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<document id="D0B7940B9B075241102C64971E014A05" ID-DOI="10.5281/zenodo.277431" ID-GBIF-Dataset="7b1e0f87-7063-4230-840a-2a8b1afe14a0" ID-ISSN="1175-5326" ID-Zenodo-Dep="277431" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460303384262" checkinUser="plazi" docAuthor="Lim, Jongok, Lee, Jongwook, Koh, Sanghyun, Lee, Bongwoo, Azevedo, Celso O. &amp; Lee, Seunghwan" docDate="2011" docId="03A9FE14D37CFFB5FF6DF902FB3218FA" docLanguage="en" docName="zt02866p038.pdf" docOrigin="Zootaxa 2866" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Epyris hangunensis Terayama 2005" docType="treatment" docVersion="7" lastPageNumber="8" masterDocId="FF90866CD37AFFB2FFFAFFFAFFEF1A7E" masterDocTitle="Taxonomy of Epyris Westwood (Hymenoptera: Bethylidae) from Korea, with the descriptions of ten new species" masterLastPageNumber="38" masterPageNumber="1" pageNumber="7" updateTime="1698252956732" updateUser="plazi">
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<mods:title id="0D78292E609A841B19963AAE0E729E71">Taxonomy of Epyris Westwood (Hymenoptera: Bethylidae) from Korea, with the descriptions of ten new species</mods:title>
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<mods:namePart id="F05F96E7A3D8A7534DAE6F9905C96153">Lim, Jongok</mods:namePart>
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<mods:namePart id="6F995136B99C0665EE0D1E108DFF3AF2">Lee, Jongwook</mods:namePart>
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<mods:namePart id="1B49545E27947AE021175AAFD2C13BD2">Koh, Sanghyun</mods:namePart>
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<mods:namePart id="1FED26A7C2D03C26C8DC7F7F2A4898E8">Lee, Bongwoo</mods:namePart>
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<mods:namePart id="52FE91F3DEE6B0527F4FACA3CD26689C">Azevedo, Celso O.</mods:namePart>
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<mods:namePart id="94858E4D5A20561537858A0F3B0D08D0">Lee, Seunghwan</mods:namePart>
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<mods:date id="FF747B67B063227F2F754679675C46B5">2011</mods:date>
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<treatment id="03A9FE14D37CFFB5FF6DF902FB3218FA" ID-DOI="http://doi.org/10.5281/zenodo.6187620" ID-GBIF-Taxon="119400101" ID-Zenodo-Dep="6187620" LSID="urn:lsid:plazi:treatment:03A9FE14D37CFFB5FF6DF902FB3218FA" httpUri="http://treatment.plazi.org/id/03A9FE14D37CFFB5FF6DF902FB3218FA" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<subSubSection id="C31A1C89D37CFFB4FF6DF902FD831D6C" box="[151,620,1784,1811]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BBF4F02D37CFFB4FF6DF902FD831D6C" blockId="6.[151,620,1784,1844]" box="[151,620,1784,1811]" pageId="6" pageNumber="7">
<heading id="D0F7F86ED37CFFB4FF6DF902FD831D6C" bold="true" box="[151,620,1784,1811]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="B9749310D37CFFB4FF6DF902FD831D6C" bold="true" box="[151,620,1784,1811]" pageId="6" pageNumber="7">
3.
