465 lines
70 KiB
XML
465 lines
70 KiB
XML
<document id="7EB4A0588F19487CBD0E9075A012B181" ENCODING="UTF-8" ID-GBIF-Dataset="55d6386c-4e4e-4f94-a5c1-752b6601e83f" ModsDocID="z01040p049" checkinTime="1247058950468" checkinUser="thomas" docAuthor="Wolmar B. Wosiacki" docDate="2005" docId="318F4901348EAA861AA5C73C3D8FBE86" docLanguage="en" docName="2005_Wosiacki_gg1.xml" docOrigin="Zootaxa 1040" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:19E95D15-4A14-421D-88F0-6E75B6585D50" docTitle="Trichomycterus iheringi" docType="treatment" docVersion="12" lastPageNumber="61" masterDocId="CF9DBF59DC5CFC9BB834F546D982B567" masterDocTitle="A new species of Trichomycterus (Siluriformes: Trichomycteridae) from south Brazil and redescription of T. iheringi (Eigenmann)." masterLastPageNumber="64" masterPageNumber="49" pageNumber="56" updateTime="1701314188202" updateUser="plazi">
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<mods:titleInfo id="1506F9D0160E29714FC402213107D6BC">
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<mods:title id="C710056CBDAAC61CD58D63CA8995C8CB">A new species of Trichomycterus (Siluriformes: Trichomycteridae) from south Brazil and redescription of T. iheringi (Eigenmann).</mods:title>
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<mods:namePart id="428D8A18EA906BCE0FDA0A8F4958642F">Wolmar B. Wosiacki</mods:namePart>
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<mods:typeOfResource id="B1480453CC0DF8323C73B6785BBC7D98">text</mods:typeOfResource>
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<mods:title id="4632D55105DFBE9F7E9D503DEF031260">Zootaxa</mods:title>
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<mods:part id="0A5F7414629C845481928558C9224824">
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<mods:date id="3B3603AA63640423A353D1D7C941DB37">2005</mods:date>
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<mods:number id="1DEC3CABFE3BEDB9D69924596E599934">1040</mods:number>
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<mods:start id="6E2034229296AE0BE5378241687032AC">49</mods:start>
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<mods:classification id="F3B5265EE702A9265F275D6601CD06C7">journal article</mods:classification>
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<mods:identifier id="13F492EBB38EE3FE0FA16A168C00F16E" type="Plazi-Custom">z01040p049</mods:identifier>
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<mods:identifier id="A211A4C3A2536DB3712CFCBFC32B5990" type="ZooBank">19E95D15-4A14-421D-88F0-6E75B6585D50</mods:identifier>
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<treatment id="318F4901348EAA861AA5C73C3D8FBE86" ID-DOI="http://doi.org/10.5281/zenodo.6266782" ID-GBIF-Taxon="100124139" ID-Zenodo-Dep="6266782" LSID="urn:lsid:plazi:treatment:318F4901348EAA861AA5C73C3D8FBE86" httpUri="http://treatment.plazi.org/id/318F4901348EAA861AA5C73C3D8FBE86" lastPageNumber="61" pageNumber="56">
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<subSubSection id="2221BB13A1B46C427854FFBDE35EF57D" type="nomenclature">
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<paragraph id="2EDFC04617575BF5E7C309C4831A2D0E" pageNumber="56">
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<taxonomicName id="C338C9C91792A49D4A423BE1247F44C8" ID-CoL="58FHK" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="iheringi">Trichomycterus iheringi (Eigenmann, 1917)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="1254913C42D70D255C822DC71ECECF94" type="description">
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<paragraph id="00F52E4E7C65BD7F8DB70B768B9C2C72" pageNumber="56">(Figs. 3, 4; Table 1)</paragraph>
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</subSubSection>
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<subSubSection id="9E0B15A263AEBD4415D6361947008537" type="reference_group">
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<paragraph id="1AE3D9F5239494D35BF62BB89E2E3C53" pageNumber="56">
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<taxonomicName id="73FEC890D892B9F8FDDDEEA30D6A9294" LSID-ZBK="urn:lsid:zoobank.org:act:10ADC134-AEC4-4D94-AFE1-9E9D423FFFA9" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pygidium iheringi Eigenmann 1917:697" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="iheringi">Pygidium iheringi Eigenmann, 1917</taxonomicName>
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:697 (Brazil,
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<normalizedToken id="3B8EB9F30ECAB11409AA65A5AE678DF5" originalValue="São">Sao</normalizedToken>
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Paulo). Eigenmann, 1918:330. Henn, 1928:80. Gosline, 1945:60. Ibarra & Stewart, 1987:73.
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</paragraph>
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<paragraph id="368EAFCB18EE0BCE3AA36DF9380A6A70" pageNumber="56">
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<taxonomicName id="BD89192F4119AE404A2D9808B08D1DCA" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="iheringi">Trichomycterus iheringi</taxonomicName>
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- Miranda-Ribeiro, 1918:725. Fowler 1954:27. Burgess, 1989:322. Bizerril, 1994:623. de Pinna & Wosiacki, 2003:282. Costa, 1992:103. Eschmeyer, 1998:760. Barbosa & Costa, 2003:287. Bockmann & Sazima, 2004:61. Bockmann, Casatti & de Pinna, 2004:226. Triques & Vono, 2004:162. Wosiacki, 2004:9. Wosiacki & Garavello, 2004:9.
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</paragraph>
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</subSubSection>
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<subSubSection id="11FFD616148303B0A5DFC7E63309B773" type="nomenclature">
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<paragraph id="196D531C4B1C0BC61BC632936961A55A" pageNumber="56">
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<materialsCitation id="70E938036D0E0ADCE3A01CDFE726D3BC" ID-GBIF-Occurrence="923893617" country="Brazil" stateProvince="São Paulo">
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<typeStatus id="44CFFEDDD2362120430E7F52D75177C4">Holotype</typeStatus>
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:
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<collectionCode id="4AFA43EC92E2CBDC2C40937CB69FA687" collectionName="USA, California, San Francisco, California Academy of Sciences">CAS</collectionCode>
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64585 (Ex. IU 10785), 139.6mm SL, Santos,
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<collectingRegion id="B0805922062F82F98DF3C96BFE0D8D93">
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<normalizedToken id="2EFD56840C0E69504DBB71059B9826D3" originalValue="São">Sao</normalizedToken>
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Paulo
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</collectingRegion>
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,
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<collectingCountry id="A51E0C1A8D844A5166051CC2CDE32B31">Brazil</collectingCountry>
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, collected by R. von Ihering.
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</materialsCitation>
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</paragraph>
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<paragraph id="FCD85782901ADB7CFCEEA1ED757F7677" pageNumber="56">
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<materialsCitation id="8E968B580A40CA853147E56CBC655055" ID-GBIF-Occurrence="923893613" country="Brazil" stateProvince="São Paulo">
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<typeStatus id="182857B65CBC854125D6870C04623880">Paratypes</typeStatus>
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:
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<collectionCode id="4B17B711E9A3E9F90510141AB0971C91" collectionName="USA, California, San Francisco, California Academy of Sciences">CAS</collectionCode>
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64586, 3 ex. (117.6 128mm SL). Orig. n. IU 10785, collected with holotype;
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</materialsCitation>
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<materialsCitation id="23083F814F4B1BA6FA170944864C6479" ID-GBIF-Occurrence="923893614" country="Brazil" stateProvince="São Paulo">
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<collectionCode id="50A1C9FF13A09156BA9A747D32E4B3CB" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)">FMNH</collectionCode>
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58074, 2 ex. (125.7-134.1mm, X-rays), Sapina,
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<collectingRegion id="F968E80470801F1ED428918BDA70A854">
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<normalizedToken id="8E1D78C862E6CAA27E8FD34DE3D4740B" originalValue="São">Sao</normalizedToken>
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Paulo
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</collectingRegion>
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,
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<collectingCountry id="788F9847BBC21E2D54F4FB9920F475B4">Brazil</collectingCountry>
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, collected by J.D.Haseman,
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<date id="312DFFA84C785363894BE52E1409EBEE" day="23" month="07" value="1908-07-23" year="1908">23 July 1908</date>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="0A787F56C423D18DEA932CC4A9BED76F" type="diagnosis">
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<paragraph id="CC615E8CE9A06C47D5AB93A88FAB3589" pageNumber="57">
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Diagnosis: No uniquely derived features were found to diagnose
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<taxonomicName id="C3023289233A356FA754C1DF4C97603D" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="iheringi">T. iheringi</taxonomicName>
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.
