164 lines
17 KiB
XML
164 lines
17 KiB
XML
<document id="66653A3CA77DAFD8D75241386BC81FDE" ID-CLB-Dataset="264199" ID-DOI="10.11646/zootaxa.5332.1.1" ID-GBIF-Dataset="66e0ec51-d494-43d4-965e-a2cd1462ef54" ID-ISSN="1175-5326" ID-Zenodo-Dep="8261502" ID-ZooBank="424F7439-4095-46A5-93E3-C4130E3B6D9A" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1692350092877" checkinUser="plazi" docAuthor="POORANI, J." docDate="2023" docId="03C7416214CC46A3BDDF36EBFA48FC90" docLanguage="en" docName="zootaxa.5332.1.1.pdf" docOrigin="Zootaxa 5332 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5332.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heteroneda Crotch" docType="treatment" docVersion="8" lastPageNumber="149" masterDocId="FFFE391A14584637BD48367CFFCDFF9B" masterDocTitle="An illustrated guide to lady beetles (Coleoptera: Coccinellidae) of the Indian Subcontinent. Part 1. Tribe Coccinellini" masterLastPageNumber="307" masterPageNumber="1" pageNumber="149" updateTime="1699593964221" updateUser="ExternalLinkService">
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<mods:title id="6848C0CDE63397E8D0042A2B07434AC8">An illustrated guide to lady beetles (Coleoptera: Coccinellidae) of the Indian Subcontinent. Part 1. Tribe Coccinellini</mods:title>
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<mods:namePart id="97DB7E3D09A1C7FA32D66BDB70D48031">POORANI, J.</mods:namePart>
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<mods:typeOfResource id="E452382E1B1AB771274DB15747A1EE4D">text</mods:typeOfResource>
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<mods:title id="4756B9DC583F8EEEF287D4AC2ECE5DE5">Zootaxa</mods:title>
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<mods:part id="910EE4DB701750DDA5243A1043821043">
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<mods:date id="97EFABF01E248B016797E9E6C500634F">2023</mods:date>
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<mods:number id="6EDFB4C44586C2EA02DB15830F24EDFF">2023-08-18</mods:number>
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<mods:number id="5BD4032C1A7956B467995BECF247A93B">5332</mods:number>
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<mods:classification id="9E77637DB0002CDF38F48492E243BA9A">journal article</mods:classification>
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<mods:identifier id="07C77860EF20D8620D09AF2D8BCA0D06" type="CLB-Dataset">264199</mods:identifier>
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<mods:identifier id="F20DFB76BDDB9974A7B8996697EF6823" type="DOI">10.11646/zootaxa.5332.1.1</mods:identifier>
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<treatment id="03C7416214CC46A3BDDF36EBFA48FC90" ID-DOI="http://doi.org/10.5281/zenodo.8273795" ID-GBIF-Taxon="212450243" ID-Zenodo-Dep="8273795" LSID="urn:lsid:plazi:treatment:03C7416214CC46A3BDDF36EBFA48FC90" httpUri="http://treatment.plazi.org/id/03C7416214CC46A3BDDF36EBFA48FC90" lastPageNumber="149" pageId="148" pageNumber="149">
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<subSubSection id="C374A3FF14CC46A3BDDF36EBFE17FF2A" box="[151,474,151,178]" pageId="148" pageNumber="149" type="nomenclature">
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<paragraph id="8BD1F07414CC46A3BDDF36EBFE17FF2A" blockId="148.[151,474,151,178]" box="[151,474,151,178]" pageId="148" pageNumber="149">
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<emphasis id="B91A2C6614CC46A3BDDF36EBFE17FF2A" bold="true" box="[151,474,151,178]" pageId="148" pageNumber="149">
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Genus
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<taxonomicName id="4C6E8BF714CC46A3BDA636EBFE17FF2A" authority="Crotch" authorityName="Crotch" box="[238,474,151,178]" class="Insecta" family="Coccinellidae" genus="Heteroneda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="genus">
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<emphasis id="B91A2C6614CC46A3BDA636EBFEB1FF29" bold="true" box="[238,380,151,178]" italics="true" pageId="148" pageNumber="149">Heteroneda</emphasis>
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Crotch
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</taxonomicName>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C374A3FF14CC46A3BDDF36A0FCB3FE89" pageId="148" pageNumber="149" type="reference_group">
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<paragraph id="8BD1F07414CC46A3BDDF36A0FCB3FE89" blockId="148.[151,1437,219,274]" pageId="148" pageNumber="149">
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<treatmentCitationGroup id="AB7ED75A14CC46A3BDDF36A0FCB3FE89" pageId="148" pageNumber="149">
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<taxonomicName id="4C6E8BF714CC46A3BDDF36A0FE74FF68" authority="Crotch, 1871: 6" authorityName="Crotch" authorityPageNumber="6" authorityYear="1871" box="[151,441,219,244]" class="Insecta" family="Coccinellidae" genus="Heteroneda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="genus">
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<emphasis id="B91A2C6614CC46A3BDDF36A0FEC3FF6F" box="[151,270,220,244]" italics="true" pageId="148" pageNumber="149">Heteroneda</emphasis>
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Crotch, 1871: 6
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</taxonomicName>
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.