<taxonomicName id="4C003481D37CFFB4FF4EF903FD831D6C" authority="Terayama 2005" authorityName="Terayama" authorityYear="2005" box="[180,620,1784,1811]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hangunensis">
<emphasis id="B9749310D37CFFB4FF4EF903FE4A1D6D" bold="true" box="[180,421,1785,1811]" italics="true" pageId="6" pageNumber="7">Epyris hangunensis</emphasis>
<bibRefCitation id="EF9132F3D37CFFB4FE56F902FD831D6C" author="Terayama" box="[428,620,1784,1810]" pageId="6" pageNumber="7" refString="Terayama, M. (2005) Descriptions of new taxa and distribution records of the family Bethylidae (Insecta, Hymenoptera). III. Subfamily Epyrinae. Liberal Arts, Bulletin of Kanto Gakuen University, 13, 51 - 112." type="journal article" year="2005">Terayama 2005</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31A1C89D37CFFB4FF6DF8E6FEC81D4A" box="[151,295,1820,1844]" pageId="6" pageNumber="7" type="description">
<paragraph id="8BBF4F02D37CFFB4FF6DF8E6FEC81D4A" blockId="6.[151,620,1784,1844]" box="[151,295,1820,1844]" pageId="6" pageNumber="7">
(
<figureCitation id="133B5387D37CFFB4FF65F8E6FEF01D4A" box="[159,287,1820,1844]" captionStart="FIGURES 21 27" captionStartId="7.[151,263,1972,1995]" captionTargetBox="[338,1252,685,1958]" captionTargetId="figure@7.[338,1253,685,1958]" captionTargetPageId="7" captionText="FIGURES 21 27. Epyris hangunensis (male). 21. Habitus; 22. Head; 23. Mandible and clypeus; 24. Antenna; 25. Pronotum; 26. Fore wing; 27. Propodeum. Scale bar: 0.20 mm for 22, 23, 25, 27; 0.30 mm for 24, 26; 1.00 mm for 21." httpUri="https://zenodo.org/record/277434/files/figure.png" pageId="6" pageNumber="7">Figs 2127</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31A1C89D37CFFB4FF6DF89CFD0D1D04" box="[151,738,1891,1915]" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="8BBF4F02D37CFFB4FF6DF89CFD0D1D04" blockId="6.[151,738,1891,1915]" box="[151,738,1891,1915]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="AB10682CD37CFFB4FF6DF89CFD0D1D04" box="[151,738,1891,1915]" pageId="6" pageNumber="7">
<taxonomicName id="4C003481D37CFFB4FF6DF89CFDC51D04" authority="Terayama 2005: 72" authorityName="Terayama" authorityPageNumber="72" authorityYear="2005" box="[151,554,1892,1915]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hangunensis">
<emphasis id="B9749310D37CFFB4FF6DF89CFEB11D05" box="[151,350,1893,1915]" italics="true" pageId="6" pageNumber="7">Epyris hangunensis</emphasis>
<treatmentCitation id="0AA16913D37CFFB4FE9EF89FFDC51D04" author="Terayama" box="[356,554,1892,1914]" page="72" pageId="6" pageNumber="7" year="2005">
<bibRefCitation id="EF9132F3D37CFFB4FE9EF89FFDEB1D04" author="Terayama" box="[356,516,1892,1914]" pageId="6" pageNumber="7" refString="Terayama, M. (2005) Descriptions of new taxa and distribution records of the family Bethylidae (Insecta, Hymenoptera). III. Subfamily Epyrinae. Liberal Arts, Bulletin of Kanto Gakuen University, 13, 51 - 112." type="journal article" year="2005">Terayama 2005</bibRefCitation>
: 72
</treatmentCitation>
</taxonomicName>
.
<figureCitation id="133B5387D37CFFB4FDCEF89EFD511D04" box="[564,702,1892,1914]" captionStart="FIGURES 106 115" captionStartId="21.[151,263,1632,1655]" captionTargetBox="[328,1241,320,1616]" captionTargetId="figure@21.[328,1259,320,1616]" captionTargetPageId="21" captionText="FIGURES 106 115. Epyris yamatonis (male). 106. Habitus; 107. Head; 108. Mandible and clypeus; 109. Antenna; 110. Pronotum and mesonotum; 111. Fore wing; 112. Propodeum; 113. Metasoma; 114. Genitalia, ventral view; 115. Hypopygium. Scale bar: 0.10 mm for 114, 115; 0.20 mm for 107 110, 112; 0.50 mm for 111, 113; 1.00 mm for 106." httpUri="https://zenodo.org/record/277443/files/figure.png" pageId="6" pageNumber="7">Figs 111, 112</figureCitation>
,
<figureCitation id="133B5387D37CFFB4FD32F899FD331D05" box="[712,732,1891,1915]" captionStart="FIGURES 1 10" captionStartId="4.[151,263,1762,1785]" captionTargetBox="[328,1259,450,1741]" captionTargetId="figure@4.[328,1259,445,1741]" captionTargetPageId="4" captionText="FIGURES 1 10. Epyris aequalis sp. nov. (male). 1. Habitus; 2. Head; 3. Mandible and clypeus; 4. Antenna; 5. Pronotum and mesonotum; 6. Fore wing; 7. Propodeum; 8. Metasoma; 9. Genitalia, ventral view; 10. Hypopygium, ventral view. Scale bar: 0.10 mm for 9, 10; 0.20 mm for 2, 3, 5, 7; 0.30 mm for 6, 8; 0.50 mm for 1, 4." httpUri="https://zenodo.org/record/277432/files/figure.png" pageId="6" pageNumber="7">3</figureCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31A1C89D37CFFB4FF6DF85DFDE51D9A" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BBF4F02D37CFFB4FF6DF85DFDE51D9A" blockId="6.[151,1436,1959,2020]" pageId="6" pageNumber="7">
<emphasis id="B9749310D37CFFB4FF6DF85DFE961DBE" bold="true" box="[151,377,1959,1984]" pageId="6" pageNumber="7">Material examined</emphasis>
.
