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<document id="B8201ABBBFD87B4E289DC77CB110E69D" ID-CLB-Dataset="28091" ID-DOI="10.11646/zootaxa.4504.1.3" ID-GBIF-Dataset="0b803003-244e-4511-aaf4-5e9c3f863861" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606164" ID-ZooBank="52BA28B6-4280-4BD0-8B38-C794F04E4A66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553577945224" checkinUser="plazi" docAuthor="Abdalla, Ishtiag H., Deschodt, Christian M., Scholtz, Clarke H. &amp; Sole, Catherine L." docDate="2018" docId="B276DC1EFFA0FFE5FF0DF922FAF306E0" docLanguage="en" docName="zootaxa.4504.1.3.pdf" docOrigin="Zootaxa 4504 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macroderes amplior Frolov &amp; Scholtz 2005" docType="treatment" docVersion="8" lastPageNumber="44" masterDocId="4E4FA466FFA2FFE6FF9AFFF0FFD20545" masterDocTitle="An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa" masterLastPageNumber="75" masterPageNumber="41" pageNumber="43" updateTime="1698712718888" updateUser="ExternalLinkService">
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<mods:title id="C1A931B35E35BF3394F352BA07EDA985">An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa</mods:title>
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<mods:namePart id="FDEF2F77638F6393268815C6C05C5D4F">Abdalla, Ishtiag H.</mods:namePart>
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<mods:namePart id="973BD8EA978B2DA354B3D936D0C09F1F">Deschodt, Christian M.</mods:namePart>
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<mods:namePart id="6D53DBCCFAA22C90959A010B7A19F7D6">Scholtz, Clarke H.</mods:namePart>
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<mods:namePart id="5826D0DB53012A56F7364D621A7A3F0D">Sole, Catherine L.</mods:namePart>
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<mods:title id="5AF1C0B2CCC200791B208B7256E548ED">Zootaxa</mods:title>
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<mods:part id="17ED7859D262AC476B542F9B0982E7B9">
<mods:date id="D451003788A63FD6FF3E9776201673F4">2018</mods:date>
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<mods:number id="B46DC1AC3F272DB2DF5CEF933A27E4FD">2018-10-23</mods:number>
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<mods:classification id="FBB2F781E5000B8B9E9FBA06895E2A8A">journal article</mods:classification>
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<treatment id="B276DC1EFFA0FFE5FF0DF922FAF306E0" ID-DOI="http://doi.org/10.5281/zenodo.5996720" ID-GBIF-Taxon="154494822" ID-Zenodo-Dep="5996720" LSID="urn:lsid:plazi:treatment:B276DC1EFFA0FFE5FF0DF922FAF306E0" httpUri="http://treatment.plazi.org/id/B276DC1EFFA0FFE5FF0DF922FAF306E0" lastPageId="3" lastPageNumber="44" pageId="2" pageNumber="43">
<subSubSection id="72C53E83FFA0FFE4FF0DF922FD6303A8" box="[151,689,1746,1773]" pageId="2" pageNumber="43" type="nomenclature">
<paragraph id="3A606D08FFA0FFE4FF0DF922FD6303A8" blockId="2.[151,689,1746,1806]" box="[151,689,1746,1773]" pageId="2" pageNumber="43">
<heading id="6128DA64FFA0FFE4FF0DF922FD6303A8" bold="true" box="[151,689,1746,1773]" fontSize="11" level="1" pageId="2" pageNumber="43" reason="1">
<taxonomicName id="FDDF168BFFA0FFE4FF0DF922FD6303A8" ID-CoL="3X4TC" authority="Frolov &amp; Scholtz, 2005" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[151,689,1746,1773]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="amplior">
<emphasis id="08ABB11AFFA0FFE4FF0DF922FD6303A8" bold="true" box="[151,689,1746,1773]" pageId="2" pageNumber="43">
<emphasis id="08ABB11AFFA0FFE4FF0DF922FE5903A9" bold="true" box="[151,395,1746,1772]" italics="true" pageId="2" pageNumber="43">Macroderes amplior</emphasis>
<bibRefCitation id="5E4E10F9FFA0FFE4FE09F923FD6303A8" author="Frolov, A. V. &amp; Scholtz, C. H." box="[403,689,1747,1773]" pageId="2" pageNumber="43" pagination="373 - 393" refId="ref21694" refString="Frolov, A. V. &amp; Scholtz, C. H. (2005) Revision of the southern African genus Macroderes Westwood (Coleoptera: Scarabaeidae: Scarabaeinae). Annales de la Societe Entomologique de France, 40 (3 - 4), 373 - 393. https: // doi. org / 10.1080 / 00379271.2004.10697429" type="journal article" year="2005">Frolov &amp; Scholtz, 2005</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="72C53E83FFA0FFE4FF0DF905FE98024B" box="[151,330,1781,1806]" pageId="2" pageNumber="43" type="description">
<paragraph id="3A606D08FFA0FFE4FF0DF905FE98024B" blockId="2.[151,689,1746,1806]" box="[151,330,1781,1806]" pageId="2" pageNumber="43">
(
<figureCitation id="A2E4718DFFA0FFE4FF05F905FF3F024B" box="[159,237,1781,1806]" captionStart="FIGURES 3–6" captionStartId="4.[151,264,1319,1342]" captionTargetBox="[333,1214,234,1266]" captionTargetId="figure@4.[323,1283,195,1362]" captionTargetPageId="4" captionText="FIGURES 3–6. Habitus of Macroderes species. 3, M. amplior Frolov &amp; Scholtz, 2005; 4, M. arrowi Janssens, 1939; 5, M. cornutus Frolov &amp; Scholtz, 2005; 6, M. endroedyi Frolov &amp; Scholtz, 2005." httpUri="https://zenodo.org/record/2606168/files/figure.png" pageId="2" pageNumber="43">Figs. 3</figureCitation>
,
<figureCitation id="A2E4718DFFA0FFE4FF60F906FECB024B" box="[250,281,1782,1806]" captionStart="FIGURES 24–39" captionStartId="10.[151,264,1343,1366]" captionTargetBox="[323,1250,205,1286]" captionTargetId="figure@10.[323,1283,195,1303]" captionTargetPageId="10" captionText="FIGURES 24–39. Internal sac of aedeagus (24), aedeagus (25) and internal sac sclerite (26-39) of Macroderes species. 26, M. endroedyi Frolov &amp; Scholtz, 2005; 27, M. amplior Frolov &amp; Scholtz, 2005; 28, M. bias (Olivier, 1789); 29, M. cederbergensis Abdalla &amp; Deschodt, new species; 30, M. cornutus Frolov &amp; Scholtz, 2005; 31, M. foveatus Frolov &amp; Scholtz, 2005; 32, M. gifboomi Abdalla &amp; Scholtz, new species; 33, M. leipoldti Abdalla &amp; Deschodt, new species; 34, M. mutilans Kolbe, 1908; 35, M. namakwanus Frolov &amp; Scholtz, 2005; 36, M. oreatus Abdalla &amp; Deschodt, new species; 37, M. porselinus Abdalla, new species; 38, M. soleiana Abdalla &amp; Deschodt, new species; 39, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606178/files/figure.png" pageId="2" pageNumber="43">27</figureCitation>
,
<figureCitation id="A2E4718DFFA0FFE4FEBFF905FE90024B" box="[293,322,1781,1806]" captionStart="FIGURES 64–69" captionStartId="17.[151,264,1917,1940]" captionTargetBox="[158,1427,474,1901]" captionTargetId="figure@17.[158,1427,474,1901]" captionTargetPageId="17" captionText="FIGURES 64–69. Punctuation of the base of pronotum (64, 65) and sculpture of second elytra intervals (66–69) of Macroderes species. 64, M. amplior Frolov &amp; Scholtz, 2005; 65, 67, M. fornicatus Sharp, 1880; 66, M. bias (Olivier, 1789); 68, M. undulatus Preudhomme de Borre, 1880; 69, M. nitidus Harold, 1877." httpUri="https://zenodo.org/record/2606190/files/figure.png" pageId="2" pageNumber="43">64</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFA0FFE4FF0DF8CDFD470216" box="[151,661,1853,1875]" pageId="2" pageNumber="43" type="reference_group">
<paragraph id="3A606D08FFA0FFE4FF0DF8CDFD470216" blockId="2.[151,661,1853,1875]" box="[151,661,1853,1875]" pageId="2" pageNumber="43">
<treatmentCitationGroup id="1ACF4A26FFA0FFE4FF0DF8CDFD470216" box="[151,661,1853,1875]" pageId="2" pageNumber="43">
<taxonomicName id="FDDF168BFFA0FFE4FF0DF8CDFD430216" ID-CoL="3X4TC" authority="Frolov &amp; Scholtz, 2005: 383" authorityName="Frolov &amp; Scholtz" authorityPageNumber="383" authorityYear="2005" box="[151,657,1853,1875]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="amplior">
<emphasis id="08ABB11AFFA0FFE4FF0DF8CDFEB70216" box="[151,357,1853,1875]" italics="true" pageId="2" pageNumber="43">Macroderes amplior</emphasis>
<treatmentCitation id="BB7E4B19FFA0FFE4FEF6F8CDFD430216" author="Frolov, A. V. &amp; Scholtz, C. H." box="[364,657,1853,1875]" page="383" pageId="2" pageNumber="43" year="2005">
<bibRefCitation id="5E4E10F9FFA0FFE4FEF6F8CDFD8D0216" author="Frolov, A. V. &amp; Scholtz, C. H." box="[364,607,1853,1875]" pageId="2" pageNumber="43" pagination="373 - 393" refId="ref21694" refString="Frolov, A. V. &amp; Scholtz, C. H. (2005) Revision of the southern African genus Macroderes Westwood (Coleoptera: Scarabaeidae: Scarabaeinae). Annales de la Societe Entomologique de France, 40 (3 - 4), 373 - 393. https: // doi. org / 10.1080 / 00379271.2004.10697429" type="journal article" year="2005">Frolov &amp; Scholtz, 2005</bibRefCitation>
: 383
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
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<subSubSection id="72C53E83FFA0FFE5FF0DF88FFF180789" lastPageId="3" lastPageNumber="44" pageId="2" pageNumber="43" type="materials_examined">
<paragraph id="3A606D08FFA0FFE4FF0DF88FFB1302DD" blockId="2.[151,1436,1919,2016]" box="[151,1217,1919,1944]" pageId="2" pageNumber="43">
<materialsCitation id="8AB76755FFA0FFE4FF0DF88FFB6F02DD" ID-GBIF-Occurrence="2028137728" box="[151,1213,1919,1944]" country="South Africa" latitude="-30.976828" location="Km" longLatPrecision="1" longitude="17.998802" municipality="Prov" pageId="2" pageNumber="43" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
<emphasis id="08ABB11AFFA0FFE4FF0DF88FFEE702DD" bold="true" box="[151,309,1919,1944]" pageId="2" pageNumber="43">
<typeStatus id="E564D3AAFFA0FFE4FF0DF88FFF0002DD" box="[151,210,1919,1944]" pageId="2" pageNumber="43">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="42C82D98FFA0FFE4FEA6F870FEA702DD" box="[316,373,1920,1944]" name="South Africa" pageId="2" pageNumber="43">RSA</collectingCountry>
,
<collectingRegion id="F81BA3EAFFA0FFE4FE1AF870FDCE02DD" box="[384,540,1920,1944]" country="South Africa" name="Western Cape" pageId="2" pageNumber="43">Western Cape</collectingRegion>
<collectingMunicipality id="DA04F772FFA0FFE4FDB9F870FD8B02D2" box="[547,601,1920,1943]" pageId="2" pageNumber="43">Prov</collectingMunicipality>
, 5
<location id="3F003BD3FFA0FFE4FDE3F870FD7102D2" LSID="urn:lsid:plazi:treatment:B276DC1EFFA0FFE5FF0DF922FAF306E0:3F003BD3FFA0FFE4FDE3F870FD7102D2" box="[633,675,1920,1943]" country="South Africa" latitude="-30.976828" longLatPrecision="1" longitude="17.998802" municipality="Prov" name="Km" pageId="2" pageNumber="43" stateProvince="Western Cape">Km</location>
<locationDeviation id="2BD2742AFFA0FFE4FD30F870FC8502DD" box="[682,855,1919,1944]" location="Rietpoort" pageId="2" pageNumber="43">W of Rietpoort</locationDeviation>
,
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<geoCoordinate id="5FEB0BCFFFA0FFE4FB93F88FFB6F02DD" box="[1033,1213,1919,1944]" direction="east" orientation="longitude" pageId="2" pageNumber="43" precision="1" value="17.998802">017°59'55.69''E</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="3A606D08FFA0FFE5FF5DF854FED10778" blockId="2.[151,1436,1919,2016]" lastBlockId="3.[151,1437,151,933]" lastPageId="3" lastPageNumber="44" pageId="2" pageNumber="43">
<emphasis id="08ABB11AFFA0FFE4FF5DF854FE2602F8" bold="true" box="[199,500,1956,1981]" pageId="2" pageNumber="43">Type material examined.</emphasis>
<materialsCitation id="8AB76755FFA0FFE5FE64F854FBB205F5" ID-GBIF-Occurrence="2028137707" collectingDate="2003-09-03" collectingDateMax="2003-09-11" collectingDateMin="2003-09-03" collectionCode="TMSA" collectorName="A. Frolov &amp; C. Deschodt" country="South Africa" lastPageId="3" lastPageNumber="44" latitude="-30.976828" location="Rietpoort" longLatPrecision="1" longitude="17.998802" municipality="Prov" pageId="2" pageNumber="43" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="holotype">
<typeStatus id="E564D3AAFFA0FFE4FE64F854FDBB02F8" box="[510,617,1956,1981]" pageId="2" pageNumber="43" type="holotype">Holotype</typeStatus>
,
<specimenCount id="2CD9A681FFA0FFE4FDEDF855FD5E02F9" box="[631,652,1957,1980]" pageId="2" pageNumber="43" type="male">♂</specimenCount>
(
<collectionCode id="5CCEF5CDFFA0FFE4FD04F854FD2102F9" box="[670,755,1956,1980]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="2" pageNumber="43">TMSA</collectionCode>
): “
<collectingCountry id="42C82D98FFA0FFE4FC88F854FC9902F9" box="[786,843,1956,1980]" name="South Africa" pageId="2" pageNumber="43">RSA</collectingCountry>
/
<collectingRegion id="F81BA3EAFFA0FFE4FCFCF855FBD702F9" box="[870,1029,1956,1980]" country="South Africa" name="Western Cape" pageId="2" pageNumber="43">Western Cape</collectingRegion>
<collectingMunicipality id="DA04F772FFA0FFE4FB95F855FB9602F9" box="[1039,1092,1957,1980]" pageId="2" pageNumber="43">Prov</collectingMunicipality>
/
<quantity id="FD27C0EDFFA0FFE4FBC5F854FB4902F8" box="[1119,1179,1956,1981]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="43" unit="km" value="5.0">5 km</quantity>
W of /
<location id="3F003BD3FFA0FFE4FB66F854FAB402F8" LSID="urn:lsid:plazi:treatment:B276DC1EFFA0FFE5FF0DF922FAF306E0:3F003BD3FFA0FFE4FB66F854FAB402F8" box="[1276,1382,1956,1981]" country="South Africa" latitude="-30.976828" longLatPrecision="1" longitude="17.998802" municipality="Prov" name="Rietpoort" pageId="2" pageNumber="43" stateProvince="Western Cape">Rietpoort</location>
/
<collectorName id="972A08DEFFA0FFE4FA1BF854FF3002A5" pageId="2" pageNumber="43">A. Frolov</collectorName>
&amp;
<collectorName id="972A08DEFFA0FFE4FE9CF838FE4302A5" box="[262,401,1991,2016]" pageId="2" pageNumber="43">C. Deschodt</collectorName>
leg [printed] // AF–0035(15) /
<date id="4E614BC8FFA0FFE4FD6AF838FC4B02A5" box="[752,921,1992,2016]" pageId="2" pageNumber="43" value="2003-09-03" valueMax="2003-09-11" valueMin="2003-09-03">
<collectingDate id="5E25B220FFA0FFE4FD6AF838FC4B02A5" box="[752,921,1992,2016]" pageId="2" pageNumber="43" value="2003-09-03" valueMax="2003-09-11" valueMin="2003-09-03">3–11. IX. 2003</collectingDate>
</date>
/
<geoCoordinate id="5FEB0BCFFFA0FFE4FC35F838FBB402A5" box="[943,1126,1991,2016]" direction="south" orientation="latitude" pageId="2" pageNumber="43" precision="1" value="-30.976828">S 30° 58' 36.58''</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFA0FFE4FBF7F838FAE002A5" box="[1133,1330,1991,2016]" direction="east" orientation="longitude" pageId="2" pageNumber="43" precision="1" value="17.998802">E 017° 59' 55.69''</geoCoordinate>
[printed] //
<typeStatus id="E564D3AAFFA1FFE5FF34FF68FE8605F5" box="[174,340,152,176]" pageId="3" pageNumber="44" type="holotype">HOLOTYPUS</typeStatus>
/
<taxonomicName id="FDDF168BFFA1FFE5FEF0FF68FD8205F5" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[362,592,152,176]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="amplior">
<emphasis id="08ABB11AFFA1FFE5FEF0FF68FD8205F5" box="[362,592,152,176]" italics="true" pageId="3" pageNumber="44">Macroderes amplior</emphasis>
</taxonomicName>
/ Frolov &amp; Scholtz / 2003 [red label, printed]”
</materialsCitation>
.
<materialsCitation id="8AB76755FFA1FFE5FBF1FF68FE460590" ID-GBIF-Occurrence="2028137708" collectingDate="2003-09-03" collectingDateMax="2003-09-11" collectingDateMin="2003-09-03" collectionCode="TMSA" collectorName="A. Frolov &amp; C. Deschodt" country="South Africa" latitude="-30.976828" location="Rietpoort" longLatPrecision="1" longitude="17.998802" municipality="Prov" pageId="3" pageNumber="44" specimenCount="11" specimenCount-female="10" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
<typeStatus id="E564D3AAFFA1FFE5FBF1FF68FB0E05F5" box="[1131,1244,152,176]" pageId="3" pageNumber="44" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="2CD9A681FFA1FFE5FB7CFF68FAC305EA" box="[1254,1297,152,176]" pageId="3" pageNumber="44" type="male">1 ♂</specimenCount>
and
<specimenCount id="2CD9A681FFA1FFE5FAD2FF68FA4405EA" box="[1352,1430,152,176]" pageId="3" pageNumber="44" type="female">10 ♀♀</specimenCount>
, same data as holotype
</materialsCitation>
,
<specimenCount id="2CD9A681FFA1FFE5FE3AFF4CFE1E0591" box="[416,460,188,212]" pageId="3" pageNumber="44" type="male">1 ♂</specimenCount>
and
<specimenCount id="2CD9A681FFA1FFE5FD9FFF4DFD970591" box="[517,581,189,212]" pageId="3" pageNumber="44" type="female">7 ♀♀</specimenCount>
in UPSA,
<specimenCount id="2CD9A681FFA1FFE5FD58FF4CFCD10591" box="[706,771,188,212]" pageId="3" pageNumber="44" type="female">3 ♀♀</specimenCount>
in TMSA;
<materialsCitation id="8AB76755FFA1FFE5FC1DFF4CFED20405" ID-GBIF-Occurrence="2028137709" collectingDate="2003-09-03" collectingDateMax="2003-09-11" collectingDateMin="2003-09-03" collectionCode="TMSA" collectorName="C. Deschodt" country="South Africa" latitude="-30.980137" location="Rietpoort" longLatPrecision="1" longitude="17.993439" municipality="Prov" pageId="3" pageNumber="44" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="2CD9A681FFA1FFE5FC1DFF4CFC600591" box="[903,946,188,212]" pageId="3" pageNumber="44" type="male">1 ♂</specimenCount>
and
<specimenCount id="2CD9A681FFA1FFE5FC71FF4CFBF90591" box="[1003,1067,188,212]" pageId="3" pageNumber="44" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFA1FFE5FBA1FF4CFB420591" box="[1083,1168,188,212]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="3" pageNumber="44">TMSA</collectionCode>
): “
<collectingCountry id="42C82D98FFA1FFE5FB34FF4CFB350591" box="[1198,1255,188,212]" name="South Africa" pageId="3" pageNumber="44">RSA</collectingCountry>
/
<collectingRegion id="F81BA3EAFFA1FFE5FB65FF4DFA4E0591" box="[1279,1436,188,212]" country="South Africa" name="Western Cape" pageId="3" pageNumber="44">Western Cape</collectingRegion>
<collectingMunicipality id="DA04F772FFA1FFE5FF0DFF10FF1E05B2" box="[151,204,224,247]" pageId="3" pageNumber="44">Prov</collectingMunicipality>
/
<quantity id="FD27C0EDFFA1FFE5FF79FF2FFECE05BD" box="[227,284,223,248]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="44" unit="km" value="5.0">5 km</quantity>
W of /
<location id="3F003BD3FFA1FFE5FEE9FF2FFE0F05BD" LSID="urn:lsid:plazi:treatment:B276DC1EFFA0FFE5FF0DF922FAF306E0:3F003BD3FFA1FFE5FEE9FF2FFE0F05BD" box="[371,477,223,248]" country="South Africa" latitude="-30.980137" longLatPrecision="1" longitude="17.993439" municipality="Prov" name="Rietpoort" pageId="3" pageNumber="44" stateProvince="Western Cape">Rietpoort</location>
/ A. Frolov &amp;
<collectorName id="972A08DEFFA1FFE5FD1CFF10FCC305BD" box="[646,785,223,248]" pageId="3" pageNumber="44">C. Deschodt</collectorName>
leg [printed] // AF-0036 (16) /
<date id="4E614BC8FFA1FFE5FBEFFF10FACD05BD" box="[1141,1311,224,248]" pageId="3" pageNumber="44" value="2003-09-03" valueMax="2003-09-11" valueMin="2003-09-03">
<collectingDate id="5E25B220FFA1FFE5FBEFFF10FACD05BD" box="[1141,1311,224,248]" pageId="3" pageNumber="44" value="2003-09-03" valueMax="2003-09-11" valueMin="2003-09-03">3–11. IX. 2003</collectingDate>
</date>
/
<geoCoordinate id="5FEB0BCFFFA1FFE5FAAFFF10FF320459" direction="south" orientation="latitude" pageId="3" pageNumber="44" precision="1" value="-30.980137">S 30° 58' 48.49''</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFA1FFE5FF70FEF5FE6A0458" box="[234,440,260,285]" direction="east" orientation="longitude" pageId="3" pageNumber="44" precision="1" value="17.993439">E 017° 59' 36.38''</geoCoordinate>
[printed] //
<typeStatus id="E564D3AAFFA1FFE5FDDCFEF4FD310459" box="[582,739,260,284]" pageId="3" pageNumber="44" type="paratype">PARATYPUS</typeStatus>
/
<taxonomicName id="FDDF168BFFA1FFE5FD65FEF4FC350459" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[767,999,260,284]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="amplior">
<emphasis id="08ABB11AFFA1FFE5FD65FEF4FC350459" box="[767,999,260,284]" italics="true" pageId="3" pageNumber="44">Macroderes amplior</emphasis>
</taxonomicName>
/ Frolov &amp; Scholtz / 2003 [red label, printed]”
</materialsCitation>
;
<materialsCitation id="8AB76755FFA1FFE5FE91FED8FD9204CD" ID-GBIF-Occurrence="2028137727" collectionCode="TMSA" country="South Africa" county="Afr" location="Rietpoort" municipality="Namaquland" pageId="3" pageNumber="44" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="2CD9A681FFA1FFE5FE91FED8FEE7047A" box="[267,309,296,320]" pageId="3" pageNumber="44" type="male">1 ♂</specimenCount>
(
<collectionCode id="5CCEF5CDFFA1FFE5FEDEFED8FE4B0405" box="[324,409,296,320]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="3" pageNumber="44">TMSA</collectionCode>
): “S.
<collectingCounty id="D3011584FFA1FFE5FE4FFED7FE2E0405" box="[469,508,295,320]" pageId="3" pageNumber="44">Afr</collectingCounty>
/
<collectingMunicipality id="DA04F772FFA1FFE5FD8BFED7FD720405" box="[529,672,295,320]" pageId="3" pageNumber="44">Namaquland</collectingMunicipality>
/
<location id="3F003BD3FFA1FFE5FD2CFED7FCF20405" LSID="urn:lsid:plazi:treatment:B276DC1EFFA0FFE5FF0DF922FAF306E0:3F003BD3FFA1FFE5FD2CFED7FCF20405" box="[694,800,295,320]" country="South Africa" county="Afr" municipality="Namaquland" name="Rietpoort" pageId="3" pageNumber="44" stateProvince="Western Cape">Rietpoort</location>
farm / 30°. 59' S–18°. 06' E / [printed] //
<typeStatus id="E564D3AAFFA1FFE5FB6AFED8FA5C0405" box="[1264,1422,296,320]" pageId="3" pageNumber="44" type="paratype">PARATYPUS</typeStatus>
/
<taxonomicName id="FDDF168BFFA1FFE5FF0DFEBCFEAF0421" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[151,381,332,356]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="amplior">
<emphasis id="08ABB11AFFA1FFE5FF0DFEBCFEAF0421" box="[151,381,332,356]" italics="true" pageId="3" pageNumber="44">Macroderes amplior</emphasis>
</taxonomicName>
/ Frolov &amp; Scholtz / 2003 [red label, printed] //
<typeStatus id="E564D3AAFFA1FFE5FC05FEBCFBEF0421" box="[927,1085,332,356]" pageId="3" pageNumber="44" type="paratype">PARATYPUS</typeStatus>
/
<taxonomicName id="FDDF168BFFA1FFE5FBCEFEBCFAE80421" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[1108,1338,332,356]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="amplior">
<emphasis id="08ABB11AFFA1FFE5FBCEFEBCFAE80421" box="[1108,1338,332,356]" italics="true" pageId="3" pageNumber="44">Macroderes amplior</emphasis>
</taxonomicName>
/ Frolov &amp; Scholtz / 2003 [red label, printed]”
</materialsCitation>
;
<materialsCitation id="8AB76755FFA1FFE5FDD1FE80FB900495" ID-GBIF-Occurrence="2028137729" collectingDate="1981-09-09" collectionCode="SANC" country="South Africa" county="Red" location="Karro" municipality="Gravel Mountain" pageId="3" pageNumber="44" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="2CD9A681FFA1FFE5FDD1FE80FDA404C2" box="[587,630,368,392]" pageId="3" pageNumber="44" type="female">1 ♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFA1FFE5FD1FFE80FD0404CD" box="[645,726,368,392]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="3" pageNumber="44" type="Museum">SANC</collectionCode>
): “KOMAGGAS / NW / CP / (
<quantity id="FD27C0EDFFA1FFE5FBAFFE80FBBD04CD" box="[1077,1135,367,392]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="44" unit="km" value="7.0">7 km</quantity>
N) /
<date id="4E614BC8FFA1FFE5FB31FE80FADD04CD" box="[1195,1295,368,392]" pageId="3" pageNumber="44" value="1981-09-09">
<collectingDate id="5E25B220FFA1FFE5FB31FE80FADD04CD" box="[1195,1295,368,392]" pageId="3" pageNumber="44" value="1981-09-09">9. IX. 81</collectingDate>
</date>
/
<collectingCounty id="D3011584FFA1FFE5FABFFE9FFA8004CD" box="[1317,1362,367,392]" pageId="3" pageNumber="44">Red</collectingCounty>
sand /
<collectingMunicipality id="DA04F772FFA1FFE5FF0DFE64FE8804E8" box="[151,346,404,429]" pageId="3" pageNumber="44">Gravel Mountain</collectingMunicipality>
/
<location id="3F003BD3FFA1FFE5FEECFE65FE6504E9" LSID="urn:lsid:plazi:treatment:B276DC1EFFA0FFE5FF0DF922FAF306E0:3F003BD3FFA1FFE5FEECFE65FE6504E9" box="[374,439,405,428]" country="South Africa" county="Red" municipality="Gravel Mountain" name="Karro" pageId="3" pageNumber="44" stateProvince="Western Cape">Karro</location>
/
<location id="3F003BD3FFA1FFE5FE4EFE64FDFF04E8" LSID="urn:lsid:plazi:treatment:B276DC1EFFA0FFE5FF0DF922FAF306E0:3F003BD3FFA1FFE5FE4EFE64FDFF04E8" box="[468,557,404,429]" country="South Africa" county="Red" municipality="Gravel Mountain" name="Donkey" pageId="3" pageNumber="44" stateProvince="Western Cape">Donkey</location>
&amp; ALV.
<location id="3F003BD3FFA1FFE5FD01FE64FD0E04E8" LSID="urn:lsid:plazi:treatment:B276DC1EFFA0FFE5FF0DF922FAF306E0:3F003BD3FFA1FFE5FD01FE64FD0E04E8" box="[667,732,404,429]" country="South Africa" county="Red" municipality="Gravel Mountain" name="Davis" pageId="3" pageNumber="44" stateProvince="Western Cape">Davis</location>
[printed] //
<emphasis id="08ABB11AFFA1FFE5FCF6FE64FC2304E9" box="[876,1009,404,428]" italics="true" pageId="3" pageNumber="44">
<location id="3F003BD3FFA1FFE5FCF6FE64FC2304E9" LSID="urn:lsid:plazi:treatment:B276DC1EFFA0FFE5FF0DF922FAF306E0:3F003BD3FFA1FFE5FCF6FE64FC2304E9" box="[876,1009,404,428]" country="South Africa" county="Red" municipality="Gravel Mountain" name="Macorderes" pageId="3" pageNumber="44" stateProvince="Western Cape">Macorderes</location>
</emphasis>
sp / det. CSIRO. DBRU [printed] //
<typeStatus id="E564D3AAFFA1FFE5FF0DFE48FEE70495" box="[151,309,440,464]" pageId="3" pageNumber="44" type="paratype">PARATYPUS</typeStatus>
/
<taxonomicName id="FDDF168BFFA1FFE5FED1FE48FDE20495" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[331,560,440,464]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="amplior">
<emphasis id="08ABB11AFFA1FFE5FED1FE48FDE20495" box="[331,560,440,464]" italics="true" pageId="3" pageNumber="44">Macroderes amplior</emphasis>
</taxonomicName>
/ Frolov &amp; Scholtz / 2003 [red label, printed]”
</materialsCitation>
;
<materialsCitation id="8AB76755FFA1FFE5FBD7FE48FED20778" ID-GBIF-Occurrence="2028137717" collectingDate="2003-11-03" collectionCode="SAMC" collectorName="C. Deschodt" country="South Africa" county="Prov" latitude="-30.980137" location="Rietpoort" longLatPrecision="1" longitude="17.993439" municipality="Km" pageId="3" pageNumber="44" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype">
<specimenCount id="2CD9A681FFA1FFE5FBD7FE48FBA5048A" box="[1101,1143,440,464]" pageId="3" pageNumber="44" type="male">1 ♂</specimenCount>
and
<specimenCount id="2CD9A681FFA1FFE5FB35FE48FB030495" box="[1199,1233,440,464]" pageId="3" pageNumber="44" type="female">1♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFA1FFE5FB7BFE48FAE50495" box="[1249,1335,440,464]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="3" pageNumber="44" type="Museum">SAMC</collectionCode>
): “
<collectingCountry id="42C82D98FFA1FFE5FACEFE48FA5F0495" box="[1364,1421,440,464]" name="South Africa" pageId="3" pageNumber="44">RSA</collectingCountry>
/
<collectingRegion id="F81BA3EAFFA1FFE5FF0DFE2DFEE004B1" box="[151,306,476,500]" country="South Africa" name="Western Cape" pageId="3" pageNumber="44">Western Cape</collectingRegion>
<collectingCounty id="D3011584FFA1FFE5FEA3FE2DFEBC04B1" box="[313,366,477,500]" pageId="3" pageNumber="44">Prov</collectingCounty>
/ 5
<collectingMunicipality id="DA04F772FFA1FFE5FE14FE2CFE6804B0" box="[398,442,476,501]" pageId="3" pageNumber="44">Km</collectingMunicipality>
W of /
<location id="3F003BD3FFA1FFE5FD94FE2CFDAA04B0" LSID="urn:lsid:plazi:treatment:B276DC1EFFA0FFE5FF0DF922FAF306E0:3F003BD3FFA1FFE5FD94FE2CFDAA04B0" box="[526,632,476,501]" country="South Africa" county="Prov" latitude="-30.980137" longLatPrecision="1" longitude="17.993439" municipality="Km" name="Rietpoort" pageId="3" pageNumber="44" stateProvince="Western Cape">Rietpoort</location>
/ A. Frolov &amp;
<collectorName id="972A08DEFFA1FFE5FC87FE2CFC7504B0" box="[797,935,476,501]" pageId="3" pageNumber="44">C. Deschodt</collectorName>
leg [printed] // AF-0036 (16) /
<date id="4E614BC8FFA1FFE5FA9EFE2CFAAA04B1" box="[1284,1400,476,500]" pageId="3" pageNumber="44" value="2003-11-03">
<collectingDate id="5E25B220FFA1FFE5FA9EFE2CFAAA04B1" box="[1284,1400,476,500]" pageId="3" pageNumber="44" value="2003-11-03">3-11-2003</collectingDate>
</date>
/
<geoCoordinate id="5FEB0BCFFFA1FFE5FA16FE2CFEEB075D" direction="south" orientation="latitude" pageId="3" pageNumber="44" precision="1" value="-30.980137">S 30° 58' 48.49''</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFA1FFE5FEDBFDF0FDD9075D" box="[321,523,511,536]" direction="east" orientation="longitude" pageId="3" pageNumber="44" precision="1" value="17.993439">E 017° 59' 36.38''</geoCoordinate>
[printed] //
<typeStatus id="E564D3AAFFA1FFE5FD0FFDF0FCD1075D" box="[661,771,512,536]" pageId="3" pageNumber="44" type="paratype">Paratypus</typeStatus>
/
<taxonomicName id="FDDF168BFFA1FFE5FC86FDF0FBD1075D" authorityName="Frolov &amp; Scholtz" authorityYear="2005" box="[796,1027,512,536]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="amplior">
<emphasis id="08ABB11AFFA1FFE5FC86FDF0FBD1075D" box="[796,1027,512,536]" italics="true" pageId="3" pageNumber="44">Macroderes amplior</emphasis>
</taxonomicName>
/ Frolov &amp; Scholtz 2003 [red label, printed]”
</materialsCitation>
.
</paragraph>
<paragraph id="3A606D08FFA1FFE5FF5DFDB7FE2007C0" blockId="3.[151,1437,151,933]" pageId="3" pageNumber="44">
<emphasis id="08ABB11AFFA1FFE5FF5DFDB7FDED0725" bold="true" box="[199,575,583,608]" pageId="3" pageNumber="44">Additional material examined.</emphasis>
<materialsCitation id="8AB76755FFA1FFE5FDD1FDB8FE3C07C0" ID-GBIF-Occurrence="2028137716" collectingDate="2004-08-30" collectionCode="UPSA" collectorName="C. Deschodt &amp; M. Deschodt" latitude="-30.97729" location="Rietpoort" longLatPrecision="1" longitude="18.00026" pageId="3" pageNumber="44" specimenCount="1" specimenCount-female="1">
<specimenCount id="2CD9A681FFA1FFE5FDD1FDB8FDA8071A" box="[587,634,584,608]" pageId="3" pageNumber="44" type="female">1 ♀</specimenCount>
(
<collectionCode id="5CCEF5CDFFA1FFE5FD14FDB8FD0E0725" box="[654,732,584,608]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="3" pageNumber="44">UPSA</collectionCode>
): “
<location id="3F003BD3FFA1FFE5FD65FDB7FCB90725" LSID="urn:lsid:plazi:treatment:B276DC1EFFA0FFE5FF0DF922FAF306E0:3F003BD3FFA1FFE5FD65FDB7FCB90725" box="[767,875,583,608]" latitude="-30.97729" longLatPrecision="1" longitude="18.00026" name="Rietpoort" pageId="3" pageNumber="44">Rietpoort</location>
/
<geoCoordinate id="5FEB0BCFFFA1FFE5FC16FDB8FBC90725" box="[908,1051,584,608]" direction="south" orientation="latitude" pageId="3" pageNumber="44" precision="1" value="-30.97729">S 30.97729°</geoCoordinate>
<geoCoordinate id="5FEB0BCFFFA1FFE5FBB2FDB8FB680725" box="[1064,1210,583,608]" direction="east" orientation="longitude" pageId="3" pageNumber="44" precision="1" value="18.00026">E 18.00026°</geoCoordinate>
/
<date id="4E614BC8FFA1FFE5FB40FDB7FAB00725" box="[1242,1378,583,608]" pageId="3" pageNumber="44" value="2004-08-30">
<collectingDate id="5E25B220FFA1FFE5FB40FDB7FAB00725" box="[1242,1378,583,608]" pageId="3" pageNumber="44" value="2004-08-30">30.viii.2004</collectingDate>
</date>
/
<collectorName id="972A08DEFFA1FFE5FA18FDB8FED307C0" pageId="3" pageNumber="44">C. Deschodt</collectorName>
&amp;
<collectorName id="972A08DEFFA1FFE5FEBEFD9DFE6707C0" box="[292,437,620,645]" pageId="3" pageNumber="44">M. Deschodt</collectorName>
leg”
</materialsCitation>
.
</paragraph>
<paragraph id="3A606D08FFA1FFE5FF5DFD7FFF180789" blockId="3.[151,1437,151,933]" pageId="3" pageNumber="44">
<emphasis id="08ABB11AFFA1FFE5FF5DFD7FFE9907ED" bold="true" box="[199,331,655,680]" pageId="3" pageNumber="44">Size range.</emphasis>
Males length:
<quantity id="FD27C0EDFFA1FFE5FE6CFD60FD4907E2" box="[502,667,656,680]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3199999999999998" metricValueMin="1.18" pageId="3" pageNumber="44" unit="mm" value="12.5" valueMax="13.2" valueMin="11.8">11.8–13.2 mm</quantity>
, width: 8.5–9.0 mm; females length:
<quantity id="FD27C0EDFFA1FFE5FBDFFD7FFB3907E2" box="[1093,1259,655,680]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" metricValueMax="1.25" metricValueMin="1.0699999999999998" pageId="3" pageNumber="44" unit="mm" value="11.6" valueMax="12.5" valueMin="10.7">10.7–12.5 mm</quantity>
, width:
<quantity id="FD27C0EDFFA1FFE5FAD2FD7FFF140789" metricMagnitude="-3" metricUnit="m" metricValue="9.1" metricValueMax="9.6" metricValueMin="8.6" pageId="3" pageNumber="44" unit="mm" value="9.1" valueMax="9.6" valueMin="8.6">8.6–9.6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFA1FFE5FF5DFD27FC680619" pageId="3" pageNumber="44" type="diagnosis">
<paragraph id="3A606D08FFA1FFE5FF5DFD27FC680619" blockId="3.[151,1437,151,933]" pageId="3" pageNumber="44">
<emphasis id="08ABB11AFFA1FFE5FF5DFD27FE0307B5" bold="true" box="[199,465,727,752]" pageId="3" pageNumber="44">Differential diagnosis.</emphasis>
Resemble
<taxonomicName id="FDDF168BFFA1FFE5FDCCFD29FD3A07B5" authority="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole, 2018" authorityName="Abdalla &amp; Deschodt &amp; Scholtz &amp; Sole" authorityYear="2018" box="[598,744,729,752]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="tortuosus" status="sp. nov.">
<emphasis id="08ABB11AFFA1FFE5FDCCFD29FD3A07B5" box="[598,744,729,752]" italics="true" pageId="3" pageNumber="44">M. tortuosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13980C61FFA1FFE5FD69FD28FCAC07B5" box="[755,894,727,752]" pageId="3" pageNumber="44" rank="species">new species</taxonomicNameLabel>
in having pronotum with elongated punctures (
<figureCitation id="A2E4718DFFA1FFE5FF05FD0CFF230650" box="[159,241,764,789]" captionStart="FIGURES 64–69" captionStartId="17.[151,264,1917,1940]" captionTargetBox="[158,1427,474,1901]" captionTargetId="figure@17.[158,1427,474,1901]" captionTargetPageId="17" captionText="FIGURES 64–69. Punctuation of the base of pronotum (64, 65) and sculpture of second elytra intervals (66–69) of Macroderes species. 64, M. amplior Frolov &amp; Scholtz, 2005; 65, 67, M. fornicatus Sharp, 1880; 66, M. bias (Olivier, 1789); 68, M. undulatus Preudhomme de Borre, 1880; 69, M. nitidus Harold, 1877." httpUri="https://zenodo.org/record/2606190/files/figure.png" pageId="3" pageNumber="44">Fig. 64</figureCitation>
) and
<taxonomicName id="FDDF168BFFA1FFE5FEAAFD0DFE670651" authorityName="Kolbe" authorityYear="1908" box="[304,437,764,788]" class="Insecta" family="Scarabaeidae" genus="Macroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="mutilans">
<emphasis id="08ABB11AFFA1FFE5FEAAFD0DFE670651" box="[304,437,764,788]" italics="true" pageId="3" pageNumber="44">M. mutilans</emphasis>
</taxonomicName>
in the general body features, however, it can be distinguished from the former by having impunctate lateral borders of the pronotum and from the later by having rather undulate surface of elytra and also by having very long lateral process of internal sac of aedeagus (
<figureCitation id="A2E4718DFFA1FFE5FCFAFCB4FC600619" box="[864,946,836,861]" captionStart="FIGURES 24–39" captionStartId="10.[151,264,1343,1366]" captionTargetBox="[323,1250,205,1286]" captionTargetId="figure@10.[323,1283,195,1303]" captionTargetPageId="10" captionText="FIGURES 24–39. Internal sac of aedeagus (24), aedeagus (25) and internal sac sclerite (26-39) of Macroderes species. 26, M. endroedyi Frolov &amp; Scholtz, 2005; 27, M. amplior Frolov &amp; Scholtz, 2005; 28, M. bias (Olivier, 1789); 29, M. cederbergensis Abdalla &amp; Deschodt, new species; 30, M. cornutus Frolov &amp; Scholtz, 2005; 31, M. foveatus Frolov &amp; Scholtz, 2005; 32, M. gifboomi Abdalla &amp; Scholtz, new species; 33, M. leipoldti Abdalla &amp; Deschodt, new species; 34, M. mutilans Kolbe, 1908; 35, M. namakwanus Frolov &amp; Scholtz, 2005; 36, M. oreatus Abdalla &amp; Deschodt, new species; 37, M. porselinus Abdalla, new species; 38, M. soleiana Abdalla &amp; Deschodt, new species; 39, M. tourtosus Abdalla &amp; Scholtz, new species." httpUri="https://zenodo.org/record/2606178/files/figure.png" pageId="3" pageNumber="44">Fig. 27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="72C53E83FFA1FFE5FF5DFC97FAF306E0" pageId="3" pageNumber="44" type="description">
<paragraph id="3A606D08FFA1FFE5FF5DFC97FAF306E0" blockId="3.[151,1437,151,933]" pageId="3" pageNumber="44">
<emphasis id="08ABB11AFFA1FFE5FF5DFC97FE2506C5" bold="true" box="[199,503,871,896]" pageId="3" pageNumber="44">Habitat and distribution.</emphasis>
Rietpoort and Komaggas are two localities in Namaqualand region (
<figureCitation id="A2E4718DFFA1FFE5FA88FC97FA8B06C5" box="[1298,1369,871,896]" captionStart="FIGURE 2" captionStartId="2.[151,250,1574,1596]" captionText="FIGURE 2. Distribution map of Macroderes species." pageId="3" pageNumber="44">Fig. 2</figureCitation>
). The area is dry and hot; characterised by low annual rainfall and a wide distribution of succulent vegetation.
</paragraph>
</subSubSection>
</treatment>
</document>
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