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<typeStatus id="54D54ED614CC46A3BC8C36A0FE3BFF6F" box="[452,502,220,244]" pageId="148" pageNumber="149">Type</typeStatus>
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species:
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<taxonomicName id="4C6E8BF714CC46A3BF1E36A0FC19FF6F" authority="Fabricius, 1801" authorityName="Fabricius" authorityYear="1801" box="[598,980,220,244]" class="Insecta" family="Coccinellidae" genus="Coccinella" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="species" species="reticulata">
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<emphasis id="B91A2C6614CC46A3BF1E36A0FCE0FF6F" box="[598,813,220,244]" italics="true" pageId="148" pageNumber="149">Coccinella reticulata</emphasis>
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Fabricius, 1801
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</taxonomicName>
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(by subsequent designation of Crotch 1874: 162) (=
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<taxonomicName id="4C6E8BF714CC46A3BC5B3687FDEAFE89" authority="Crotch, 1871" authorityName="Crotch" authorityYear="1871" box="[275,551,250,274]" class="Insecta" family="Coccinellidae" genus="Heteroneda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="species" species="billardieri">
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<emphasis id="B91A2C6614CC46A3BC5B3687FE54FE89" box="[275,409,250,274]" italics="true" pageId="148" pageNumber="149">H. billardieri</emphasis>
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Crotch, 1871
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</taxonomicName>
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;
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<treatmentCitation id="0ACFD66514CC46A3BF793686FCBFFE89" author="Iablokoff-Khnzorian, S. M." box="[561,882,250,274]" page="538" pageId="148" pageNumber="149" year="1982">
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<bibRefCitation id="EFFF8D8514CC46A3BF793686FCBFFE89" author="Iablokoff-Khnzorian, S. M." box="[561,882,250,274]" pageId="148" pageNumber="149" refId="ref114042" refString="Iablokoff-Khnzorian, S. M. (1982) Les Coccinelles Coleopteres-Coccinellidae. Tribu Coccinellini des regions Palearctique et Orientale. Boubee, Paris, 568 pp. [in French]" type="book" year="1982">Iablokoff-Khnzorian 1982: 538</bibRefCitation>
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</treatmentCitation>
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).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C374A3FF14CC46A3BDDF373CFB8BFDCC" pageId="148" pageNumber="149" type="diagnosis">
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<paragraph id="8BD1F07414CC46A3BDDF373CFB8BFDCC" blockId="148.[151,1437,320,781]" pageId="148" pageNumber="149">
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<emphasis id="B91A2C6614CC46A3BDDF373CFEDFFEC1" bold="true" box="[151,274,320,346]" pageId="148" pageNumber="149">Diagnosis.</emphasis>
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Elytra moderately to strongly convex, glabrous. Anterior clypeal border straight between lateral projections. Antenna 11-segmented; slightly shorter than head capsule, with a weak three-segmented club; terminal antennomere longer than penultimate, rounded apically. Terminal maxillary palpomere securiform. Pronotal disc evenly convex transversely, with narrowly upturned or slightly thickened external borders. Prothoracic hypomeron without foveae near anterolateral corners; prosternal process with distinct carinae extending forward to the middle of the prosternum. Anterior margin of mesoventrite distinctly, deeply emarginate medially. Lateral margins of elytra with complete beads; epipleuron without foveae. Tibial spur formula 0-2-2. Abdominal postcoxal line not recurved, incomplete laterally, without an oblique associated line (From
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<bibRefCitation id="EFFF8D8514CC46A3BE1C3442FBF7FDCC" author="Slipinski, A. & Li, J. & Pang, H." box="[852,1082,572,601]" pageId="148" pageNumber="149" refId="ref122780" refString="Slipinski, A., Li, J. & Pang, H. (2020) Ladybird Beetles of the Australo-Pacific Region Coleoptera: Coccinellidae: Coccinellini. CSIRO Publishing, Clayton, Victoria, 240 pp. https: // doi. org / 10.1071 / 9781486303885" type="book" year="2020">
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Ślipiński
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<emphasis id="B91A2C6614CC46A3BEF63441FC22FDCD" box="[958,1007,572,598]" italics="true" pageId="148" pageNumber="149">et al</emphasis>
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. 2020
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C374A3FF14CC46A3BD8F341CFCDCFDE1" box="[199,785,608,634]" pageId="148" pageNumber="149" type="distribution">
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<paragraph id="8BD1F07414CC46A3BD8F341CFCDCFDE1" blockId="148.[151,1437,320,781]" box="[199,785,608,634]" pageId="148" pageNumber="149">
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<emphasis id="B91A2C6614CC46A3BD8F341CFEACFDE1" bold="true" box="[199,353,608,634]" pageId="148" pageNumber="149">Distribution.</emphasis>
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<collectingRegion id="49AA3E9614CC46A3BC2F341CFE0EFDE1" box="[359,451,608,634]" country="Saudi Arabia" name="Ash Sharqiyah" pageId="148" pageNumber="149">Oriental</collectingRegion>
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and Australo-Pacific regions.
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</paragraph>
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</subSubSection>
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<subSubSection id="C374A3FF14CC46A3BD8F34F8FABAFD7C" pageId="148" pageNumber="149" type="discussion">
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<paragraph id="8BD1F07414CC46A3BD8F34F8FABAFD7C" blockId="148.[151,1437,320,781]" pageId="148" pageNumber="149">
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<emphasis id="B91A2C6614CC46A3BD8F34F8FEF0FD05" bold="true" box="[199,317,644,670]" pageId="148" pageNumber="149">Affinities.</emphasis>
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<bibRefCitation id="EFFF8D8514CC46A3BC0934F8FDA5FD04" author="Tomaszewska, W. & Escalona, H. E. & Hartley, D. & Li, J. & Wang, X. & Li, H. S. & Pang, H. & Slipinski, A. & Zwick, A." box="[321,616,644,671]" pageId="148" pageNumber="149" pagination="611 - 631" refId="ref123461" refString="Tomaszewska, W., Escalona, H. E., Hartley, D., Li, J., Wang, X., Li, H. S., Pang, H., Slipinski, A. & Zwick, A. (2021) Phylogeny of true ladybird beetles (Coccinellidae: Coccinellini) reveals pervasive convergent evolution and a rapid Cenozoic radiation. Systematic Entomology, 46, 611 - 631. https: // doi. org / 10.1111 / syen. 12479" type="journal article" year="2021">
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Tomaszewska
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<emphasis id="B91A2C6614CC46A3BCAB34F9FDDFFD05" box="[483,530,644,670]" italics="true" pageId="148" pageNumber="149">et al</emphasis>
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. (2021)
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</bibRefCitation>
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included it in the
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<emphasis id="B91A2C6614CC46A3BE6134F8FC61FD05" box="[809,940,644,670]" italics="true" pageId="148" pageNumber="149">Synonycha-</emphasis>
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group of genera and recovered it as a strongly supported genus in a clade comprising
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<taxonomicName id="4C6E8BF714CC46A3BF0434D4FD5AFD59" box="[588,663,680,706]" class="Insecta" family="Coccinellidae" genus="Calvia" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="genus">
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<emphasis id="B91A2C6614CC46A3BF0434D4FD5AFD59" box="[588,663,680,706]" italics="true" pageId="148" pageNumber="149">Calvia</emphasis>
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</taxonomicName>
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(part),
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<taxonomicName id="4C6E8BF714CC46A3BFA234D4FCA7FD59" box="[746,874,680,706]" class="Insecta" family="Coccinellidae" genus="Singhikalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="genus">
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<emphasis id="B91A2C6614CC46A3BFA234D4FCA7FD59" box="[746,874,680,706]" italics="true" pageId="148" pageNumber="149">Singhikalia</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C6E8BF714CC46A3BE3F34D4FBC8FD59" box="[887,1029,680,706]" class="Insecta" family="Coccinellidae" genus="Phrynocaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="genus">
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<emphasis id="B91A2C6614CC46A3BE3F34D4FBC8FD59" box="[887,1029,680,706]" italics="true" pageId="148" pageNumber="149">Phrynocaria</emphasis>
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</taxonomicName>
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(including part of
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<taxonomicName id="4C6E8BF714CC46A3B99F34D4FA90FD59" box="[1239,1373,680,706]" class="Insecta" family="Coccinellidae" genus="Coelophora" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="genus">
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<emphasis id="B91A2C6614CC46A3B99F34D4FA90FD59" box="[1239,1373,680,706]" italics="true" pageId="148" pageNumber="149">Coelophora</emphasis>
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</taxonomicName>
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), and
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<taxonomicName id="4C6E8BF714CC46A3BDDF34B1FEE6FD7D" authorityName="Crotch" authorityYear="1874" box="[151,299,717,742]" class="Insecta" family="Coccinellidae" genus="Docimocaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="genus">
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<emphasis id="B91A2C6614CC46A3BDDF34B1FEE6FD7D" box="[151,299,717,742]" italics="true" pageId="148" pageNumber="149">Docimocaria</emphasis>
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</taxonomicName>
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. Their analyses suggest
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<taxonomicName id="4C6E8BF714CC46A3BF7534B0FD0DFD7D" box="[573,704,716,742]" class="Insecta" family="Coccinellidae" genus="Heteroneda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="genus">
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<emphasis id="B91A2C6614CC46A3BF7534B0FD0DFD7D" box="[573,704,716,742]" italics="true" pageId="148" pageNumber="149">Heteroneda</emphasis>
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</taxonomicName>
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is closely related to
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<taxonomicName id="4C6E8BF714CC46A3BEEF34B0FBF8FD7D" box="[935,1077,716,742]" class="Insecta" family="Coccinellidae" genus="Phrynocaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="genus">
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<emphasis id="B91A2C6614CC46A3BEEF34B0FBF8FD7D" box="[935,1077,716,742]" italics="true" pageId="148" pageNumber="149">Phrynocaria</emphasis>
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</taxonomicName>
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, “albeit with weak support”.
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</paragraph>
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</subSubSection>
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<subSubSection id="C374A3FF14CC46A3BD8F348DFA48FC90" box="[199,1413,752,781]" pageId="148" pageNumber="149" type="taxon_list">
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<paragraph id="8BD1F07414CC46A3BD8F348DFA48FC90" blockId="148.[151,1437,320,781]" box="[199,1413,752,781]" pageId="148" pageNumber="149">
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<emphasis id="B91A2C6614CC46A3BD8F348DFE5FFC91" bold="true" box="[199,402,752,779]" pageId="148" pageNumber="149">Included species.</emphasis>
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The
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<typeStatus id="54D54ED614CC46A3BC83348DFE36FC90" box="[459,507,753,779]" pageId="148" pageNumber="149">type</typeStatus>
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species,
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<taxonomicName id="4C6E8BF714CC46A3BF29348DFC86FC91" authority="Crotch" authorityName="Crotch" box="[609,843,752,778]" class="Insecta" family="Coccinellidae" genus="Heteroneda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="species" species="billardieri">
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<emphasis id="B91A2C6614CC46A3BF29348DFD38FC91" box="[609,757,752,778]" italics="true" pageId="148" pageNumber="149">H. billardieri</emphasis>
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Crotch
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</taxonomicName>
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, is distributed in this region (
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<bibRefCitation id="EFFF8D8514CC46A3B9D9348EFAB7FC90" author="Slipinski, A. & Li, J. & Pang, H." box="[1169,1402,752,781]" pageId="148" pageNumber="149" refId="ref122780" refString="Slipinski, A., Li, J. & Pang, H. (2020) Ladybird Beetles of the Australo-Pacific Region Coleoptera: Coccinellidae: Coccinellini. CSIRO Publishing, Clayton, Victoria, 240 pp. https: // doi. org / 10.1071 / 9781486303885" type="book" year="2020">
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Ślipiński
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<emphasis id="B91A2C6614CC46A3B9B6348DFAE2FC91" box="[1278,1327,752,778]" italics="true" pageId="148" pageNumber="149">et al</emphasis>
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. 2020
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |