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<document id="07AC88467D1643749BD4ADF31101CB67" ID-CLB-Dataset="28091" ID-DOI="10.11646/zootaxa.4504.1.3" ID-GBIF-Dataset="0b803003-244e-4511-aaf4-5e9c3f863861" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606164" ID-ZooBank="52BA28B6-4280-4BD0-8B38-C794F04E4A66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553577945224" checkinUser="plazi" docAuthor="Abdalla, Ishtiag H., Deschodt, Christian M., Scholtz, Clarke H. &amp; Sole, Catherine L." docDate="2018" docId="B276DC1EFFAEFFE8FF0DF89DFB3905BD" docLanguage="en" docName="zootaxa.4504.1.3.pdf" docOrigin="Zootaxa 4504 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macroderes foveatus Frolov &amp; Scholtz 2005" docType="treatment" docVersion="8" lastPageNumber="55" masterDocId="4E4FA466FFA2FFE6FF9AFFF0FFD20545" masterDocTitle="An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa" masterLastPageNumber="75" masterPageNumber="41" pageNumber="53" updateTime="1698712718888" updateUser="ExternalLinkService">
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<mods:title id="011CD6DCF34894ECF8992A8240F00FFE">An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa</mods:title>
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<mods:namePart id="D4516989B181A163CBBF4AFFF321DF8B">Abdalla, Ishtiag H.</mods:namePart>
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<mods:namePart id="06851A2227548CD28A0EC451D7117AC0">Deschodt, Christian M.</mods:namePart>
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<treatment id="B276DC1EFFAEFFE8FF0DF89DFB3905BD" ID-DOI="http://doi.org/10.5281/zenodo.5996734" ID-GBIF-Taxon="154494815" ID-Zenodo-Dep="5996734" LSID="urn:lsid:plazi:treatment:B276DC1EFFAEFFE8FF0DF89DFB3905BD" httpUri="http://treatment.plazi.org/id/B276DC1EFFAEFFE8FF0DF89DFB3905BD" lastPageId="14" lastPageNumber="55" pageId="12" pageNumber="53">
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<paragraph id="3A606D08FFAEFFEAFF0DF89DFD6A02C2" blockId="12.[151,696,1901,1961]" box="[151,696,1901,1927]" pageId="12" pageNumber="53">
<heading id="6128DA64FFAEFFEAFF0DF89DFD6A02C2" bold="true" box="[151,696,1901,1927]" fontSize="11" level="1" pageId="12" pageNumber="53" reason="1">
<taxonomicName id="FDDF168BFFAEFFEAFF0DF89DFD6A02C2" ID-CoL="3X4TL" authority="Frolov &amp; Scholtz, 2005" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[151,696,1901,1927]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="53" phylum="Arthropoda" rank="species" species="foveatus">
<emphasis id="08ABB11AFFAEFFEAFF0DF89DFD6A02C2" bold="true" box="[151,696,1901,1927]" pageId="12" pageNumber="53">
<emphasis id="08ABB11AFFAEFFEAFF0DF89DFE4002C2" bold="true" box="[151,402,1901,1927]" italics="true" pageId="12" pageNumber="53">Macroderes foveatus</emphasis>
<bibRefCitation id="5E4E10F9FFAEFFEAFE00F89DFD6A02C2" author="Frolov, A. V. &amp; Scholtz, C. H." box="[410,696,1901,1927]" pageId="12" pageNumber="53" pagination="373 - 393" refId="ref21694" refString="Frolov, A. V. &amp; Scholtz, C. H. (2005) Revision of the southern African genus Macroderes Westwood (Coleoptera: Scarabaeidae: Scarabaeinae). Annales de la Societe Entomologique de France, 40 (3 - 4), 373 - 393. https: // doi. org / 10.1080 / 00379271.2004.10697429" type="journal article" year="2005">Frolov &amp; Scholtz, 2005</bibRefCitation>
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<subSubSection id="72C53E83FFAEFFEAFF0DF860FE2302ED" box="[151,497,1936,1961]" pageId="12" pageNumber="53" type="description">
<paragraph id="3A606D08FFAEFFEAFF0DF860FE2302ED" blockId="12.[151,696,1901,1961]" box="[151,497,1936,1961]" pageId="12" pageNumber="53">
(
<figureCitation id="A2E4718DFFAEFFEAFF05F860FF3F02ED" box="[159,237,1936,1961]" captionStart="FIGURES 710" captionStartId="5.[151,264,1812,1835]" captionTargetBox="[330,1235,720,1729]" captionTargetId="figure@5.[319,1279,700,1780]" captionTargetPageId="5" captionText="FIGURES 710. Habitus of Macroderes species. 7, M. fornicatus Sharp, 1880; 8, M. foveatus Frolov &amp; Scholtz, 2005; 9, M. bias (Olivier, 1789); 10, M. greeni (Kirby, 1818)." httpUri="https://zenodo.org/record/2606170/files/figure.png" pageId="12" pageNumber="53">Figs. 8</figureCitation>
,
<figureCitation id="A2E4718DFFAEFFEAFF60F860FECA02ED" box="[250,280,1936,1960]" captionStart="FIGURES 2439" captionStartId="10.[151,264,1343,1366]" captionTargetBox="[323,1250,205,1286]" captionTargetId="figure@10.[323,1283,195,1303]" captionTargetPageId="10" captionText="FIGURES 2439. Internal sac of aedeagus (24), aedeagus (25) and internal sac sclerite (26-39) of Macroderes species. 26, M. endroedyi Frolov &amp; Scholtz, 2005; 27, M. amplior Frolov &amp; Scholtz, 2005; 28, M. bias (Olivier, 1789); 29, M. cederbergensis Abdalla &amp; Deschodt, new species; 30, M. cornutus Frolov &amp; Scholtz, 2005; 31, M. foveatus Frolov &amp; Scholtz, 2005; 32, M. gifboomi Abdalla &amp; Scholtz, new species; 33, M. leipoldti Abdalla &amp; Deschodt, new species; 34, M. mutilans Kolbe, 1908; 35, M. namakwanus Frolov &amp; Scholtz, 2005; 36, M. oreatus Abdalla &amp; Deschodt, new species; 37, M. porselinus Abdalla, new species; 38, M. soleiana Abdalla &amp; Deschodt, new species; 39, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606178/files/figure.png" pageId="12" pageNumber="53">31</figureCitation>
,
<figureCitation id="A2E4718DFFAEFFEAFEBFF860FE9102ED" box="[293,323,1936,1960]" captionStart="FIGURES 4045" captionStartId="12.[151,264,1736,1759]" captionTargetBox="[306,1266,609,1690]" captionTargetId="figure@12.[306,1266,609,1690]" captionTargetPageId="12" captionText="FIGURES 4045. Pronotum of Macroderes species. 40, M. foveatus Frolov &amp; Scholtz, 2005; 41, M. cornutus Frolov &amp; Scholtz, 2005; 42, M. namakwanus Frolov &amp; Scholtz, 2005; 43, M. politulus Preudhomme de Borre, 1880; 44, M. greeni (Kirby, 1818); 45, M. endroedyi Frolov &amp; Scholtz, 2005." httpUri="https://zenodo.org/record/2606180/files/figure.png" pageId="12" pageNumber="53">40</figureCitation>
,
<figureCitation id="A2E4718DFFAEFFEAFED4F860FE1202EC" box="[334,448,1936,1961]" captionStart="FIGURES 4651" captionStartId="13.[151,264,1948,1971]" captionTargetBox="[311,1265,844,1918]" captionTargetId="figure@13.[311,1265,844,1918]" captionTargetPageId="13" captionText="FIGURES 4651. Lateral side of pronotum (46, 47) and dorsal and lateral side of the head (4851) of Macroderes species. 46, 49, 51, M. foveatus Frolov &amp; Scholtz, 2005; 47, 48, 50, M. cornutus Frolov &amp; Scholtz, 2005." httpUri="https://zenodo.org/record/2606184/files/figure.png" pageId="12" pageNumber="53">46, 49, 51</figureCitation>
,
<figureCitation id="A2E4718DFFAEFFEAFE56F860FE3B02ED" box="[460,489,1936,1960]" captionStart="FIGURES 7075" captionStartId="18.[151,264,1624,1647]" captionTargetBox="[159,1435,197,1611]" captionTargetId="figure@18.[159,1435,197,1611]" captionTargetPageId="18" captionText="FIGURES 7075. Sculpture of second elytra intervals of Macroderes species. 70, M. arrowi Janssens, 1939; 71, M. cornutus Frolov &amp; Scholtz, 2005; 72, M. endroedyi Frolov &amp; Scholtz, 2005;73, M. foveatus Frolov &amp; Scholtz, 2005; 74, M. mutilans Kolbe, 1908; 75, M. namakwanus Frolov &amp; Scholtz, 2005." httpUri="https://zenodo.org/record/2606196/files/figure.png" pageId="12" pageNumber="53">73</figureCitation>
)
</paragraph>
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<paragraph id="3A606D08FFAEFFEBFF0DF828FC7605F5" blockId="12.[151,666,2008,2030]" lastBlockId="13.[151,1437,151,789]" lastPageId="13" lastPageNumber="54" pageId="12" pageNumber="53">
<treatmentCitationGroup id="1ACF4A26FFAEFFEBFF0DF828FC7605F5" lastPageId="13" lastPageNumber="54" pageId="12" pageNumber="53">
<taxonomicName id="FDDF168BFFAEFFEAFF0DF828FD4402AB" ID-CoL="3X4TL" authority="Frolov &amp; Scholtz, 2005: 390" authorityName="Frolov &amp; Scholtz" authorityPageNumber="390" authorityYear="2005" box="[151,662,2008,2030]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="53" phylum="Arthropoda" rank="species" species="foveatus">
<emphasis id="08ABB11AFFAEFFEAFF0DF828FEBB02AB" box="[151,361,2008,2030]" italics="true" pageId="12" pageNumber="53">Macroderes foveatus</emphasis>
<treatmentCitation id="BB7E4B19FFAEFFEAFEEAF828FD4402AB" author="Frolov, A. V. &amp; Scholtz, C. H." box="[368,662,2008,2030]" page="390" pageId="12" pageNumber="53" year="2005">
<bibRefCitation id="5E4E10F9FFAEFFEAFEEAF828FDB102AB" author="Frolov, A. V. &amp; Scholtz, C. H." box="[368,611,2008,2030]" pageId="12" pageNumber="53" pagination="373 - 393" refId="ref21694" refString="Frolov, A. V. &amp; Scholtz, C. H. (2005) Revision of the southern African genus Macroderes Westwood (Coleoptera: Scarabaeidae: Scarabaeinae). Annales de la Societe Entomologique de France, 40 (3 - 4), 373 - 393. https: // doi. org / 10.1080 / 00379271.2004.10697429" type="journal article" year="2005">Frolov &amp; Scholtz, 2005</bibRefCitation>
: 390
</treatmentCitation>
</taxonomicName>
.
<emphasis id="08ABB11AFFAFFFEBFF0DFF67FEE705F5" bold="true" box="[151,309,151,176]" pageId="13" pageNumber="54">
<typeStatus id="E564D3AAFFAFFFEBFF0DFF67FF0005F5" box="[151,210,151,176]" pageId="13" pageNumber="54">Type</typeStatus>
locality.
</emphasis>
S. Afr., SW
<collectingRegion id="F81BA3EAFFAFFFEBFE5FFF68FDD305F5" box="[453,513,152,176]" country="South Africa" name="Western Cape" pageId="13" pageNumber="54">Cape</collectingRegion>
, 3 Km E of Veldrif,
<geoCoordinate id="5FEB0BCFFFAFFFEBFD7FFF67FCEF05F5" box="[741,829,151,176]" direction="south" orientation="latitude" pageId="13" pageNumber="54" precision="925" value="-32.766666">32°46'S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFAFFFEBFCD9FF68FC4D05F5" box="[835,927,152,176]" direction="east" orientation="longitude" pageId="13" pageNumber="54" precision="925" value="18.233334">18°14'E</geoCoordinate>
.
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</paragraph>
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<emphasis id="08ABB11AFFAFFFEBFF5DFF4CFE230590" bold="true" box="[199,497,188,213]" pageId="13" pageNumber="54">Type material examined.</emphasis>
<materialsCitation id="8AB76755FFAFFFEBFE62FF4CFA4B0458" ID-GBIF-Occurrence="2028137780" collectingDate="1981-08-31" collectionCode="TMSA" collectorName="S. Afr &amp; Endrody-Younga" latitude="-32.766666" location="Veldrif" longLatPrecision="1209" longitude="18.233334" pageId="13" pageNumber="54" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="holotype">
<typeStatus id="E564D3AAFFAFFFEBFE62FF4CFDB10590" box="[504,611,188,213]" pageId="13" pageNumber="54" type="holotype">Holotype</typeStatus>
,
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(
<collectionCode id="5CCEF5CDFFAFFFEBFD0EFF4CFD3B0591" box="[660,745,188,212]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="13" pageNumber="54">TMSA</collectionCode>
): “
<collectorName id="972A08DEFFAFFFEBFC93FF4CFC9C0590" box="[777,846,188,213]" pageId="13" pageNumber="54">S. Afr</collectorName>
/ SW
<collectingRegion id="F81BA3EAFFAFFFEBFC0DFF4CFC030591" box="[919,977,188,212]" country="South Africa" name="Western Cape" pageId="13" pageNumber="54">Cape</collectingRegion>
/
<location id="3F003BD3FFAFFFEBFC72FF4CFBE40590" LSID="urn:lsid:plazi:treatment:B276DC1EFFAEFFE8FF0DF89DFB3905BD:3F003BD3FFAFFFEBFC72FF4CFBE40590" box="[1000,1078,188,213]" latitude="-32.766666" longLatPrecision="1209" longitude="18.233334" name="Veldrif" pageId="13" pageNumber="54" stateProvince="Western Cape">Veldrif</location>
/ 3Km E /
<geoCoordinate id="5FEB0BCFFFAFFFEBFB2CFF4CFACC0591" box="[1206,1310,188,213]" direction="south" orientation="latitude" pageId="13" pageNumber="54" precision="925" value="-32.766666">32° 46' S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFAFFFEBFAAFFF4CFA4E0591" box="[1333,1436,188,212]" direction="east" orientation="longitude" pageId="13" pageNumber="54" precision="925" value="18.233334">18° 14' E</geoCoordinate>
[printed] //
<date id="4E614BC8FFAFFFEBFE86FF10FE5905BD" box="[284,395,224,248]" pageId="13" pageNumber="54" value="1981-08-31">
<collectingDate id="5E25B220FFAFFFEBFE86FF10FE5905BD" box="[284,395,224,248]" pageId="13" pageNumber="54" value="1981-08-31">31.8.1981</collectingDate>
</date>
/ EY: 1870 /
<collectingMethod id="E39E151FFFAFFFEBFDA6FF2FFD1805B2" box="[572,714,223,248]" pageId="13" pageNumber="54">ground traps</collectingMethod>
/ 59 days /
<collectorName id="972A08DEFFAFFFEBFCC3FF2FFBF905BD" box="[857,1067,223,248]" pageId="13" pageNumber="54">Endrödy—Younga</collectorName>
leg [printed] // groundtrap with meat bait [printed] //
<typeStatus id="E564D3AAFFAFFFEBFE07FEF4FD910459" box="[413,579,260,284]" pageId="13" pageNumber="54" type="holotype">HOLOTYPUS</typeStatus>
/
<taxonomicName id="FDDF168BFFAFFFEBFDFFFEF4FC860459" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[613,852,260,284]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="foveatus">
<emphasis id="08ABB11AFFAFFFEBFDFFFEF4FC860459" box="[613,852,260,284]" italics="true" pageId="13" pageNumber="54">Macroderes foveatus</emphasis>
</taxonomicName>
/ Frolov &amp; Scholtz / 2003 [red label, printed]”
</materialsCitation>
.
<materialsCitation id="8AB76755FFAFFFEBFF0DFED8FD7F0405" ID-GBIF-Occurrence="2028137795" box="[151,685,295,320]" collectingDate="1981-08-31" collectionCode="TMSA" collectorName="S. Afr &amp; Endrody-Younga" latitude="-32.766666" location="Veldrif" longLatPrecision="1209" longitude="18.233334" pageId="13" pageNumber="54" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape" typeStatus="paratype">
<typeStatus id="E564D3AAFFAFFFEBFF0DFED8FEDB0405" box="[151,265,296,320]" pageId="13" pageNumber="54" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="2CD9A681FFAFFFEBFE8EFED8FEED047A" box="[276,319,296,320]" pageId="13" pageNumber="54" type="female">1 ♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFAFFFEBFED5FED8FE720405" box="[335,416,296,320]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="13" pageNumber="54">TMSA</collectionCode>
) same data as holotype
</materialsCitation>
;
<materialsCitation id="8AB76755FFAFFFEBFD20FED8FB7204CD" ID-GBIF-Occurrence="2028137814" collectingDate="1983-08-23" collectionCode="TMSA" collectorName="S. Afr &amp; Penrith" latitude="-32.933334" location="Brakfontein" longLatPrecision="1207" longitude="18.25" pageId="13" pageNumber="54" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="2CD9A681FFAFFFEBFD20FED8FD34047A" box="[698,742,296,320]" pageId="13" pageNumber="54" type="female">1 ♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFAFFFEBFD6CFED8FC990405" box="[758,843,296,320]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="13" pageNumber="54">TMSA</collectionCode>
): “
<collectorName id="972A08DEFFAFFFEBFCF6FED8FC630405" box="[876,945,295,320]" pageId="13" pageNumber="54">S. Afr</collectorName>
/ S.W.
<collectingRegion id="F81BA3EAFFAFFFEBFB9DFED8FB930405" box="[1031,1089,296,320]" country="South Africa" name="Western Cape" pageId="13" pageNumber="54">Cape</collectingRegion>
/
<location id="3F003BD3FFAFFFEBFBC2FED7FB0C0405" LSID="urn:lsid:plazi:treatment:B276DC1EFFAEFFE8FF0DF89DFB3905BD:3F003BD3FFAFFFEBFBC2FED7FB0C0405" box="[1112,1246,295,320]" latitude="-32.933334" longLatPrecision="1207" longitude="18.25" name="Brakfontein" pageId="13" pageNumber="54" stateProvince="Western Cape">Brakfontein</location>
farm /
<geoCoordinate id="5FEB0BCFFFAFFFEBFAAEFED8FA490405" box="[1332,1435,295,320]" direction="south" orientation="latitude" pageId="13" pageNumber="54" precision="925" value="-32.933334">32° 56' S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFAFFFEBFF0DFEBCFF2C0421" box="[151,254,332,357]" direction="east" orientation="longitude" pageId="13" pageNumber="54" precision="925" value="18.25">18° 15' E</geoCoordinate>
[printed] //
<date id="4E614BC8FFAFFFEBFE1EFEBCFE260421" box="[388,500,332,356]" pageId="13" pageNumber="54" value="1983-08-23">
<collectingDate id="5E25B220FFAFFFEBFE1EFEBCFE260421" box="[388,500,332,356]" pageId="13" pageNumber="54" value="1983-08-23">23.8.1983</collectingDate>
</date>
/ EY: 1967 / groundtraps / 72 days / Endroeydi,
<collectorName id="972A08DEFFAFFFEBFBBBFEBCFBA00420" box="[1057,1138,332,357]" pageId="13" pageNumber="54">Penrith</collectorName>
leg [printed] // groundtrap with faeces bait [printed] //
<typeStatus id="E564D3AAFFAFFFEBFE57FE80FDB804CD" box="[461,618,368,392]" pageId="13" pageNumber="54" type="paratype">PARATYPUS</typeStatus>
/
<taxonomicName id="FDDF168BFFAFFFEBFD1AFE80FCB804CD" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[640,874,368,392]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="foveatus">
<emphasis id="08ABB11AFFAFFFEBFD1AFE80FCB804CD" box="[640,874,368,392]" italics="true" pageId="13" pageNumber="54">Macroderes foveatus</emphasis>
</taxonomicName>
/ Frolov &amp; Scholtz / 2003”
</materialsCitation>
.
</paragraph>
<paragraph id="3A606D08FFAFFFEBFF5DFE64FEF3075D" blockId="13.[151,1437,151,789]" pageId="13" pageNumber="54">
<emphasis id="08ABB11AFFAFFFEBFF5DFE64FDE504E8" bold="true" box="[199,567,404,429]" pageId="13" pageNumber="54">Additional material examined.</emphasis>
<materialsCitation id="8AB76755FFAFFFEBFDA5FE64FAB00495" ID-GBIF-Occurrence="2028137791" collectingDate="2016-07-19" collectionCode="UPSA" collectorName="C. Deschodt &amp; W. P. Strumpher" country="South Africa" latitude="-32.793938" location="Veldrif" longLatPrecision="1" longitude="18.248173" pageId="13" pageNumber="54" specimenCount="2" specimenCount-male="2" stateProvince="Western Cape">
<specimenCount id="2CD9A681FFAFFFEBFDA5FE64FD5204E9" box="[575,640,404,428]" pageId="13" pageNumber="54" type="male">2 ♂♂</specimenCount>
(
<collectionCode id="5CCEF5CDFFAFFFEBFD0BFE64FD0D04E9" box="[657,735,404,428]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="13" pageNumber="54">UPSA</collectionCode>
): “
<collectingCountry id="42C82D98FFAFFFEBFD67FE64FCE404E9" box="[765,822,404,428]" name="South Africa" pageId="13" pageNumber="54">RSA</collectingCountry>
/
<collectingRegion id="F81BA3EAFFAFFFEBFCD5FE65FB8B04E8" box="[847,1113,404,429]" country="South Africa" name="Western Cape" pageId="13" pageNumber="54">Western Cape Province</collectingRegion>
/
<location id="3F003BD3FFAFFFEBFBE8FE64FB1204E8" LSID="urn:lsid:plazi:treatment:B276DC1EFFAEFFE8FF0DF89DFB3905BD:3F003BD3FFAFFFEBFBE8FE64FB1204E8" box="[1138,1216,404,429]" country="South Africa" latitude="-32.793938" longLatPrecision="1" longitude="18.248173" name="Veldrif" pageId="13" pageNumber="54" stateProvince="Western Cape">Veldrif</location>
/ 7 Km E /
<date id="4E614BC8FFAFFFEBFAD6FE64FF1D0495" pageId="13" pageNumber="54" value="2016-07-19">
<collectingDate id="5E25B220FFAFFFEBFAD6FE64FF1D0495" pageId="13" pageNumber="54" value="2016-07-19">19. vii. 2016</collectingDate>
</date>
/
<geoCoordinate id="5FEB0BCFFFAFFFEBFF70FE48FE7A0495" box="[234,424,440,464]" direction="south" orientation="latitude" pageId="13" pageNumber="54" precision="1" value="-32.793938">32° 47' 38.18'' S</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFAFFFEBFE2BFE48FDA2048A" box="[433,624,439,464]" direction="east" orientation="longitude" pageId="13" pageNumber="54" precision="1" value="18.248173">18° 14' 53.42'' E</geoCoordinate>
//
<collectingMethod id="E39E151FFFAFFFEBFD09FE47FC8F048A" box="[659,861,439,464]" pageId="13" pageNumber="54">baited pitfall trap</collectingMethod>
: pig dung /
<collectorName id="972A08DEFFAFFFEBFC70FE48FBAA0495" box="[1002,1144,439,464]" pageId="13" pageNumber="54">C. Deschodt</collectorName>
&amp;
<collectorName id="972A08DEFFAFFFEBFB3AFE48FA8A0495" box="[1184,1368,439,464]" pageId="13" pageNumber="54">W.P. Strümpher</collectorName>
</materialsCitation>
;
<materialsCitation id="8AB76755FFAFFFEBFAEAFE48FECF075D" ID-GBIF-Occurrence="2028137788" collectingDate="2015-08-14" collectionCode="UPSA" collectorName="C. Deschodt &amp; W. P. Strumpher" country="South Africa" latitude="-31.50317" location="Kommandokraal" longLatPrecision="1" longitude="18.20939" pageId="13" pageNumber="54" specimenCount="1" specimenCount-female="1">
<specimenCount id="2CD9A681FFAFFFEBFAEAFE48FA4E048A" box="[1392,1436,440,464]" pageId="13" pageNumber="54" type="female">1 ♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFAFFFEBFF05FE2CFF3F04B1" box="[159,237,476,500]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="13" pageNumber="54">UPSA</collectionCode>
): “
<collectingCountry id="42C82D98FFAFFFEBFE96FE2CFE7204B0" box="[268,416,476,501]" name="South Africa" pageId="13" pageNumber="54">South Africa</collectingCountry>
/
<location id="3F003BD3FFAFFFEBFE20FE2CFDA404B0" LSID="urn:lsid:plazi:treatment:B276DC1EFFAEFFE8FF0DF89DFB3905BD:3F003BD3FFAFFFEBFE20FE2CFDA404B0" box="[442,630,476,501]" country="South Africa" latitude="-31.50317" longLatPrecision="1" longitude="18.20939" name="Kommandokraal" pageId="13" pageNumber="54">Kommandokraal</location>
farm /
<geoCoordinate id="5FEB0BCFFFAFFFEBFD54FE2CFC8804B0" box="[718,858,476,501]" direction="south" orientation="latitude" pageId="13" pageNumber="54" precision="1" value="-31.50317">S 31.50317°</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFAFFFEBFCFEFE2DFC2D04B1" box="[868,1023,476,500]" direction="east" orientation="longitude" pageId="13" pageNumber="54" precision="1" value="18.20939">E 018.20939°</geoCoordinate>
/
<date id="4E614BC8FFAFFFEBFB80FE2CFB4104B0" box="[1050,1171,476,501]" pageId="13" pageNumber="54" value="2015-08-14">
<collectingDate id="5E25B220FFAFFFEBFB80FE2CFB4104B0" box="[1050,1171,476,501]" pageId="13" pageNumber="54" value="2015-08-14">14.8. 2015</collectingDate>
</date>
/
<collectorName id="972A08DEFFAFFFEBFB37FE2CFAEB04B0" box="[1197,1337,476,501]" pageId="13" pageNumber="54">C. Deschodt</collectorName>
&amp;
<collectorName id="972A08DEFFAFFFEBFAFBFE2DFEC1075D" pageId="13" pageNumber="54">W. P. Strümpher</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="3A606D08FFAFFFEBFF5DFDD4FF11071A" blockId="13.[151,1437,151,789]" pageId="13" pageNumber="54">
<emphasis id="08ABB11AFFAFFFEBFF5DFDD4FE9E0778" bold="true" box="[199,332,548,573]" pageId="13" pageNumber="54">Size range.</emphasis>
Males length:
<quantity id="FD27C0EDFFAFFFEBFE60FDD4FD730779" box="[506,673,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" metricValueMax="1.04" metricValueMin="1.02" pageId="13" pageNumber="54" unit="mm" value="10.3" valueMax="10.4" valueMin="10.2">10.210.4 mm</quantity>
, width:
<quantity id="FD27C0EDFFAFFFEBFD65FDD4FC5E0779" box="[767,908,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="7.4" metricValueMin="6.2" pageId="13" pageNumber="54" unit="mm" value="6.8" valueMax="7.4" valueMin="6.2">6.27.4 mm</quantity>
; females length:
<quantity id="FD27C0EDFFAFFFEBFBCBFDD4FB380779" box="[1105,1258,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="9.45" metricValueMax="10.1" metricValueMin="8.8" pageId="13" pageNumber="54" unit="mm" value="9.45" valueMax="10.1" valueMin="8.8">8.810.1 mm</quantity>
, width:
<quantity id="FD27C0EDFFAFFFEBFAD2FDD4FF11071A" metricMagnitude="-3" metricUnit="m" metricValue="6.199999999999999" metricValueMax="7.1" metricValueMin="5.3" pageId="13" pageNumber="54" unit="mm" value="6.199999999999999" valueMax="7.1" valueMin="5.3">5.37.1 mm</quantity>
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFAFFFEBFF5DFD9CFEA60650" pageId="13" pageNumber="54" type="diagnosis">
<paragraph id="3A606D08FFAFFFEBFF5DFD9CFEA60650" blockId="13.[151,1437,151,789]" pageId="13" pageNumber="54">
<emphasis id="08ABB11AFFAFFFEBFF5DFD9CFE1F07C0" bold="true" box="[199,461,620,645]" pageId="13" pageNumber="54">Differential diagnosis.</emphasis>
This species differs from all
<taxonomicName id="FDDF168BFFAFFFEBFC8EFD9CFC4B07C1" authorityName="Westwood" authorityYear="1842" box="[788,921,620,644]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="08ABB11AFFAFFFEBFC8EFD9CFC4B07C1" box="[788,921,620,644]" italics="true" pageId="13" pageNumber="54">Macroderes</emphasis>
</taxonomicName>
species except
<taxonomicName id="FDDF168BFFAFFFEBFBDDFD9DFB1C07C1" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[1095,1230,621,644]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="08ABB11AFFAFFFEBFBDDFD9DFB1C07C1" box="[1095,1230,621,644]" italics="true" pageId="13" pageNumber="54">M. cornutus</emphasis>
</taxonomicName>
in that the base of the pronotum bears very deep triangular concavity (
<figureCitation id="A2E4718DFFAFFFEBFD40FD7FFCFB07ED" box="[730,809,655,680]" captionStart="FIGURES 710" captionStartId="5.[151,264,1812,1835]" captionTargetBox="[330,1235,720,1729]" captionTargetId="figure@5.[319,1279,700,1780]" captionTargetPageId="5" captionText="FIGURES 710. Habitus of Macroderes species. 7, M. fornicatus Sharp, 1880; 8, M. foveatus Frolov &amp; Scholtz, 2005; 9, M. bias (Olivier, 1789); 10, M. greeni (Kirby, 1818)." httpUri="https://zenodo.org/record/2606170/files/figure.png" pageId="13" pageNumber="54">Figs. 8</figureCitation>
,
<figureCitation id="A2E4718DFFAFFFEBFCACFD60FC8607ED" box="[822,852,656,680]" captionStart="FIGURES 4045" captionStartId="12.[151,264,1736,1759]" captionTargetBox="[306,1266,609,1690]" captionTargetId="figure@12.[306,1266,609,1690]" captionTargetPageId="12" captionText="FIGURES 4045. Pronotum of Macroderes species. 40, M. foveatus Frolov &amp; Scholtz, 2005; 41, M. cornutus Frolov &amp; Scholtz, 2005; 42, M. namakwanus Frolov &amp; Scholtz, 2005; 43, M. politulus Preudhomme de Borre, 1880; 44, M. greeni (Kirby, 1818); 45, M. endroedyi Frolov &amp; Scholtz, 2005." httpUri="https://zenodo.org/record/2606180/files/figure.png" pageId="13" pageNumber="54">40</figureCitation>
) however, it can be separated by the combination of the following characteristics: lack of the rounded tubercle in the frontal suture that characterises
<taxonomicName id="FDDF168BFFAFFFEBFB57FD45FA860789" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[1229,1364,693,716]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="08ABB11AFFAFFFEBFB57FD45FA860789" box="[1229,1364,693,716]" italics="true" pageId="13" pageNumber="54">M. cornutus</emphasis>
</taxonomicName>
(
<figureCitation id="A2E4718DFFAFFFEBFAFEFD44FF0D07B5" captionStart="FIGURES 4651" captionStartId="13.[151,264,1948,1971]" captionTargetBox="[311,1265,844,1918]" captionTargetId="figure@13.[311,1265,844,1918]" captionTargetPageId="13" captionText="FIGURES 4651. Lateral side of pronotum (46, 47) and dorsal and lateral side of the head (4851) of Macroderes species. 46, 49, 51, M. foveatus Frolov &amp; Scholtz, 2005; 47, 48, 50, M. cornutus Frolov &amp; Scholtz, 2005." httpUri="https://zenodo.org/record/2606184/files/figure.png" pageId="13" pageNumber="54">Figs. 49, 51</figureCitation>
), the internal sac sclerite bears small lateral processes (
<figureCitation id="A2E4718DFFAFFFEBFCD4FD27FC7307B5" box="[846,929,727,752]" captionStart="FIGURES 2439" captionStartId="10.[151,264,1343,1366]" captionTargetBox="[323,1250,205,1286]" captionTargetId="figure@10.[323,1283,195,1303]" captionTargetPageId="10" captionText="FIGURES 2439. Internal sac of aedeagus (24), aedeagus (25) and internal sac sclerite (26-39) of Macroderes species. 26, M. endroedyi Frolov &amp; Scholtz, 2005; 27, M. amplior Frolov &amp; Scholtz, 2005; 28, M. bias (Olivier, 1789); 29, M. cederbergensis Abdalla &amp; Deschodt, new species; 30, M. cornutus Frolov &amp; Scholtz, 2005; 31, M. foveatus Frolov &amp; Scholtz, 2005; 32, M. gifboomi Abdalla &amp; Scholtz, new species; 33, M. leipoldti Abdalla &amp; Deschodt, new species; 34, M. mutilans Kolbe, 1908; 35, M. namakwanus Frolov &amp; Scholtz, 2005; 36, M. oreatus Abdalla &amp; Deschodt, new species; 37, M. porselinus Abdalla, new species; 38, M. soleiana Abdalla &amp; Deschodt, new species; 39, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606178/files/figure.png" pageId="13" pageNumber="54">Fig. 31</figureCitation>
), less elongated punctures at each side of the pronotum (
<figureCitation id="A2E4718DFFAFFFEBFE89FD0CFEB50650" box="[275,359,764,789]" captionStart="FIGURES 4651" captionStartId="13.[151,264,1948,1971]" captionTargetBox="[311,1265,844,1918]" captionTargetId="figure@13.[311,1265,844,1918]" captionTargetPageId="13" captionText="FIGURES 4651. Lateral side of pronotum (46, 47) and dorsal and lateral side of the head (4851) of Macroderes species. 46, 49, 51, M. foveatus Frolov &amp; Scholtz, 2005; 47, 48, 50, M. cornutus Frolov &amp; Scholtz, 2005." httpUri="https://zenodo.org/record/2606184/files/figure.png" pageId="13" pageNumber="54">Fig. 46</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="6EA03D80FFAFFFEBFF0DF86CFB960293" httpUri="https://zenodo.org/record/2606184/files/figure.png" pageId="13" pageNumber="54" startId="13.[151,264,1948,1971]" targetBox="[311,1265,844,1918]" targetPageId="13">
<paragraph id="3A606D08FFAFFFEBFF0DF86CFB960293" blockId="13.[151,1436,1948,2006]" pageId="13" pageNumber="54">
<emphasis id="08ABB11AFFAFFFEBFF0DF86CFE9F02F7" bold="true" box="[151,333,1948,1971]" pageId="13" pageNumber="54">FIGURES 4651</emphasis>
. Lateral side of pronotum (46, 47) and dorsal and lateral side of the head (4851) of
<taxonomicName id="FDDF168BFFAFFFEBFB32F86DFAF302F6" authorityName="Westwood" authorityYear="1842" box="[1192,1313,1949,1971]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="08ABB11AFFAFFFEBFB32F86DFAF302F6" box="[1192,1313,1949,1971]" italics="true" pageId="13" pageNumber="54">Macroderes</emphasis>
</taxonomicName>
species. 46, 49, 51,
<taxonomicName id="FDDF168BFFAFFFEBFF79F831FD830293" authority="Frolov &amp; Scholtz, 2005" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[227,593,1984,2006]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="foveatus">
<emphasis id="08ABB11AFFAFFFEBFF79F831FE8A0293" box="[227,344,1984,2006]" italics="true" pageId="13" pageNumber="54">M. foveatus</emphasis>
Frolov &amp; Scholtz, 2005
</taxonomicName>
; 47, 48, 50,
<taxonomicName id="FDDF168BFFAFFFEBFD57F831FB930293" authority="Frolov &amp; Scholtz, 2005" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[717,1089,1984,2006]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="08ABB11AFFAFFFEBFD57F831FC9A0293" box="[717,840,1985,2006]" italics="true" pageId="13" pageNumber="54">M. cornutus</emphasis>
Frolov &amp; Scholtz, 2005
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="72C53E83FFACFFE8FF5DFF67FB3905BD" pageId="14" pageNumber="55" type="description">
<paragraph id="3A606D08FFACFFE8FF5DFF67FB3905BD" blockId="14.[151,1436,151,248]" pageId="14" pageNumber="55">
<emphasis id="08ABB11AFFACFFE8FF5DFF67FE2005F5" bold="true" box="[199,498,151,176]" pageId="14" pageNumber="55">Habitat and distribution.</emphasis>
This species is recorded from two localities in the southern western part of
<collectingRegion id="F81BA3EAFFACFFE8FADBFF68FEEE0590" country="South Africa" name="Western Cape" pageId="14" pageNumber="55">Western Cape Province</collectingRegion>
along the west coast of
<collectingCountry id="42C82D98FFACFFE8FDD6FF4CFD0E0590" box="[588,732,188,213]" name="South Africa" pageId="14" pageNumber="55">South Africa</collectingCountry>
(
<figureCitation id="A2E4718DFFACFFE8FD76FF4CFCE30591" box="[748,817,188,213]" captionStart="FIGURE 2" captionStartId="2.[151,250,1574,1596]" captionText="FIGURE 2. Distribution map of Macroderes species." pageId="14" pageNumber="55">Fig. 2</figureCitation>
). Generally, the area is characterised by Mediterranean climate with warm, dry summers and moderate humid, rainy winters (
<bibRefCitation id="5E4E10F9FFACFFE8FC3FFF2FFB0D05BD" author="Mucina, L. &amp; Rutherford, M. C." box="[933,1247,223,248]" pageId="14" pageNumber="55" pagination="1 - 807" refId="ref21930" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The Vegetation of South Africa, Lesotho and Swaziland. Strelitzia, 19, 1 - 807." type="journal article" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>