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<taxonomicName id="50B7C1BF21B25B07DD1108808AB7EA63" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="iheringi">Trichomycterus iheringi</taxonomicName>
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shares with
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<taxonomicName id="173DDEDCE16E33D3CBC83731A883D8BF" LSID-ZBK="urn:lsid:zoobank.org:act:0ED36072-3C3B-41A0-81EA-AF017739D4DB" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus castroi de Pinna 1992:90" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="castroi">T. castroi</taxonomicName>
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,
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<taxonomicName id="627D43E8FA121E38B03F61541728C906" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="davisi">T. davisi</taxonomicName>
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,
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<taxonomicName id="22FFBFB0540511981FC279EC775D17DF" LSID-ZBK="urn:lsid:zoobank.org:act:78F93270-ED22-48B6-8329-CB52BBCD4664" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus guaraquessaba Wosiacki 2005" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="guaraquessaba">T. guaraquessaba</taxonomicName>
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,
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<taxonomicName id="A901CE7684DA8B3B29EAFF4CD59FCEEC" LSID-ZBK="urn:lsid:zoobank.org:act:656BBD86-BAC9-4255-B723-0776CFBF111F" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus mboycy Wosiacki & Garavello 2004:8" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="mboycy">T. mboycy</taxonomicName>
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,
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<taxonomicName id="9130F6DB5398BFBABC8B2F8EA6C763D1" LSID-ZBK="urn:lsid:zoobank.org:act:D11FBE09-11EC-43EF-8BD7-21BE3F735BC1" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus naipi Wosiacki & Garavello 2004:2" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="naipi">T. naipi</taxonomicName>
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,
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<taxonomicName id="9407841725EA39E91AC37EC56615914A" LSID-ZBK="urn:lsid:zoobank.org:act:5E60557F-9A8F-4467-BF3B-8C7E55D2FB7F" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus papilliferus Wosiacki & Garavello 2004:5" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="papilliferus">T. papilliferus</taxonomicName>
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,
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<taxonomicName id="28CE767557345049782D47FAF58AD33A" LSID-ZBK="urn:lsid:zoobank.org:act:090F10B3-41E6-426A-BE70-89E47F702B19" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus plumbeus Wosiacki & Garavello 2004:13" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="plumbeus">T. plumbeus</taxonomicName>
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,
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<taxonomicName id="2D46C307BA891A02093AF1E921BA4EC5" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="stawiarski">T. stawiarski</taxonomicName>
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, and
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<taxonomicName id="02EC3BAEC884CC5719AE0084E9F62F69" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="zonatus">T. zonatus</taxonomicName>
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the first pectoral-fin ray not prolonged as a filament and differs from all other congeners from south and southeastern Brazil (
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<taxonomicName id="D2EDB85E3A09A9A5FA3FCC3D6DDBC222" LSID-ZBK="urn:lsid:zoobank.org:act:D510F712-BBB5-4761-BC3A-B3F198A1B84E" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus albinotatus Costa 1992:102" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="albinotatus">T. albinotatus</taxonomicName>
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,
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<taxonomicName id="E9B45AA210E8EAE9D657D809A149D787" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="alternatus">T. alternatus</taxonomicName>
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,
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<taxonomicName id="5B25046C4F654CC0C302FF3527BD35F8" LSID-ZBK="urn:lsid:zoobank.org:act:5EF275E1-6327-4FEA-BE29-9E571D6DACF7" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus auroguttatus Costa 1992:105" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="auroguttatus">T. auroguttatus</taxonomicName>
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,
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<taxonomicName id="606F5B24DFA128F197F3F3F8F80C24C7" LSID-ZBK="urn:lsid:zoobank.org:act:89D8ECF9-B9D9-440E-98A1-9EDD49B8DBC5" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus bahianus Costa 1992:105" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="bahianus">T. bahianus</taxonomicName>
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,
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<taxonomicName id="2092EFFF6BEA457E63B9C7C7A9855966" LSID-ZBK="urn:lsid:zoobank.org:act:84D84BFD-1592-4583-B2D4-0BB6B004D9C3" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus brasiliensis Lütken & Reinhardt 1874:29 [1]" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="brasiliensis">T. brasiliensis</taxonomicName>
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,
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<taxonomicName id="D47D669628EDC2348C12766F8B78CF9C" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="candidu">T. candidu</taxonomicName>
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,
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<taxonomicName id="77D5B2C0D2CFCDBFEB60ACF9E81A55BE" LSID-ZBK="urn:lsid:zoobank.org:act:35245433-EEF9-4FD8-8A63-CDC3C190B8EE" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus caudofasciatus Alencar & Costa 2004:5" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="caudofasciatus">T. caudofasciatus</taxonomicName>
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,
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<taxonomicName id="7760C6B85E290B467C4A454513D7D5E7" LSID-ZBK="urn:lsid:zoobank.org:act:C4969FB6-8D9B-4A86-84F0-625DBAC0B4B2" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus concolor Costa 1992:107" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="concolor">T. concolor</taxonomicName>
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,
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<taxonomicName id="0E104A7D1DE2D18664782F772A01A53D" LSID-ZBK="urn:lsid:zoobank.org:act:197E097C-2451-43DE-B902-111021AAEB5D" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus giganteus Lima & Costa 2004:2" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="giganteus">T. giganteus</taxonomicName>
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,
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<taxonomicName id="41B9357C5EF8AC776F7BC21E97E6622C" LSID-ZBK="urn:lsid:zoobank.org:act:89A16C5C-802B-438F-8649-96750C9A2864" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus goeldii Boulenger 1896:154" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="goeldii">T. goeldii</taxonomicName>
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,
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<taxonomicName id="B065637EBB22CA6D4AD0AA23D7472AB9" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="immaculatus">T. immaculatus</taxonomicName>
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,
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<taxonomicName id="9F4D7DD7A35F3DC71410F37EF93833C2" LSID-ZBK="urn:lsid:zoobank.org:act:7EF24AEB-4DFD-4085-AE13-2BEE3B431248" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus itacambirussu Triques & Vono 2004:165" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="itacambirussu">T. itacambirussu</taxonomicName>
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,
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<taxonomicName id="FC79D1DAE3623C170A3FE3A8E25E0A56" LSID-ZBK="urn:lsid:zoobank.org:act:4B83205A-2285-4A82-9F41-E15DD605A259" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus brasiliensis itatiayae Miranda-Ribeiro 1906:p.? [15]" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="itatiayae">T. itatiayae</taxonomicName>
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,
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<taxonomicName id="4402ED931EEAC3D773409EDB59960805" LSID-ZBK="urn:lsid:zoobank.org:act:7254A8B5-08D6-4B2E-9237-0F939A14DD75" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus jequitinhonhae Triques & Vono 2004:167" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="jequitinhonhae">T. jequitinhonhae</taxonomicName>
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,
|
|
<taxonomicName id="79F71BC389F1519E066DD10C49FFC338" LSID-ZBK="urn:lsid:zoobank.org:act:4733DBED-E135-4493-B4F6-1C35E826A5E2" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus landinga Triques & Vono 2004:162" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="landinga">T. landinga</taxonomicName>
|
|
,
|
|
<taxonomicName id="A9E718205CCF8DDA6D52D3AFD963848B" LSID-ZBK="urn:lsid:zoobank.org:act:C143F305-A280-49C3-821C-B414A05BCCE7" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus longibarbatus Costa 1992:104" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="longibarbatus">T. longibarbatus</taxonomicName>
|
|
,
|
|
<taxonomicName id="9D1194DBC5341C0CAB8F0940340D0A31" LSID-ZBK="urn:lsid:zoobank.org:act:1F83BB4D-68B6-4DCC-B9D7-75BCBBFC34D0" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus maracaya Bockmann & Sazima 2004:63" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="maracaya">T. maracaya</taxonomicName>
|
|
,
|
|
<taxonomicName id="B8346A68550EF48A7EDC67882EFC2252" LSID-ZBK="urn:lsid:zoobank.org:act:D85474E7-5A0D-48AD-B35B-A0FB33DD0370" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus mimonha Costa 1992:106" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="mimonha">T. mimonha</taxonomicName>
|
|
,
|
|
<taxonomicName id="32048A88304DF1C7082F69B4217AE577" LSID-ZBK="urn:lsid:zoobank.org:act:B81066A2-E359-4A20-9D82-D3C7BAC03970" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus mirissumba Costa 1992:107" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="mirissumba">T. mirissumba</taxonomicName>
|
|
,
|
|
<taxonomicName id="13490F9F7A927A184F0AE31D971D99F4" LSID-ZBK="urn:lsid:zoobank.org:act:1CDFD373-4DC7-4347-8567-24DC017A1F1A" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus nigricans Valenciennes 1832:348" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="nigricans">T. nigricans</taxonomicName>
|
|
,
|
|
<taxonomicName id="B1970143C5BB6F076FE96BE9BB248956" LSID-ZBK="urn:lsid:zoobank.org:act:F76E50AB-DAC7-4056-853D-1F1DCBCA1F26" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus pantherinus Alencar & Costa 2004:2" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="pantherinus">T. pantherinus</taxonomicName>
|
|
,
|
|
<taxonomicName id="95B4A7188258CB1E86D9605D9E3F50D9" LSID-ZBK="urn:lsid:zoobank.org:act:2BB25233-9646-40EF-B848-F881B7C9B717" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus potschi Barbosa & Costa 2003:282" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="potschi">T. potschi</taxonomicName>
|
|
,
|
|
<taxonomicName id="FE9D82EBB2FF9744A72FAC02B7B875DF" LSID-ZBK="urn:lsid:zoobank.org:act:129BCA0E-8B83-4876-893D-42ADAA932D19" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus taroba Wosiacki & Garavello 2004:10" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="taroba">T. taroba</taxonomicName>
|
|
,
|
|
<taxonomicName id="72314B13638A20917161D98442FE4769" LSID-ZBK="urn:lsid:zoobank.org:act:0DFF9225-2629-424C-BEB9-074B4F16E67F" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus trefauti Wosiacki 2004:3" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="trefauti">T. trefauti</taxonomicName>
|
|
,
|
|
<taxonomicName id="0C7C7E17A53797AC180A20585A97559C" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="reinhardti">T. reinhardti</taxonomicName>
|
|
,
|
|
<taxonomicName id="7EAD689310A782B2E50F692AF177083E" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="triguttatus">T. triguttatus</taxonomicName>
|
|
,
|
|
<taxonomicName id="EEA8B6670EC9AA948FE082090BB61D43" LSID-ZBK="urn:lsid:zoobank.org:act:9E1679CD-2562-4664-843B-C3F48192607F" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus variegatus Costa 1992:103" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="variegatus">T. variegatus</taxonomicName>
|
|
, and
|
|
<taxonomicName id="2846E82A69A36D05F5EDFB58D7AF1855" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="vermiculatus">T. vermiculatus</taxonomicName>
|
|
) by the presence of the first pectoral-fin ray prolonged as a filament.
|
|
<taxonomicName id="D12B45DAE6009BE60630668B8880EC65" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="iheringi">Trichomycterus iheringi</taxonomicName>
|
|
shares with the larger assemblage of species congeners from south and southeastern Brazil the presence of two pores s6 paired at the interorbital space (vs. one symphyseal pore s6 in
|
|
<taxonomicName id="904F27D5E706533BF0E3EF9812F06841" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="alternatus">T. alternatus</taxonomicName>
|
|
,
|
|
<taxonomicName id="5A22AF195CAB6E9AEC588A3F2DB72F24" LSID-ZBK="urn:lsid:zoobank.org:act:197E097C-2451-43DE-B902-111021AAEB5D" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus giganteus Lima & Costa 2004:2" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="giganteus">T. giganteus</taxonomicName>
|
|
,
|
|
<taxonomicName id="525345C27E13872B1FF52C07362D465C" LSID-ZBK="urn:lsid:zoobank.org:act:1CDFD373-4DC7-4347-8567-24DC017A1F1A" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus nigricans Valenciennes 1832:348" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="nigricans">T. nigricans</taxonomicName>
|
|
, and
|
|
<taxonomicName id="F44280D010C8A3228655CC05DA5980E1" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="paquequerensis">T. paquequerensis</taxonomicName>
|
|
).
|
|
<taxonomicName id="73123C2D3DE6FFC1D78A2168A96751FC" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="iheringi">Trichomycterus iheringi</taxonomicName>
|
|
shares with
|
|
<taxonomicName id="825B09FC361D9EDECC8759C8EB7FCFEF" LSID-ZBK="urn:lsid:zoobank.org:act:0ED36072-3C3B-41A0-81EA-AF017739D4DB" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus castroi de Pinna 1992:90" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="castroi">T. castroi</taxonomicName>
|
|
,
|
|
<taxonomicName id="F88A4E2A259A80EF4CAB40366207EFA1" LSID-ZBK="urn:lsid:zoobank.org:act:78F93270-ED22-48B6-8329-CB52BBCD4664" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus guaraquessaba Wosiacki 2005" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="guaraquessaba">T. guaraquessaba</taxonomicName>
|
|
,
|
|
<taxonomicName id="FBD89F3F38DEB49156728E15D1404249" LSID-ZBK="urn:lsid:zoobank.org:act:090F10B3-41E6-426A-BE70-89E47F702B19" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus plumbeus Wosiacki & Garavello 2004:13" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="plumbeus">T. plumbeus</taxonomicName>
|
|
, and
|
|
<taxonomicName id="AAF2457BB367A4A1B4475EB4744BBE28" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="stawiarski">T. stawiarski</taxonomicName>
|
|
I-7 pectoral-fin rays (vs. I-5 in
|
|
<taxonomicName id="723FF12F59504DCDCE7FFEACBC945B3D" LSID-ZBK="urn:lsid:zoobank.org:act:656BBD86-BAC9-4255-B723-0776CFBF111F" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus mboycy Wosiacki & Garavello 2004:8" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="mboycy">T. mboycy</taxonomicName>
|
|
,
|
|
<taxonomicName id="7EC40161ACC544AF2B729002FD9D8D01" LSID-ZBK="urn:lsid:zoobank.org:act:D11FBE09-11EC-43EF-8BD7-21BE3F735BC1" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus naipi Wosiacki & Garavello 2004:2" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="naipi">T. naipi</taxonomicName>
|
|
, and I-6 in
|
|
<taxonomicName id="FFB8F3ACC068B56A6EBE3C10E634E225" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="davisi">T. davisi</taxonomicName>
|
|
,
|
|
<taxonomicName id="27B4D66D9F90C5CCE8AAEC2CE7FE535A" LSID-ZBK="urn:lsid:zoobank.org:act:5E60557F-9A8F-4467-BF3B-8C7E55D2FB7F" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus papilliferus Wosiacki & Garavello 2004:5" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="papilliferus">T. papilliferus</taxonomicName>
|
|
, and
|
|
<taxonomicName id="1E8BD62E8D7102F606ACC21CEE7D0A13" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="zonatus">T. zonatus</taxonomicName>
|
|
).
|
|
<taxonomicName id="54BF3F559C28C37E3619C9400663A856" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="iheringi">Trichomycterus iheringi</taxonomicName>
|
|
is readily from
|
|
<taxonomicName id="E9A48D1B5CA9A72709F09993D6555A88" LSID-ZBK="urn:lsid:zoobank.org:act:0ED36072-3C3B-41A0-81EA-AF017739D4DB" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus castroi de Pinna 1992:90" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="castroi">T. castroi</taxonomicName>
|
|
and
|
|
<taxonomicName id="15215EC3A8939D09B4D3E55C0A163C41" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="stawiarski">T. stawiarski</taxonomicName>
|
|
by the uniform light tan color pattern with numerous, poorly defined small spots over the dorsal and lateral portions of the head, trunk and caudal peduncle (vs. large and well defined spots and stripes). Furthermore,
|
|
<taxonomicName id="C6156C16C6E7194D281D2AB31107D923" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="iheringi">T. iheringi</taxonomicName>
|
|
can be distinguished from
|
|
<taxonomicName id="E661D2276CA26365691E01C0D11CD2B9" LSID-ZBK="urn:lsid:zoobank.org:act:78F93270-ED22-48B6-8329-CB52BBCD4664" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus guaraquessaba Wosiacki 2005" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="guaraquessaba">T. guaraquessaba</taxonomicName>
|
|
,
|
|
<taxonomicName id="F59651525A3C25C6ED7AC0C52F29F452" LSID-ZBK="urn:lsid:zoobank.org:act:090F10B3-41E6-426A-BE70-89E47F702B19" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus plumbeus Wosiacki & Garavello 2004:13" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="plumbeus">T. plumbeus</taxonomicName>
|
|
, and other congeners by the combination of pelvic-fin margin distance from the urogenital opening equal to half of its the fins length (vs. in contact with urogenital opening or covering it), caudal fin rounded (vs. truncate as in
|
|
<taxonomicName id="EB4A558B970EA82F0FAF6EE48D963DEA" LSID-ZBK="urn:lsid:zoobank.org:act:78F93270-ED22-48B6-8329-CB52BBCD4664" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus guaraquessaba Wosiacki 2005" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="guaraquessaba">T. guaraquessaba</taxonomicName>
|
|
), and 9-10 branched rays in the dorsal fin (vs. 6-8).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9209E1E58B945D6CECBC3EB2A27DAF0A" type="description">
|
|
<paragraph id="CBF4B8657B9EAD9E3BEE09864621DABA" pageNumber="57">
|
|
Redescription: Morphometric data for
|
|
<typeStatus id="0F5ED34AF3CC8F7B166695538E653E83">holotype</typeStatus>
|
|
and
|
|
<typeStatus id="104F6D808E660C2A2967D8D2285309CB">paratypes</typeStatus>
|
|
are given in Table 1.
|
|
</paragraph>
|
|
<paragraph id="71FB57182159260BF1CE84931AD58D70" pageNumber="57">“T1”</paragraph>
|
|
<paragraph id="BEC3A3676A46776FBF3828BF2F3A0DFE" pageNumber="57">Body elongate, roughly cylindrical close to head and gradually more compressed in trunk towards caudal fin. Profiles of trunk straight dorsally and slightly convex ventrally. Dorsal and ventral profiles of caudal peduncle straight. Integument thick, especially over base of dorsal and anal fins. Small papillae on lips and scattered over lateral surface of head.</paragraph>
|
|
<paragraph id="76285CB99713EBD0BD910644FBEFC292" lastPageNumber="58" pageNumber="57">Head wide, rounded and depressed, slightly longer than wide, the transversal section at posterior tip of opercle slightly wider than anteriorly at nostril, anterior margin rounded. Lateral region of eye slightly swollen by jaw muscles in both large and small specimens. Profiles of head straight dorsally and convex ventrally. Eyes rounded, well defined rim, dorsally oriented, covered by thin skin, distinctly separated from surface of eyeball. Ocular structures readily visible on surface of skin, not deeply sunken. Orbital rim not free. Anterior nostril same size as posterior one, surrounded by fleshy flap of integument. Posterior nostril with thin flap of integument anteriorly. Anterior and posterior nostrils half diameter of eye. Gill membranes thick, united to isthmus only at anteriormost point. Gill openings not constricted. Five branchiostegal rays visible externally, 7-8 in X-rayed specimens (Fig. 4). Mouth subterminal, its corners laterally oriented. Lower lip with conspicuous lateral fleshy lobes, internal to origin of rictal barbels. Anterior margin of upper lip rounded. Small papillae on external surface of upper lip; large papillae continuous inside of mouth to region of teeth attachment. Upper lip continuous with dorsal surface of head. Barbels long with large bases and gradually narrowing towards tip. Tips of nasal barbels reaching midway between posterior rim of eye and base of anterior opercular odontodes; tips of maxillary barbels reaching tip of posterior opercular odontodes; tips of rictal barbels reaching base of posterior most interopercular odontodes. Origin of nasal barbels on posterolateral portion of integument flap around anterior nostril. Interopercular patch of odontodes long, 20-22 conical odontodes covered by thick integument, anteriormost odontodes smaller and slightly curved medially, gradually longer and more curved distally. Opercular patch of odontodes rounded, covering posterior edge of opercle, 25-27 conical odontodes, anterior ones smaller and straight, gradually longer and curved distally. Supraorbital canal complete, infraorbital incomplete. Infraorbital anterior section pores i1 and i3, and posterior section pores i10 and i11. Supraorbital pores s1, s2 and s6 paired.</paragraph>
|
|
<paragraph id="3A6AAF513F4E3CDAD923FD125D9C42AA" pageNumber="59">
|
|
Pectoral fin with rounded margin, I-7 rays, first short and thickest, third and fourth longest. Dorsal-fin with margin semicircular when expanded, II-III/9-10 (
|
|
<typeStatus id="999353298E3C52EFC9C9EB1351D8E521">Holotype</typeStatus>
|
|
, II/9) rays, fourth the longest. Anal fin slightly elongate in overall shape, slightly smaller than dorsal fin, I/5-II/5 (
|
|
<typeStatus id="B5400A60FED965B6CE6A83ABE09133F1">Holotype</typeStatus>
|
|
, II/5) rays, fourth the longest, origin at vertical through seventh dorsal-fin ray. Pelvic-fin origin anterior to dorsal-fin origin, rounded margin, not reaching anal and urogenital openings, I/4 rays, second and third longest. Caudal fin rounded, margin distinctly wider than remaining caudal region, I/11/I principal rays, branched rays splitting three times. Only the first dorsal and ventral caudal-fin accessory rays visible. Anal and urogenital openings closer to anal-fin origin than pelvic-fin base.