<typeStatus id="54BBF1A0D37CFFB4FE7CF852FE011DBE" box="[390,494,1960,1984]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
3,
<collectingCountry id="F3170F92D37CFFB4FDECF853FDB01DBE" box="[534,607,1961,1984]" name="South Korea" pageId="6" pageNumber="7">Korea</collectingCountry>
: Suwon-si (labeled as Suigen), GG,
<date id="FFBE69C2D37CFFB4FC14F852FB9F1DBE" box="[1006,1136,1960,1984]" pageId="6" pageNumber="7" value="1925-09-16">16.IX.1925</date>
, K. Sato leg.
<typeStatus id="54BBF1A0D37CFFB4FAFEF853FA891DBF" box="[1284,1382,1961,1985]" pageId="6" pageNumber="7" type="paratype">Paratype</typeStatus>
1 3, same collection data as
<typeStatus id="54BBF1A0D37CFFB4FE5BF836FDE81D9A" box="[417,519,1996,2020]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A1C89D37DFFB5FF3DFF6DFB391BB1" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BBF4F02D37DFFB5FF3DFF6DFB391BB1" blockId="7.[151,1436,151,644]" pageId="7" pageNumber="8">
<emphasis id="B9749310D37DFFB5FF3DFF6DFED51ACE" bold="true" box="[199,314,151,176]" pageId="7" pageNumber="8">Diagnosis</emphasis>
. Body length ~
<quantity id="4CF8E2E7D37DFFB5FE1DFF6DFDD01AD1" box="[487,575,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="7" pageNumber="8" unit="mm" value="4.4">4.4 mm</quantity>
(
<figureCitation id="133B5387D37DFFB5FDB4FF6DFD4E1AD1" box="[590,673,151,175]" captionStart="FIGURES 21 27" captionStartId="7.[151,263,1972,1995]" captionTargetBox="[338,1252,685,1958]" captionTargetId="figure@7.[338,1253,685,1958]" captionTargetPageId="7" captionText="FIGURES 21 27. Epyris hangunensis (male). 21. Habitus; 22. Head; 23. Mandible and clypeus; 24. Antenna; 25. Pronotum; 26. Fore wing; 27. Propodeum. Scale bar: 0.20 mm for 22, 23, 25, 27; 0.30 mm for 24, 26; 1.00 mm for 21." httpUri="https://zenodo.org/record/277434/files/figure.png" pageId="7" pageNumber="8">Fig. 21</figureCitation>
); LFW ~3.0 mm (
<figureCitation id="133B5387D37DFFB5FC94FF6DFC2D1AD1" box="[878,962,151,175]" captionStart="FIGURES 21 27" captionStartId="7.[151,263,1972,1995]" captionTargetBox="[338,1252,685,1958]" captionTargetId="figure@7.[338,1253,685,1958]" captionTargetPageId="7" captionText="FIGURES 21 27. Epyris hangunensis (male). 21. Habitus; 22. Head; 23. Mandible and clypeus; 24. Antenna; 25. Pronotum; 26. Fore wing; 27. Propodeum. Scale bar: 0.20 mm for 22, 23, 25, 27; 0.30 mm for 24, 26; 1.00 mm for 21." httpUri="https://zenodo.org/record/277434/files/figure.png" pageId="7" pageNumber="8">Fig. 26</figureCitation>
). Antenna and legs dark castaneous. Head 1.1 × as long as wide (
<figureCitation id="133B5387D37DFFB5FE61FF46FE1F1AAA" box="[411,496,188,212]" captionStart="FIGURES 21 27" captionStartId="7.[151,263,1972,1995]" captionTargetBox="[338,1252,685,1958]" captionTargetId="figure@7.[338,1253,685,1958]" captionTargetPageId="7" captionText="FIGURES 21 27. Epyris hangunensis (male). 21. Habitus; 22. Head; 23. Mandible and clypeus; 24. Antenna; 25. Pronotum; 26. Fore wing; 27. Propodeum. Scale bar: 0.20 mm for 22, 23, 25, 27; 0.30 mm for 24, 26; 1.00 mm for 21." httpUri="https://zenodo.org/record/277434/files/figure.png" pageId="7" pageNumber="8">Fig. 22</figureCitation>
), anterior half densely punctuate. Vertex broadly almost straight with rounded corners. Temple profile almost parallel. Mandible with single sharpened tooth (
