430 lines
45 KiB
XML
430 lines
45 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.62335" ID-GBIF-Dataset="c99cf6eb-9d09-4e1d-898a-b872c047a07e" ID-PMC="PMC8390884" ID-Pensoft-Pub="1313-2970-1044-339" ID-Pensoft-UUID="D0F25BD8394A5582A19FC1A27947BE26" ID-PubMed="34512089" ID-ZooBank="8264C99932014A3BB39C222790AC0192" ModsDocID="1313-2970-1044-339" checkinTime="1623875413266" checkinUser="pensoft" docAuthor="Liebherr, James K., Porch, Nick, Shaw, Matthew, Sinclair, Bronte E. & Maddison, David R." docDate="2021" docId="472CE67146395350A146D7C06A1AB419" docLanguage="en" docName="ZooKeys 1044: 339-373" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.62335" docTitle="Theprisa convexa" docType="treatment" docVersion="4" id="D0F25BD8394A5582A19FC1A27947BE26" lastPageNumber="339" masterDocId="D0F25BD8394A5582A19FC1A27947BE26" masterDocTitle="Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae)" masterLastPageNumber="373" masterPageNumber="339" pageNumber="339" updateTime="1668150451991" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Liebherr, James K.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9831-884X</mods:nameIdentifier>
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<mods:affiliation>Department of Entomology, John H. and Anna B. Comstock Hall, 129 Garden Ave., Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
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<mods:nameIdentifier type="email">jkl5@cornell.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Porch, Nick</mods:namePart>
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<mods:affiliation>School of Life & Environmental Sciences & Centre for Integrated Ecology, Deakin University, Geelong, VIC 3216, Australia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaw, Matthew</mods:namePart>
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<mods:affiliation>South Australian Museum, South Terrace, Adelaide, SA 5000, Australia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sinclair, Bronte E.</mods:namePart>
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<mods:affiliation>Australian National Insect Collection, Building 101, Clunies Ross St., Black Mountain, ACT 2601, Australia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Maddison, David R.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7152-3824</mods:nameIdentifier>
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<mods:affiliation>Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-06-16</mods:number>
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<mods:detail type="volume">
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<mods:number>1044</mods:number>
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<mods:extent unit="page">
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<mods:start>339</mods:start>
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<mods:end>373</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1044.62335</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1044.62335</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1044-339</mods:identifier>
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<mods:identifier type="ZooBank">8264C99932014A3BB39C222790AC0192</mods:identifier>
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<mods:identifier type="Pensoft-UUID">D0F25BD8394A5582A19FC1A27947BE26</mods:identifier>
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<treatment ID-GBIF-Taxon="182754044" LSID="urn:lsid:plazi:treatment:472CE67146395350A146D7C06A1AB419" httpUri="http://treatment.plazi.org/id/472CE67146395350A146D7C06A1AB419" lastPageNumber="339" pageId="0" pageNumber="339">
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<subSubSection pageId="0" pageNumber="339" type="nomenclature">
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<paragraph pageId="0" pageNumber="339">
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<taxonomicName LSID="472CE671-4639-5350-A146-D7C06A1AB419" authority="(Sloane)" baseAuthorityName="Sloane" baseAuthorityYear="1920" class="Insecta" family="Carabidae" genus="Theprisa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theprisa convexa" order="Coleoptera" pageId="0" pageNumber="339" phylum="Arthropoda" rank="species" species="convexa">Theprisa convexa (Sloane)</taxonomicName>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus illustrations of Theprisa spp.; range of standardized body lengths indicated A T. otway female B T. convexa female C T. montana female." figureDoi="10.3897/zookeys.1044.62335.figure1" httpUri="https://binary.pensoft.net/fig/553850" pageId="0" pageNumber="339">Figures 1B</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male aedeagus including parameres of Theprisa spp., right, or anatomically ventral view A T. otway B T. convexa C T. montana D T. australis E T. darlingtoni. Abbreviations: ad, apical divot; af, apical face; fl, flagellum; lp, left paramere; mf, microtrichial field; rp, right paramere." figureDoi="10.3897/zookeys.1044.62335.figure3" httpUri="https://binary.pensoft.net/fig/553852" pageId="0" pageNumber="339">, 3B</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male aedeagus internal sac in everted condition for Theprisa spp. A T. otway, right view B T. convexa C T. montana D T. australis E T. darlingtoni. Abbreviations: fl, flagellum; mf, microtrichial field; rp, right paramere. Scale bars: 0.5 