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<document id="690FD0B9EDA1C019273ABD4FDFA5AF0A" ID-DOI="10.5281/zenodo.185971" ID-GBIF-Dataset="1db39382-b4f0-46fa-ae08-dd5e57e0e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="185971" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460034098030" checkinUser="plazi" docAuthor="Pimvichai, Piyatida, Enghoff, Henrik &amp; Panha, Somsak" docDate="2009" docId="1644D538F44CFFCDFF49FB689B9EFE2A" docLanguage="en" docName="zt02016p050.pdf" docOrigin="Zootaxa 2016" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Thyropygus implicatus Demange 1961, n. comb." docType="treatment" docVersion="11" lastPageNumber="32" masterDocId="EA7DAD40F442FFC2FFDEFFED9F66FFCD" masterDocTitle="A revision of the Thyropygus allevatus group. Part 1: the T. opinatus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)" masterLastPageNumber="50" masterPageNumber="17" pageNumber="31" updateTime="1698234296060" updateUser="plazi">
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<mods:title id="FC40873222E6F1996A8AD2FC5CA58DA6">A revision of the Thyropygus allevatus group. Part 1: the T. opinatus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)</mods:title>
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<mods:namePart id="B8C1A77F270234039BC8CCC65BA95903">Pimvichai, Piyatida</mods:namePart>
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<mods:namePart id="545F8D4B99AE96A1A060A67E47E463F7">Enghoff, Henrik</mods:namePart>
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<mods:namePart id="8DA6367482158346E834377EB78DD315">Panha, Somsak</mods:namePart>
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<treatment id="1644D538F44CFFCDFF49FB689B9EFE2A" ID-DOI="http://doi.org/10.5281/zenodo.6218620" ID-GBIF-Taxon="119382578" ID-Zenodo-Dep="6218620" LSID="urn:lsid:plazi:treatment:1644D538F44CFFCDFF49FB689B9EFE2A" httpUri="http://treatment.plazi.org/id/1644D538F44CFFCDFF49FB689B9EFE2A" lastPageId="15" lastPageNumber="32" pageId="14" pageNumber="31">
<subSubSection id="D6F737A5F44CFFCCFF49FB689E0BFB0F" pageId="14" pageNumber="31" type="nomenclature">
<paragraph id="9E52642EF44CFFCCFF49FB689D9BFB52" blockId="14.[151,765,1157,1218]" box="[151,765,1157,1183]" pageId="14" pageNumber="31">
<heading id="C51AD342F44CFFCCFF49FB689D9BFB52" bold="true" box="[151,765,1157,1183]" fontSize="11" level="1" pageId="14" pageNumber="31" reason="1">
<emphasis id="AC99B83CF44CFFCCFF49FB689D9BFB52" bold="true" box="[151,765,1157,1183]" pageId="14" pageNumber="31">
<taxonomicName id="59ED1FADF44CFFCCFF49FB689DE3FB52" ID-CoL="56T2H" authority="Demange, 1961" authorityName="Demange" authorityYear="1961" box="[151,645,1157,1183]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="31" phylum="Arthropoda" rank="species" species="implicatus" status="comb. nov.">
<emphasis id="AC99B83CF44CFFCCFF49FB689ECCFB52" bold="true" box="[151,426,1157,1183]" italics="true" pageId="14" pageNumber="31">Thyropygus implicatus</emphasis>
(
<bibRefCitation id="FA7C19DFF44CFFCCFE64FB689D1BFB52" author="Demange" box="[442,637,1157,1183]" pageId="14" pageNumber="50" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange, 1961</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="B7AA0547F44CFFCCFD4CFB689D9BFB52" box="[658,765,1157,1183]" pageId="14" pageNumber="31">n. comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9E52642EF44CFFCCFF49FB459E0BFB0F" blockId="14.[151,765,1157,1218]" box="[151,365,1192,1218]" pageId="14" pageNumber="31">
(
<figureCitation id="06D678ABF44CFFCCFF41FB459F95FB0F" box="[159,243,1192,1218]" captionStart="FIGURE 2" captionStartId="6.[151,257,1579,1603]" captionTargetBox="[188,1398,194,1553]" captionTargetId="figure@6.[188,1398,194,1555]" captionTargetPageId="6" captionText="FIGURE 2. Thyropygus implicatus, SEM. A: apical view, left telopodite removed. B: posterior view, left telopodite removed. C: lateral view, with right telopodite. D: left coxa, lateral view." httpUri="https://zenodo.org/record/185973/files/figure.png" pageId="14" pageNumber="31">Figs. 2</figureCitation>
,
<figureCitation id="06D678ABF44CFFCCFF21FB459E69FB0F" box="[255,271,1192,1218]" captionStart="FIGURE 3" captionStartId="7.[151,255,976,1000]" captionTargetBox="[415,1171,390,951]" captionTargetId="figure@7.[415,1171,390,952]" captionTargetPageId="7" captionText="FIGURE 3. Thyropygus implicatus, SEM. Spatulate lobe (sl), palette (pa), blepharochaetae (bp)." httpUri="https://zenodo.org/record/185974/files/figure.png" pageId="14" pageNumber="31">3</figureCitation>
,
<figureCitation id="06D678ABF44CFFCCFEC2FB459E4FFB0F" box="[284,297,1192,1218]" captionStart="FIGURE 8" captionStartId="15.[151,255,1929,1953]" captionTargetBox="[237,1346,556,1897]" captionTargetId="figure@15.[226,1360,545,1906]" captionTargetPageId="15" captionText="FIGURE 8. Thyropygus implicatus, specimen from Thale Ban National Park, gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C: left telopodite, anterior view. D: left telopodite, posterior view." httpUri="https://zenodo.org/record/185979/files/figure.png" pageId="14" pageNumber="31">8</figureCitation>
AD)
</paragraph>
</subSubSection>
<subSubSection id="D6F737A5F44CFFCCFF49FB189A74FAC1" box="[151,1298,1269,1292]" pageId="14" pageNumber="31" type="reference_group">
<paragraph id="9E52642EF44CFFCCFF49FB189A74FAC1" blockId="14.[151,1298,1269,1292]" box="[151,1298,1269,1292]" pageId="14" pageNumber="31">
<treatmentCitationGroup id="BEFD4300F44CFFCCFF49FB189A74FAC1" box="[151,1298,1269,1292]" pageId="14" pageNumber="31">
<taxonomicName id="59ED1FADF44CFFCCFF49FB189D1FFAC1" ID-CoL="YGDB" authority="Demange, 1961: 181" authorityName="Demange" authorityPageNumber="181" authorityYear="1961" box="[151,633,1269,1292]" class="Diplopoda" family="Harpagophoridae" genus="Cornugonus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="31" phylum="Arthropoda" rank="species" species="implicatus">
<emphasis id="AC99B83CF44CFFCCFF49FB189EF6FAC1" box="[151,400,1269,1292]" italics="true" pageId="14" pageNumber="31">Cornugonus implicatus</emphasis>
<treatmentCitation id="1F4C423FF44CFFCCFE49FB189D1FFAC1" author="Demange" box="[407,633,1269,1292]" page="181" pageId="14" pageNumber="31" year="1961">
<bibRefCitation id="FA7C19DFF44CFFCCFE49FB189D22FAC1" author="Demange" box="[407,580,1269,1292]" pageId="14" pageNumber="50" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange, 1961</bibRefCitation>
: 181
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="4156DA8CF44CFFCCFD5DFB189D80FAC1" box="[643,742,1269,1292]" pageId="14" pageNumber="31" type="holotype">Holotype</typeStatus>
(not seen) in the Natural History Museum, London.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D6F737A5F44CFFCCFF49FAD69B91FAB1" pageId="14" pageNumber="31" type="materials_examined">
<paragraph id="9E52642EF44CFFCCFF49FAD69B91FAB1" blockId="14.[151,1437,1339,2023]" pageId="14" pageNumber="31">
<emphasis id="AC99B83CF44CFFCCFF49FAD69E68FA98" bold="true" box="[151,270,1339,1365]" pageId="14" pageNumber="31">Material:</emphasis>
<specimenCount id="88EBAFA7F44CFFCCFECBFAD19EE7FA9B" box="[277,385,1340,1366]" pageId="14" pageNumber="31" type="male">10 males</specimenCount>
,
<specimenCount id="88EBAFA7F44CFFCCFE53FAD19D67FA9B" box="[397,513,1340,1366]" pageId="14" pageNumber="31" type="female">5 females</specimenCount>
, and
<specimenCount id="88EBAFA7F44CFFCCFDE0FAD19DD5FA9B" box="[574,691,1340,1366]" pageId="14" pageNumber="31" type="juvenile">1 juvenile</specimenCount>
<collectingCountry id="E6FA24BEF44CFFCCFD64FAD19C33FA9B" box="[698,853,1340,1366]" name="Thailand" pageId="14" pageNumber="31">THAILAND</collectingCountry>
, Satun Province, Thale Ban National Park, 6° 42ˏ 29˝ N, 100° 10ˏ 46˝ E.
<date id="EA5342EEF44CFFCCFE7BFA8F9D08FAB1" box="[421,622,1378,1404]" pageId="14" pageNumber="31" value="1991-10-15">15 October 1991</date>
. M. Andersen, O. Martin and N. Scharff leg., (
<collectionCode id="F8FCFCEBF44CFFCCFB4EFA8F9B8AFAB1" box="[1168,1260,1378,1404]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="14" pageNumber="31">ZMUC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D6F737A5F44CFFCCFF1BFA659D56F9DA" pageId="14" pageNumber="31" type="diagnosis">
<paragraph id="9E52642EF44CFFCCFF1BFA659D56F9DA" blockId="14.[151,1437,1339,2023]" pageId="14" pageNumber="31">
<emphasis id="AC99B83CF44CFFCCFF1BFA659E36FA6F" bold="true" box="[197,336,1416,1442]" pageId="14" pageNumber="31">Diagnosis:</emphasis>
A species of the
<taxonomicName id="59ED1FADF44CFFCCFDEEFA649DFAFA6F" box="[560,668,1417,1442]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="31" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="AC99B83CF44CFFCCFDEEFA649DFAFA6F" box="[560,668,1417,1442]" italics="true" pageId="14" pageNumber="31">opinatus</emphasis>
</taxonomicName>
subgroup. Spatulate lobe (
<emphasis id="AC99B83CF44CFFCCFC2DFA649B6EFA6F" box="[1011,1032,1417,1442]" italics="true" pageId="14" pageNumber="31">sl</emphasis>
) at the apical part of telopodite terminating in a sharp brown spine. Similar in this respect to
<taxonomicName id="59ED1FADF44CFFCCFCA0FA5D9B65FA04" box="[894,1027,1456,1481]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="31" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="AC99B83CF44CFFCCFCA0FA5D9B65FA04" box="[894,1027,1456,1481]" italics="true" pageId="14" pageNumber="31">T. opinatus</emphasis>
</taxonomicName>
,
<taxonomicName id="59ED1FADF44CFFCCFBCAFA5D9BEEFA04" box="[1044,1160,1456,1481]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="31" phylum="Arthropoda" rank="species" species="floweri">
<emphasis id="AC99B83CF44CFFCCFBCAFA5D9BEEFA04" box="[1044,1160,1456,1481]" italics="true" pageId="14" pageNumber="31">T. floweri</emphasis>
</taxonomicName>
and
<taxonomicName id="59ED1FADF44CFFCCFB1AFA5D9A53FA04" box="[1220,1333,1456,1481]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="31" phylum="Arthropoda" rank="species" species="erectus">
<emphasis id="AC99B83CF44CFFCCFB1AFA5D9A53FA04" box="[1220,1333,1456,1481]" italics="true" pageId="14" pageNumber="31">T. erectus</emphasis>
</taxonomicName>
. Differs from these species by the having the lateral process of the anterior coxal fold (
<emphasis id="AC99B83CF44CFFCCFB87FA3A9BE6FA3D" box="[1113,1152,1495,1520]" italics="true" pageId="14" pageNumber="31">alp</emphasis>
) apically bent abruptly mesad, its tip with serrate margins.
</paragraph>
</subSubSection>
<subSubSection id="D6F737A5F44CFFCDFF1BF9CE9A50FEEB" lastPageId="15" lastPageNumber="32" pageId="14" pageNumber="31" type="description">
<paragraph id="9E52642EF44CFFCCFF1BF9CE9D66F915" blockId="14.[151,1437,1339,2023]" pageId="14" pageNumber="31">
<emphasis id="AC99B83CF44CFFCCFF1BF9CE9E05F9F0" bold="true" box="[197,355,1571,1597]" pageId="14" pageNumber="31">Description:</emphasis>
Adult males with 6370 podous rings, no apodous rings. Length ca.
<quantity id="5915C9CBF44CFFCCFB78F9C99A46F9F3" box="[1190,1312,1572,1598]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="14" pageNumber="31" unit="cm" value="12.5" valueMax="13.0" valueMin="12.0">1213 cm</quantity>
, width ca. 7.48.0 mm. Adult females with 6580 podous rings, no apodous rings. Length ca.
<quantity id="5915C9CBF44CFFCCFB7FF9A79A78F9A9" box="[1185,1310,1610,1636]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" metricValueMax="2.2" metricValueMin="1.2" pageId="14" pageNumber="31" unit="cm" value="17.0" valueMax="22.0" valueMin="12.0">1222 cm</quantity>
, width ca. 8.011.0 mm. Color in life unknown; preserved specimens with head, antennae, legs, prozona, middorsal metazona, epiproct, paraprocts and hyproproct brown; metazona, margins of epiproct and paraprocts, and the tip of hypoproct darker brown.
</paragraph>
<paragraph id="9E52642EF44CFFCDFF1BF9089A50FEEB" blockId="14.[151,1437,1339,2023]" lastBlockId="15.[151,1437,152,487]" lastPageId="15" lastPageNumber="32" pageId="14" pageNumber="31">
Gonopods (
<figureCitation id="06D678ABF44CFFCCFE8EF9089EC5F932" box="[336,419,1765,1791]" captionStart="FIGURE 8" captionStartId="15.[151,255,1929,1953]" captionTargetBox="[237,1346,556,1897]" captionTargetId="figure@15.[226,1360,545,1906]" captionTargetPageId="15" captionText="FIGURE 8. Thyropygus implicatus, specimen from Thale Ban National Park, gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C: left telopodite, anterior view. D: left telopodite, posterior view." httpUri="https://zenodo.org/record/185979/files/figure.png" pageId="14" pageNumber="31">Figs. 8</figureCitation>
AD): Anterior coxal fold (
<emphasis id="AC99B83CF44CFFCCFD2EF9089C6AF933" box="[752,780,1765,1790]" italics="true" pageId="14" pageNumber="31">ac</emphasis>
) (
<figureCitation id="06D678ABF44CFFCCFCF6F9089C09F932" box="[808,879,1765,1791]" captionStart="FIGURE 8" captionStartId="15.[151,255,1929,1953]" captionTargetBox="[237,1346,556,1897]" captionTargetId="figure@15.[226,1360,545,1906]" captionTargetPageId="15" captionText="FIGURE 8. Thyropygus implicatus, specimen from Thale Ban National Park, gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C: left telopodite, anterior view. D: left telopodite, posterior view." httpUri="https://zenodo.org/record/185979/files/figure.png" pageId="14" pageNumber="31">Fig. 8</figureCitation>
A): lateral process (
<emphasis id="AC99B83CF44CFFCCFB80F9089BE2F933" box="[1118,1156,1765,1790]" italics="true" pageId="14" pageNumber="31">alp</emphasis>
) apically bent abruptly mesad, tip with serrate margins, posterior side with a horizontal crest; mesal process (
<emphasis id="AC99B83CF44CFFCCFB5AF8E19BD1F8E8" box="[1156,1207,1804,1829]" italics="true" pageId="14" pageNumber="31">amp</emphasis>
) much shorter than lateral process, directed meso-distad, pointed. Posterior coxal fold (
<emphasis id="AC99B83CF44CFFCCFC70F8DE9CACF881" box="[942,970,1843,1868]" italics="true" pageId="14" pageNumber="31">pc</emphasis>
) (
<figureCitation id="06D678ABF44CFFCCFC3DF8DF9B4EF881" box="[995,1064,1842,1868]" captionStart="FIGURE 8" captionStartId="15.[151,255,1929,1953]" captionTargetBox="[237,1346,556,1897]" captionTargetId="figure@15.[226,1360,545,1906]" captionTargetPageId="15" captionText="FIGURE 8. Thyropygus implicatus, specimen from Thale Ban National Park, gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C: left telopodite, anterior view. D: left telopodite, posterior view." httpUri="https://zenodo.org/record/185979/files/figure.png" pageId="14" pageNumber="31">Fig. 8</figureCitation>
B) with lateral paracoxites (
<emphasis id="AC99B83CF44CFFCCFAB1F8DE9AEDF881" box="[1391,1419,1843,1868]" italics="true" pageId="14" pageNumber="31">px</emphasis>
), digitiform; mesal process (
<emphasis id="AC99B83CF44CFFCCFE05F8B49D68F8BF" box="[475,526,1881,1906]" italics="true" pageId="14" pageNumber="31">pmp</emphasis>
) complex, strongly developed along an anterior-posterior axis, with a large posterior broad lobe (
<emphasis id="AC99B83CF44CFFCCFE79F86D9EA9F854" box="[423,463,1920,1945]" italics="true" pageId="14" pageNumber="31">pbl</emphasis>
) and an anterior slender digitiform process (
<emphasis id="AC99B83CF44CFFCCFBDDF86D9B57F854" box="[1027,1073,1920,1945]" italics="true" pageId="14" pageNumber="31">adp</emphasis>
); lateral process (
<emphasis id="AC99B83CF44CFFCCFAC8F86D9A58F854" box="[1302,1342,1920,1945]" italics="true" pageId="14" pageNumber="31">plp</emphasis>
) small, pointed. Telopodite (
<figureCitation id="06D678ABF44CFFCCFE4CF84B9E83F80D" box="[402,485,1958,1984]" captionStart="FIGURE 8" captionStartId="15.[151,255,1929,1953]" captionTargetBox="[237,1346,556,1897]" captionTargetId="figure@15.[226,1360,545,1906]" captionTargetPageId="15" captionText="FIGURE 8. Thyropygus implicatus, specimen from Thale Ban National Park, gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C: left telopodite, anterior view. D: left telopodite, posterior view." httpUri="https://zenodo.org/record/185979/files/figure.png" pageId="14" pageNumber="31">Figs. 8</figureCitation>
CD) leaving coxite between
<emphasis id="AC99B83CF44CFFCCFC8EF84A9CE5F80D" box="[848,899,1959,1984]" italics="true" pageId="14" pageNumber="31">pmp</emphasis>
and
<emphasis id="AC99B83CF44CFFCCFC61F84A9C83F80D" box="[959,997,1959,1984]" italics="true" pageId="14" pageNumber="31">plp</emphasis>
; femoral spine (
<emphasis id="AC99B83CF44CFFCCFB70F84A9BA5F80D" box="[1198,1219,1959,1984]" italics="true" pageId="14" pageNumber="31">fe</emphasis>
) basally massive, ending in sharp point, directed almost straightly mesad,
<emphasis id="AC99B83CF44CFFCCFCF0F8209C11F82B" box="[814,887,1997,2022]" italics="true" pageId="14" pageNumber="31">in situ</emphasis>
resting against posterior surface of
<emphasis id="AC99B83CF44CFFCCFAC1F8209A22F82B" box="[1311,1348,1997,2022]" italics="true" pageId="14" pageNumber="31">alp</emphasis>
, fitting under its horizontal crest, crossing over with
<emphasis id="AC99B83CF44DFFCDFD1CFF759DBEFF7C" box="[706,728,152,177]" italics="true" pageId="15" pageNumber="32">fe</emphasis>
of the opposite side; telopodite distally to
<emphasis id="AC99B83CF44DFFCDFB3BFF759B9DFF7C" box="[1253,1275,152,177]" italics="true" pageId="15" pageNumber="32">fe</emphasis>
with a large, round lobe (
<emphasis id="AC99B83CF44DFFCDFEF3FF529E23FF15" box="[301,325,191,216]" italics="true" pageId="15" pageNumber="32">lo</emphasis>
) projecting distolaterally; tibial spine (
<emphasis id="AC99B83CF44DFFCDFCF6FF529C5FFF15" box="[808,825,191,216]" italics="true" pageId="15" pageNumber="32">ti</emphasis>
) long, slender, curving in horizontal plane, its tip upturned and resting against anterior surface of
<emphasis id="AC99B83CF44DFFCDFD3EFF089C6BFF33" box="[736,781,229,254]" italics="true" pageId="15" pageNumber="32">adp</emphasis>
; apical part: spatulate lobe (
<emphasis id="AC99B83CF44DFFCDFBB4FF089B18FF33" box="[1130,1150,229,254]" italics="true" pageId="15" pageNumber="32">sl</emphasis>
) terminating in a sharp brown spine; palette (
<emphasis id="AC99B83CF44DFFCDFE4BFEE19ED4FEE8" box="[405,434,268,293]" italics="true" pageId="15" pageNumber="32">pa</emphasis>
) simple, gutter-like, distally with about nine brownish blepharochaetae (
<emphasis id="AC99B83CF44DFFCDFAD6FEE19A43FEE8" box="[1288,1317,268,293]" italics="true" pageId="15" pageNumber="32">bp</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="D6F737A5F44DFFCDFF1BFEDF9B9EFE2A" pageId="15" pageNumber="32" type="distribution">
<paragraph id="9E52642EF44DFFCDFF1BFEDF9B9EFE2A" blockId="15.[151,1437,152,487]" pageId="15" pageNumber="32">
<emphasis id="AC99B83CF44DFFCDFF1BFEDF9E0AFE81" bold="true" box="[197,364,306,332]" pageId="15" pageNumber="32">Distribution</emphasis>
(
<figureCitation id="06D678ABF44DFFCDFEA1FEDF9E84FE81" box="[383,482,306,332]" captionStart="FIGURE 20" captionStartId="32.[151,255,1208,1232]" captionText="FIGURE 20. Known distribution of the species of the T. opinatus subgroup." pageId="15" pageNumber="32">Fig. 20</figureCitation>
): Described from
<collectingCountry id="E6FA24BEF44DFFCDFD15FEDF9C16FE81" box="[715,880,306,332]" name="Malaysia" pageId="15" pageNumber="32">MALAYSIA</collectingCountry>
, Penang Hill (5° 25ˏ 29˝ N, 100° 16ˏ 7˝ E) (
<bibRefCitation id="FA7C19DFF44DFFCDFF41FEB49E03FEBE" author="Demange" box="[159,357,345,371]" pageId="15" pageNumber="50" refString="Demange, J. M. (1961) Materiaux pour servir a une revision des Harpagophoridae. Memoires du Museum National d'Histoire Naturelle, Ser. A (Zool.), 24, 1 - 274." type="journal article" year="1961">Demange, 1961</bibRefCitation>
), based on specimens (including
<typeStatus id="4156DA8CF44DFFCDFD20FEB49C0DFEBE" box="[766,875,345,371]" pageId="15" pageNumber="32" type="holotype">holotype</typeStatus>
) belonging to the Natural History Museum in London, and further specimens from
<collectingCountry id="E6FA24BEF44DFFCDFD8EFE6D9D8AFE57" box="[592,748,384,410]" name="Malaysia" pageId="15" pageNumber="32">MALAYSIA</collectingCountry>
,
<collectingCountry id="E6FA24BEF44DFFCDFD26FE6D9C39FE57" box="[760,863,384,410]" name="Malaysia" pageId="15" pageNumber="32">Malacca</collectingCountry>
, Sultanate of Kedah (2° 27ˏ 16˝ N, 102° 25ˏ 47˝ E) belonging to Naturalis, Leiden. The species was recorded as new for the
<collectingCountry id="E6FA24BEF44DFFCDFBEFFE4B9B0EFE0D" box="[1073,1128,422,448]" name="Thailand" pageId="15" pageNumber="32">Thai</collectingCountry>
fauna by
<bibRefCitation id="FA7C19DFF44DFFCDFB3CFE4B9AFAFE0D" author="Enghoff" box="[1250,1436,422,448]" pageId="15" pageNumber="50" refString="Enghoff, H. (2005) The millipedes of Thailand (Diplopoda). Steenstrupia, 29 (1), 87 - 103." type="journal article" year="2005">Enghoff (2005)</bibRefCitation>
based on the specimens studied by us (
<collectingCountry id="E6FA24BEF44DFFCDFD85FE209D91FE2A" box="[603,759,461,487]" name="Thailand" pageId="15" pageNumber="32">THAILAND</collectingCountry>
, Satun Province, Thale Ban National Park).
</paragraph>
</subSubSection>
</treatment>
</document>