Road" municipality="Bukit Timah" pageId="1" pageNumber="54" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D00FD1EEA3C293DCFECB9311FC3DB11" box="[917,1000,1097,1119]" pageId="1" pageNumber="54" type="female">1 female</specimenCount>
(
<collectionCode id="ED17AE52EA3C293DCF8DB9311803DB11" box="[1012,1064,1097,1119]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="54">ZRC</collectionCode>
):
<location id="8ED9604CEA3C293DC84EB93118C9DB11" LSID="urn:lsid:plazi:treatment:03AF8781EA3C293ECF3ABF641E5ADCFB:8ED9604CEA3C293DC84EB93118C9DB11" box="[1079,1250,1097,1119]" country="Singapore" municipality="Bukit Timah" name="Rifle Range Road" pageId="1" pageNumber="54">Rifle Range Road</location>
, east of
<collectingMunicipality id="6BDDACEDEA3C293DC957B9311983DB11" box="[1326,1448,1097,1119]" pageId="1" pageNumber="54">Bukit Timah</collectingMunicipality>
, coll.
<collectorName id="26F35341EA3C293DCF25B91E1FE2DB35" box="[860,969,1126,1147]" pageId="1" pageNumber="54">C. M. Yang</collectorName>
et al.,
<date id="FFB81057EA3C293DC87EB91E18ADDB35" box="[1031,1158,1126,1147]" pageId="1" pageNumber="54" value="1993-05-27">
<collectingDate id="EFFCE9BFEA3C293DC87EB91E18ADDB35" box="[1031,1158,1126,1147]" pageId="1" pageNumber="54" value="1993-05-27">27 May 1993</collectingDate>
</date>
</materialsCitation>
“417 ZRC.6.18870 (NS102)”;
<materialsCitation id="3B6E3CCAEA3C293DCF54B9FA1836DBFA" collectingDate="1992-04-16" collectionCode="ZRC" collectorName="D. H. Murphy &amp; C. M. Yang" country="Singapore" location="Nee Soon Swamp Forest" pageId="1" pageNumber="54" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D00FD1EEA3C293DCF54B9FA1FAADBD6" box="[813,897,1154,1176]" pageId="1" pageNumber="54" type="female">1 female</specimenCount>
(
<collectionCode id="ED17AE52EA3C293DCFE9B9FA1FEFDBD6" box="[912,964,1154,1176]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="54">ZRC</collectionCode>
):
<location id="8ED9604CEA3C293DCFAFB9FB18E1DBD6" LSID="urn:lsid:plazi:treatment:03AF8781EA3C293ECF3ABF641E5ADCFB:8ED9604CEA3C293DCFAFB9FB18E1DBD6" box="[982,1226,1154,1176]" country="Singapore" name="Nee Soon Swamp Forest" pageId="1" pageNumber="54">Nee Soon Swamp Forest</location>
, coll.
<collectorName id="26F35341EA3C293DC97EB9FB19BADBD6" box="[1287,1425,1154,1176]" pageId="1" pageNumber="54">D. H. Murphy</collectorName>
&amp;
<collectorName id="26F35341EA3C293DCF54B9E71FB6DBFB" box="[813,925,1183,1205]" pageId="1" pageNumber="54">C. M. Yang</collectorName>
,
<date id="FFB81057EA3C293DCFDFB9E71836DBFA" box="[934,1053,1183,1205]" pageId="1" pageNumber="54" value="1992-04-16">
<collectingDate id="EFFCE9BFEA3C293DCFDFB9E71836DBFA" box="[934,1053,1183,1205]" pageId="1" pageNumber="54" value="1992-04-16">16 Apr.1992</collectingDate>
</date>
</materialsCitation>
“Y790”;
<materialsCitation id="3B6E3CCAEA3C293DC803B9E718D3DB9F" collectingDate="1992-05-06" collectionCode="NMW, ZRC" collectorName="K. L. Yeo" country="Singapore" location="Nee Soon Swamp Forest" pageId="1" pageNumber="54" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D00FD1EEA3C293DC803B9E71896DBFB" box="[1146,1213,1183,1205]" pageId="1" pageNumber="54" type="male">1 male</specimenCount>
,
<specimenCount id="9D00FD1EEA3C293DC8BFB9E71932DBFB" box="[1222,1305,1183,1205]" pageId="1" pageNumber="54" type="female">1 female</specimenCount>
(
<collectionCode id="ED17AE52EA3C293DC95FB9E7194EDBFA" box="[1318,1381,1183,1204]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="54">NMW</collectionCode>
,
<collectionCode id="ED17AE52EA3C293DC917B9E7198ADBFB" box="[1390,1441,1183,1205]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="54">ZRC</collectionCode>
): same locality, but coll.
<collectorName id="26F35341EA3C293DC86BB9C4185EDB9F" box="[1042,1141,1212,1233]" pageId="1" pageNumber="54">K. L. Yeo</collectorName>
,
<date id="FFB81057EA3C293DC8F9B9C418D3DB9F" box="[1152,1272,1212,1234]" pageId="1" pageNumber="54" value="1992-05-06">
<collectingDate id="EFFCE9BFEA3C293DC8F9B9C418D3DB9F" box="[1152,1272,1212,1234]" pageId="1" pageNumber="54" value="1992-05-06">6 May 1992</collectingDate>
</date>
</materialsCitation>
“424 Y794D”.
</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA3C293ECF54B9821E63DDFB" lastPageId="2" lastPageNumber="55" pageId="1" pageNumber="54" type="diagnosis">
<paragraph id="8BB93697EA3C293DCF54B98219A1DA3C" blockId="1.[813,1451,1274,1394]" pageId="1" pageNumber="54">
<emphasis id="B972EA85EA3C293DCF54B9821F89DA5C" bold="true" box="[813,930,1274,1298]" italics="true" pageId="1" pageNumber="54">Diagnosis.</emphasis>
— Habitus and colouration as in
<taxonomicName id="4C064D14EA3C293DC95EB9831FD1DA7C" authority="Jach &amp; Diaz, 1998" authorityName="Jach &amp; Diaz" authorityYear="1998" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="undulata">
<emphasis id="B972EA85EA3C293DC95EB9831980DA5C" box="[1319,1451,1275,1298]" italics="true" pageId="1" pageNumber="54">H. undulata</emphasis>
Jäch &amp; Díaz, 1998
</taxonomicName>
(see Jäch &amp; Díaz, 1998:
<figureCitation id="133D2A12EA3C293DC96FB8621973DA7C" box="[1302,1368,1306,1330]" captionStart="Fig" captionStartId="6.[136,171,2060,2082]" captionTargetBox="[166,1446,190,2011]" captionTargetId="figure-8@6.[43,1493,170,2020]" captionTargetPageId="6" captionText="Fig. 7. Hydraena yangae: a, aedeagus in lateral view; b, aedeagus, dorsal view (setae of left paramere omitted in a, b); c, aedeagus, ventral view (setae of right paramere omitted); d, male sternite X and spiculum; e, gonocoxite; f, female tergite X; g, h, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351606" httpUri="https://zenodo.org/record/5351606/files/figure.png" pageId="1" pageNumber="54">Fig. 7</figureCitation>
),
<quantity id="4CFE9B72EA3C293DC916B8621FB9DA1C" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="1" pageNumber="54" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.40– 1.50 mm</quantity>
long. See original description of the latter for details of external characters (Jäch &amp; Díaz, 1998: 167).
</paragraph>
<paragraph id="8BB93697EA3C293DCF54B8E21815DABC" blockId="1.[813,1451,1434,1522]" pageId="1" pageNumber="54">
Male terminal sternite and spiculum (
<figureCitation id="133D2A12EA3C293DC89CB8E21910DAFC" box="[1253,1339,1434,1458]" captionStart="Fig" captionStartId="8.[136,171,2037,2059]" captionTargetBox="[186,1444,192,1986]" captionTargetId="figure-8@8.[125,1459,174,2004]" captionTargetPageId="8" captionText="Fig. 9. Hydraena formula: a, aedeagus in lateral view (setae of both parameres omitted); b, aedeagus, dorsal view; c, aedeagus, ventral view (setae of left paramere omitted in b, c); d, left paramere; e, male sternite X and spiculum; f, gonocoxite; g, female tergite X; h, i, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351610" httpUri="https://zenodo.org/record/5351610/files/figure.png" pageId="1" pageNumber="54">Fig. 9e</figureCitation>
): Sternite firmly connected with spiculum, resembling a contrabass; apex slightly emarginate.
</paragraph>
<paragraph id="8BB93697EA3C293DCF54BB6218B9D9FC" blockId="1.[813,1452,1562,1714]" pageId="1" pageNumber="54">
Aedeagus (
<figureCitation id="133D2A12EA3C293DCFD1BB621838D97C" box="[936,1043,1562,1586]" captionStart="Fig" captionStartId="8.[136,171,2037,2059]" captionTargetBox="[186,1444,192,1986]" captionTargetId="figure-8@8.[125,1459,174,2004]" captionTargetPageId="8" captionText="Fig. 9. Hydraena formula: a, aedeagus in lateral view (setae of both parameres omitted); b, aedeagus, dorsal view; c, aedeagus, ventral view (setae of left paramere omitted in b, c); d, left paramere; e, male sternite X and spiculum; f, gonocoxite; g, female tergite X; h, i, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351610" httpUri="https://zenodo.org/record/5351610/files/figure.png" pageId="1" pageNumber="54">Fig. 9a–c</figureCitation>
): Very similar to that of
<taxonomicName id="4C064D14EA3C293DC950BB631987D97C" authorityName="Jach &amp; Diaz" authorityYear="1998" box="[1321,1452,1563,1586]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="undulata">
<emphasis id="B972EA85EA3C293DC950BB631987D97C" box="[1321,1452,1563,1586]" italics="true" pageId="1" pageNumber="54">H. undulata</emphasis>
</taxonomicName>
from
<collectingCountry id="F3117607EA3C293DCF1EBB421F9ED91C" box="[871,949,1594,1618]" name="Taiwan" pageId="1" pageNumber="54">Taiwan</collectingCountry>
(see Jäch &amp; Díaz, 1998: Fig. 20a–d). It differs from the latter mainly in the following characteristics: Distal lobe with spiny apex. Both parameres shorter, right one more setose than in
<taxonomicName id="4C064D14EA3C293DC873BBE318A7D9FC" authorityName="Jach &amp; Diaz" authorityYear="1998" box="[1034,1164,1691,1714]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="undulata">
<emphasis id="B972EA85EA3C293DC873BBE318A7D9FC" box="[1034,1164,1691,1714]" italics="true" pageId="1" pageNumber="54">H. undulata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB93697EA3C293DCF54BBA2183FD81C" blockId="1.[813,1452,1754,1874]" pageId="1" pageNumber="54">
Gonocoxite (
<figureCitation id="133D2A12EA3C293DCFC5BBA21822D9BC" box="[956,1033,1754,1778]" captionStart="Fig" captionStartId="8.[136,171,2037,2059]" captionTargetBox="[186,1444,192,1986]" captionTargetId="figure-8@8.[125,1459,174,2004]" captionTargetPageId="8" captionText="Fig. 9. Hydraena formula: a, aedeagus in lateral view (setae of both parameres omitted); b, aedeagus, dorsal view; c, aedeagus, ventral view (setae of left paramere omitted in b, c); d, left paramere; e, male sternite X and spiculum; f, gonocoxite; g, female tergite X; h, i, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351610" httpUri="https://zenodo.org/record/5351610/files/figure.png" pageId="1" pageNumber="54">Fig. 9f</figureCitation>
) and female tergite X (
<figureCitation id="133D2A12EA3C293DC968BBA21949D9BC" box="[1297,1378,1754,1778]" captionStart="Fig" captionStartId="8.[136,171,2037,2059]" captionTargetBox="[186,1444,192,1986]" captionTargetId="figure-8@8.[125,1459,174,2004]" captionTargetPageId="8" captionText="Fig. 9. Hydraena formula: a, aedeagus in lateral view (setae of both parameres omitted); b, aedeagus, dorsal view; c, aedeagus, ventral view (setae of left paramere omitted in b, c); d, left paramere; e, male sternite X and spiculum; f, gonocoxite; g, female tergite X; h, i, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351610" httpUri="https://zenodo.org/record/5351610/files/figure.png" pageId="1" pageNumber="54">
Fig.
<quantity id="4CFE9B72EA3C293DC93EBBA21949D9BC" box="[1351,1378,1754,1778]" metricMagnitude="-3" metricUnit="kg" metricValue="9.0" pageId="1" pageNumber="54" unit="g" value="9.0">9g</quantity>
</figureCitation>
) more or less as in
<taxonomicName id="4C064D14EA3C293DCFBFBB831864D85C" authorityName="Jach &amp; Diaz" authorityYear="1998" box="[966,1103,1787,1810]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="undulata">
<emphasis id="B972EA85EA3C293DCFBFBB831864D85C" box="[966,1103,1787,1810]" italics="true" pageId="1" pageNumber="54">H. undulata</emphasis>
</taxonomicName>
(see Jäch &amp; Díaz, 1998: Fig. 20e). Cavea of inner plate of gonocoxite undivided in the specimens examined.
</paragraph>
<paragraph id="8BB93697EA3C293DCF54BA021F9ED8FC" blockId="1.[813,1451,1914,1970]" pageId="1" pageNumber="54">
Spermatheca (
<figureCitation id="133D2A12EA3C293DCFB5BA02181FD8DC" box="[972,1076,1914,1938]" captionStart="Fig" captionStartId="8.[136,171,2037,2059]" captionTargetBox="[186,1444,192,1986]" captionTargetId="figure-8@8.[125,1459,174,2004]" captionTargetPageId="8" captionText="Fig. 9. Hydraena formula: a, aedeagus in lateral view (setae of both parameres omitted); b, aedeagus, dorsal view; c, aedeagus, ventral view (setae of left paramere omitted in b, c); d, left paramere; e, male sternite X and spiculum; f, gonocoxite; g, female tergite X; h, i, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5351610" httpUri="https://zenodo.org/record/5351610/files/figure.png" pageId="1" pageNumber="54">Fig. 9h–i</figureCitation>
) not distinguishable from that of
<taxonomicName id="4C064D14EA3C293DCF54BAE31F85D8FC" authorityName="Jach &amp; Diaz" authorityYear="1998" box="[813,942,1947,1970]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="undulata">
<emphasis id="B972EA85EA3C293DCF54BAE31F85D8FC" box="[813,942,1947,1970]" italics="true" pageId="1" pageNumber="54">H. undulata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB93697EA3C293ECF54BAA21E99DEDB" blockId="1.[813,1451,2010,2098]" lastBlockId="2.[136,775,189,405]" lastPageId="2" lastPageNumber="55" pageId="1" pageNumber="54">
Secondary sexual characters: Male forecoxa with a very small projection directed ventrad. Male mesoventrite slightly more deeply impressed between mesoventral disc and mesoventral process. Male mesoventral process more slender. Male metaventral plaques (
<figureCitation id="133D2A12EA3F293ECE55BDA51EA9DFBB" box="[556,642,221,245]" captionStart="Figs" captionStartId="12.[137,181,2091,2113]" captionTargetBox="[138,1476,1428,2061]" captionTargetId="figure-8@12.[134,1479,1425,2064]" captionTargetPageId="12" captionText="Figs. 12–14. Meso- and metaventrite of: 12, Hydraena hendrichi; 13, H. singaporensis; and 14, H. formula." figureDoi="http://doi.org/10.5281/zenodo.5351616" httpUri="https://zenodo.org/record/5351616/files/figure.png" pageId="2" pageNumber="55">Fig. 14</figureCitation>
) very thin, carinate. Male tergite X widely and asymmetrically excavate apically (excavation distinctly deeper on left side); dorsal apical margin asymmetrically and sinuously emarginate (more strongly emarginate on left side); corners subacute, right corner hardly noticeably longer than left one.
</paragraph>
<paragraph id="8BB93697EA3F293ECCF1BCC51ECADD1B" blockId="2.[136,775,445,598]" pageId="2" pageNumber="55">
Variability: In the specimens from Sumatra, the apical segment of the maxillary palpi is brownish subapically. In most of the few specimens examined from
<collectingCountry id="F3117607EA3F293ECEE1BC851F2DDD5B" box="[664,774,509,533]" name="Singapore" pageId="2" pageNumber="55">Singapore</collectingCountry>
the apical segment of the maxillary palpi is more or less unicoloured yellowish, which may be due to tenerality.
</paragraph>
<paragraph id="8BB93697EA3F293ECCF1BF061E63DDFB" blockId="2.[136,775,637,693]" pageId="2" pageNumber="55">In some of the males, the hind tibia is hardly noticeably curved subapically and widened apically.</paragraph>
</subSubSection>
<subSubSection id="C31C651CEA3F293ECCF1BFA41E5ADCFB" pageId="2" pageNumber="55" type="distribution">
<paragraph id="8BB93697EA3F293ECCF1BFA41CDCDC7B" blockId="2.[136,775,732,821]" pageId="2" pageNumber="55">
<emphasis id="B972EA85EA3F293ECCF1BFA41D3EDDBB" bold="true" box="[136,277,732,757]" italics="true" pageId="2" pageNumber="55">Distribution.</emphasis>
— This species is obviously wide-spread in the Oriental Region, from
<collectingCountry id="F3117607EA3F293ECDE7BF851DF5DC5B" box="[414,478,765,789]" name="Nepal" pageId="2" pageNumber="55">Nepal</collectingCountry>
to South
<collectingCountry id="F3117607EA3F293ECE3BBF851EA9DC5B" box="[578,642,765,789]" name="China" pageId="2" pageNumber="55">China</collectingCountry>
and western
<collectingCountry id="F3117607EA3F293ECCF1BE651CD8DC7B" box="[136,243,797,821]" name="Indonesia" pageId="2" pageNumber="55">Indonesia</collectingCountry>
.
</paragraph>
<paragraph id="8BB93697EA3F293ECCF1BE261E5ADCFB" blockId="2.[136,775,861,949]" pageId="2" pageNumber="55">
In
<collectingCountry id="F3117607EA3F293ECCDDBE251D3BDC3B" box="[164,272,861,885]" name="Singapore" pageId="2" pageNumber="55">Singapore</collectingCountry>
(
<figureCitation id="133D2A12EA3F293ECD64BE251D40DC3B" box="[285,363,861,885]" captionStart="Fig" captionStartId="14.[137,172,2090,2112]" captionTargetBox="[137,1470,1208,2063]" captionTargetId="figure-8@14.[137,1478,1202,2067]" captionTargetPageId="14" captionText="Fig. 15. Geographical map of Singapore, showing distribution of Hydraenidae." figureDoi="http://doi.org/10.5281/zenodo.5351618" httpUri="https://zenodo.org/record/5351618/files/figure.png" pageId="2" pageNumber="55">Fig. 15</figureCitation>
),
<taxonomicName id="4C064D14EA3F293ECD04BE261DDADC3B" authorityName="Orchymont" authorityYear="1932" box="[381,497,862,885]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="formula">
<emphasis id="B972EA85EA3F293ECD04BE261DDADC3B" box="[381,497,862,885]" italics="true" pageId="2" pageNumber="55">H. formula</emphasis>
</taxonomicName>
was collected in the Bukit Timah Reserve, near the Rifle Range Road (east of Bukit Timah), and in the Nee Soon Swamp Forest.
</paragraph>
</subSubSection>
</treatment>
</document>
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