mm" figureDoi="10.3897/zookeys.1044.62335.figure4" httpUri="https://binary.pensoft.net/fig/553853" pageId="0" pageNumber="339">, 4B</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Parameres of male aedeagus for Theprisa spp., outside lateral view. Right paramere shown above left paramere A T. otway B T. convexa C T. montana D T. australis E T. darlingtoni. Abbreviations: lp, left paramere; rp, right paramere." figureDoi="10.3897/zookeys.1044.62335.figure5" httpUri="https://binary.pensoft.net/fig/553854" pageId="0" pageNumber="339">, 5B</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Structures associated with abdominal segments VIII and IX for Theprisa spp., dorsal view A T. otway B T. convexa C T. montana D T. australis E T. darlingtoni. Abbreviations: VIII, antecostal apodeme of abdominal segment VIII; IX, antecostal apodeme of abdominal segment IX; sp, spiracle of segment VIII; t, tergite of segment IX. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1044.62335.figure6" httpUri="https://binary.pensoft.net/fig/553855" pageId="0" pageNumber="339">, 6B</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female reproductive tract of Theprisa spp., ventral view A T. otway B T. convexa C T. montana D T. australis. Abbreviations: bc, bursa copulatrix; co, common oviduct; dgd, defensive gland efferent duct; gc, gonocoxa; hg, hindgut; hs, helminthoid sclerite; sd, spermathecal duct; sg, spermathecal gland; sgd, spermathecal gland duct; sgs, spermathecal gland stem; sp, spermatheca; v, vagina. Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1044.62335.figure7" httpUri="https://binary.pensoft.net/fig/553856" pageId="0" pageNumber="339">, 7B</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Left gonocoxa, ventral view of Theprisa spp., ventral view A T. otway B T. convexa C T. montana D T. australis E T. darlingtoni. Abbreviations: afs, apical fringe seta (e) of gonocoxite 1; ans, apical nematiform setae; des, dorsal ensiform seta; gc 1, basal gonocoxite 1; gc 2, apical gonocoxite 2; les, lateral ensiform setae; r, ramus." figureDoi="10.3897/zookeys.1044.62335.figure8" httpUri="https://binary.pensoft.net/fig/553857" pageId="0" pageNumber="339">, 8B</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distributional records for Theprisa spp.: T. otway (black circle); T. convexa (black square); T. montana (white circle)." figureDoi="10.3897/zookeys.1044.62335.figure9" httpUri="https://binary.pensoft.net/fig/553858" pageId="0" pageNumber="339">, 9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="339" type="reference_group">
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<paragraph pageId="0" pageNumber="339">
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<taxonomicName authorityName="Sloane" authorityYear="1920" class="Insecta" family="Carabidae" genus="Phersita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phersita convexa" order="Coleoptera" pageId="0" pageNumber="339" phylum="Arthropoda" rank="species" species="convexa">Phersita convexa</taxonomicName>
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Sloane, 1920: 158.
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</paragraph>
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<paragraph pageId="0" pageNumber="339">
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<taxonomicName baseAuthorityName="Sloane" baseAuthorityYear="1920" class="Insecta" family="Carabidae" genus="Theprisa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theprisa convexa" order="Coleoptera" pageId="0" pageNumber="339" phylum="Arthropoda" rank="species" species="convexa">Theprisa convexa</taxonomicName>
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Moore, 1963b: 285.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="339" type="types">
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<paragraph pageId="0" pageNumber="339">Types.</paragraph>
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<paragraph pageId="0" pageNumber="339">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="339">Lectotype</emphasis>
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male (SAMA) labelled by P. J. Darlington, Jr. hereby designated: card-mounted specimen with right antenna broken off distad pedicel, left mesoleg absent // Type // T. / Zeehan // Phersita / convexa Sl. / Id. by T. G. Sloane // Lectoholo. / P. convexa / PJD Sl. [red paper label] // Phersita 9.11564 / convexa Sl. / Tasmania / TYPE [red ink vertically at right end of label] // SAMA Database No. 25-035536 // LECTOTYPE ♂ / Phersita / convexa / Sloane / M. Shaw & J. K.
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<bibRefCitation DOI="https://doi.org/10.1071/IS19012" author="Liebherr, JK" journalOrPublisher="Invertebrate Systematics" pageId="0" pageNumber="339" pagination="1 - 33" refId="B29" refString="Liebherr, JK, 2020. Phylogenetic placement of the Australian Pharetis, gen. nov., and Spherita, gen. nov., in a revised classification of the circum-Antarctic Moriomorphini (Coleoptera: Carabidae). Invertebrate Systematics 34: 1 - 33, DOI: https://doi.org/10.1071/IS19012" title="Phylogenetic placement of the Australian Pharetis, gen. nov., and Spherita, gen. nov., in a revised classification of the circum-Antarctic Moriomorphini (Coleoptera: Carabidae)." url="https://doi.org/10.1071/IS19012" volume="34" year="2020">Liebherr 2020</bibRefCitation>
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[black-margined red label]. By this action Zeehan, Tasmania is designated type locality.
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</paragraph>
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<paragraph pageId="0" pageNumber="339">
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Sloane stated that this species was described from specimens inhabiting "Zeehan (Simson, No. 2123); Strahan and Waratah (Carter and Lea). Eleven specimens have been examined. (
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.19535" author="Sloane, TG" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="0" pageNumber="339" pagination="113 - 178" refId="B43" refString="Sloane, TG, 1920. The Carabidae of Tasmania. Proceedings of the Linnean Society of New South Wales 45: 113 - 178, DOI: https://doi.org/10.5962/bhl.part.19535" title="The Carabidae of Tasmania." url="https://doi.org/10.5962/bhl.part.19535" volume="45" year="1920">Sloane 1920</bibRefCitation>
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: 158)." We account below for the 10 paralectotypes to accompany the lectotype in the type series to ensure that this species did not include a specimen of the Tasmanian
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<taxonomicName lsidName="T. darlingtoni" pageId="0" pageNumber="339" rank="species" species="darlingtoni">
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<emphasis italics="true" pageId="0" pageNumber="339">T. darlingtoni</emphasis>
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</taxonomicName>
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, sp. nov., described below. Paralectotypes are numbered to correspond with a gallery of images of the specimens and labels (Suppl. material 1). The first three paralectotypes are deposited in the Sloane Collection (ANIC):
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<emphasis bold="true" pageId="0" pageNumber="339">1</emphasis>
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and
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<emphasis bold="true" pageId="0" pageNumber="339">2</emphasis>
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, male (left) and female (right) mounted on single card] // Strahan / Tas: Lea / & Carter // Phersita / convexa Sl. / Id. by T. G. Sloane. // Co-type //
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<emphasis bold="true" italics="true" pageId="0" pageNumber="339">Paratype</emphasis>
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[blue label of no significance as specimen is part of syntype series] // red rectangle with three holes //
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<emphasis bold="true" italics="true" pageId="0" pageNumber="339">Paralectotype</emphasis>
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♀, ♂ / Phersita / convexa Sloane / B. Sinclair & J. K. /
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<bibRefCitation DOI="https://doi.org/10.1071/IS19012" author="Liebherr, JK" journalOrPublisher="Invertebrate Systematics" pageId="0" pageNumber="339" pagination="1 - 33" refId="B29" refString="Liebherr, JK, 2020. Phylogenetic placement of the Australian Pharetis, gen. nov., and Spherita, gen. nov., in a revised classification of the circum-Antarctic Moriomorphini (Coleoptera: Carabidae). Invertebrate Systematics 34: 1 - 33, DOI: https://doi.org/10.1071/IS19012" title="Phylogenetic placement of the Australian Pharetis, gen. nov., and Spherita, gen. nov., in a revised classification of the circum-Antarctic Moriomorphini (Coleoptera: Carabidae)." url="https://doi.org/10.1071/IS19012" volume="34" year="2020">Liebherr 2020</bibRefCitation>
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[black-bordered red label];
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<emphasis bold="true" pageId="0" pageNumber="339">3</emphasis>
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, female: Zeehan / 12/91 [on reverse] // Phersita / convexa Sl. / Id. by T. G. Sloane. // Co-type //
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<emphasis bold="true" italics="true" pageId="0" pageNumber="339">Paratype</emphasis>
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//
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<emphasis bold="true" italics="true" pageId="0" pageNumber="339">Paralectotype</emphasis>
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♀ [as above]. The other seven paralectotypes that accompany the lectotype (SAMA) include:
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<emphasis bold="true" pageId="0" pageNumber="339">4</emphasis>
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, male card mounted // Zeehan // Phersita / convexa Sl. / Id. by T. G. Sloane // SAMA Database / No. 25-039812;
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<emphasis bold="true" pageId="0" pageNumber="339">5</emphasis>
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and
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<emphasis bold="true" pageId="0" pageNumber="339">6</emphasis>
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, two teneral females separately card mounted, on single pin // 3123 [blue paper label] // Tasmania / A. Simson (2 labels) // Phersita / convexa Sl. / Id. by T. G. Sloane // SAMA Database / No. 25-039813 // SAMA Database / No. 25-039814;
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<emphasis bold="true" pageId="0" pageNumber="339">7</emphasis>
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and
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<emphasis bold="true" pageId="0" pageNumber="339">8</emphasis>
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, two specimens, gender not determined, separately card mounted, on single pin // Tasmania / A. Simson (2 labels) // 3123 [blue paper label] // Phersita 19594 / convexa Sl. / Id. by T. G. Sloane / Cotype [red ink vertically on right end of label] // SAMA Database / No. 25-039819 // SAMA Database / No. 25-039820;
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<emphasis bold="true" pageId="0" pageNumber="339">9</emphasis>
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and
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<emphasis bold="true" pageId="0" pageNumber="339">10</emphasis>
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, two specimens mounted on one card, gender not determined // Waratah / Tas: Lea / & Carter (2 labels) // Phersita / convexa Sl. / Id. by T. G. Sloane // Co-type // // SAMA Database / No. 25-039821 // SAMA Database / No. 25-039822. All paralectotypes in SAMA also bear a bottom label, one per pin:
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<emphasis bold="true" pageId="0" pageNumber="339">
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<emphasis italics="true" pageId="0" pageNumber="339">Paralectotype</emphasis>
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(
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<emphasis italics="true" pageId="0" pageNumber="339">s</emphasis>
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)
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</emphasis>
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/ Phersita / convexa Sloane / M. Shaw & J. K.
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<bibRefCitation DOI="https://doi.org/10.1071/IS19012" author="Liebherr, JK" journalOrPublisher="Invertebrate Systematics" pageId="0" pageNumber="339" pagination="1 - 33" refId="B29" refString="Liebherr, JK, 2020. Phylogenetic placement of the Australian Pharetis, gen. nov., and Spherita, gen. nov., in a revised classification of the circum-Antarctic Moriomorphini (Coleoptera: Carabidae). Invertebrate Systematics 34: 1 - 33, DOI: https://doi.org/10.1071/IS19012" title="Phylogenetic placement of the Australian Pharetis, gen. nov., and Spherita, gen. nov., in a revised classification of the circum-Antarctic Moriomorphini (Coleoptera: Carabidae)." url="https://doi.org/10.1071/IS19012" volume="34" year="2020">Liebherr 2020</bibRefCitation>
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[black margined red label]. The discrepancy in recorded lot numbers 2123 in the published description versus 3123 on the specimen labels is adjudicated in favor of the specimen labels. Looking through registers at SAMA, we have not been able to locate this number, nor several similar missing numbers that also relate to various Tasmanian beetles from Simson.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="339" type="extended diagnosis">
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<paragraph pageId="0" pageNumber="339">Extended diagnosis</paragraph>
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<paragraph pageId="0" pageNumber="339">
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(n = 5). This species is aptly named due to the convex, domed elytra with depressed scutellar area (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus illustrations of Theprisa spp.; range of standardized body lengths indicated A T. otway female B T. convexa female C T. montana female." figureDoi="10.3897/zookeys.1044.62335.figure1" httpUri="https://binary.pensoft.net/fig/553850" pageId="0" pageNumber="339">1B</figureCitation>
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). The elytra are narrowed basally, with the angulate humeri constricted laterally; HuW/MEW = 0.61. In keeping with the basally constricted elytra, the pronotum is more cordate than in the other species (MPW/BPW = 1.23-1.31) and the lateral margins are sinuate anterad the pronotal basal seta articulatory socket. The pronotal base is also unique in the genus, with the marginal bead continuously marked across its breadth, and the lateral portions of the basal margin posterad the laterobasal depressions angled anteriorly relative to the extended median base. The laterobasal depression is broadly quadrate, with a moderately upraised, punctate tubercle bordered medially by a deep impression laterad the median base, and laterally by a narrow longitudinal depression inside the broadly elevated lateral margin. The body surface is glossy, with the vertex covered with indistinct transverse lines joined into an elongate transverse mesh in parts, the pronotal disc glossy with micropunctures visible across the disc, and the elytral disc and apex subiridescent due to indistinct transverse line microsculpture. Apical abdominal ventrite of male with two setae each side along margin plus two widely spaced medial subapical setae nearly in line with the outer four apical setae, female apical ventrite with two setae each side plus a transverse line of three or four medial subapical setae. Standardized body length is 6.2-7.2 mm.
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</paragraph>
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<caption doi="10.3897/zookeys.1044.62335.figure3" httpUri="https://binary.pensoft.net/fig/553852" pageId="0" pageNumber="339" start="Figure 3" startId="F3">
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<paragraph pageId="0" pageNumber="339">
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<emphasis bold="true" pageId="0" pageNumber="339">Figure 3.</emphasis>
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Male aedeagus including parameres of
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<taxonomicName authorityName="Moore" authorityYear="1963" class="Insecta" family="Carabidae" genus="Theprisa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theprisa" order="Orthoptera" pageId="0" pageNumber="339" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="339">Theprisa</emphasis>
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</taxonomicName>
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spp., right, or anatomically ventral view
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<emphasis bold="true" pageId="0" pageNumber="339">A</emphasis>
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<taxonomicName lsidName="T. otway" pageId="0" pageNumber="339" rank="species" species="otway">
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<emphasis italics="true" pageId="0" pageNumber="339">T. otway</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="339">B</emphasis>
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<taxonomicName lsidName="T. convexa" pageId="0" pageNumber="339" rank="species" species="convexa">
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<emphasis italics="true" pageId="0" pageNumber="339">T. convexa</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="339">C</emphasis>
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<taxonomicName lsidName="T. montana" pageId="0" pageNumber="339" rank="species" species="montana">
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<emphasis italics="true" pageId="0" pageNumber="339">T. montana</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="339">D</emphasis>
|
||
<taxonomicName lsidName="T. australis" pageId="0" pageNumber="339" rank="species" species="australis">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. australis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">E</emphasis>
|
||
<taxonomicName lsidName="T. darlingtoni" pageId="0" pageNumber="339" rank="species" species="darlingtoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. darlingtoni</emphasis>
|
||
</taxonomicName>
|
||
. Abbreviations: ad, apical divot; af, apical face; fl, flagellum; lp, left paramere; mf, microtrichial field; rp, right paramere.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="339">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="339">Male genitalia</emphasis>
|
||
(n = 6). Aedeagal median lobe robust, base broadly open on right side, basal margin extended as a sagittal crest dorsad basal opening (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male aedeagus including parameres of Theprisa spp., right, or anatomically ventral view A T. otway B T. convexa C T. montana D T. australis E T. darlingtoni. Abbreviations: ad, apical divot; af, apical face; fl, flagellum; lp, left paramere; mf, microtrichial field; rp, right paramere." figureDoi="10.3897/zookeys.1044.62335.figure3" httpUri="https://binary.pensoft.net/fig/553852" pageId="0" pageNumber="339">3B</figureCitation>
|
||
); median lobe apex broadly and evenly rounded, extended only slightly beyond ostium, lateral surfaces of apex densely covered with large pits; internal sac bearing an apically narrowed lanceolate flagellum (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male aedeagus internal sac in everted condition for Theprisa spp. A T. otway, right view B T. convexa C T. montana D T. australis E T. darlingtoni. Abbreviations: fl, flagellum; mf, microtrichial field; rp, right paramere. Scale bars: 0.5 mm" figureDoi="10.3897/zookeys.1044.62335.figure4" httpUri="https://binary.pensoft.net/fig/553853" pageId="0" pageNumber="339">4B</figureCitation>
|
||
), its broadly sclerotized base extremely evident in uneverted specimens (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male aedeagus including parameres of Theprisa spp., right, or anatomically ventral view A T. otway B T. convexa C T. montana D T. australis E T. darlingtoni. Abbreviations: ad, apical divot; af, apical face; fl, flagellum; lp, left paramere; mf, microtrichial field; rp, right paramere." figureDoi="10.3897/zookeys.1044.62335.figure3" httpUri="https://binary.pensoft.net/fig/553852" pageId="0" pageNumber="339">3B</figureCitation>
|
||
); right paramere elongate, evenly curved along length, gradually narrowed in apical half to tightly rounded tip (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Parameres of male aedeagus for Theprisa spp., outside lateral view. Right paramere shown above left paramere A T. otway B T. convexa C T. montana D T. australis E T. darlingtoni. Abbreviations: lp, left paramere; rp, right paramere." figureDoi="10.3897/zookeys.1044.62335.figure5" httpUri="https://binary.pensoft.net/fig/553854" pageId="0" pageNumber="339">5B</figureCitation>
|
||
), bearing 19-29 short setae along ventral margin in apical half, 2-9 setae on dorsal surface near apex, and with apex bearing 0-2 setae; left paramere slightly broader near midlength, apex slightly constricted before rounded apex, ventral margin usually glabrous (one individual with a single short ventral seta, a second with nine short setae), dorsal surface with 0-5 setae near apex, and apex with 0-2 setae present; antecostal apodeme of abdominal segment IX rounded distally, the apical juncture of lateral arms broad (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Structures associated with abdominal segments VIII and IX for Theprisa spp., dorsal view A T. otway B T. convexa C T. montana D T. australis E T. darlingtoni. Abbreviations: VIII, antecostal apodeme of abdominal segment VIII; IX, antecostal apodeme of abdominal segment IX; sp, spiracle of segment VIII; t, tergite of segment IX. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1044.62335.figure6" httpUri="https://binary.pensoft.net/fig/553855" pageId="0" pageNumber="339">6B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1044.62335.figure4" httpUri="https://binary.pensoft.net/fig/553853" pageId="0" pageNumber="339" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="339">
|
||
<emphasis bold="true" pageId="0" pageNumber="339">Figure 4.</emphasis>
|
||
Male aedeagus internal sac in everted condition for
|
||
<taxonomicName authorityName="Moore" authorityYear="1963" class="Insecta" family="Carabidae" genus="Theprisa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theprisa" order="Orthoptera" pageId="0" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">Theprisa</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
<emphasis bold="true" pageId="0" pageNumber="339">A</emphasis>
|
||
<taxonomicName lsidName="T. otway" pageId="0" pageNumber="339" rank="species" species="otway">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. otway</emphasis>
|
||
</taxonomicName>
|
||
, right view
|
||
<emphasis bold="true" pageId="0" pageNumber="339">B</emphasis>
|
||
<taxonomicName lsidName="T. convexa" pageId="0" pageNumber="339" rank="species" species="convexa">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. convexa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">C</emphasis>
|
||
<taxonomicName lsidName="T. montana" pageId="0" pageNumber="339" rank="species" species="montana">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. montana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">D</emphasis>
|
||
<taxonomicName lsidName="T. australis" pageId="0" pageNumber="339" rank="species" species="australis">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. australis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">E</emphasis>
|
||
<taxonomicName lsidName="T. darlingtoni" pageId="0" pageNumber="339" rank="species" species="darlingtoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. darlingtoni</emphasis>
|
||
</taxonomicName>
|
||
. Abbreviations: fl, flagellum; mf, microtrichial field; rp, right paramere. Scale bars: 0.5 mm
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1044.62335.figure5" httpUri="https://binary.pensoft.net/fig/553854" pageId="0" pageNumber="339" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="339">
|
||
<emphasis bold="true" pageId="0" pageNumber="339">Figure 5.</emphasis>
|
||
Parameres of male aedeagus for
|
||
<taxonomicName authorityName="Moore" authorityYear="1963" class="Insecta" family="Carabidae" genus="Theprisa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theprisa" order="Orthoptera" pageId="0" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">Theprisa</emphasis>
|
||
</taxonomicName>
|
||
spp., outside lateral view. Right paramere shown above left paramere
|
||
<emphasis bold="true" pageId="0" pageNumber="339">A</emphasis>
|
||
<taxonomicName lsidName="T. otway" pageId="0" pageNumber="339" rank="species" species="otway">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. otway</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">B</emphasis>
|
||
<taxonomicName lsidName="T. convexa" pageId="0" pageNumber="339" rank="species" species="convexa">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. convexa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">C</emphasis>
|
||
<taxonomicName lsidName="T. montana" pageId="0" pageNumber="339" rank="species" species="montana">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. montana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">D</emphasis>
|
||
<taxonomicName lsidName="T. australis" pageId="0" pageNumber="339" rank="species" species="australis">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. australis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">E</emphasis>
|
||
<taxonomicName lsidName="T. darlingtoni" pageId="0" pageNumber="339" rank="species" species="darlingtoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. darlingtoni</emphasis>
|
||
</taxonomicName>
|
||
. Abbreviations: lp, left paramere; rp, right paramere.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1044.62335.figure6" httpUri="https://binary.pensoft.net/fig/553855" pageId="0" pageNumber="339" start="Figure 6" startId="F6">
|
||
<paragraph pageId="0" pageNumber="339">
|
||
<emphasis bold="true" pageId="0" pageNumber="339">Figure 6.</emphasis>
|
||
Structures associated with abdominal segments VIII and IX for
|
||
<taxonomicName authorityName="Moore" authorityYear="1963" class="Insecta" family="Carabidae" genus="Theprisa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theprisa" order="Orthoptera" pageId="0" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">Theprisa</emphasis>
|
||
</taxonomicName>
|
||
spp., dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="339">A</emphasis>
|
||
<taxonomicName lsidName="T. otway" pageId="0" pageNumber="339" rank="species" species="otway">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. otway</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">B</emphasis>
|
||
<taxonomicName lsidName="T. convexa" pageId="0" pageNumber="339" rank="species" species="convexa">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. convexa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">C</emphasis>
|
||
<taxonomicName lsidName="T. montana" pageId="0" pageNumber="339" rank="species" species="montana">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. montana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">D</emphasis>
|
||
<taxonomicName lsidName="T. australis" pageId="0" pageNumber="339" rank="species" species="australis">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. australis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">E</emphasis>
|
||
<taxonomicName lsidName="T. darlingtoni" pageId="0" pageNumber="339" rank="species" species="darlingtoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. darlingtoni</emphasis>
|
||
</taxonomicName>
|
||
. Abbreviations: VIII, antecostal apodeme of abdominal segment VIII; IX, antecostal apodeme of abdominal segment IX; sp, spiracle of segment VIII; t, tergite of segment IX. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="339">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="339">Female reproductive tract</emphasis>
|
||
(n = 1). Bursa copulatrix of vase-like configuration, vaginal area constricted relative to broader distal portion of bursa, length 1.25
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
maximum breadth compressed under microslide (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female reproductive tract of Theprisa spp., ventral view A T. otway B T. convexa C T. montana D T. australis. Abbreviations: bc, bursa copulatrix; co, common oviduct; dgd, defensive gland efferent duct; gc, gonocoxa; hg, hindgut; hs, helminthoid sclerite; sd, spermathecal duct; sg, spermathecal gland; sgd, spermathecal gland duct; sgs, spermathecal gland stem; sp, spermatheca; v, vagina. Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1044.62335.figure7" httpUri="https://binary.pensoft.net/fig/553856" pageId="0" pageNumber="339">7B</figureCitation>
|
||
); helminthoid sclerite present, rounded apically, not extended beyond juncture with spermathecal duct; spermathecal duct stout, sinuously recurved to meet spermatheca, length twice that of annulated spermathecal reservoir; spermathecal gland duct very thin, length half that of spermathecal reservoir which it joins at reservoir base; spermathecal gland comprising sclerotized stem plus membranous reservoir bearing numerous ductules; gonocoxa bipartite, basal gonocoxite 1 with single apical fringe seta, median surface glabrous, membranous ramus present (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Left gonocoxa, ventral view of Theprisa spp., ventral view A T. otway B T. convexa C T. montana D T. australis E T. darlingtoni. Abbreviations: afs, apical fringe seta (e) of gonocoxite 1; ans, apical nematiform setae; des, dorsal ensiform seta; gc 1, basal gonocoxite 1; gc 2, apical gonocoxite 2; les, lateral ensiform setae; r, ramus." figureDoi="10.3897/zookeys.1044.62335.figure8" httpUri="https://binary.pensoft.net/fig/553857" pageId="0" pageNumber="339">8B</figureCitation>
|
||
); apical gonocoxite 2 with base narrow, lateral margin arcuate, apex acuminate; two narrow lateral ensiform setae and one dorsal ensiform present; two apical nematiform setae set in fossa at apical 1/4 of apical gonocoxite length.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1044.62335.figure7" httpUri="https://binary.pensoft.net/fig/553856" pageId="0" pageNumber="339" start="Figure 7" startId="F7">
|
||
<paragraph pageId="0" pageNumber="339">
|
||
<emphasis bold="true" pageId="0" pageNumber="339">Figure 7.</emphasis>
|
||
Female reproductive tract of
|
||
<taxonomicName authorityName="Moore" authorityYear="1963" class="Insecta" family="Carabidae" genus="Theprisa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theprisa" order="Orthoptera" pageId="0" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">Theprisa</emphasis>
|
||
</taxonomicName>
|
||
spp., ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="339">A</emphasis>
|
||
<taxonomicName lsidName="T. otway" pageId="0" pageNumber="339" rank="species" species="otway">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. otway</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">B</emphasis>
|
||
<taxonomicName lsidName="T. convexa" pageId="0" pageNumber="339" rank="species" species="convexa">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. convexa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">C</emphasis>
|
||
<taxonomicName lsidName="T. montana" pageId="0" pageNumber="339" rank="species" species="montana">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. montana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">D</emphasis>
|
||
<taxonomicName lsidName="T. australis" pageId="0" pageNumber="339" rank="species" species="australis">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. australis</emphasis>
|
||
</taxonomicName>
|
||
. Abbreviations: bc, bursa copulatrix; co, common oviduct; dgd, defensive gland efferent duct; gc, gonocoxa; hg, hindgut; hs, helminthoid sclerite; sd, spermathecal duct; sg, spermathecal gland; sgd, spermathecal gland duct; sgs, spermathecal gland stem; sp, spermatheca; v, vagina. Scale bars: 0.50 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1044.62335.figure8" httpUri="https://binary.pensoft.net/fig/553857" pageId="0" pageNumber="339" start="Figure 8" startId="F8">
|
||
<paragraph pageId="0" pageNumber="339">
|
||
<emphasis bold="true" pageId="0" pageNumber="339">Figure 8.</emphasis>
|
||
Left gonocoxa, ventral view of
|
||
<taxonomicName authorityName="Moore" authorityYear="1963" class="Insecta" family="Carabidae" genus="Theprisa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theprisa" order="Orthoptera" pageId="0" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">Theprisa</emphasis>
|
||
</taxonomicName>
|
||
spp., ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="339">A</emphasis>
|
||
<taxonomicName lsidName="T. otway" pageId="0" pageNumber="339" rank="species" species="otway">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. otway</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">B</emphasis>
|
||
<taxonomicName lsidName="T. convexa" pageId="0" pageNumber="339" rank="species" species="convexa">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. convexa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">C</emphasis>
|
||
<taxonomicName lsidName="T. montana" pageId="0" pageNumber="339" rank="species" species="montana">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. montana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">D</emphasis>
|
||
<taxonomicName lsidName="T. australis" pageId="0" pageNumber="339" rank="species" species="australis">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. australis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="339">E</emphasis>
|
||
<taxonomicName lsidName="T. darlingtoni" pageId="0" pageNumber="339" rank="species" species="darlingtoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. darlingtoni</emphasis>
|
||
</taxonomicName>
|
||
. Abbreviations: afs, apical fringe seta(e) of gonocoxite 1; ans, apical nematiform setae; des, dorsal ensiform seta; gc1, basal gonocoxite 1; gc2, apical gonocoxite 2; les, lateral ensiform setae; r, ramus.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="339" type="distribution">
|
||
<paragraph pageId="0" pageNumber="339">Distribution and habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="339">
|
||
<taxonomicName baseAuthorityName="Sloane" baseAuthorityYear="1920" class="Insecta" family="Carabidae" genus="Theprisa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theprisa convexa" order="Orthoptera" pageId="0" pageNumber="339" phylum="Arthropoda" rank="species" species="convexa">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">Theprisa convexa</emphasis>
|
||
</taxonomicName>
|
||
is known from the mountainous western portions of Tasmania (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distributional records for Theprisa spp.: T. otway (black circle); T. convexa (black square); T. montana (white circle)." figureDoi="10.3897/zookeys.1044.62335.figure9" httpUri="https://binary.pensoft.net/fig/553858" pageId="0" pageNumber="339">9</figureCitation>
|
||
), within the West, King, Central Highlands, and Northern Slopes biogeographic regions (
|
||
<bibRefCitation author="IBRA" journalOrPublisher="Cladistics" pageId="0" pageNumber="339" publicationUrl="https://www.environment.gov.au/system/files/pages/5b3d2d31-2355-4b60-820c-e370572b2520/files/bioregions-new.pdf" refId="B23" refString="IBRA, 2012. Interim Biogeographic Regionalisation for Australia (IBRA) version 7. Commonwealth of Australia, ERIN, Australian Government Department of the Environment and Energy, Canberra, October 2016. https://www.environment.gov.au/system/files/pages/5b3d2d31-2355-4b60-820c-e370572b2520/files/bioregions-new.pdf" title="Interim Biogeographic Regionalisation for Australia (IBRA) version 7. Commonwealth of Australia, ERIN, Australian Government Department of the Environment and Energy, Canberra, October 2016." url="https://www.environment.gov.au/system/files/pages/5b3d2d31-2355-4b60-820c-e370572b2520/files/bioregions-new.pdf" year="2012">IBRA 2012</bibRefCitation>
|
||
). The beetles are terrestrial, with specimens recorded from under decaying logs, in leaf litter associated with
|
||
<taxonomicName class="Rosopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Nothofagus cunninghamii" order="Fagales" pageId="0" pageNumber="339" phylum="Magnoliophyta" rank="species" species="cunninghamii">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">Nothofagus cunninghamii</emphasis>
|
||
</taxonomicName>
|
||
(Hooker) (
|
||
<taxonomicName class="Rosopsida" family="Nothofagaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Fagales" pageId="0" pageNumber="339" phylum="Magnoliophyta" rank="family">Nothofagaceae</taxonomicName>
|
||
), and from riverine forest with
|
||
<taxonomicName authorityName="Cavanilles" authorityYear="1798" class="Magnoliopsida" family="Cunoniaceae" genus="Eucryphia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Eucryphia" order="Oxalidales" pageId="0" pageNumber="339" phylum="Magnoliophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">Eucryphia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="Cavanilles" authorityYear="1798" class="Magnoliopsida" family="Cunoniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Oxalidales" pageId="0" pageNumber="339" phylum="Magnoliophyta" rank="family">Cunoniaceae</taxonomicName>
|
||
),
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" genus="Richea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Richea" order="Ericales" pageId="0" pageNumber="339" phylum="Angiospermae" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">Richea</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Ericales" pageId="0" pageNumber="339" phylum="Angiospermae" rank="family">Ericaceae</taxonomicName>
|
||
),
|
||
<taxonomicName lsidName="N. cunninghamii" pageId="0" pageNumber="339" rank="species" species="cunninghamii">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">N. cunninghamii</emphasis>
|
||
</taxonomicName>
|
||
, and tree ferns. Recorded habitats range 80-1200 m elevation. We have not had the opportunity to examine specimens reported by
|
||
<bibRefCitation DOI="https://doi.org/10.3897/subtbiol.9.2516" author="Eberhard, S" journalOrPublisher="Subterranean Biology" pageId="0" pageNumber="339" pagination="1 - 72" refId="B17" refString="Eberhard, S, Giachino, PM, 2011. Tasmanian Trechinae and Psydrinae (Coleoptera: Carabidae): a taxonomic and biogeographic synthesis, with descriptions of new species and evaluation of the impact of Quaternary climatic changes on evolution of the subterranean fauna. Subterranean Biology 9: 1 - 72, DOI: https://doi.org/10.3897/subtbiol.9.2516" title="Tasmanian Trechinae and Psydrinae (Coleoptera: Carabidae): a taxonomic and biogeographic synthesis, with descriptions of new species and evaluation of the impact of Quaternary climatic changes on evolution of the subterranean fauna." url="https://doi.org/10.3897/subtbiol.9.2516" volume="9" year="2011">Eberhard and Giachino (2011)</bibRefCitation>
|
||
, however the geographical expanse of collecting sites they reported agrees totally with our data, with the exception of additional outlying records herein from Christmas Hills, 35 km SW Smithton (Kethley, FMNH), and Trowutta (Madden, TMAG), northwest Tasmania (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distributional records for Theprisa spp.: T. otway (black circle); T. convexa (black square); T. montana (white circle)." figureDoi="10.3897/zookeys.1044.62335.figure9" httpUri="https://binary.pensoft.net/fig/553858" pageId="0" pageNumber="339">9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1044.62335.figure9" httpUri="https://binary.pensoft.net/fig/553858" pageId="0" pageNumber="339" start="Figure 9" startId="F9">
|
||
<paragraph pageId="0" pageNumber="339">
|
||
<emphasis bold="true" pageId="0" pageNumber="339">Figure 9.</emphasis>
|
||
Distributional records for
|
||
<taxonomicName authorityName="Moore" authorityYear="1963" class="Insecta" family="Carabidae" genus="Theprisa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theprisa" order="Orthoptera" pageId="0" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">Theprisa</emphasis>
|
||
</taxonomicName>
|
||
spp.:
|
||
<taxonomicName lsidName="T. otway" pageId="0" pageNumber="339" rank="species" species="otway">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. otway</emphasis>
|
||
</taxonomicName>
|
||
(black circle);
|
||
<taxonomicName lsidName="T. convexa" pageId="0" pageNumber="339" rank="species" species="convexa">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. convexa</emphasis>
|
||
</taxonomicName>
|
||
(black square);
|
||
<taxonomicName lsidName="T. montana" pageId="0" pageNumber="339" rank="species" species="montana">
|
||
<emphasis italics="true" pageId="0" pageNumber="339">T. montana</emphasis>
|
||
</taxonomicName>
|
||
(white circle).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |