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<document id="D1A152B310E7D389FED80BBBBAB12B18" ID-CLB-Dataset="24986" ID-DOI="10.11646/zootaxa.4691.5.5" ID-GBIF-Dataset="864e2b8d-c2da-4a20-9fbf-02bcb20be24a" ID-ISSN="1175-5326" ID-Zenodo-Dep="3527562" ID-ZooBank="0684AE18-0510-4F7B-B75D-AE5177FBF2A2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1572858058763" checkinUser="plazi" docAuthor="Moreno, Antonio, Roszkowska, Milena, García, Mario Alberto, Flores, José Juan &amp; Kaczmarek, Łukasz" docDate="2019" docId="038242511102FFBA3DECFE59FA98FCD7" docLanguage="en" docName="zootaxa.4691.5.5.pdf" docOrigin="Zootaxa 4691 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Astatumen trinacriae" docType="treatment" docVersion="11" lastPageNumber="514" masterDocId="FFBB3A29110FFFB73D7BFF84FFC8FFEC" masterDocTitle="Current knowledge on Mexican tardigrades with a description of Milnesium cassandrae sp. nov. (Eutardigrada: Milnesiidae) and discussion on the taxonomic value of dorsal pseudoplates in the genus Milnesium Doyère, 1840" masterLastPageNumber="524" masterPageNumber="501" pageNumber="514" updateTime="1698761973636" updateUser="ExternalLinkService">
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<mods:title id="AEED2A9C719D1A798658BCCDEB28E7F1">Current knowledge on Mexican tardigrades with a description of Milnesium cassandrae sp. nov. (Eutardigrada: Milnesiidae) and discussion on the taxonomic value of dorsal pseudoplates in the genus Milnesium Doyère, 1840</mods:title>
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<mods:namePart id="D1070995561A38CB7A4ACD449D2B57B9">Moreno, Antonio</mods:namePart>
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<mods:namePart id="1D2F698A513120F330D2A1D4FE88AC74">Roszkowska, Milena</mods:namePart>
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<mods:namePart id="F0A4CCE5AF33C4B0246AB01A1F7F346B">García, Mario Alberto</mods:namePart>
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<mods:namePart id="4AE0339E1FE0918E50F79903C1CAA9B0">Flores, José Juan</mods:namePart>
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<mods:namePart id="21350FC9048E74C5E700947E1F69521F">Kaczmarek, Łukasz</mods:namePart>
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<mods:title id="8765EB2D1B51781923EA9E2FDE6B4863">Zootaxa</mods:title>
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<mods:date id="614833DA77E87DDAE946829C3257B740">2019</mods:date>
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<mods:number id="2D3A3DD3072977C1CF31E321B1B96DF1">2019-11-04</mods:number>
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<mods:number id="18423033F2F312C651EA036046D01FC4">4691</mods:number>
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<treatment id="038242511102FFBA3DECFE59FA98FCD7" ID-DOI="http://doi.org/10.5281/zenodo.5586400" ID-GBIF-Taxon="160127063" ID-Zenodo-Dep="5586400" LSID="urn:lsid:plazi:treatment:038242511102FFBA3DECFE59FA98FCD7" httpUri="http://treatment.plazi.org/id/038242511102FFBA3DECFE59FA98FCD7" lastPageNumber="514" pageId="13" pageNumber="514">
<subSubSection id="C331A0CC1102FFBA3DECFE59FE28FDF7" pageId="13" pageNumber="514" type="nomenclature">
<paragraph id="8B94F3471102FFBA3DECFE59FDB6FE1B" blockId="13.[151,1436,476,827]" box="[151,638,476,503]" pageId="13" pageNumber="514">
<heading id="D0DC442B1102FFBA3DECFE59FDB6FE1B" bold="true" box="[151,638,476,503]" fontSize="11" level="1" pageId="13" pageNumber="514" reason="1">
<taxonomicName id="4C2B88C41102FFBA3DECFE59FDB6FE1B" ID-CoL="HNL4" authority="(Arcidiacono, 1962)" baseAuthorityName="Arcidiacono" baseAuthorityYear="1962" box="[151,638,476,503]" class="Eutardigrada" family="Hypsibiidae" genus="Astatumen" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="514" phylum="Tardigrada" rank="species" species="trinacriae">
<emphasis id="B95F2F551102FFBA3DECFE59FDB6FE1B" bold="true" box="[151,638,476,503]" pageId="13" pageNumber="514">
<emphasis id="B95F2F551102FFBA3DECFE59FE45FE1B" bold="true" box="[151,397,477,503]" italics="true" pageId="13" pageNumber="514">Astatumen trinacriae</emphasis>
(
<bibRefCitation id="EFBA8EB61102FFBA3CE0FE58FDBEFE1B" author="Arcidiacono, R." box="[411,630,476,503]" pageId="13" pageNumber="514" pagination="123 - 134" refId="ref13580" refString="Arcidiacono, R. (1962) Contributo alla conoscenza dei Tardigradi dei Monti Nebrodi e descrizione di una nuova specie di Itaquascon. Bollettino delle Sedute della Accademia Gioenia di Scienze Naturali in Catania, 3, 123 - 134." type="journal article" year="1962">Arcidiacono, 1962</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B94F3471102FFBA3DECFD85FE28FDF7" blockId="13.[151,1436,476,827]" box="[151,480,512,539]" pageId="13" pageNumber="514">
<emphasis id="B95F2F551102FFBA3DECFD85FEFCFDF7" bold="true" box="[151,308,512,539]" pageId="13" pageNumber="514">Terra typica:</emphasis>
<collectingCountry id="F33CB3D71102FFBA3C40FD85FEA6FDF7" box="[315,366,513,539]" name="Italy" pageId="13" pageNumber="514">Italy</collectingCountry>
(Europe).
</paragraph>
</subSubSection>
<subSubSection id="C331A0CC1102FFBA3DECFDA0FE54FD23" pageId="13" pageNumber="514" type="materials_examined">
<paragraph id="8B94F3471102FFBA3DECFDA0FE54FD23" blockId="13.[151,1436,476,827]" pageId="13" pageNumber="514">
<emphasis id="B95F2F551102FFBA3DECFDA0FE4AFDD3" bold="true" box="[151,386,548,575]" pageId="13" pageNumber="514">Material examined:</emphasis>
<materialsCitation id="3B43F91A1102FFBA3CF2FDA1FAD0FD8F" ID-GBIF-Occurrence="2446790926" collectingDate="2015-02-09" collectorName="Santiago &amp; Moss &amp; Mario Alberto Garcia Aranda" country="Mexico" elevation="2370" latitude="25.345642" location="La Penita" longLatPrecision="1" longitude="-100.36812" pageId="13" pageNumber="514" specimenCount="27" stateProvince="Nuevo Leon">
<specimenCount id="9D2D38CE1102FFBA3CF2FDA1FDECFDD3" box="[393,548,549,575]" pageId="13" pageNumber="514" type="generic">27 specimens</specimenCount>
:
<quantity id="4CD35EA21102FFBA3F55FDA1FDB4FDD3" box="[558,636,549,575]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="514" unit="km" value="1.2">1.2 km</quantity>
E of
<location id="8EF4A59C1102FFBA3FC3FDA1FCEAFDD3" LSID="urn:lsid:plazi:treatment:038242511102FFBA3DECFE59FA98FCD7:8EF4A59C1102FFBA3FC3FDA1FCEAFDD3" box="[696,802,549,575]" country="Mexico" latitude="25.345642" longLatPrecision="1" longitude="-100.36812" name="La Penita" pageId="13" pageNumber="514" stateProvince="Nuevo Leon">La Peñita</location>
(
<geoCoordinate id="EE1F95801102FFBA3E4BFDA0FC15FDD3" box="[816,989,548,575]" degrees="25" direction="north" minutes="20" orientation="latitude" pageId="13" pageNumber="514" precision="1" seconds="44.31" value="25.345642">25°2044.31N</geoCoordinate>
,
<geoCoordinate id="EE1F95801102FFBA3E93FDA0FB6EFDD3" box="[1000,1190,548,575]" degrees="100" direction="west" minutes="22" orientation="longitude" pageId="13" pageNumber="514" precision="1" seconds="05.22" value="-100.36812">100°2205.22W</geoCoordinate>
,
<emphasis id="B95F2F551102FFBA39CBFDA2FB1AFDD3" box="[1200,1234,550,575]" italics="true" pageId="13" pageNumber="514">ca.</emphasis>
<elevation id="000614741102FFBA39A3FDA1FA9DFDAC" box="[1240,1365,549,576]" metricMagnitude="3" metricUnit="m" metricValue="2.37" pageId="13" pageNumber="514" unit="m" value="2370.0">
<quantity id="4CD35EA21102FFBA39A3FDA1FAE5FDD3" box="[1240,1325,549,575]" metricMagnitude="3" metricUnit="m" metricValue="2.37" pageId="13" pageNumber="514" unit="m" value="2370.0">2370 m</quantity>
asl
</elevation>
),
<collectorName id="26DE96911102FFBA3812FDA1FED6FD8F" pageId="13" pageNumber="514">Santiago</collectorName>
,
<collectingRegion id="49EF3DA51102FFBA3C52FDCDFE70FD8F" box="[297,440,585,611]" country="Mexico" name="Nuevo Leon" pageId="13" pageNumber="514">Nuevo Leon</collectingRegion>
,
<collectingCountry id="F33CB3D71102FFBA3CBFFDCDFDD5FD8F" box="[452,541,585,611]" name="Mexico" pageId="13" pageNumber="514">Mexico</collectingCountry>
.
<collectorName id="26DE96911102FFBA3F53FDCDFDADFD8F" box="[552,613,585,611]" pageId="13" pageNumber="514">Moss</collectorName>
on soil, coll.
<collectorName id="26DE96911102FFBA3E7BFDCDFB86FD8F" box="[768,1102,585,611]" pageId="13" pageNumber="514">Mario Alberto Garcia Aranda</collectorName>
,
<date id="FF95D5871102FFBA3921FDCDFAD0FD8F" box="[1114,1304,584,611]" pageId="13" pageNumber="514" value="2015-02-09">
<collectingDate id="EFD12C6F1102FFBA3921FDCDFAD0FD8F" box="[1114,1304,584,611]" pageId="13" pageNumber="514" value="2015-02-09">9 February 2015</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B43F91A1102FFBA3858FDCDFE51FD23" ID-GBIF-Occurrence="2446790930" collectingDate="2016-12-19" collectorName="Ocampo &amp; Moss &amp; Jonas A. Delgadillo Villalobos" country="Mexico" county="Sierra Maderas del Carmen" elevation="2263" latitude="28.961931" location="One" longLatPrecision="1" longitude="-102.578674" municipality="Frijoles" pageId="13" pageNumber="514" specimenCount="1" stateProvince="Coahuila de Zaragoza">
<location id="8EF4A59C1102FFBA3858FDCDFA9AFD8F" LSID="urn:lsid:plazi:treatment:038242511102FFBA3DECFE59FA98FCD7:8EF4A59C1102FFBA3858FDCDFA9AFD8F" box="[1315,1362,585,611]" country="Mexico" county="Sierra Maderas del Carmen" latitude="28.961931" longLatPrecision="1" longitude="-102.578674" municipality="Frijoles" name="One" pageId="13" pageNumber="514" stateProvince="Coahuila de Zaragoza">One</location>
specimen:
<locationDeviation id="9A26EA651102FFBA3C59FDE9FE1CFD6B" box="[290,468,621,647]" location="Agua" pageId="13" pageNumber="514">
<quantity id="4CD35EA21102FFBA3C59FDE9FE93FD6B" box="[290,347,621,647]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="514" unit="km" value="7.0">7 km</quantity>
E of Agua
</locationDeviation>
de los
<collectingMunicipality id="6BF0693D1102FFBA3F5FFDE9FDB2FD6B" box="[548,634,621,647]" pageId="13" pageNumber="514">Frijoles</collectingMunicipality>
(
<geoCoordinate id="EE1F95801102FFBA3FF1FDE8FC84FD6B" box="[650,844,620,647]" degrees="28" direction="north" minutes="57" orientation="latitude" pageId="13" pageNumber="514" precision="1" seconds="42.952" value="28.961931">28°57 42.952”N</geoCoordinate>
,
<geoCoordinate id="EE1F95801102FFBA3E23FDE9FBE3FD6B" box="[856,1067,621,647]" degrees="102" direction="west" minutes="34" orientation="longitude" pageId="13" pageNumber="514" precision="1" seconds="43.228" value="-102.578674">102°34 43.228”W</geoCoordinate>
,
<emphasis id="B95F2F551102FFBA394CFDEAFB90FD6B" box="[1079,1112,622,647]" italics="true" pageId="13" pageNumber="514">ca.</emphasis>
<elevation id="000614741102FFBA3924FDE9FB15FD64" box="[1119,1245,621,648]" metricMagnitude="3" metricUnit="m" metricValue="2.263" pageId="13" pageNumber="514" unit="m" value="2263.0">
<quantity id="4CD35EA21102FFBA3924FDE9FB7CFD6B" box="[1119,1204,621,647]" metricMagnitude="3" metricUnit="m" metricValue="2.263" pageId="13" pageNumber="514" unit="m" value="2263.0">2263 m</quantity>
asl
</elevation>
),
<collectingCounty id="62F58BCB1102FFBA3989FDE9FEAFFD47" pageId="13" pageNumber="514">Sierra Maderas del Carmen</collectingCounty>
,
<collectorName id="26DE96911102FFBA3C09FD15FE1CFD47" box="[370,468,657,683]" pageId="13" pageNumber="514">Ocampo</collectorName>
,
<collectingRegion id="49EF3DA51102FFBA3CA5FD15FD8EFD47" box="[478,582,657,683]" country="Mexico" name="Coahuila de Zaragoza" pageId="13" pageNumber="514">Coahuila</collectingRegion>
,
<collectingCountry id="F33CB3D71102FFBA3F2AFD15FD62FD47" box="[593,682,657,683]" name="Mexico" pageId="13" pageNumber="514">Mexico</collectingCountry>
.
<collectorName id="26DE96911102FFBA3FCFFD15FD39FD47" box="[692,753,657,683]" pageId="13" pageNumber="514">Moss</collectorName>
on tree (
<taxonomicName id="4C2B88C41102FFBA3E28FD15FC79FD47" box="[851,945,657,683]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="514" phylum="Tracheophyta" rank="genus">
<emphasis id="B95F2F551102FFBA3E28FD15FC79FD47" box="[851,945,657,683]" italics="true" pageId="13" pageNumber="514">Quercus</emphasis>
</taxonomicName>
sp.) coll.
<collectorName id="26DE96911102FFBA3964FD15FABEFD47" box="[1055,1398,657,683]" pageId="13" pageNumber="514">Jonas A. Delgadillo Villalobos</collectorName>
,
<date id="FF95D5871102FFBA38FBFD15FE51FD23" pageId="13" pageNumber="514" value="2016-12-19">
<collectingDate id="EFD12C6F1102FFBA38FBFD15FE51FD23" pageId="13" pageNumber="514" value="2016-12-19">19 December 2016</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C331A0CC1102FFBA3DECFD5CFA98FCD7" pageId="13" pageNumber="514" type="discussion">
<paragraph id="8B94F3471102FFBA3DECFD5CFA98FCD7" blockId="13.[151,1436,476,827]" pageId="13" pageNumber="514">
<emphasis id="B95F2F551102FFBA3DECFD5CFEEAFD1F" bold="true" box="[151,290,728,755]" pageId="13" pageNumber="514">Comments:</emphasis>
Our specimens correspond well with the definition of this species proposed by
<bibRefCitation id="EFBA8EB61102FFBA39E9FD5DFA8AFD1F" author="Dastych, H." box="[1170,1346,729,755]" pageId="13" pageNumber="514" pagination="1 - 255" refId="ref14328" refString="Dastych, H. (1988) The Tardigrada of Poland. Monografie Fauny Polski, 16, 1 - 255." type="journal article" year="1988">Dastych (1988)</bibRefCitation>
.
<taxonomicName id="4C2B88C41102FFBA3830FD5DFE41FCFB" baseAuthorityName="Arcidiacono" baseAuthorityYear="1962" class="Eutardigrada" family="Hypsibiidae" genus="Astatumen" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="514" phylum="Tardigrada" rank="species" species="trinacriae">
<emphasis id="B95F2F551102FFBA3830FD5DFE41FCFB" italics="true" pageId="13" pageNumber="514">Astatumen trinacriae</emphasis>
</taxonomicName>
has a mainly Holarctic distribution (
<bibRefCitation id="EFBA8EB61102FFBA3E52FD79FBE7FCFB" author="Kaczmarek, L. &amp; Michalczyk, L. &amp; McInnes, S. J." box="[809,1071,765,791]" pageId="13" pageNumber="514" pagination="1 - 62" refId="ref15501" refString="Kaczmarek, L., Michalczyk, L. &amp; McInnes, S. J. (2014) Annotated zoogeography of non - marine Tardigrada. Part I: Central America. Zootaxa, 3763 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 3763.1.1" type="journal article" year="2014">
Kaczmarek
<emphasis id="B95F2F551102FFBA3EC9FD7AFC2CFCFB" box="[946,996,765,791]" italics="true" pageId="13" pageNumber="514">et al</emphasis>
. 2014
</bibRefCitation>
;
<bibRefCitation id="EFBA8EB61102FFBA3940FD79FB15FCFB" author="McInnes, S. J." box="[1083,1245,765,791]" pageId="13" pageNumber="514" pagination="257 - 352" refId="ref16129" refString="McInnes, S. J. (1994) Zoogeographic distribution of terrestrial / freshwater tardigrades from current literature. Journal of Natural History, 28, 257 - 352. https: // doi. org / 10.1080 / 00222939400770131" type="journal article" year="1994">McInnes 1994</bibRefCitation>
) but can be confused with
<taxonomicName id="4C2B88C41102FFBA3C24FCA5FC1FFCD7" authority="(Weglarska, 1959) (Zawierucha et al. 2014)" authorityName="Weglarska" authorityYear="1959" baseAuthorityName="Weglarska" baseAuthorityYear="1959" box="[351,983,800,827]" pageId="13" pageNumber="514" rank="species" species="bartosi">
<emphasis id="B95F2F551102FFBA3C24FCA5FE28FCD7" box="[351,480,801,827]" italics="true" pageId="13" pageNumber="514">Ast. bartosi</emphasis>
(
<bibRefCitation id="EFBA8EB61102FFBA3C94FCA5FD67FCD7" author="Weglarska, B." box="[495,687,800,827]" pageId="13" pageNumber="514" pagination="354 - 357" refId="ref19702" refString="Weglarska, B. (1959) Tardigraden Polens II. Vestnik Ceskoslovenske spolecnosti Zoologicke, 23, 354 - 357." type="journal article" year="1959">Węglarska, 1959</bibRefCitation>
) (
<bibRefCitation id="EFBA8EB61102FFBA3FBDFCA5FC07FCD7" author="Zawierucha, K. &amp; Dziamiecki, J. &amp; Jakubowska, N. &amp; Michalczyk, L. &amp; Kaczmarek, L." box="[710,975,801,827]" pageId="13" pageNumber="514" pagination="81 - 105" refId="ref19804" refString="Zawierucha, K., Dziamiecki, J., Jakubowska, N., Michalczyk, L. &amp; Kaczmarek, L. (2014) New tardigrade records for the Baltic states with a description of Minibiotus formosus sp. n. (Eutardigrada, Macrobiotidae). ZooKeys, 408, 81 - 105. https: // doi. org / 10.3897 / zookeys. 408.661 2" type="journal article" year="2014">
Zawierucha
<emphasis id="B95F2F551102FFBA3E28FCA6FC43FCD7" box="[851,907,801,827]" italics="true" pageId="13" pageNumber="514">et al.</emphasis>
2014
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