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<document id="96708CD5480E4E1DD3BB1CE82B9D5270" ID-CLB-Dataset="20988" ID-DOI="10.5852/ejt.2020.713" ID-GBIF-Dataset="1d5fdead-d612-4c66-b482-4f051f990351" ID-Zenodo-Dep="4020752" ID-ZooBank="A6CB58F5-1ECC-47F0-AA07-798844AF80A7" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1599662866733" checkinUser="valdenar" docAuthor="Antić, Dragan Ž. &amp; Reip, Hans S." docDate="2020" docId="03A963622812FFFD2E5B1E2F0CF3FEBA" docLanguage="en" docName="ejt-713_antic_reip.pdf" docOrigin="European Journal of Taxonomy 713" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Martvilia Antić &amp; Reip 2020, gen. nov." docType="treatment" docUuid="2B37D666-2887-45E0-AC8D-ECE226C08BD8" docUuidSource="ZooBank" docVersion="11" lastPageNumber="81" masterDocId="FF901B1A285DFFAD2C24183E0F08FFEE" masterDocTitle="The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)" masterLastPageNumber="106" masterPageNumber="1" pageNumber="80" updateTime="1698845858502" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6C0229135A7D3B72FA0F70279358039C">The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)</mods:title>
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<mods:namePart id="77A9FC8E83DC88F5B6461EC06E7E8BB6">Antić, Dragan Ž.</mods:namePart>
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<mods:affiliation id="EE71D3AF931B94879E30B2FEEB7E13F9">University of Belgrade - Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia. Department of Soil Zoology, Senckenberg Museum of Natural History Görlitz, Am Museum 1, 02826 Görlitz, Germany. Thüringer Höhlenverein e. V., Johann-Sebastian-Bach-Str. 16 a, 98529 Suhl, Germany.</mods:affiliation>
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<mods:namePart id="3E49DB2FBCDFF6633BB161AAA083D375">Reip, Hans S.</mods:namePart>
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<mods:title id="2C9E98A3058698497C6E6A14609795F0">European Journal of Taxonomy</mods:title>
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<mods:date id="D2D9C1EBD5B2F8B655C558BA9C426F58">2020</mods:date>
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<paragraph id="8BBFD2742812FFE22E5B1E2F0CBCF9C5" blockId="79.[363,1224,1552,1650]" box="[639,948,1552,1579]" pageId="79" pageNumber="80">
<heading id="D0F765182812FFE22E5B1E2F0CBCF9C5" box="[639,948,1552,1579]" centered="true" fontSize="11" level="2" pageId="79" pageNumber="80" reason="2">
Genus
<taxonomicName id="4C00A9F72812FFE22EF61E2E0C4AF9C5" ID-CoL="845HQ" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[722,834,1552,1579]" class="Diplopoda" family="Julidae" genus="Martvilia" kingdom="Animalia" order="Julida" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9740E662812FFE22EF61E2E0C4AF9C5" bold="true" box="[722,834,1552,1579]" italics="true" pageId="79" pageNumber="80">Martvilia</emphasis>
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<taxonomicNameLabel id="A247B31D2812FFE22F6E1E2F0CBCF9C5" box="[842,948,1553,1579]" pageId="79" pageNumber="80" rank="genus">gen. nov.</taxonomicNameLabel>
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<paragraph id="8BBFD2742812FFE22D4F1E0A0BC0F9A1" blockId="79.[363,1224,1552,1650]" box="[363,1224,1588,1615]" pageId="79" pageNumber="80">
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urn:lsid:zoobank.org:act:
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<paragraph id="8BBFD2742812FFE22E881E690C8FF99C" blockId="79.[363,1224,1552,1650]" box="[684,903,1623,1650]" pageId="79" pageNumber="80">
<figureCitation id="133BCEF12812FFE22E881E690C3AF99C" box="[684,818,1623,1650]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="81.[189,232,1894,1920]" captionStartId-1="82.[189,232,1844,1870]" captionTargetBox-0="[206,1382,265,1868]" captionTargetBox-1="[209,1374,548,1815]" captionTargetId-0="figure@81.[206,1382,265,1868]" captionTargetId-1="figure@82.[189,1399,538,1819]" captionTargetPageId-0="81" captionTargetPageId-1="82" captionText-0="Fig. 50. Martvilia parva gen. et sp. nov. (AC, E: holotype ♂ from Jortsku Cave (ZMUM); D, F: non-type ♀ from Motena Cave (IZB)). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 7, lateral view. F. Telson, posterior view. Arrow indicates metazonal seta. Scale bars: 1 mm." captionText-1="Fig. 51. Martvilia parva gen. et sp. nov., type ♂♂ (ZMUM). A. Holotype, left gonopods, mesal view. B. Holotype, left gonopods, lateral view. C. Paratype, right promere, posteromesal view. D. Paratype, leg-pair 1, anterior view. E. Paratype, penes, posterior view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bar: 0.3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4020862" figureDoi-1="http://doi.org/10.5281/zenodo.4020864" httpUri-0="https://zenodo.org/record/4020862/files/figure.png" httpUri-1="https://zenodo.org/record/4020864/files/figure.png" pageId="79" pageNumber="80">Figs 5051</figureCitation>
,
<figureCitation id="133BCEF12812FFE22F1A1E690C56F99C" box="[830,862,1623,1650]" captionStart="Fig" captionStartId="96.[189,232,1921,1947]" captionTargetBox="[189,1399,979,1893]" captionTargetId="figure@96.[189,1399,979,1893]" captionTargetPageId="96" captionText="Fig. 56. General distribution of the Caucasian leucogeorgiinines. The white dashed line hypothetically indicates the boundary between the northwestern and southeastern groups of species of the genus Leucogeorgia Verhoeff, 1930." figureDoi="http://doi.org/10.5281/zenodo.4020873" httpUri="https://zenodo.org/record/4020873/files/figure.png" pageId="79" pageNumber="80">56</figureCitation>
,
<figureCitation id="133BCEF12812FFE22F4E1E690C8FF99C" box="[874,903,1623,1650]" captionStart="Fig" captionStartId="98.[189,232,1964,1990]" captionTargetBox="[189,1399,1026,1935]" captionTargetId="figure@98.[189,1399,1026,1935]" captionTargetPageId="98" captionText="Fig. 58. Distribution of the southeastern group of the genus Leucogeorgia Verhoeff, 1930 and the species Martvilia parva gen. et sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4020877" httpUri="https://zenodo.org/record/4020877/files/figure.png" pageId="79" pageNumber="80">58</figureCitation>
</paragraph>
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<subSubSection id="C31A81FF2812FFE22C991EA60D89F8A5" pageId="79" pageNumber="80" type="diagnosis">
<paragraph id="8BBFD2742812FFE22C991EA60E3EF95C" blockId="79.[189,310,1688,1714]" box="[189,310,1688,1714]" pageId="79" pageNumber="80">
<heading id="D0F765182812FFE22C991EA60E3EF95C" bold="true" box="[189,310,1688,1714]" fontSize="11" level="3" pageId="79" pageNumber="80" reason="3">
<emphasis id="B9740E662812FFE22C991EA60E3EF95C" bold="true" box="[189,310,1688,1714]" pageId="79" pageNumber="80">Diagnosis</emphasis>
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<paragraph id="8BBFD2742812FFE22C991EF90D89F8A5" blockId="79.[189,1399,1735,1867]" pageId="79" pageNumber="80">
Differs from all other members of the tribe
<taxonomicName id="4C00A9F72812FFE22EC81EF90C97F90F" authorityName="Verhoeff" authorityYear="1930" box="[748,927,1735,1761]" class="Diplopoda" family="Julidae" kingdom="Animalia" order="Julida" pageId="79" pageNumber="80" phylum="Arthropoda" rank="tribe" tribe="Leucogeorgiini">Leucogeorgiini</taxonomicName>
by having an almost completely free mesomere separated by a deep incision from the mesomeral lamella. These two structures are connected only in the basal third in mesal view. Like some members of the genus
<taxonomicName id="4C00A9F72812FFE2280F1F300BC5F8C9" authorityName="Verhoeff" authorityYear="1930" box="[1067,1229,1806,1831]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B9740E662812FFE2280F1F300BC5F8C9" box="[1067,1229,1806,1831]" italics="true" pageId="79" pageNumber="80">Leucogeorgia</emphasis>
</taxonomicName>
, this genus is characterized by modified mouthparts.
</paragraph>
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<subSubSection id="C31A81FF2812FFE22C991F4C0E67F830" pageId="79" pageNumber="80" type="etymology">
<paragraph id="8BBFD2742812FFE22C991F4C0E4CF862" blockId="79.[189,324,1906,1932]" box="[189,324,1906,1932]" pageId="79" pageNumber="80">
<heading id="D0F765182812FFE22C991F4C0E4CF862" bold="true" box="[189,324,1906,1932]" fontSize="11" level="3" pageId="79" pageNumber="80" reason="3">
<emphasis id="B9740E662812FFE22C991F4C0E4CF862" bold="true" box="[189,324,1906,1932]" pageId="79" pageNumber="80">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBFD2742812FFE22C991F9F0E67F830" blockId="79.[189,1398,1953,2014]" pageId="79" pageNumber="80">
This new genus is named after the
<typeStatus id="54BB6CD62812FFE22E4A1F9F0DA9F855" box="[622,673,1953,1979]" pageId="79" pageNumber="80">type</typeStatus>
locality of its
<typeStatus id="54BB6CD62812FFE22F7C1F9F0C83F855" box="[856,907,1953,1979]" pageId="79" pageNumber="80">type</typeStatus>
species, Martvili village. The name is a feminine noun.
</paragraph>
</subSubSection>
<subSubSection id="C31A81FF280DFFFD2C9919320CF3FEBA" pageId="80" pageNumber="81" type="type_taxon">
<paragraph id="8BBFD274280DFFFD2C9919320E50FECB" blockId="80.[189,344,267,294]" box="[189,344,267,294]" pageId="80" pageNumber="81">
<heading id="D0F76518280DFFFD2C9919320E50FECB" bold="true" box="[189,344,267,294]" fontSize="11" level="3" pageId="80" pageNumber="81" reason="3">
<emphasis id="B9740E66280DFFFD2C9919320E50FECB" bold="true" box="[189,344,267,294]" pageId="80" pageNumber="81">
<typeStatus id="54BB6CD6280DFFFD2C9919320FF2FEC8" box="[189,250,268,294]" pageId="80" pageNumber="81">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBFD274280DFFFD2C9919040CF3FEBA" blockId="80.[189,1019,314,340]" box="[189,1019,314,340]" pageId="80" pageNumber="81">
<taxonomicName id="4C00A9F7280DFFFD2C9919040E7FFEBA" ID-CoL="849YJ" authority="Antić &amp; Reip, 2020" authorityName="Antić &amp; Reip" authorityYear="2020" box="[189,375,314,340]" class="Diplopoda" family="Julidae" genus="Martvilia" kingdom="Animalia" order="Julida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="parva" status="gen. et sp. nov.">
<emphasis id="B9740E66280DFFFD2C9919040E7FFEBA" box="[189,375,314,340]" italics="true" pageId="80" pageNumber="81">Martvilia parva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B31D280DFFFD2D5B19040D39FEBA" box="[383,561,314,340]" pageId="80" pageNumber="81" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, by present designation and monotypy.
</paragraph>
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</treatment>
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