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<document id="0CDA3513135BA8220B194D3A7C800DB9" ID-DOI="10.11646/zootaxa.3587.1.1" ID-GBIF-Dataset="20d58797-2815-434b-a9c5-5786e926af9d" ID-ISSN="1175-5326" ID-Zenodo-Dep="283115" ID-ZooBank="91FCB387-5D4F-4F12-ABDC-B06D7F60A271" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1460563276739" checkinUser="plazi" docAuthor="Boehme, Nicole F., Tanner, David A., Williams, Kevin A. &amp; Pitts, James P." docDate="2012" docId="038187E51616FFB7FF09EDF4FC6DF97A" docLanguage="en" docName="zt03587p045.pdf" docOrigin="Zootaxa 3587" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Sphaeropthalma marpesia Blake 1879" docType="treatment" docVersion="15" lastPageNumber="31" masterDocId="FFB8FF9D1608FFA9FF9EED63FFB0FF9D" masterDocTitle="Faunal study of velvet ants (Hymenoptera: Mutillidae) and their activity patterns and habitat preference at Ash Meadows National Wildlife Refuge, Nye County, Nevada, USA" masterLastPageNumber="45" masterPageNumber="1" pageNumber="31" updateTime="1698320574733" updateUser="plazi">
<mods:mods id="774CD3B2AE6102313A63D221ACE2E8C8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="09D59E2D7E5660C97149B86A5757EBA9">
<mods:title id="E22ABCE1D5DD9BD24DADBE9D7AB928DC">Faunal study of velvet ants (Hymenoptera: Mutillidae) and their activity patterns and habitat preference at Ash Meadows National Wildlife Refuge, Nye County, Nevada, USA</mods:title>
</mods:titleInfo>
<mods:name id="1D8395E02CDA532B70F7D590F5F06937" type="personal">
<mods:role id="20D079715F544E19513BDF1F5FE541A0">
<mods:roleTerm id="F7843848DB1D30B936BC99826E0BB716">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6BCCD6570B94938A1C3C053A856FDD40">Boehme, Nicole F.</mods:namePart>
</mods:name>
<mods:name id="3C43554116392B81BDD39FBBC9A96203" type="personal">
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</mods:role>
<mods:namePart id="2254AFB98B20C90941BE24DC337F48E7">Tanner, David A.</mods:namePart>
</mods:name>
<mods:name id="B2AD9540E34004C254B57343868138F9" type="personal">
<mods:role id="F11A0C9583BFE374AA242A9037BDAA36">
<mods:roleTerm id="64197252A6BEA5A4CCEB74B734E228E1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="484BD617B9554F8B7FE56DBC2D61BCFB">Williams, Kevin A.</mods:namePart>
</mods:name>
<mods:name id="6C007EA2547E38FDED394186BC9F367F" type="personal">
<mods:role id="2D75746149077C7BF207D9817DD62E6B">
<mods:roleTerm id="6458D82009B60267FE9E66AE9D7725B9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B8722BA43A09DF6410BA39246B0F896C">Pitts, James P.</mods:namePart>
</mods:name>
<mods:typeOfResource id="8888DA3ACB3BBF57C289C22A06084944">text</mods:typeOfResource>
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<mods:title id="24DF21C0D5BD5B5E7D5B72A78A52366C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A5F2BB377A5E3BAEC75A30901C292E0A">
<mods:date id="D54EE3521F27D0BA3F00389CCC92233B">2012</mods:date>
<mods:detail id="8BC0AD5FED2BBF9D71569F1FA7120381" type="pubDate">
<mods:number id="EB1A64127C4CAF1C59AFA24E1206A6F6">2012-12-17</mods:number>
</mods:detail>
<mods:detail id="5CACBA78545787395EE2AA457A5EA05E" type="volume">
<mods:number id="3C1EF6C82D9FCF10DA15A9A7E502DAAD">3587</mods:number>
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<mods:start id="FD327495E492867C6526F7CC226AA44E">1</mods:start>
<mods:end id="723942DE360932291AD6AE740D7F1462">45</mods:end>
</mods:extent>
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<mods:classification id="D39CF7F93E2EF3333FAE7886882B37BA">journal article</mods:classification>
<mods:identifier id="48248B40CD0ED6631C27F5078425DA77" type="DOI">10.11646/zootaxa.3587.1.1</mods:identifier>
<mods:identifier id="5FD8B8AB49145DF9906ED6AF12675285" type="GBIF-Dataset">20d58797-2815-434b-a9c5-5786e926af9d</mods:identifier>
<mods:identifier id="408C6E9FCEB87E4FFE5E3E597069798B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3EA6C130DC0096E0A9D12A9B3C552358" type="Zenodo-Dep">283115</mods:identifier>
<mods:identifier id="53A91D6123FEA182408A8E40C508C6E0" type="ZooBank">91FCB387-5D4F-4F12-ABDC-B06D7F60A271</mods:identifier>
</mods:mods>
<treatment id="038187E51616FFB7FF09EDF4FC6DF97A" ID-DOI="http://doi.org/10.5281/zenodo.5627556" ID-GBIF-Taxon="119557100" ID-Zenodo-Dep="5627556" LSID="urn:lsid:plazi:treatment:038187E51616FFB7FF09EDF4FC6DF97A" httpUri="http://treatment.plazi.org/id/038187E51616FFB7FF09EDF4FC6DF97A" lastPageNumber="31" pageId="30" pageNumber="31">
<subSubSection id="C33265781616FFB7FF09EDF4FD31FF2C" box="[151,641,151,177]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8B9736F31616FFB7FF09EDF4FD31FF2C" blockId="30.[151,641,151,177]" box="[151,641,151,177]" pageId="30" pageNumber="31">
<heading id="D0DF819F1616FFB7FF09EDF4FD31FF2C" bold="true" box="[151,641,151,177]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<taxonomicName id="4C284D701616FFB7FF09EDF4FD31FF2C" authority="Blake, 1879" authorityName="Blake" authorityYear="1879" box="[151,641,151,177]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="marpesia">
<emphasis id="B95CEAE11616FFB7FF09EDF4FD31FF2C" bold="true" box="[151,641,151,177]" pageId="30" pageNumber="31">
<emphasis id="B95CEAE11616FFB7FF09EDF4FE65FF2C" bold="true" box="[151,469,151,177]" italics="true" pageId="30" pageNumber="31">Sphaeropthalma marpesia</emphasis>
(
<bibRefCitation id="EFB94B021616FFB7FE7BEDF4FDC8FF2C" author="Blake, C. A." box="[485,632,151,177]" firstAuthor="Blake" journalOrPublisher="Transaction of the American Entomological Society" pageId="30" pageNumber="31" pagination="243 - 254" part="7" refId="ref57193" refString="Blake, C. A. (1879) Catalogue of the Mutillidae of North America, with descriptions of new species. Transaction of the American Entomological Society, 7, 243 - 254." title="Catalogue of the Mutillidae of North America, with descriptions of new species" type="journal article" year="1879">Blake, 1879</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33265781616FFB7FF09EDBCFDB3FF69" box="[151,515,222,245]" pageId="30" pageNumber="31" type="reference_group">
<paragraph id="8B9736F31616FFB7FF09EDBCFDB3FF69" blockId="30.[151,1341,222,368]" box="[151,515,222,245]" pageId="30" pageNumber="31">
<taxonomicName id="4C284D701616FFB7FF09EDBCFE4FFF69" authority="Marpesia Blake, 1879: 247" authorityName="Marpesia Blake" authorityPageNumber="247" authorityYear="1879" box="[151,511,222,245]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B95CEAE11616FFB7FF09EDBCFEF9FF68" box="[151,329,223,245]" italics="true" pageId="30" pageNumber="31">Mutilla Marpesia</emphasis>
<treatmentCitation id="0A8910E21616FFB7FED1EDBDFE4FFF69" author="Blake" box="[335,511,222,244]" page="247" pageId="30" pageNumber="31" year="1879">
<bibRefCitation id="EFB94B021616FFB7FED1EDBDFE4FFF69" author="Blake, C. A." box="[335,511,222,244]" firstAuthor="Blake" journalOrPublisher="Transaction of the American Entomological Society" page="247" pageId="30" pageNumber="31" pagination="243 - 254" part="7" refId="ref57193" refString="Blake, C. A. (1879) Catalogue of the Mutillidae of North America, with descriptions of new species. Transaction of the American Entomological Society, 7, 243 - 254." title="Catalogue of the Mutillidae of North America, with descriptions of new species" type="journal article" year="1879">Blake, 1879: 247</bibRefCitation>
</treatmentCitation>
</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="C33265781616FFB7FD97EDBCFC49FF68" box="[521,1017,222,245]" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8B9736F31616FFB7FD97EDBCFC49FF68" blockId="30.[151,1341,222,368]" box="[521,1017,222,245]" pageId="30" pageNumber="31">
♀.
<typeStatus id="549388511616FFB7FDB7EDBCFD3EFF69" box="[553,654,223,244]" pageId="30" pageNumber="31" type="lectotype">Lectotype</typeStatus>
(designated here): Kansas (
<collectionCode id="ED39AE361616FFB7FC39EDBCFC5EFF68" LSID="urn:lsid:biocol.org:col:34784" box="[935,1006,223,245]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="30" pageNumber="31">ANSP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C33265781616FFB7FF09ED9EFD3CFE8E" box="[151,652,253,275]" pageId="30" pageNumber="31" type="reference_group">
<paragraph id="8B9736F31616FFB7FF09ED9EFD3CFE8E" blockId="30.[151,1341,222,368]" box="[151,652,253,275]" pageId="30" pageNumber="31">
<taxonomicName id="4C284D701616FFB7FF09ED9EFD38FE8E" authority="Blake, 1886: 235" authorityName="Blake" authorityPageNumber="235" authorityYear="1886" box="[151,648,253,275]" class="Insecta" family="Mutillidae" genus="Sphaerophthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="luteola">
<emphasis id="B95CEAE11616FFB7FF09ED9EFEFAFE8E" box="[151,330,253,275]" italics="true" pageId="30" pageNumber="31">Sphaerophthalma</emphasis>
[sic!]
<emphasis id="B95CEAE11616FFB7FE12ED9EFE62FE8E" box="[396,466,253,275]" italics="true" pageId="30" pageNumber="31">luteola</emphasis>
<treatmentCitation id="0A8910E21616FFB7FE46ED9EFD38FE8E" author="Blake" box="[472,648,253,275]" page="235" pageId="30" pageNumber="31" year="1886">
<bibRefCitation id="EFB94B021616FFB7FE46ED9EFD38FE8E" author="Blake, C. A." box="[472,648,253,275]" firstAuthor="Blake" journalOrPublisher="Transactions of the American Entomological Society" page="235" pageId="30" pageNumber="31" pagination="179 - 286" part="13" refId="ref57229" refString="Blake, C. A. (1886) Monograph of the Mutillidae of North America. Transactions of the American Entomological Society, 13, 179 - 286." title="Monograph of the Mutillidae of North America" type="journal article" year="1886">Blake, 1886: 235</bibRefCitation>
</treatmentCitation>
</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="C33265781616FFB7FD0DED9DFBDAFE8E" box="[659,1130,253,275]" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8B9736F31616FFB7FD0DED9DFBDAFE8E" blockId="30.[151,1341,222,368]" box="[659,1130,253,275]" pageId="30" pageNumber="31">
♀.
<typeStatus id="549388511616FFB7FD2CED9DFCA7FE8E" box="[690,791,254,275]" pageId="30" pageNumber="31" type="lectotype">Lectotype</typeStatus>
(designated here): Utah (
<collectionCode id="ED39AE361616FFB7FB87ED9EFBEFFE8E" LSID="urn:lsid:biocol.org:col:34784" box="[1049,1119,253,275]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="30" pageNumber="31">ANSP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C33265781616FFB7FF09EC7FFC27FEF2" pageId="30" pageNumber="31" type="reference_group">
<paragraph id="8B9736F31616FFB7FF09EC7FFB5BFEAF" blockId="30.[151,1341,222,368]" box="[151,1259,283,306]" pageId="30" pageNumber="31">
<taxonomicName id="4C284D701616FFB7FF09EC7FFC96FEAC" authority="Viereck, 1906: 189" authorityName="Viereck" authorityPageNumber="189" authorityYear="1906" box="[151,806,283,306]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="imperialiformis" subGenus="Photopsis">
<emphasis id="B95CEAE11616FFB7FF09EC7FFE8DFEAF" box="[151,317,284,306]" italics="true" pageId="30" pageNumber="31">Sphaeropthalma</emphasis>
(
<emphasis id="B95CEAE11616FFB7FED2EC7FFE00FEAF" box="[332,432,284,306]" italics="true" pageId="30" pageNumber="31">Photopsis</emphasis>
)
<emphasis id="B95CEAE11616FFB7FE20EC7FFDECFEAF" box="[446,604,284,306]" italics="true" pageId="30" pageNumber="31">imperialiformis</emphasis>
<treatmentCitation id="0A8910E21616FFB7FDFCEC78FC96FEAC" author="Viereck" box="[610,806,283,305]" page="189" pageId="30" pageNumber="31" year="1906">
<bibRefCitation id="EFB94B021616FFB7FDFCEC78FC96FEAC" author="Viereck, H. L." box="[610,806,283,305]" firstAuthor="Viereck" journalOrPublisher="Transactions of the American Entomological Society" page="189" pageId="30" pageNumber="31" pagination="185 - 189" part="32" refId="ref59161" refString="Viereck, H. L. (1906) Notes and Descriptions of Hymenoptera from the Western United States. Transactions of the American Entomological Society, 32, 185 - 189." title="Notes and Descriptions of Hymenoptera from the Western United States" type="journal article" year="1906">Viereck, 1906: 189</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, 3.
<typeStatus id="549388511616FFB7FCCEEC78FC02FEAC" box="[848,946,283,305]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
: Kansas, Morton Co. (SEMC).
</paragraph>
<paragraph id="8B9736F31616FFB7FF09EC59FC5FFECD" blockId="30.[151,1341,222,368]" box="[151,1007,314,336]" pageId="30" pageNumber="31">
<taxonomicName id="4C284D701616FFB7FF09EC59FC7CFECD" authority="Schuster, 1958: 34" authorityName="Schuster" authorityPageNumber="34" authorityYear="1958" box="[151,972,314,336]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="imperialiformis" subGenus="Photopsis" subSpecies="imperialiformis">
<emphasis id="B95CEAE11616FFB7FF09EC59FE8DFECD" box="[151,317,314,336]" italics="true" pageId="30" pageNumber="31">Sphaeropthalma</emphasis>
(
<emphasis id="B95CEAE11616FFB7FED2EC59FE00FECD" box="[332,432,314,336]" italics="true" pageId="30" pageNumber="31">Photopsis</emphasis>
)
<emphasis id="B95CEAE11616FFB7FE20EC59FCB0FECD" box="[446,768,314,336]" italics="true" pageId="30" pageNumber="31">imperialiformis imperialiformis</emphasis>
:
<treatmentCitation id="0A8910E21616FFB7FC93EC59FC7CFECD" author="Schuster" box="[781,972,314,336]" page="34" pageId="30" pageNumber="31" year="1958">
<bibRefCitation id="EFB94B021616FFB7FC93EC59FC7CFECD" author="Schuster, R. M." box="[781,972,314,336]" firstAuthor="Schuster" journalOrPublisher="Entomologica Americana" page="34" pageId="30" pageNumber="31" pagination="1 - 130" part="37" refId="ref58869" refString="Schuster, R. M. (1958) A revision of the sphaeropthalmine Mutillidae of America north of Mexico. Volume II. Entomologica Americana, 37, 1 - 130." title="A revision of the sphaeropthalmine Mutillidae of America north of Mexico. Volume II" type="journal article" year="1958">Schuster, 1958: 34</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, 3.
</paragraph>
<paragraph id="8B9736F31616FFB7FF09EC3AFC27FEF2" blockId="30.[151,1341,222,368]" box="[151,919,345,367]" pageId="30" pageNumber="31">
<taxonomicName id="4C284D701616FFB7FF09EC3AFC24FEF2" authority="Schuster, 1958: 34" authorityName="Schuster" authorityPageNumber="34" authorityYear="1958" box="[151,916,345,367]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="imperialiformis" subGenus="Photopsis" subSpecies="maricopae">
<emphasis id="B95CEAE11616FFB7FF09EC3AFD7EFEF2" box="[151,718,345,367]" italics="true" pageId="30" pageNumber="31">Sphaeropthalma (Photopsis) imperialiformis maricopae</emphasis>
<treatmentCitation id="0A8910E21616FFB7FD4BEC3AFC24FEF2" author="Schuster" box="[725,916,345,367]" page="34" pageId="30" pageNumber="31" year="1958">
<bibRefCitation id="EFB94B021616FFB7FD4BEC3AFC24FEF2" author="Schuster, R. M." box="[725,916,345,367]" firstAuthor="Schuster" journalOrPublisher="Entomologica Americana" page="34" pageId="30" pageNumber="31" pagination="1 - 130" part="37" refId="ref58869" refString="Schuster, R. M. (1958) A revision of the sphaeropthalmine Mutillidae of America north of Mexico. Volume II. Entomologica Americana, 37, 1 - 130." title="A revision of the sphaeropthalmine Mutillidae of America north of Mexico. Volume II" type="journal article" year="1958">Schuster, 1958: 34</bibRefCitation>
</treatmentCitation>
</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="C33265781616FFB7FC00EC3AFA8DFEED" box="[926,1341,345,368]" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8B9736F31616FFB7FC00EC3AFA8DFEED" blockId="30.[151,1341,222,368]" box="[926,1341,345,368]" pageId="30" pageNumber="31">
3.
<typeStatus id="549388511616FFB7FC23EC3AFBAFFEF2" box="[957,1055,345,367]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
: Arizona, Phoenix (
<collectionCode id="ED39AE361616FFB7FB79EC39FA82FEED" box="[1255,1330,346,368]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="30" pageNumber="31">UMSP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C33265781616FFB7FF09ECF8FEC1FC81" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="8B9736F31616FFB7FF09ECF8FEC1FC81" blockId="30.[151,1437,411,1768]" pageId="30" pageNumber="31">
<emphasis id="B95CEAE11616FFB7FF09ECF8FEA1FE29" bold="true" box="[151,273,411,436]" pageId="30" pageNumber="31">Diagnosis.</emphasis>
MALE. The male of
<taxonomicName id="4C284D701616FFB7FD84ECFEFD13FE29" box="[538,675,413,436]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="marpesia">
<emphasis id="B95CEAE11616FFB7FD84ECFEFD13FE29" box="[538,675,413,436]" italics="true" pageId="30" pageNumber="31">S. marpesia</emphasis>
</taxonomicName>
can be separated from all other nocturnal species by its lack of mesosternal processes and by its coloration; the integument is black throughout except metasomal segment 3–6 are orangish and by the setal coloration of the vertex, pronotum, mesonotum and metasomal segment 2 that varies from silver to orange. The mandible is moderately dilated, distally little or scarcely wider than at tooth, the ventral basal tooth of the mandible is small, and the apex is vertical (see
<bibRefCitation id="EFB94B021616FFB7FCB9EF4FFC13FDD9" author="Pitts, J. P." box="[807,931,556,580]" firstAuthor="Pitts" journalOrPublisher="Zootaxa" pageId="30" pageNumber="31" pagination="1 - 20" part="1248" refId="ref58473" refString="Pitts, J. P. (2006) Review of the Sphaeropthalma imperialis species-group (Hymenoptera: Mutillidae), with descriptions of females and taxonomic notes. Zootaxa, 1248, 1 - 20." title="Review of the Sphaeropthalma imperialis species-group (Hymenoptera: Mutillidae), with descriptions of females and taxonomic notes" type="journal article" year="2006">Pitts, 2006</bibRefCitation>
:
<figureCitation id="13132A761616FFB7FC33EF4FFC5EFDD9" box="[941,1006,556,580]" captionStart="FIGURES 2 – 11" captionStartId="8.[151,264,1614,1637]" captionTargetBox="[169,1430,419,1583]" captionTargetId="figure@8.[150,1436,411,1593]" captionTargetPageId="8" captionText="FIGURES 2 – 11. Male genitalia. 2. Odontophotopsis acmaea: dorsal view (left) and internal view (right); 3. O. armata: dorsal view and lateral view of cuspis (inset); 4. O. bellona: dorsal view; 5. O. mamata: dorsal view and lateral view of cuspis (inset); 6. O. microdonta: dorsal view (right) and ventral view (left); 7. O. piute: dorsal view; 8. O. serca: dorsal view and lateral view of cuspis (inset); 9. Sphaeropthalma nana: internal view; 10. S. orestes: dorsal view (left) and ventral view (right); and 11. S. parkeri: dorsal view (left) and ventral view (right)." httpUri="https://zenodo.org/record/283117/files/figure.png" pageId="30" pageNumber="31">Fig. 6</figureCitation>
) Also, the head is quadrate posteriorly being long and parallel behind the eyes (see
<bibRefCitation id="EFB94B021616FFB7FD0AEF2CFCBAFDFA" author="Pitts, J. P." box="[660,778,591,615]" firstAuthor="Pitts" journalOrPublisher="Zootaxa" pageId="30" pageNumber="31" pagination="1 - 20" part="1248" refId="ref58473" refString="Pitts, J. P. (2006) Review of the Sphaeropthalma imperialis species-group (Hymenoptera: Mutillidae), with descriptions of females and taxonomic notes. Zootaxa, 1248, 1 - 20." title="Review of the Sphaeropthalma imperialis species-group (Hymenoptera: Mutillidae), with descriptions of females and taxonomic notes" type="journal article" year="2006">Pitts 2006</bibRefCitation>
:
<figureCitation id="13132A761616FFB7FC88EF2CFCEBFDFA" box="[790,859,591,615]" captionStart="FIGURES 2 – 11" captionStartId="8.[151,264,1614,1637]" captionTargetBox="[169,1430,419,1583]" captionTargetId="figure@8.[150,1436,411,1593]" captionTargetPageId="8" captionText="FIGURES 2 – 11. Male genitalia. 2. Odontophotopsis acmaea: dorsal view (left) and internal view (right); 3. O. armata: dorsal view and lateral view of cuspis (inset); 4. O. bellona: dorsal view; 5. O. mamata: dorsal view and lateral view of cuspis (inset); 6. O. microdonta: dorsal view (right) and ventral view (left); 7. O. piute: dorsal view; 8. O. serca: dorsal view and lateral view of cuspis (inset); 9. Sphaeropthalma nana: internal view; 10. S. orestes: dorsal view (left) and ventral view (right); and 11. S. parkeri: dorsal view (left) and ventral view (right)." httpUri="https://zenodo.org/record/283117/files/figure.png" pageId="30" pageNumber="31">Fig. 2</figureCitation>
), the clypeus is deeply depressed below the dorsal mandibular margin, the sternal felt line is present, and by characteristic genitalic morphology (see
<bibRefCitation id="EFB94B021616FFB7FB75EF17FAD0FD11" author="Pitts, J. P." box="[1259,1376,628,652]" firstAuthor="Pitts" journalOrPublisher="Zootaxa" pageId="30" pageNumber="31" pagination="1 - 20" part="1248" refId="ref58473" refString="Pitts, J. P. (2006) Review of the Sphaeropthalma imperialis species-group (Hymenoptera: Mutillidae), with descriptions of females and taxonomic notes. Zootaxa, 1248, 1 - 20." title="Review of the Sphaeropthalma imperialis species-group (Hymenoptera: Mutillidae), with descriptions of females and taxonomic notes" type="journal article" year="2006">Pitts 2006</bibRefCitation>
:
<figureCitation id="13132A761616FFB7FAF5EF17FF51FD32" captionStart="FIGURES 12–21" captionStartId="12.[151,264,1978,2001]" captionTargetBox="[241,1347,1204,1948]" captionTargetId="figure@12.[237,1349,1191,1957]" captionTargetPageId="12" captionText="FIGURES 12–21. Male mandibles: 12. Odontophotopsis armata; 13. O. clypeata; 14. O. microdonta; 15. O. piute; 16. O. serca; 17. O. setifera; 18. Sphaeropthalma edwardsii; 19. S. orestes; 20. S. parkeri; and 21. S. uro." pageId="30" pageNumber="31">Figs 14–16</figureCitation>
). FEMALE. The female of this species is easily recognized by its unique color pattern (see
<bibRefCitation id="EFB94B021616FFB7FB70EFF4FAD4FD32" author="Pitts, J. P." box="[1262,1380,663,687]" firstAuthor="Pitts" journalOrPublisher="Zootaxa" pageId="30" pageNumber="31" pagination="1 - 20" part="1248" refId="ref58473" refString="Pitts, J. P. (2006) Review of the Sphaeropthalma imperialis species-group (Hymenoptera: Mutillidae), with descriptions of females and taxonomic notes. Zootaxa, 1248, 1 - 20." title="Review of the Sphaeropthalma imperialis species-group (Hymenoptera: Mutillidae), with descriptions of females and taxonomic notes" type="journal article" year="2006">Pitts 2006</bibRefCitation>
: Fig. 25). Other useful characters include the petiolate metasomal segment 1, the small ventral angulation located basally on the mandible, the granulate pygidium, and the presence of plumose setae especially on the fringes of the metasomal tergites.
</paragraph>
</subSubSection>
<subSubSection id="C33265781616FFB7FF59EE44FD9AFAAA" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8B9736F31616FFB7FF59EE44FD9AFAAA" blockId="30.[151,1437,411,1768]" pageId="30" pageNumber="31">
<emphasis id="B95CEAE11616FFB7FF59EE44FE05FCDD" bold="true" box="[199,437,807,832]" pageId="30" pageNumber="31">Material examined.</emphasis>
<typeStatus id="549388511616FFB7FE21EE4BFE44FCDD" box="[447,500,808,832]" pageId="30" pageNumber="31">Type</typeStatus>
material.
<materialsCitation id="3B403CAE1616FFB7FDEEEE4BFAA7FCA2" ID-GBIF-Occurrence="3744625182" box="[624,1303,807,832]" pageId="30" pageNumber="31" typeStatus="lectotype">
<typeStatus id="549388511616FFB7FDEEEE4BFD59FCA2" box="[624,745,808,831]" pageId="30" pageNumber="31" type="lectotype">
<emphasis id="B95CEAE11616FFB7FDEEEE4BFD59FCA2" bold="true" box="[624,745,808,831]" italics="true" pageId="30" pageNumber="31">Lectotypes</emphasis>
</typeStatus>
:
<taxonomicName id="4C284D701616FFB7FD62EE4AFC3EFCDD" box="[764,910,809,832]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="marpesia">
<emphasis id="B95CEAE11616FFB7FD62EE4AFC3EFCDD" box="[764,910,809,832]" italics="true" pageId="30" pageNumber="31">M. marpesia</emphasis>
</taxonomicName>
: Kansas,
<typeStatus id="549388511616FFB7FB9BEE4BFB8DFCDD" box="[1029,1085,808,832]" pageId="30" pageNumber="31">Type</typeStatus>
no. 4542 (
<collectionCode id="ED39AE361616FFB7FB5FEE44FABFFCA2" LSID="urn:lsid:biocol.org:col:34784" box="[1217,1295,807,831]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="30" pageNumber="31">ANSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B403CAE1616FFB7FAB8EE4BFE59FCF9" ID-GBIF-Occurrence="3744625243" pageId="30" pageNumber="31">
<taxonomicName id="4C284D701616FFB7FAB8EE4BFA24FCDD" box="[1318,1428,808,832]" class="Insecta" family="Mutillidae" genus="Sphaerophthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="luteola">
<emphasis id="B95CEAE11616FFB7FAB8EE4BFA24FCDD" box="[1318,1428,808,832]" italics="true" pageId="30" pageNumber="31">S. luteola</emphasis>
</taxonomicName>
: Utah,
<typeStatus id="549388511616FFB7FF43EE2EFEA4FCF8" box="[221,276,845,869]" pageId="30" pageNumber="31">Type</typeStatus>
no. 4543 (
<collectionCode id="ED39AE361616FFB7FE0DEE2FFE51FCF9" LSID="urn:lsid:biocol.org:col:34784" box="[403,481,844,868]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="30" pageNumber="31">ANSP</collectionCode>
)
</materialsCitation>
.
<typeStatus id="549388511616FFB7FE68EE28FDD9FCF9" box="[502,617,843,868]" pageId="30" pageNumber="31" type="holotype">
<emphasis id="B95CEAE11616FFB7FE68EE28FDD9FCF9" bold="true" box="[502,617,843,868]" italics="true" pageId="30" pageNumber="31">Holotypes</emphasis>
</typeStatus>
:
<materialsCitation id="3B403CAE1616FFB7FDE4EE2EFF49FC1A" ID-GBIF-Occurrence="3744624638" pageId="30" pageNumber="31">
<taxonomicName id="4C284D701616FFB7FDE4EE2EFCF6FCF9" box="[634,838,844,868]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="imperialiformis">
<emphasis id="B95CEAE11616FFB7FDE4EE2EFCF6FCF9" box="[634,838,844,868]" italics="true" pageId="30" pageNumber="31">S. imperialiformis</emphasis>
</taxonomicName>
: Kansas, Morton Co., 3200’,
<date id="FF9610331616FFB7FB01EE2EFAA6FCF9" box="[1183,1302,844,868]" pageId="30" pageNumber="31" value="1902-06">June 1902</date>
, F.H. Snow (
<collectionCode id="ED39AE361616FFB7FF01EE0CFF41FC1A" box="[159,241,879,903]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="30" pageNumber="31">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B403CAE1616FFB7FE97EE13FAFDFC1A" ID-GBIF-Occurrence="3744625220" box="[265,1357,879,904]" pageId="30" pageNumber="31">
<taxonomicName id="4C284D701616FFB7FE97EE13FDEAFC15" box="[265,602,880,904]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="imperialiformis" subSpecies="maricopae">
<emphasis id="B95CEAE11616FFB7FE97EE13FDEAFC15" box="[265,602,880,904]" italics="true" pageId="30" pageNumber="31">S. imperialiformis maricopae</emphasis>
</taxonomicName>
: Arizona, Phoenix,
<date id="FF9610331616FFB7FCD7EE0CFB9FFC1A" box="[841,1071,879,903]" pageId="30" pageNumber="31" value="1935-09-25">25 September 1935</date>
, R.H. Crandall (
<collectionCode id="ED39AE361616FFB7FB6DEE0CFAF5FC1A" box="[1267,1349,879,903]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="30" pageNumber="31">UMSP</collectionCode>
)
</materialsCitation>
. Other material.
<materialsCitation id="3B403CAE1616FFB7FE9DEEF7FB80FC31" ID-GBIF-Occurrence="3744625079" box="[259,1072,916,941]" pageId="30" pageNumber="31">
<emphasis id="B95CEAE11616FFB7FE9DEEF7FED4FC31" bold="true" box="[259,356,916,940]" pageId="30" pageNumber="31">Nevada,</emphasis>
Nye Co., AMNWR: Non-dune site 1: 3 ♂, LT,
<date id="FF9610331616FFB7FC13EEF7FB9CFC31" box="[909,1068,916,940]" pageId="30" pageNumber="31" value="2009-05-26" valueMax="2009-05-28" valueMin="2009-05-26">26–28.V.2009</date>
,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FBA4EEF6FA27FC31" ID-GBIF-Occurrence="3744624871" box="[1082,1431,916,940]" pageId="30" pageNumber="31">
7 ♂, LT,
<date id="FF9610331616FFB7FB3AEEF7FAE0FC31" box="[1188,1360,916,940]" pageId="30" pageNumber="31" value="2009-06-23" valueMax="2009-06-25" valueMin="2009-06-23">23–25.VI.2009</date>
, NFB
</materialsCitation>
;
<materialsCitation id="3B403CAE1616FFB7FF09EED4FDEEFC52" ID-GBIF-Occurrence="3744625475" box="[151,606,951,976]" pageId="30" pageNumber="31">
Non-dune site 2: 3 ♂, LT,
<date id="FF9610331616FFB7FE25EED4FDEAFC52" box="[443,602,951,975]" pageId="30" pageNumber="31" value="2009-05-26" valueMax="2009-05-28" valueMin="2009-05-26">26–28.V.2009</date>
,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FDFBEED4FCC9FC52" ID-GBIF-Occurrence="3744625143" box="[613,889,951,976]" pageId="30" pageNumber="31">
1 ♂, LT,
<date id="FF9610331616FFB7FD57EED4FCC5FC52" box="[713,885,951,975]" pageId="30" pageNumber="31" value="2009-06-23" valueMax="2009-06-25" valueMin="2009-06-23">23–25.VI.2009</date>
,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FC1EEED4FB72FC52" ID-GBIF-Occurrence="3744625097" box="[896,1218,951,976]" pageId="30" pageNumber="31">
2 ♂, LT,
<date id="FF9610331616FFB7FC7DEED4FBCDFC52" box="[995,1149,951,975]" pageId="30" pageNumber="31" value="2009-07-06" valueMax="2009-07-08" valueMin="2009-07-06">6–8.VII.2009</date>
, NFB
</materialsCitation>
;
<materialsCitation id="3B403CAE1616FFB7FB53EED4FE24FC69" ID-GBIF-Occurrence="3744624483" pageId="30" pageNumber="31">
Non-dune site 3: 1 ♀, PT
<date id="FF9610331616FFB7FF6FEEBFFE20FC69" box="[241,400,988,1012]" pageId="30" pageNumber="31" value="2009-05-12" valueMax="2009-05-14" valueMin="2009-05-12">12–14.V.2009</date>
,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FE3DEEBFFCA7FC69" ID-GBIF-Occurrence="3744625130" box="[419,791,988,1012]" pageId="30" pageNumber="31">
1 ♂, LT,
<date id="FF9610331616FFB7FD80EEBFFD7AFC69" box="[542,714,988,1012]" pageId="30" pageNumber="31" value="2009-06-23" valueMax="2009-06-25" valueMin="2009-06-23">23–25.VI.2009</date>
, NFB
</materialsCitation>
;
<materialsCitation id="3B403CAE1616FFB7FCB4EEBFFA91FC69" ID-GBIF-Occurrence="3744624545" box="[810,1313,988,1012]" pageId="30" pageNumber="31">
Non-dune site 4: 2 ♂, LT,
<date id="FF9610331616FFB7FBE3EEBFFAADFC69" box="[1149,1309,988,1012]" pageId="30" pageNumber="31" value="2009-05-26" valueMax="2009-05-28" valueMin="2009-05-26">26–28.V.2009</date>
,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FAAEEEBFFE39FB8A" ID-GBIF-Occurrence="3744624722" pageId="30" pageNumber="31">
4 ♂, LT,
<date id="FF9610331616FFB7FF09EE9CFEF3FB8A" box="[151,323,1023,1047]" pageId="30" pageNumber="31" value="2009-06-23" valueMax="2009-06-25" valueMin="2009-06-23">23–25.VI.2009</date>
, NFB
</materialsCitation>
;
<materialsCitation id="3B403CAE1616FFB7FE08EE9CFCDAFB8A" ID-GBIF-Occurrence="3744625482" box="[406,874,1023,1048]" pageId="30" pageNumber="31">
Sand dune site 1: 1 ♂, LT,
<date id="FF9610331616FFB7FD59EE9CFCD7FB8A" box="[711,871,1023,1047]" pageId="30" pageNumber="31" value="2009-05-12" valueMax="2009-05-14" valueMin="2009-05-12">12–14.V.2009</date>
,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FCEDEE9CFBCDFB8A" ID-GBIF-Occurrence="3744624502" box="[883,1149,1023,1048]" pageId="30" pageNumber="31">
4 ♂, LT,
<date id="FF9610331616FFB7FC47EE9CFBC8FB8A" box="[985,1144,1023,1047]" pageId="30" pageNumber="31" value="2009-05-26" valueMax="2009-05-28" valueMin="2009-05-26">26–28.V.2009</date>
,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FB1BEE9CFF61FBA1" ID-GBIF-Occurrence="3744624507" pageId="30" pageNumber="31">
1 ♂, LT,
<date id="FF9610331616FFB7FB72EE9CFA28FB8A" box="[1260,1432,1023,1047]" pageId="30" pageNumber="31" value="2009-06-23" valueMax="2009-06-25" valueMin="2009-06-23">23–25.VI.2009</date>
, NFB
</materialsCitation>
;
<materialsCitation id="3B403CAE1616FFB7FF42E947FD16FBA1" ID-GBIF-Occurrence="3744625511" box="[220,678,1060,1084]" pageId="30" pageNumber="31">
Sand dune site 2: 1 ♂, LT,
<date id="FF9610331616FFB7FD9DE947FD12FBA1" box="[515,674,1060,1084]" pageId="30" pageNumber="31" value="2009-05-12" valueMax="2009-05-14" valueMin="2009-05-12">12–14.V.2009</date>
,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FD32E947FC46FBA1" ID-GBIF-Occurrence="3744625048" box="[684,1014,1060,1084]" pageId="30" pageNumber="31">
5 ♂, LT,
<date id="FF9610331616FFB7FC90E947FC1DFBA1" box="[782,941,1060,1084]" pageId="30" pageNumber="31" value="2009-05-26" valueMax="2009-05-28" valueMin="2009-05-26">26–28.V.2009</date>
, NFB,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FC62E947FA2CFBA1" ID-GBIF-Occurrence="3744625280" box="[1020,1436,1060,1084]" pageId="30" pageNumber="31">
3 ♂, LT,
<date id="FF9610331616FFB7FBC0E947FB4DFBA1" box="[1118,1277,1060,1084]" pageId="30" pageNumber="31" value="2009-06-08" valueMax="2009-06-15" valueMin="2009-06-08">8–15.VI.2009</date>
, NFB &amp; DAT,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FF09E92BFE0DFBC2" ID-GBIF-Occurrence="3744625180" box="[151,445,1095,1120]" pageId="30" pageNumber="31">
7 ♂, LT,
<date id="FF9610331616FFB7FE9AE924FE09FBC2" box="[260,441,1095,1119]" pageId="30" pageNumber="31" value="2009-07-21" valueMax="2009-07-23" valueMin="2009-07-21">21–23.VII.2009</date>
,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FE56E924FC8CFBC2" ID-GBIF-Occurrence="3744624470" box="[456,828,1095,1120]" pageId="30" pageNumber="31">
1 ♂, LT,
<date id="FF9610331616FFB7FDABE924FD43FBC2" box="[565,755,1095,1119]" pageId="30" pageNumber="31" value="2009-08-17" valueMax="2009-08-19" valueMin="2009-08-17">17–19.VIII.2009</date>
, NFB
</materialsCitation>
;
<materialsCitation id="3B403CAE1616FFB7FCD5E924FA9EFBC2" ID-GBIF-Occurrence="3744624932" box="[843,1326,1095,1120]" pageId="30" pageNumber="31">
Sand dune site 3: 2 ♂, LT,
<date id="FF9610331616FFB7FB15E924FA9AFBC2" box="[1163,1322,1095,1119]" pageId="30" pageNumber="31" value="2009-05-26" valueMax="2009-05-28" valueMin="2009-05-26">26–28.V.2009</date>
,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FAA7E924FEF7FB19" ID-GBIF-Occurrence="3744624830" pageId="30" pageNumber="31">
4 ♂, LT,
<date id="FF9610331616FFB7FF09E90FFEF3FB19" box="[151,323,1132,1156]" pageId="30" pageNumber="31" value="2009-06-23" valueMax="2009-06-25" valueMin="2009-06-23">23–25.VI.2009</date>
,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FED1E90FFDE1FB19" ID-GBIF-Occurrence="3744625373" box="[335,593,1132,1156]" pageId="30" pageNumber="31">
2 ♂, LT,
<date id="FF9610331616FFB7FE2AE90FFDFEFB19" box="[436,590,1132,1156]" pageId="30" pageNumber="31" value="2009-07-06" valueMax="2009-07-08" valueMin="2009-07-06">6–8.VII.2009</date>
,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FDC4E90FFCC8FB19" ID-GBIF-Occurrence="3744625412" box="[602,888,1132,1156]" pageId="30" pageNumber="31">
2 ♂, LT,
<date id="FF9610331616FFB7FD21E90FFCC4FB19" box="[703,884,1132,1156]" pageId="30" pageNumber="31" value="2009-07-21" valueMax="2009-07-23" valueMin="2009-07-21">21–23.VII.2009</date>
,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FC1EE90FFB7EFB19" ID-GBIF-Occurrence="3744625328" box="[896,1230,1132,1156]" pageId="30" pageNumber="31">
2 ♂, LT,
<date id="FF9610331616FFB7FC78E90FFB38FB19" box="[998,1160,1132,1156]" pageId="30" pageNumber="31" value="2009-08-04" valueMax="2009-08-06" valueMin="2009-08-04">4–6.VIII.2009</date>
, NFB
</materialsCitation>
;
<materialsCitation id="3B403CAE1616FFB7FB44E90FFDA9FB35" ID-GBIF-Occurrence="3744625211" pageId="30" pageNumber="31">
Sand dune site 4: 3 ♂, LT,
<date id="FF9610331616FFB7FF61E9ECFEC3FB3A" box="[255,371,1167,1191]" pageId="30" pageNumber="31" value="2008-06-09">9.VI.2008</date>
, NFB &amp; DAT,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FDBCE9ECFC9DFB3A" ID-GBIF-Occurrence="3744624807" box="[546,813,1167,1192]" pageId="30" pageNumber="31">
1 ♂, LT,
<date id="FF9610331616FFB7FD14E9ECFC9AFB3A" box="[650,810,1167,1191]" pageId="30" pageNumber="31" value="2009-05-12" valueMax="2009-05-14" valueMin="2009-05-12">12–14.V.2009</date>
,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FCA8E9ECFB38FB3A" ID-GBIF-Occurrence="3744624826" box="[822,1160,1167,1192]" pageId="30" pageNumber="31">
3 ♂, LT,
<date id="FF9610331616FFB7FC01E9ECFB8EFB3A" box="[927,1086,1167,1191]" pageId="30" pageNumber="31" value="2009-05-26" valueMax="2009-05-28" valueMin="2009-05-26">26–28.V.2009</date>
, NFB,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FB0FE9ECFE9FFB51" ID-GBIF-Occurrence="3744624916" pageId="30" pageNumber="31">
8 ♂, LT,
<date id="FF9610331616FFB7FB67E9ECFA28FB3A" box="[1273,1432,1167,1191]" pageId="30" pageNumber="31" value="2009-06-08" valueMax="2009-06-15" valueMin="2009-06-08">8–15.VI.2009</date>
, NFB &amp; DAT,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FEA8E9D7FD23FB51" ID-GBIF-Occurrence="3744625503" box="[310,659,1204,1228]" pageId="30" pageNumber="31">
8 ♂, LT,
<date id="FF9610331616FFB7FE03E9D7FDF9FB51" box="[413,585,1204,1228]" pageId="30" pageNumber="31" value="2009-06-23" valueMax="2009-06-25" valueMin="2009-06-23">23–25.VI.2009</date>
, NFB,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FD05E9D7FC68FB51" ID-GBIF-Occurrence="3744625244" box="[667,984,1204,1228]" pageId="30" pageNumber="31">
1 ♂, LT,
<date id="FF9610331616FFB7FC9FE9D7FC22FB51" box="[769,914,1204,1228]" pageId="30" pageNumber="31" value="2009-09-04" valueMax="2009-09-06" valueMin="2009-09-04">4–6.IX.2009</date>
, NFB
</materialsCitation>
;
<materialsCitation id="3B403CAE1616FFB7FC7AE9D7FED7FB72" ID-GBIF-Occurrence="3744624671" pageId="30" pageNumber="31">
Sand dune site 5: 4 ♂, LT,
<date id="FF9610331616FFB7FA8BE9D7FA27FB51" box="[1301,1431,1204,1228]" pageId="30" pageNumber="31" value="2008-06-24">24.VI.2008</date>
, NFB, DAT &amp; JPP,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FEF1E9B4FD20FB72" ID-GBIF-Occurrence="3744624727" box="[367,656,1239,1264]" pageId="30" pageNumber="31">
1 ♂, LT,
<date id="FF9610331616FFB7FE48E9B4FD3CFB72" box="[470,652,1239,1263]" pageId="30" pageNumber="31" value="2009-07-21" valueMax="2009-07-23" valueMin="2009-07-21">21–23.VII.2009</date>
,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FD06E9B4FBB5FB72" ID-GBIF-Occurrence="3744625150" box="[664,1029,1239,1264]" pageId="30" pageNumber="31">
2 ♂, LT,
<date id="FF9610331616FFB7FD61E9B4FC0EFB72" box="[767,958,1239,1263]" pageId="30" pageNumber="31" value="2009-08-17" valueMax="2009-08-19" valueMin="2009-08-17">17–19.VIII.2009</date>
, NFB
</materialsCitation>
;
<materialsCitation id="3B403CAE1616FFB7FB8FE9B4FE9BFA89" ID-GBIF-Occurrence="3744624827" pageId="30" pageNumber="31">
Copeland site: 1 ♂, LT,
<date id="FF9610331616FFB7FABCE9B4FA28FB72" box="[1314,1432,1239,1263]" pageId="30" pageNumber="31" value="2008-05-30">30.V.2008</date>
, NFB &amp; DAT
</materialsCitation>
;
<materialsCitation id="3B403CAE1616FFB7FEA8E99FFC05FA89" ID-GBIF-Occurrence="3744624819" box="[310,949,1276,1300]" pageId="30" pageNumber="31">
Mesquite site 1: 1 ♂, PT,
<date id="FF9610331616FFB7FDC7E99FFD6BFA89" box="[601,731,1276,1300]" pageId="30" pageNumber="31" value="2008-06-24">24.VI.2008</date>
, NFB, DAT &amp; JPP,
</materialsCitation>
<materialsCitation id="3B403CAE1616FFB7FC23E99FFB49FA89" ID-GBIF-Occurrence="3744624547" box="[957,1273,1276,1300]" pageId="30" pageNumber="31">
1 ♀, PT,
<date id="FF9610331616FFB7FBBDE99FFB03FA89" box="[1059,1203,1276,1300]" pageId="30" pageNumber="31" value="2008-09-02" valueMax="2008-09-03" valueMin="2008-09-02">2–3.IX.2008</date>
, NFB
</materialsCitation>
;
<materialsCitation id="3B403CAE1616FFB7FA9BE99FFD95FAAA" ID-GBIF-Occurrence="3744625429" pageId="30" pageNumber="31">
Mesquite site 3: 1 ♀, 1 ♂, PT,
<date id="FF9610331616FFB7FECEE87CFE51FAAA" box="[336,481,1311,1335]" pageId="30" pageNumber="31" value="2008-09-02" valueMax="2008-09-03" valueMin="2008-09-02">2–3.IX.2008</date>
, NFB
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33265781616FFB7FF59E820FDFCFAE2" pageId="30" pageNumber="31" type="distribution">
<paragraph id="8B9736F31616FFB7FF59E820FDFCFAE2" blockId="30.[151,1437,411,1768]" pageId="30" pageNumber="31">
<emphasis id="B95CEAE11616FFB7FF59E820FED0FAC1" bold="true" box="[199,352,1347,1372]" pageId="30" pageNumber="31">Distribution.</emphasis>
<collectingCountry id="F33F76631616FFB7FEF9E827FE2FFAC1" box="[359,415,1348,1372]" name="United States of America" pageId="30" pageNumber="31">USA</collectingCountry>
(Arizona, California, Colorado, Idaho, Kansas, Nevada, New
<collectingCountry id="F33F76631616FFB7FBC5E827FB03FAC1" box="[1115,1203,1348,1372]" name="Mexico" pageId="30" pageNumber="31">Mexico</collectingCountry>
, Oklahoma, Oregon, Texas, Utah and Washington),
<collectingCountry id="F33F76631616FFB7FE71E804FDF8FAE2" box="[495,584,1383,1407]" name="Mexico" pageId="30" pageNumber="31">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33265781616FFB7FF59E8E8FAE8FA5A" pageId="30" pageNumber="31" type="biology_ecology">
<paragraph id="8B9736F31616FFB7FF59E8E8FAE8FA5A" blockId="30.[151,1437,411,1768]" pageId="30" pageNumber="31">
<emphasis id="B95CEAE11616FFB7FF59E8E8FE9BFA39" bold="true" box="[199,299,1419,1444]" pageId="30" pageNumber="31">Activity.</emphasis>
Males were active from mid-spring through late summer (May through early September). Two females were collected in early autumn (
<date id="FF9610331616FFB7FDC2E8CCFCA6FA5A" box="[604,790,1455,1479]" pageId="30" pageNumber="31" value="2008-09">September 2008</date>
) and one was collected in early spring (
<date id="FF9610331616FFB7FB4DE8CCFAFCFA5A" box="[1235,1356,1455,1479]" pageId="30" pageNumber="31" value="2009-05">May 2009</date>
).
</paragraph>
</subSubSection>
<subSubSection id="C33265781616FFB7FF59E8B7FC6DF97A" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8B9736F31616FFB7FF59E8B7FCCFF9E0" blockId="30.[151,1437,411,1768]" pageId="30" pageNumber="31">
<emphasis id="B95CEAE11616FFB7FF59E8B7FE8BFA71" bold="true" box="[199,315,1492,1516]" pageId="30" pageNumber="31">Remarks.</emphasis>
<taxonomicName id="4C284D701616FFB7FEDFE8B7FDD8FA71" box="[321,616,1492,1516]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="marpesia">
<emphasis id="B95CEAE11616FFB7FEDFE8B7FDD8FA71" box="[321,616,1492,1516]" italics="true" pageId="30" pageNumber="31">Sphaeropthalma marpesia</emphasis>
</taxonomicName>
were distributed uniformly over sand dune and non-dune habitats (U=20, p=0.2). Three female and ninety-two male
<taxonomicName id="4C284D701616FFB7FD14E89BFCA1F98D" box="[650,785,1528,1552]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="marpesia">
<emphasis id="B95CEAE11616FFB7FD14E89BFCA1F98D" box="[650,785,1528,1552]" italics="true" pageId="30" pageNumber="31">S. marpesia</emphasis>
</taxonomicName>
were collected throughout the course of this study. The females were collected in May and September via pitfall trapping, and males were collected from May through early September via light and pitfall trapping.
<taxonomicName id="4C284D701616FFB7FD35EB23FC64F9C5" box="[683,980,1600,1624]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="marpesia">
<emphasis id="B95CEAE11616FFB7FD35EB23FC64F9C5" box="[683,980,1600,1624]" italics="true" pageId="30" pageNumber="31">Sphaeropthalma marpesia</emphasis>
</taxonomicName>
was not found at the NTS.
<bibRefCitation id="EFB94B021616FFB7FA88EB5CFA2CF9CA" author="Pitts, J. P." box="[1302,1436,1599,1623]" firstAuthor="Pitts" journalOrPublisher="Zootaxa" pageId="30" pageNumber="31" pagination="1 - 20" part="1248" refId="ref58473" refString="Pitts, J. P. (2006) Review of the Sphaeropthalma imperialis species-group (Hymenoptera: Mutillidae), with descriptions of females and taxonomic notes. Zootaxa, 1248, 1 - 20." title="Review of the Sphaeropthalma imperialis species-group (Hymenoptera: Mutillidae), with descriptions of females and taxonomic notes" type="journal article" year="2006">Pitts (2006)</bibRefCitation>
associated the females of this species and discussed the taxonomy.
</paragraph>
<paragraph id="8B9736F31616FFB7FF59EBE4FC6DF97A" blockId="30.[151,1437,411,1768]" pageId="30" pageNumber="31">
The
<typeStatus id="549388511616FFB7FF64EBE4FEDCF902" box="[250,364,1671,1695]" pageId="30" pageNumber="31" type="lectotype">lectotypes</typeStatus>
of
<taxonomicName id="4C284D701616FFB7FE0FEBEAFDAFF93D" box="[401,543,1673,1696]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="marpesia">
<emphasis id="B95CEAE11616FFB7FE0FEBEAFDAFF93D" box="[401,543,1673,1696]" italics="true" pageId="30" pageNumber="31">M. marpesia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C284D701616FFB7FDC8EBEBFD70F93D" box="[598,704,1672,1696]" class="Insecta" family="Mutillidae" genus="Sphaerophthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="luteola">
<emphasis id="B95CEAE11616FFB7FDC8EBEBFD70F93D" box="[598,704,1672,1696]" italics="true" pageId="30" pageNumber="31">S. luteola</emphasis>
</taxonomicName>
were selected based on having extruded genitalia and the quality of the specimens. The label data for
<taxonomicName id="4C284D701616FFB7FDB0EBCEFD0BF959" box="[558,699,1709,1732]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="marpeisa">
<emphasis id="B95CEAE11616FFB7FDB0EBCEFD0BF959" box="[558,699,1709,1732]" italics="true" pageId="30" pageNumber="31">M. marpeisa</emphasis>
</taxonomicName>
are as follows [Kan. Snow] [
<typeStatus id="549388511616FFB7FB9FEBCEFB8CF958" box="[1025,1084,1709,1733]" pageId="30" pageNumber="31">Type</typeStatus>
no. 4542] [
<taxonomicName id="4C284D701616FFB7FB23EBCEFA25F959" authority="Blake" authorityName="Blake" box="[1213,1429,1708,1732]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="marpesia">M. marpesia Blake</taxonomicName>
] and the label data for
<taxonomicName id="4C284D701616FFB7FE14EBB3FE44F975" box="[394,500,1744,1768]" class="Insecta" family="Mutillidae" genus="Sphaerophthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="luteola">
<emphasis id="B95CEAE11616FFB7FE14EBB3FE44F975" box="[394,500,1744,1768]" italics="true" pageId="30" pageNumber="31">S. luteola</emphasis>
</taxonomicName>
are [Utah] [
<typeStatus id="549388511616FFB7FDE4EBB3FD05F975" box="[634,693,1744,1768]" pageId="30" pageNumber="31">Type</typeStatus>
no. 4543] [
<taxonomicName id="4C284D701616FFB7FCABEBACFC62F97A" authority="Blake" authorityName="Blake" box="[821,978,1743,1767]" class="Insecta" family="Mutillidae" genus="Sphaerophthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="luteola">luteola Blake</taxonomicName>
].
</paragraph>
</subSubSection>
</treatment>
</document>
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