<figureCitation id="133B5387D37DFFB5FC0EFF25FBA61A89" box="[1012,1097,223,247]" captionStart="FIGURES 21 27" captionStartId="7.[151,263,1972,1995]" captionTargetBox="[338,1252,685,1958]" captionTargetId="figure@7.[338,1253,685,1958]" captionTargetPageId="7" captionText="FIGURES 21 27. Epyris hangunensis (male). 21. Habitus; 22. Head; 23. Mandible and clypeus; 24. Antenna; 25. Pronotum; 26. Fore wing; 27. Propodeum. Scale bar: 0.20 mm for 22, 23, 25, 27; 0.30 mm for 24, 26; 1.00 mm for 21." httpUri="https://zenodo.org/record/277434/files/figure.png" pageId="7" pageNumber="8">Fig. 23</figureCitation>
). First five antennal segments in ratio of 1.9: 1.0: 1.3: 1.9: 2.0 in length, flagellomere 1 1.3 × as long as wide, flagellomere 11 3.9 × as long as wide (
<figureCitation id="133B5387D37DFFB5FF24FEDDFEDB1B41" box="[222,308,295,319]" captionStart="FIGURES 21 27" captionStartId="7.[151,263,1972,1995]" captionTargetBox="[338,1252,685,1958]" captionTargetId="figure@7.[338,1253,685,1958]" captionTargetPageId="7" captionText="FIGURES 21 27. Epyris hangunensis (male). 21. Habitus; 22. Head; 23. Mandible and clypeus; 24. Antenna; 25. Pronotum; 26. Fore wing; 27. Propodeum. Scale bar: 0.20 mm for 22, 23, 25, 27; 0.30 mm for 24, 26; 1.00 mm for 21." httpUri="https://zenodo.org/record/277434/files/figure.png" pageId="7" pageNumber="8">Fig. 24</figureCitation>
). Clypeus with angulate median lobe. Eye hairy. HE 1.0 × OOL, WF 2.4 × WOT. Frontal angle of ocellar triangle obtuse. Pronotal disc 0.6 × as long as wide (
<figureCitation id="133B5387D37DFFB5FCD6FEB6FC6F1B1A" box="[812,896,332,356]" captionStart="FIGURES 21 27" captionStartId="7.[151,263,1972,1995]" captionTargetBox="[338,1252,685,1958]" captionTargetId="figure@7.[338,1253,685,1958]" captionTargetPageId="7" captionText="FIGURES 21 27. Epyris hangunensis (male). 21. Habitus; 22. Head; 23. Mandible and clypeus; 24. Antenna; 25. Pronotum; 26. Fore wing; 27. Propodeum. Scale bar: 0.20 mm for 22, 23, 25, 27; 0.30 mm for 24, 26; 1.00 mm for 21." httpUri="https://zenodo.org/record/277434/files/figure.png" pageId="7" pageNumber="8">Fig. 25</figureCitation>
). Scutellar pit oval, each pit separated from each other by 2.0 × as its maximum diameter. Propodeal disc 0.7 × as long as wide with three discal carinae, median carina reaching posterior carina, submedian discal carinae reaching two thirds of disc length (
<figureCitation id="133B5387D37DFFB5FB4BFE6EFAE91BD2" box="[1201,1286,404,428]" captionStart="FIGURES 21 27" captionStartId="7.[151,263,1972,1995]" captionTargetBox="[338,1252,685,1958]" captionTargetId="figure@7.[338,1253,685,1958]" captionTargetPageId="7" captionText="FIGURES 21 27. Epyris hangunensis (male). 21. Habitus; 22. Head; 23. Mandible and clypeus; 24. Antenna; 25. Pronotum; 26. Fore wing; 27. Propodeum. Scale bar: 0.20 mm for 22, 23, 25, 27; 0.30 mm for 24, 26; 1.00 mm for 21." httpUri="https://zenodo.org/record/277434/files/figure.png" pageId="7" pageNumber="8">Fig. 27</figureCitation>
). Metacarpus short, 0.4 × as long as pterostigma; transverse-median vein strongly convex posteriorly (
<figureCitation id="133B5387D37DFFB5FB8FFE4DFB261BB1" box="[1141,1225,439,463]" captionStart="FIGURES 21 27" captionStartId="7.[151,263,1972,1995]" captionTargetBox="[338,1252,685,1958]" captionTargetId="figure@7.[338,1253,685,1958]" captionTargetPageId="7" captionText="FIGURES 21 27. Epyris hangunensis (male). 21. Habitus; 22. Head; 23. Mandible and clypeus; 24. Antenna; 25. Pronotum; 26. Fore wing; 27. Propodeum. Scale bar: 0.20 mm for 22, 23, 25, 27; 0.30 mm for 24, 26; 1.00 mm for 21." httpUri="https://zenodo.org/record/277434/files/figure.png" pageId="7" pageNumber="8">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31A1C89D37DFFB5FF3DFE21FE541842" pageId="7" pageNumber="8" type="description">
<paragraph id="8BBF4F02D37DFFB5FF3DFE21FAD51869" blockId="7.[151,1436,151,644]" pageId="7" pageNumber="8">
<emphasis id="B9749310D37DFFB5FF3DFE21FED11B8D" bold="true" box="[199,318,475,500]" pageId="7" pageNumber="8">Variation.</emphasis>
<typeStatus id="54BBF1A0D37DFFB5FEBCFE27FE471B8B" box="[326,424,477,501]" pageId="7" pageNumber="8" type="paratype">Paratype</typeStatus>
with sparse hairs on eye; legs light to dark castaneous; TL
<quantity id="4CF8E2E7D37DFFB5FBBFFE26FB031B8A" box="[1093,1260,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.6" metricValueMin="4.4" pageId="7" pageNumber="8" unit="mm" value="4.5" valueMax="4.6" valueMin="4.4">4.404.60 mm</quantity>
; propodeal disc 0.70.72 × as long as wide; scutellar pit separated from each other by 1.92.1 × as its maximum diameter.
</paragraph>
<paragraph id="8BBF4F02D37DFFB5FF3DFDDEFE541842" blockId="7.[151,1436,151,644]" box="[199,443,548,572]" pageId="7" pageNumber="8">FEMALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C31A1C89D37DFFB5FF3DFDBDFE1A1821" box="[199,501,583,608]" pageId="7" pageNumber="8" type="distribution">
<paragraph id="8BBF4F02D37DFFB5FF3DFDBDFE1A1821" blockId="7.[151,1436,151,644]" box="[199,501,583,608]" pageId="7" pageNumber="8">
<emphasis id="B9749310D37DFFB5FF3DFDBDFE8F1821" bold="true" box="[199,352,583,608]" pageId="7" pageNumber="8">Distribution.</emphasis>
<collectingCountry id="F3170F92D37DFFB5FE9DFDB2FE441821" box="[359,427,584,607]" name="South Korea" pageId="7" pageNumber="8">Korea</collectingCountry>
(GG).
</paragraph>
</subSubSection>
<subSubSection id="C31A1C89D37DFFB5FF3DFD96FB3218FA" box="[199,1245,620,644]" pageId="7" pageNumber="8" type="discussion">
<paragraph id="8BBF4F02D37DFFB5FF3DFD96FB3218FA" blockId="7.[151,1436,151,644]" box="[199,1245,620,644]" pageId="7" pageNumber="8">
<emphasis id="B9749310D37DFFB5FF3DFD96FEDB18FA" bold="true" box="[199,308,620,644]" pageId="7" pageNumber="8">Remarks</emphasis>
. This species was previously recorded from Gyeonggi-do,
<collectingCountry id="F3170F92D37DFFB5FC30FD97FBE118FA" box="[970,1038,621,644]" name="South Korea" pageId="7" pageNumber="8">Korea</collectingCountry>
(
<bibRefCitation id="EF9132F3D37DFFB5FBE7FD96FB3E18FA" author="Terayama" box="[1053,1233,620,644]" pageId="7" pageNumber="8" refString="Terayama, M. (2005) Descriptions of new taxa and distribution records of the family Bethylidae (Insecta, Hymenoptera). III. Subfamily Epyrinae. Liberal Arts, Bulletin of Kanto Gakuen University, 13, 51 - 112." type="journal article" year="2005">Terayama 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>