|
|
</paragraph>
|
|
<paragraph id="BFFA8071FA0EF3BCCA9E12F56C611F18" pageNumber="59">Free vertebrae 35. Ribs 11 pairs, first one thickest, 3rd -4th longest. Dorsal-fin pterygiophores 9, first inserting in front of the neural spines of 15th -16th free vertebrae. Anal-fin pterygiophores 6, first in front of hemal spines of 21st -22nd free vertebrae. Dorsal procurrent rays 18. Ventral procurrent rays 8. Caudal skeleton composed of pleurostyle, hypurals 4+5, hypural 3, and fused parhypural and hypurals 1+2 (Fig. 4).</paragraph>
|
|
<paragraph id="D0EB4CC5AC6EBB88B0BBE08051AAE903" pageNumber="59">Color pattern: Refer to figure 2 for general view of color pattern in alcohol. The pattern is the same for smaller (117.6 mm SL) to larger (139.6 mm SL) specimens. Uniform light tan on dorsal surface of trunk, gradually lighter laterally. Numerous, poorly defined spots over dorsal and lateral portions of head, trunk and caudal peduncle. Spots coalescent over head at the occipital and interorbital space. Spots larger at dorsal trunk and caudal peduncle, gradually less numerous and smaller ventrally. Nasal, maxillary and rictal barbels unpigmented. Ventral surfaces of head, trunk and caudal peduncle unpigmented. Pectoral fin with few small spots over dorsal surface scattered from base to half of its length. Pelvic fins unpigmented. Dorsal fin with few small spots scattered from base to half of its length rays. Caudal fin with few spots over rays, larger and closer each other at base, gradually smaller and scattered to margin.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="510BB210BEF94F199B054DBC7D6F40F2" type="distribution">
|
|
<paragraph id="B39BFD19BAF2EB7E96C7179ADBC9F6D3" pageNumber="59">
|
|
Distribution: Known from the type-locality, Santos, on the costal area of
|
|
<normalizedToken id="93AC54E410876FE60F17D729EBF7840F" originalValue="São">Sao</normalizedToken>
|
|
Paulo State, and headwaters of rio
|
|
<normalizedToken id="60245C2706DF9413910D8392D9C5689A" originalValue="Tietê">Tiete</normalizedToken>
|
|
(
|
|
<normalizedToken id="4C6062081716DE4E451DA7D5FA4F340C" originalValue="Paraná">Parana</normalizedToken>
|
|
basin), southeastern Brazil. The Sapina locality cited by Eigenmann (1917 and 1918) for the
|
|
<typeStatus id="B25AE3C793691F03EED18D26F8C6FC7B">paratypes</typeStatus>
|
|
was not located. However, Eigenmann (1911) cited “Sapina, Rio
|
|
<normalizedToken id="F6847B1F331241E1AE91C54D4D3543BE" originalValue="Tieté">Tiete</normalizedToken>
|
|
, into
|
|
<normalizedToken id="74641BB8B9498F1782269BF6E3267EE6" originalValue="Paraná">Parana</normalizedToken>
|
|
. Coll. Nos. 1599-1616. Four miles from the city,
|
|
<date id="3AFCD576B47EF05BC8DDFD14FF68ECFB" day="23" month="07" value="1908-07-23" year="1908">July 23, 1908</date>
|
|
”, for the
|
|
<typeStatus id="5A9F7F15B3E7CC15344087EC7A9265BA">Paratypes</typeStatus>
|
|
. In the same paper, cited “Santos, coast of
|
|
<normalizedToken id="B8160C5C16611DB9A4D651774ADA99C0" originalValue="São">Sao</normalizedToken>
|
|
Paulo.
|
|
<date id="D76157FDE640350AB424F6A620246DDA" day="23" month="07" value="1908-07-23" year="1908">July 23, 1908</date>
|
|
” [no
|
|
<date id="9F56E1867DD0E1C0EB6C530AFC2AB618" day="03" month="07" value="1908-07-03" year="1908">July 3, 1908</date>
|
|
in Eigenmann, 1918] for the
|
|
<typeStatus id="76F4529023C5B804E08CFFAB4C210E80">Holotype</typeStatus>
|
|
. It is assumed that Sapina is a locality on the headwaters of rio
|
|
<normalizedToken id="7D2CD1A055B45B17A3BDAFF16567A8AF" originalValue="Tietê">Tiete</normalizedToken>
|
|
, close to Santos.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="2F8BE03321C024913625F59243010C6D" type="discussion">
|
|
<paragraph id="7CE93633F57417A39AB6EDEBE91B003B" pageNumber="60">
|
|
Remarks: The present redescription of
|
|
<taxonomicName id="81F232C67F62407AE0CA4E0B22E95310" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="iheringi">Trichomycterus iheringi</taxonomicName>
|
|
was based on type material of the
|
|
<collectionCode id="558F713F8CBE25FA1DEC10A1EC0DF41A" collectionName="USA, California, San Francisco, California Academy of Sciences">CAS</collectionCode>
|
|
and
|
|
<collectionCode id="1CEE8FE385A71D9FFBE579ED725B1B83" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)">FMNH</collectionCode>
|
|
. The
|
|
<typeStatus id="98EFE22235EE37C2A154E89957A71D81">holotype</typeStatus>
|
|
and
|
|
<typeStatus id="F43E9F489409C36E87B75B85C747AA89">paratypes</typeStatus>
|
|
were in a good state of preservation (Fig. 2), with the color pattern slightly pale. Eigenmann (1917) presented
|
|
<taxonomicName id="C409C4F777D8D0036EE88C5CBD4D14D6" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="iheringi">T. iheringi</taxonomicName>
|
|
in a short description based on six specimens which was adequate at the time, but incomplete by current standards. In the next year Eigenmann (1918; PL.50: 5) presented a transcription of the original description, with a very sharp drawing of the type. Eigenmann (1917, 1918), with no justification, allied
|
|
<taxonomicName id="68B84AD56ADAC28E5759AD49CF2040A2" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="iheringi">T. iheringi</taxonomicName>
|
|
to
|
|
<taxonomicName id="5BB9B9DC37009FE34BF1BDA0EEE8ACF5" LSID-ZBK="urn:lsid:zoobank.org:act:4EA47831-CCA4-48AA-BD01-65FB92181056" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus punctatissimus Castelnau 1855:49" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="punctatissimus">T. punctatissimus Castelnau</taxonomicName>
|
|
, from the rio Araguaia (Tocantins basin). Eigenmann (1918; PL. 45: 1) presented a cartoonish drawing of
|
|
<taxonomicName id="54ACE995423DBDE841EDF3C176E8CAFF" LSID-ZBK="urn:lsid:zoobank.org:act:4EA47831-CCA4-48AA-BD01-65FB92181056" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus punctatissimus Castelnau 1855:49" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="punctatissimus">T. punctatissimus</taxonomicName>
|
|
from Castelnau (1855), and a short text of the characters in the key of that paper (“Sides and back with minute dark specks and vermiculations between them; origin of dorsal over origin of ventral”). It was not possible to analyze the type material of
|
|
<taxonomicName id="ED69198BA580AEC516FCB08A389DFDE8" LSID-ZBK="urn:lsid:zoobank.org:act:4EA47831-CCA4-48AA-BD01-65FB92181056" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus punctatissimus Castelnau 1855:49" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="punctatissimus">T. punctatissimus</taxonomicName>
|
|
, and the original description (Castelnau, 1855) does not offer features to support the relationships of this species to
|
|
<taxonomicName id="39FD0B78D046C38FC86DD90940989318" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="iheringi">T. iheringi</taxonomicName>
|
|
. Only the color pattern cited by Castelnau (1855) for
|
|
<taxonomicName id="93494ABA6502CC730D7E41F58BC1A48A" LSID-ZBK="urn:lsid:zoobank.org:act:4EA47831-CCA4-48AA-BD01-65FB92181056" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus punctatissimus Castelnau 1855:49" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="punctatissimus">T. punctatissimus</taxonomicName>
|
|
(“sa couleur
|
|
<normalizedToken id="85417A0A4F767F6B65CE25E6CEFC6E21" originalValue="générale">generale</normalizedToken>
|
|
dun
|
|
<normalizedToken id="11D2C07C55E62E72ECFADF56B233DEF8" originalValue="châtain">chatain</normalizedToken>
|
|
clair. Il est
|
|
<normalizedToken id="B971AEBDA636B63B84535F621FBE6FC8" originalValue="entiérement">entierement</normalizedToken>
|
|
couvert de
|
|
<normalizedToken id="FA4335951FE7CE7323C31A51893B873D" originalValue="très">tres</normalizedToken>
|
|
petits point dun brun obscure
|
|
<normalizedToken id="36F286B95BFB248062CE62DD42BEF563" originalValue="très">tres</normalizedToken>
|
|
<normalizedToken id="3624F8CBB3BCE9C58ABD6D45556E4F07" originalValue="serrés">serres</normalizedToken>
|
|
et couvrant les nageoires
|
|
<normalizedToken id="AB9147B4C766BE72C2E8731D6A7D3226" originalValue="supérieures">superieures</normalizedToken>
|
|
. Le dessous du corps et les nageoires inferieures sont dun brun jaune”), cited by Eigenmann (1918) suggests some similarity to
|
|
<taxonomicName id="7BC8CA3545B821C92823B0E4AF8EBC5B" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="iheringi">T. iheringi</taxonomicName>
|
|
. However, the presence of several small spots covering the dorsum and sides of the head, trunk and caudal peduncle is a color pattern in many
|
|
<taxonomicName id="AB8875C1F566D0F5E05A3662D578BC51" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="genus">Trichomycterus</taxonomicName>
|
|
species (
|
|
<taxonomicName id="2BD0C995416FBD5E7B42C2C38A6F6741" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="davisi">T. davisi</taxonomicName>
|
|
,
|
|
<taxonomicName id="F380FA20C95B83853139257910909B65" LSID-ZBK="urn:lsid:zoobank.org:act:84D84BFD-1592-4583-B2D4-0BB6B004D9C3" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus brasiliensis Lütken & Reinhardt 1874:29 [1]" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="brasiliensis">T. brasiliensis</taxonomicName>
|
|
,
|
|
<taxonomicName id="17AEF5A2B6578B5CD142DCE8D1D5F87D" LSID-ZBK="urn:lsid:zoobank.org:act:89D8ECF9-B9D9-440E-98A1-9EDD49B8DBC5" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus bahianus Costa 1992:105" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="bahianus">T. bahianus</taxonomicName>
|
|
,
|
|
<taxonomicName id="D3921B15D6393B2E6F2EEBCB12675753" LSID-ZBK="urn:lsid:zoobank.org:act:D85474E7-5A0D-48AD-B35B-A0FB33DD0370" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus mimonha Costa 1992:106" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="mimonha">T. mimonha</taxonomicName>
|
|
,
|
|
<taxonomicName id="B122BC5DBD68E60022E21E642259693D" LSID-ZBK="urn:lsid:zoobank.org:act:B81066A2-E359-4A20-9D82-D3C7BAC03970" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus mirissumba Costa 1992:107" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="mirissumba">T. mirissumba</taxonomicName>
|
|
,
|
|
<taxonomicName id="4349C4B470A75E7B69C13870AFC588B0" LSID-ZBK="urn:lsid:zoobank.org:act:129BCA0E-8B83-4876-893D-42ADAA932D19" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus taroba Wosiacki & Garavello 2004:10" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="taroba">T. taroba</taxonomicName>
|
|
, and
|
|
<taxonomicName id="22C31C6357C1575CFB3FC4DD1A4B03CB" LSID-ZBK="urn:lsid:zoobank.org:act:F98023F9-1C43-481C-80CB-AAE788A523B6" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus rivulatus Valenciennes 1846:495" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="rivulatus">T. rivulatus Valenciennes</taxonomicName>
|
|
, among others), in
|
|
<taxonomicName id="1EA4A2B77A22FA6521948044E8FA69EA" LSID-ZBK="urn:lsid:zoobank.org:act:CF87DB58-3C18-4993-A8A4-E99588F45896" family="Trichomycteridae" genus="Eremophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Eremophilus mutisii Humboldt 1805:35" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="mutisii">Eremophilus mutisii Humboldt</taxonomicName>
|
|
, in
|
|
<taxonomicName id="D17AD74146485ED1E7939726C393A540" family="Trichomycteridae" genus="Ituglanis" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="amazonicus">Ituglanis amazonicus (Steindachner)</taxonomicName>
|
|
,
|
|
<taxonomicName id="0C1A8B516522C84DACA930F3903DE973" family="Trichomycteridae" genus="Ituglanis" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="eichorniarum">I. eichorniarum (Miranda-Ribeiro)</taxonomicName>
|
|
, and
|
|
<taxonomicName id="C70A34C15A4DD779EFF900CC9691C037" family="Trichomycteridae" genus="Ituglanis" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="proops">I. proops (Miranda-Ribeiro)</taxonomicName>
|
|
. Only a detailed phylogenetic analysis will allow inferences about the evolution of the color pattern in the relationships of the
|
|
<taxonomicName id="8B99BAFBF23814E663A4D511C912BCF8" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="genus">Trichomycterus</taxonomicName>
|
|
species and of the non monophyletic group “Trichomycterinae”.
|
|
</paragraph>
|
|
<paragraph id="858FD743BF2C7F9E992D5EAD9604A07C" lastPageNumber="61" pageNumber="60">
|
|
Bockmann & Sazima (2005) proposed a group of species,
|
|
<taxonomicName id="AE8A614A2633082CD809A92AE0AAF592" LSID-ZBK="urn:lsid:zoobank.org:act:1F83BB4D-68B6-4DCC-B9D7-75BCBBFC34D0" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus maracaya Bockmann & Sazima 2004:63" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="maracaya">T. maracaya</taxonomicName>
|
|
,
|
|
<taxonomicName id="C3FE2106BCED2129EF4EFC8603E76FC0" LSID-ZBK="urn:lsid:zoobank.org:act:D85474E7-5A0D-48AD-B35B-A0FB33DD0370" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus mimonha Costa 1992:106" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="mimonha">T. mimonha</taxonomicName>
|
|
,
|
|
<taxonomicName id="F8D3FEC2F93ECE52138521D1FF4394F8" LSID-ZBK="urn:lsid:zoobank.org:act:2BB25233-9646-40EF-B848-F881B7C9B717" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus potschi Barbosa & Costa 2003:282" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="potschi">T. potschi</taxonomicName>
|
|
, and
|
|
<taxonomicName id="93ADAA8373E9DB1719A343872F567B4C" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="vermiculatus">T. vermiculatus</taxonomicName>
|
|
, as members of
|
|
<taxonomicName id="EAD613A33F0721B9A027E2B73290A3AF" LSID-ZBK="urn:lsid:zoobank.org:act:84D84BFD-1592-4583-B2D4-0BB6B004D9C3" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus brasiliensis Lütken & Reinhardt 1874:29 [1]" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="brasiliensis">T. brasiliensis</taxonomicName>
|
|
species-complex based on the presence of fours longitudinal rows of well-defined blotches, plus I/5-6 pectoral-fin rays. As observed in the type material,
|
|
<taxonomicName id="6ECE49B9116B0BE7F75999E5DB703AB1" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="iheringi">T. iheringi</taxonomicName>
|
|
does not present the color pattern of four rows of well-defined blotches; hence it could not be included as a member of this group. In addition, all specimens of the type material of
|
|
<taxonomicName id="B4799FC047DFB5DED9ABA6B951144842" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="iheringi">T. iheringi</taxonomicName>
|
|
have I/7 pectoral-fin rays as observed by Eigenmann (1917 and 1918; P.8), in contrast to that described by Bockmann & Sazima (2005). In a slightly different way Wosiacki (2002) proposed, in a phylogenetic analysis of Trichomycterinae, a different group composed of species with 6 or fewer pectoral-fin rays [
|
|
<taxonomicName id="AFF72A82FD505D104547736ABF36977B" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="candidus">T. candidus</taxonomicName>
|
|
,
|
|
<taxonomicName id="9120F34EBAD9034D07B698F436875F6A" LSID-ZBK="urn:lsid:zoobank.org:act:D11FBE09-11EC-43EF-8BD7-21BE3F735BC1" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus naipi Wosiacki & Garavello 2004:2" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="naipi">T. naipi</taxonomicName>
|
|
,
|
|
<taxonomicName id="F2A077E480AD6F70CCB2130A9E71F5E2" LSID-ZBK="urn:lsid:zoobank.org:act:656BBD86-BAC9-4255-B723-0776CFBF111F" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus mboycy Wosiacki & Garavello 2004:8" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="mboycy">T. mboycy</taxonomicName>
|
|
,
|
|
<taxonomicName id="4CB850B728F1F493C11A9ED6D93B154A" LSID-ZBK="urn:lsid:zoobank.org:act:129BCA0E-8B83-4876-893D-42ADAA932D19" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus taroba Wosiacki & Garavello 2004:10" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="taroba">T. taroba</taxonomicName>
|
|
, and
|
|
<taxonomicName id="DCA71663656DA8C11B0B7D737B927E7D" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="species" species="sp. n. ("pelvic-less")">Trichomycterus sp. n. (“pelvic-less”)</taxonomicName>
|
|
]. Six or fewer pectoral-fin rays also are present in all
|
|
<taxonomicName id="71B2DF6ED92BF43218191332B878FF9C" LSID-ZBK="urn:lsid:zoobank.org:act:FDAB162D-116C-4E81-B78B-D677333D2F85" family="Trichomycteridae" genus="Ituglanis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ituglanis Costa & Bockmann 1993:44" order="Siluriformes" pageNumber="60" phylum="Chordata" rank="genus">Ituglanis</taxonomicName>
|
|
species and in the clade composed of Glanapteryginae, Sarcoglanidinae, Tridentinae, Stegophilinae, and Vandelliinae (sensu de Pinna, 1998), interpreted as the result of independent evolution (Wosiacki, 2002).
|
|
</paragraph>
|
|
<paragraph id="D0D1EF510EF4A20842B31CA0629328F5" pageNumber="61">
|
|
Bizerril (1994) related
|
|
<taxonomicName id="A0066FBC4DB5DB8F0FA4724F23E438E3" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="iheringi">T. iheringi</taxonomicName>
|
|
, apparently without an analysis of material, to an assemblage of species (
|
|
<taxonomicName id="0A4D4D258BC4B70724BD94694CDC07B5" LSID-ZBK="urn:lsid:zoobank.org:act:D85474E7-5A0D-48AD-B35B-A0FB33DD0370" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus mimonha Costa 1992:106" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="mimonha">T. mimonha</taxonomicName>
|
|
,
|
|
<taxonomicName id="8D398F98020B42E9D9A076FDDBDFFC9D" LSID-ZBK="urn:lsid:zoobank.org:act:B81066A2-E359-4A20-9D82-D3C7BAC03970" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus mirissumba Costa 1992:107" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="mirissumba">T. mirissumba</taxonomicName>
|
|
,
|
|
<taxonomicName id="A134C8C9201A36B4C3B048189E20549C" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="triguttatus">T. triguttatus</taxonomicName>
|
|
,
|
|
<taxonomicName id="2D5C7073E216B2455E2728A38AEA4477" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="vermiculatus">T. vermiculatus</taxonomicName>
|
|
, and
|
|
<taxonomicName id="F80E47ECC995B4A41F4A10FABA46BEB0" LSID-ZBK="urn:lsid:zoobank.org:act:C4969FB6-8D9B-4A86-84F0-625DBAC0B4B2" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus concolor Costa 1992:107" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="concolor">T. concolor</taxonomicName>
|
|
) based on the supraorbital pores being closer to the margin of the eyeball than to the sagittal line, as proposed by Costa (1992). However, as observed in the
|
|
<typeStatus id="0F2AA6D54954C6C88FEED0ECF5AFEA33">holotype</typeStatus>
|
|
(Fig. 2) and
|
|
<typeStatus id="65809C3D82383667F47D9D9E19AFEBAF">paratypes</typeStatus>
|
|
, the supraorbital pores of
|
|
<taxonomicName id="850D1E0FFA5233658BC95A559808E67E" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="iheringi">T. iheringi</taxonomicName>
|
|
are closer to the sagittal line than the margin of the eyeball, contrary to the proposal of Bizerril (1994).
|
|
</paragraph>
|
|
<paragraph id="12518C563BD3FB81193FA5EE344478D1" pageNumber="61">
|
|
Phylogenetic assignments:
|
|
<taxonomicName id="A47B864EE0EAC4B9F0D11AE6D21DB451" LSID-ZBK="urn:lsid:zoobank.org:act:78F93270-ED22-48B6-8329-CB52BBCD4664" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus guaraquessaba Wosiacki 2005" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="guaraquessaba">Trichomycterus guaraquessaba</taxonomicName>
|
|
and
|
|
<taxonomicName id="53E6D43AD619207CE2FE33FC16058969" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="iheringi">T. iheringi</taxonomicName>
|
|
share with all south and southeastern Brazil congeneric species the presence of s6 pores paired, one in each supraorbital canal, except
|
|
<taxonomicName id="4B67D20757E02D4F1D8395F774541B51" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="alternatus">T. alternatus</taxonomicName>
|
|
,
|
|
<taxonomicName id="29A80B8311EE37BD7471A7E3C14CDB1E" LSID-ZBK="urn:lsid:zoobank.org:act:197E097C-2451-43DE-B902-111021AAEB5D" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus giganteus Lima & Costa 2004:2" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="giganteus">T. giganteus</taxonomicName>
|
|
,
|
|
<taxonomicName id="B6298ED3A82EB1C74F5FE60A037EC5D6" LSID-ZBK="urn:lsid:zoobank.org:act:1CDFD373-4DC7-4347-8567-24DC017A1F1A" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus nigricans Valenciennes 1832:348" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="nigricans">T. nigricans</taxonomicName>
|
|
, and
|
|
<taxonomicName id="1AE80E2FEAC429D98BC228AEE9938E86" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="paquequerensis">T. paquequerensis</taxonomicName>
|
|
with one epiphyseal pore s6 fused at the sagittal line. Costa (1992) proposed two monophyletic groups for species of
|
|
<taxonomicName id="13AB29E535E42A44E56757948476BD60" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="genus">Trichomycterus</taxonomicName>
|
|
from south and southeastern Brazil: 1) Supraorbital pore s6 paired, each pore closer to the orbit than to the sagittal line; and 2) supraorbital pore s6paired, each pore closer to the sagittal line than to the orbit or fused into a single epiphyseal pore. The position of the supraorbital pore s6 is very variable intra- and interspecifically among Trichomycterinae; however, in a large assemblage of species, the pore s6 opening is directly from the supraorbital canal as in Copionodontinae and
|
|
<taxonomicName id="C3EE7B9ABE942A3DB618434426EC0CD7" LSID-ZBK="urn:lsid:zoobank.org:act:4ADF65AF-586D-406C-B474-3FF22ACF87C4" family="Trichomycteridae" genus="Trichogenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichogenes Britski & Ortega 1983:211" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="genus">Trichogenes</taxonomicName>
|
|
, the basalmost groups of Trichomycteridae. Wosiacki (2002) reported that there exists a branching from the supraorbital canal above the epiphyseal bar, in a medial direction, of which the external opening s6 either can be single and median (both sides fused) and at the end of a relatively long canal (equaling the distance between the supraorbital canal and the sagittal plane), or paired having a pore at each side of the sagittal plane between it and the supraorbital canal. The single pore condition is considered as a more derived state. The presence and length of a branch from the supraorbital canal opening either in a single or paired pore as observed in
|
|
<taxonomicName id="24743BD6CD4EF95D1D5B9D138B818DC8" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="alternatus">Trichomycterus alternatus</taxonomicName>
|
|
,
|
|
<taxonomicName id="397C9AD127844EA8E15E46F8A1E75B9B" LSID-ZBK="urn:lsid:zoobank.org:act:84D84BFD-1592-4583-B2D4-0BB6B004D9C3" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus brasiliensis Lütken & Reinhardt 1874:29 [1]" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="brasiliensis">T. brasiliensis</taxonomicName>
|
|
,
|
|
<taxonomicName id="82A1257D291364139A007419A4B1E973" LSID-ZBK="urn:lsid:zoobank.org:act:197E097C-2451-43DE-B902-111021AAEB5D" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus giganteus Lima & Costa 2004:2" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="giganteus">T. giganteus</taxonomicName>
|
|
,
|
|
<taxonomicName id="5550CFE54E88C798F70253225874315E" LSID-ZBK="urn:lsid:zoobank.org:act:B81066A2-E359-4A20-9D82-D3C7BAC03970" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus mirissumba Costa 1992:107" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="mirissumba">T. mirissumba</taxonomicName>
|
|
,
|
|
<taxonomicName id="12EF7DBC315BCABF325E3B63954A4E84" LSID-ZBK="urn:lsid:zoobank.org:act:D85474E7-5A0D-48AD-B35B-A0FB33DD0370" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus mimonha Costa 1992:106" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="mimonha">T. mimonha</taxonomicName>
|
|
,
|
|
<taxonomicName id="6BA22240370339BCFFB995A97BBF4B65" LSID-ZBK="urn:lsid:zoobank.org:act:1CDFD373-4DC7-4347-8567-24DC017A1F1A" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus nigricans Valenciennes 1832:348" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="nigricans">T. nigricans</taxonomicName>
|
|
, and
|
|
<taxonomicName id="1EB7D05573EE43038A05F9E8F61D2D0E" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="paquequerensis">T. paquequerensis</taxonomicName>
|
|
probably is a better way of indicating the position of pore s6 than the present reference to distance of the eyes. Regardless,
|
|
<taxonomicName id="E9AE53F08368B869503D3A9A671A845E" LSID-ZBK="urn:lsid:zoobank.org:act:78F93270-ED22-48B6-8329-CB52BBCD4664" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus guaraquessaba Wosiacki 2005" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="guaraquessaba">T. guaraquessaba</taxonomicName>
|
|
and
|
|
<taxonomicName id="96F32DBA555612C3F24D56EFA6F20E45" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="iheringi">T. iheringi</taxonomicName>
|
|
have the plesiomorphic state without a branching for pore s6.
|
|
<taxonomicName id="5D2E0A0032E1E10B3F168CB9AFD92C11" LSID-ZBK="urn:lsid:zoobank.org:act:78F93270-ED22-48B6-8329-CB52BBCD4664" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus guaraquessaba Wosiacki 2005" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="guaraquessaba">Trichomycterus guaraquessaba</taxonomicName>
|
|
and
|
|
<taxonomicName id="00410BE17B028C0D149C031D9AE23CB2" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="iheringi">T. iheringi</taxonomicName>
|
|
share, with an assemblage of species from south and southeastern Brazil (
|
|
<taxonomicName id="64E285C6D7EC9793E355C710CE372012" LSID-ZBK="urn:lsid:zoobank.org:act:1CDFD373-4DC7-4347-8567-24DC017A1F1A" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus nigricans Valenciennes 1832:348" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="nigricans">T. nigricans</taxonomicName>
|
|
,
|
|
<taxonomicName id="87E1F815EE66CA14E5A012897DC26D62" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="davisi">T. davisi</taxonomicName>
|
|
,
|
|
<taxonomicName id="0A0C6F35C7FDE3EF5C591786DF362575" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="stawiarski">T. stawiarski</taxonomicName>
|
|
,
|
|
<taxonomicName id="C1BA99C77922CF1D6199C7CDB27CEA1B" LSID-ZBK="urn:lsid:zoobank.org:act:0ED36072-3C3B-41A0-81EA-AF017739D4DB" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus castroi de Pinna 1992:90" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="castroi">T. castroi</taxonomicName>
|
|
,
|
|
<taxonomicName id="02F93032C42570782292526FFF84037F" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="triguttatus">T. triguttatus</taxonomicName>
|
|
,
|
|
<taxonomicName id="8F5E8663BA1D8373919750182A78BC8C" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="immaculatus">T. immaculatus</taxonomicName>
|
|
,
|
|
<taxonomicName id="5E7D60E29AAD2966E142D71BBF58DF56" LSID-ZBK="urn:lsid:zoobank.org:act:4B83205A-2285-4A82-9F41-E15DD605A259" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus brasiliensis itatiayae Miranda-Ribeiro 1906:p.? [15]" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="itatiayae">T. itatiayae</taxonomicName>
|
|
,
|
|
<taxonomicName id="B005661D73AE155759BFF04395F85A82" LSID-ZBK="urn:lsid:zoobank.org:act:B81066A2-E359-4A20-9D82-D3C7BAC03970" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus mirissumba Costa 1992:107" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="mirissumba">T. mirissumba</taxonomicName>
|
|
,
|
|
<taxonomicName id="1879C4FF41C07D42593E256C2F5C58B7" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="zonatus">T. zonatus</taxonomicName>
|
|
,
|
|
<taxonomicName id="7996E9C992B939F3908722DD6D815314" LSID-ZBK="urn:lsid:zoobank.org:act:D11FBE09-11EC-43EF-8BD7-21BE3F735BC1" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus naipi Wosiacki & Garavello 2004:2" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="naipi">T. naipi</taxonomicName>
|
|
,
|
|
<taxonomicName id="17595A2BEB03BE55CCB1A4660685E7A4" LSID-ZBK="urn:lsid:zoobank.org:act:129BCA0E-8B83-4876-893D-42ADAA932D19" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus taroba Wosiacki & Garavello 2004:10" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="taboba">T. taboba</taxonomicName>
|
|
,
|
|
<taxonomicName id="3AA711A86ED2CF4FDEFE274354367CDB" LSID-ZBK="urn:lsid:zoobank.org:act:5E60557F-9A8F-4467-BF3B-8C7E55D2FB7F" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus papilliferus Wosiacki & Garavello 2004:5" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="papilliferus">T. papilliferus</taxonomicName>
|
|
,
|
|
<taxonomicName id="129CF73B9042251F73DE19C260FEFB18" LSID-ZBK="urn:lsid:zoobank.org:act:656BBD86-BAC9-4255-B723-0776CFBF111F" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus mboycy Wosiacki & Garavello 2004:8" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="mboycy">T. mboycy</taxonomicName>
|
|
and
|
|
<taxonomicName id="C57D453DC3605F3DEA0D81AF533E2523" LSID-ZBK="urn:lsid:zoobank.org:act:090F10B3-41E6-426A-BE70-89E47F702B19" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Trichomycterus plumbeus Wosiacki & Garavello 2004:13" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="species" species="plumbeus">T. plumbeus</taxonomicName>
|
|
), the presence of a window at least half of the diameter of the optic foramen in the lateral of the orbito-sphenoid observed by Wosiacki (2002). This feature is an apormophy not observed in the Andeans and trans-Andean species of
|
|
<taxonomicName id="E24FFC93790BF3C27AE25280980458BA" family="Trichomycteridae" genus="Trichomycterus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="61" phylum="Chordata" rank="genus">Trichomycterus</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |