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<document id="3811D1ADAF96B691E2411CF12272D7DC" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627005272297" checkinUser="felipe" docAuthor="Published, First" docDate="2008" docId="03DF87D2FF8AAB129EC811F2420F3E21" docLanguage="en" docName="zt01804p456part1.pdf" docOrigin="Zootaxa 1804" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Corethrella (Corethrella) briannae Borkent 2008, new species" docType="treatment" docVersion="2" lastPageNumber="104" masterDocId="FFE6FFAAFFECAB759E5F1025465D3823" masterDocTitle="The Frog-Biting Midges of the World (Corethrellidae: Diptera)" masterLastPageNumber="456" masterPageNumber="1" pageNumber="103" updateTime="1699014769975" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9CCAA1EB7B9333C7C957FCBCA81A58CF">The Frog-Biting Midges of the World (Corethrellidae: Diptera)</mods:title>
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<mods:date id="89FDD115A7DEFFE6C78BE8089326BF27">2008</mods:date>
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<mods:number id="7518DFCD996FC450415E561640A3AA16">2008-06-16</mods:number>
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<subSubSection id="C36C654FFF8AAB139EC811F2451A39D2" box="[151,839,471,497]" pageId="102" pageNumber="103" type="nomenclature">
<paragraph id="8BC936C4FF8AAB139EC811F2451A39D2" blockId="102.[151,839,471,497]" box="[151,839,471,497]" pageId="102" pageNumber="103">
<heading id="D08181A8FF8AAB139EC811F2451A39D2" bold="true" box="[151,839,471,497]" fontSize="11" level="1" pageId="102" pageNumber="103" reason="1">
<emphasis id="B902EAD6FF8AAB139EC811F2451A39D2" bold="true" box="[151,839,471,497]" pageId="102" pageNumber="103">
<taxonomicName id="4C764D47FF8AAB139EC811F244F739D2" ID-CoL="8BL55" authority="Borkent" authorityName="Borkent" authorityYear="2008" box="[151,682,471,497]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="briannae" status="sp. nov." subGenus="Corethrella">
<emphasis id="B902EAD6FF8AAB139EC811F2477C39D2" bold="true" box="[151,289,471,497]" italics="true" pageId="102" pageNumber="103">Corethrella</emphasis>
(
<emphasis id="B902EAD6FF8AAB139F6D11F247E139D2" bold="true" box="[306,444,471,497]" italics="true" pageId="102" pageNumber="103">Corethrella</emphasis>
)
<emphasis id="B902EAD6FF8AAB139F9111F2446639D2" bold="true" box="[462,571,471,497]" italics="true" pageId="102" pageNumber="103">briannae</emphasis>
Borkent
</taxonomicName>
,
<taxonomicNameLabel id="A23157ADFF8AAB139CE811F2451A39D2" box="[695,839,471,497]" pageId="102" pageNumber="103" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF8AAB139EC8120747473BEF" pageId="102" pageNumber="103" type="diagnosis">
<paragraph id="8BC936C4FF8AAB139EC8120747473BEF" blockId="102.[151,1436,546,2015]" pageId="102" pageNumber="103">
DIAGNOSIS:
<emphasis id="B902EAD6FF8AAB139F0D120747893A18" box="[338,468,546,571]" italics="true" pageId="102" pageNumber="103">Male adult</emphasis>
: only extant species of
<taxonomicName id="4C764D47FF8AAB139CA6120745DD3A18" box="[761,896,546,571]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="102" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B902EAD6FF8AAB139CA6120745DD3A18" box="[761,896,546,571]" italics="true" pageId="102" pageNumber="103">Corethrella</emphasis>
</taxonomicName>
in the New World with a distinct midlength wing band (Fig. 62N), with the scutum mostly light brown but with the posterior anepisternum, most of the anepimeron and the katepisternum dark brown, midfemur light to medium brown and equal to that of base of hind femur, base of hind tibia darkly pigmented (contrasting with pale apex of hind femur) (as in
<figureCitation id="134D2A41FF8AAB139B7E12BF43D23A97" box="[1313,1423,666,692]" captionStart="FIGURE 47" captionStartId="353.[151,255,2007,2031]" captionTargetBox="[179,1398,197,1967]" captionTargetId="figure-24@353.[179,1408,197,1980]" captionTargetPageId="353" captionText="FIGURE 47. Thoraces and legs of female Corethrella, lateral view." pageId="102" pageNumber="103">Fig. 47A</figureCitation>
), gonocoxite pale and contrasting with medium brown tergite 9 (Fig. 78C).
<emphasis id="B902EAD6FF8AAB139A5212E742F73AF8" box="[1037,1194,706,731]" italics="true" pageId="102" pageNumber="103">Female adult</emphasis>
: only extant species of
<taxonomicName id="4C764D47FF8AAB139EE912CF47603B20" box="[182,317,746,771]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="102" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B902EAD6FF8AAB139EE912CF47603B20" box="[182,317,746,771]" italics="true" pageId="102" pageNumber="103">Corethrella</emphasis>
</taxonomicName>
in the New World with each of flagellomeres 13 moderately elongate (
<figureCitation id="134D2A41FF8AAB139AD812CF42AE3B27" box="[1159,1267,746,772]" captionStart="FIGURE 28" captionStartId="334.[151,255,1947,1971]" captionTargetBox="[208,1396,238,1917]" captionTargetId="figure-25@334.[177,1411,197,1921]" captionTargetPageId="334" captionText="FIGURE 28. Antennae of female Corethrella." pageId="102" pageNumber="103">Fig. 28G</figureCitation>
), flagellomere 3 without a sensillum coeloconicum, flagellomere 12 with 23 sensilla coeloconica, a distinct midlength wing band (Fig. 68M), with the scutum mostly light brown but with the posterior anepisternum, most of the anepimeron and the katepisternum dark brown, midfemur uniformly light to medium brown and equal to that of base of hind femur, and the base of hind tibia darkly pigmented (contrasting with pale apex of hind femur) (
<figureCitation id="134D2A41FF8AAB139EC0139747503BEF" box="[159,269,946,972]" captionStart="FIGURE 47" captionStartId="353.[151,255,2007,2031]" captionTargetBox="[179,1398,197,1967]" captionTargetId="figure-24@353.[179,1408,197,1980]" captionTargetPageId="353" captionText="FIGURE 47. Thoraces and legs of female Corethrella, lateral view." pageId="102" pageNumber="103">Fig. 47A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF8AAB129E9913FF47DA3B11" lastPageId="103" lastPageNumber="104" pageId="102" pageNumber="103" type="description">
<paragraph id="8BC936C4FF8AAB129E9913FF42123909" blockId="102.[151,1436,546,2015]" lastBlockId="103.[151,1437,152,1538]" lastPageId="103" lastPageNumber="104" pageId="102" pageNumber="103">
DESCRIPTION:
<emphasis id="B902EAD6FF8AAB139FC513FF44443BD0" box="[410,537,986,1011]" italics="true" pageId="102" pageNumber="103">Male adult</emphasis>
. Descriptive statistics: see
<tableCitation id="C6F4037FFF8AAB139D0813FF45873BD7" box="[855,986,986,1012]" captionStart-0="TABLE 2" captionStart-1="TABLE 3" captionStart-2="TABLE 4" captionStart-3="TABLE 5" captionStartId-0="269.[151,241,151,175]" captionStartId-1="271.[151,241,153,177]" captionStartId-2="273.[151,241,151,175]" captionStartId-3="275.[151,241,151,175]" captionText-0="TABLE 2. Length of wing of males of species of Corethrella. Values are in mm." captionText-1="TABLE 3. Ratio of foreleg tarsomeres 3/4 of males of species of Corethrella." captionText-2="TABLE 4. Ratio of distance of dorsomedial seta from the gonocoxite base/length of the gonocoxite of males of species" captionText-3="TABLE 5. Ratio of distance of subbasal seta from the gonostylus base/ length of the gonostylus of males of species of Corethrella." pageId="102" pageNumber="103">Tables 25</tableCitation>
.
<emphasis id="B902EAD6FF8AAB139DB313FF42723BD7" bold="true" box="[1004,1071,986,1012]" pageId="102" pageNumber="103">Head</emphasis>
: Outline in anterior view laterally elongate (as in
<figureCitation id="134D2A41FF8AAB139FD9142747BF3C3F" box="[390,482,1026,1052]" captionStart="FIGURE 9" captionStartId="315.[151,255,1947,1971]" captionTargetBox="[177,1410,239,1904]" captionTargetId="figure-23@315.[177,1411,197,1921]" captionTargetPageId="315" captionText="FIGURE 9. Head capsules of female Corethrella, anterior view." pageId="102" pageNumber="103">Fig. 9E</figureCitation>
). Two large setae on frons between ventromedial area of ommatida (as in
<figureCitation id="134D2A41FF8AAB139B31142746953C67" captionStart="FIGURE 16" captionStartId="322.[151,255,1947,1971]" captionTargetBox="[177,1410,349,1917]" captionTargetId="figure-31@322.[177,1411,197,1921]" captionTargetPageId="322" captionText="FIGURE 16. Dorsal portion of head capsules of female Corethrella, anterior view." pageId="102" pageNumber="103">Fig. 16F</figureCitation>
). Antenna light brown; pedicel with at least one distinctive, more elongate, stout, dorsal or dorsolateral seta; flagellomeres as in
<figureCitation id="134D2A41FF8AAB139FE0147744763C4F" box="[447,555,1106,1132]" captionStart="FIGURE 21" captionStartId="327.[151,255,1947,1971]" captionTargetBox="[197,1407,238,1921]" captionTargetId="figure-20@327.[177,1411,197,1921]" captionTargetPageId="327" captionText="FIGURE 21. Antennae of male Corethrella." pageId="102" pageNumber="103">Fig. 21H</figureCitation>
, sensilla coeloconica distributed as in
<tableCitation id="C6F4037FFF8AAB139DA9147742123C4F" box="[1014,1103,1106,1132]" captionStart="TABLE 1" captionStartId="265.[151,241,151,175]" captionText="TABLE 1. Pattern of sensilla coeloconica (= s.c.) on antennal flagellomeres of species of Corethrella. Numbers refer to the flagellomere with sensilla coeloconica, with those in brackets referring to flagellomeres with or without sensilla coeloconica. Mis. (missing) refers to damaged specimens, unknown refers to missing stages, uncertain refers to specimens in which visibility was too poor to be certain of whether sensilla coeloconica were present or not. Those species names with a “†” are those of fossils. Corethrella librata, known only as immature stages, is not included." pageId="102" pageNumber="103">Table 1</tableCitation>
; flagellomere 13 with welldeveloped apical bifurcation. Palpus pale; segment 3 somewhat ovoid, swollen at or near midlength.
<emphasis id="B902EAD6FF8AAB139B60145F43C13CB7" bold="true" box="[1343,1436,1146,1172]" pageId="102" pageNumber="103">Thorax</emphasis>
(as in
<figureCitation id="134D2A41FF8AAB139EBB148747093C9F" box="[228,340,1186,1212]" captionStart="FIGURE 47" captionStartId="353.[151,255,2007,2031]" captionTargetBox="[179,1398,197,1967]" captionTargetId="figure-24@353.[179,1408,197,1980]" captionTargetPageId="353" captionText="FIGURE 47. Thoraces and legs of female Corethrella, lateral view." pageId="102" pageNumber="103">Fig. 47A</figureCitation>
): Scutum light brown, with very anterolateral area medium brown; scutellum, mediotergite light brown; pleura mostly light brown with posterior portion of anterior anepisternum, most of posterior anepisternum, most of anepimeron, all of katepisternum dark brown, pale sclerites around base of wing. Posterior portion of dorsocentral row with group of about 6 elongate setae. Prescutal suture elongate, interrupted by area of pale cuticle. Anterior anepisternum divided diagonally by sinuous suture, dorsal portion about equal to ventral portion. Ventral portion of posterior anepisternum triangular, uniformly brown, with anterodorsal margin thick.
<emphasis id="B902EAD6FF8AAB139FD015B7478E3D8F" bold="true" box="[399,467,1426,1452]" pageId="102" pageNumber="103">Wing</emphasis>
(Fig. 62N): Apex of R
<subScript id="17F23481FF8AAB139CBB158744B13D93" attach="left" box="[740,748,1442,1456]" fontSize="6" pageId="102" pageNumber="103">2</subScript>
basal to apex of M
<subScript id="17F23481FF8AAB139D93158745893D93" attach="left" box="[972,980,1442,1456]" fontSize="6" pageId="102" pageNumber="103">1</subScript>
. Anterior margin with differently, discretely pigmented scales (indicating anterior margin of midlength band), with midlength band, some with more basal darker scales on C, A; veins (other than costa and wing margin) with well-developed scales. Halter pale.
<emphasis id="B902EAD6FF8AAB139F55162847183E04" bold="true" box="[266,325,1549,1575]" pageId="102" pageNumber="103">Legs</emphasis>
(as in
<figureCitation id="134D2A41FF8AAB139FC5162844573E04" box="[410,522,1549,1575]" captionStart="FIGURE 47" captionStartId="353.[151,255,2007,2031]" captionTargetBox="[179,1398,197,1967]" captionTargetId="figure-24@353.[179,1408,197,1980]" captionTargetPageId="353" captionText="FIGURE 47. Thoraces and legs of female Corethrella, lateral view." pageId="102" pageNumber="103">Fig. 47A</figureCitation>
): Mostly light brown, with slightly darker pigmentation on apical portion of foretibia, base and apex midtibia, hind femur with apical 2/3 pale, hind tibia with basal and apical dark brown discrete pigmentation, at least mid-, hind leg tarsomeres 24 with banding. With only slender setae, lacking scales (except for some in patch of whip-like setae on posterior portion of hind tibia). Midleg with thick, subapical setae on each of at least tarsomeres 13. Apices of fore-, midleg fifth tarsomeres undivided, with claws slightly subapical to apical (as in Fig. 75F). Claw of foreleg longer than those of mid-, hind leg. Each claw without inner tooth. Anterior claws of each leg without a basal prong. Foreleg claws unequal. Midleg claws equal. Foreleg third tarsomere shorter than fourth tarsomere. Empodia slender.
<emphasis id="B902EAD6FF8AAB139A0F170042973F1C" bold="true" box="[1104,1226,1829,1855]" pageId="102" pageNumber="103">Abdomen</emphasis>
(Fig. 78C): Light brown, with sternites 12 dark brown, tergite 9 medium brown.
<emphasis id="B902EAD6FF8AAB139DE31768426D3F44" bold="true" box="[956,1072,1869,1895]" pageId="102" pageNumber="103">Genitalia</emphasis>
(Fig. 89D): Gonocoxite pale, strongly tapering; anteromedial area with spicules similar in length to those elsewhere on gonocoxite; with well-defined dorsal row of setae, with seta 2 or setae 24 thicker than others, with row restricted to dorsal portion of gonocoxite. With one dorsomedial stout seta, more or less of even thickness for most of length, taper- ing near apex. Gonostylus (partially extended) straight for most of length, curved apically, slender, slightly tapering for most of length, somewhat expanded apically, apex pointed; one somewhat elongate, slender subbasal seta, situated anteriorly or anteroventrally; apical seta slender, elongate, simple. Aedeagus slender, elongate, tapering from base, pointed apically, with lateral margins fused at apex.
</paragraph>
<paragraph id="8BC936C4FF8BAB129E99111D47993A61" blockId="103.[151,1437,152,1538]" pageId="103" pageNumber="104">
<emphasis id="B902EAD6FF8BAB129E99111D47393972" box="[198,356,312,337]" italics="true" pageId="103" pageNumber="104">Female adult</emphasis>
. Descriptive statistics: see
<tableCitation id="C6F4037FFF8BAB129CEB111D451A3971" box="[692,839,312,338]" captionStart-0="TABLE 6" captionStart-1="TABLE 7" captionStart-2="TABLE 8" captionStart-3="TABLE 9" captionStart-4="TABLE 10" captionStart-5="TABLE 11" captionStartId-0="277.[151,241,153,177]" captionStartId-1="280.[151,241,153,177]" captionStartId-2="283.[151,241,153,177]" captionStartId-3="286.[151,241,151,175]" captionStartId-4="289.[151,239,153,177]" captionStartId-5="292.[151,241,153,177]" captionText-0="TABLE 6. Length of wing of females of species of Corethrella. Values are in mm." captionText-1="TABLE 7. Ratio of length of R1/wing length of females of species of Corethrella." captionText-2="TABLE 8. Ratio of head capsule width/height of females of species of Corethrella." captionText-3="TABLE 9. Number of setae on the clypeus of females of species of Corethrella. The number does not include those" captionText-4="TABLE 10. Number of posterior anepisternal setae of females of species of Corethrella." captionText-5="TABLE 11. Number of anepimeral setae of females of species of Corethrella." pageId="103" pageNumber="104">Tables 611</tableCitation>
. As for male, with following differences.
<emphasis id="B902EAD6FF8BAB129B0E111243C93972" bold="true" box="[1361,1428,311,337]" pageId="103" pageNumber="104">Head</emphasis>
: Coronal suture short, extending ventrally about midway along area between ommatidia (as in
<figureCitation id="134D2A41FF8BAB129B7C114543D23959" box="[1315,1423,352,378]" captionStart="FIGURE 16" captionStartId="322.[151,255,1947,1971]" captionTargetBox="[177,1410,349,1917]" captionTargetId="figure-31@322.[177,1411,197,1921]" captionTargetPageId="322" captionText="FIGURE 16. Dorsal portion of head capsules of female Corethrella, anterior view." pageId="103" pageNumber="104">Fig. 16F</figureCitation>
). Antennal flagellomeres as in
<figureCitation id="134D2A41FF8BAB129FA611AD443E3981" box="[505,611,392,418]" captionStart="FIGURE 28" captionStartId="334.[151,255,1947,1971]" captionTargetBox="[208,1396,238,1917]" captionTargetId="figure-25@334.[177,1411,197,1921]" captionTargetPageId="334" captionText="FIGURE 28. Antennae of female Corethrella." pageId="103" pageNumber="104">Fig. 28G</figureCitation>
, sensilla coeloconica distributed as in
<tableCitation id="C6F4037FFF8BAB129A6C11AD42D33981" box="[1075,1166,392,418]" captionStart="TABLE 1" captionStartId="265.[151,241,151,175]" captionText="TABLE 1. Pattern of sensilla coeloconica (= s.c.) on antennal flagellomeres of species of Corethrella. Numbers refer to the flagellomere with sensilla coeloconica, with those in brackets referring to flagellomeres with or without sensilla coeloconica. Mis. (missing) refers to damaged specimens, unknown refers to missing stages, uncertain refers to specimens in which visibility was too poor to be certain of whether sensilla coeloconica were present or not. Those species names with a “†” are those of fossils. Corethrella librata, known only as immature stages, is not included." pageId="103" pageNumber="104">Table 1</tableCitation>
. Clypeus (
<figureCitation id="134D2A41FF8BAB129B4411AD432D3981" box="[1307,1392,392,418]" captionStart="FIGURE 17" captionStartId="323.[151,255,1947,1971]" captionTargetBox="[177,1408,258,1907]" captionTargetId="figure-28@323.[177,1411,197,1921]" captionTargetPageId="323" captionText="FIGURE 17. Clypei of female Corethrella. Lateral setae, even if present, are not shown." pageId="103" pageNumber="104">Fig. 17</figureCitation>
AJ) squarish. Mandible with small, pointed teeth. Palpus as in
<figureCitation id="134D2A41FF8BAB129D3E1195459A39E9" box="[865,967,432,458]" captionStart="FIGURE 34" captionStartId="340.[151,255,1947,1971]" captionText="FIGURE 34. Right palpi of female Corethrella." pageId="103" pageNumber="104">
Fig.
<specimenCount id="9D70FD4DFF8BAB129DC71195459A39E9" box="[920,967,432,458]" pageId="103" pageNumber="104" type="generic">34F</specimenCount>
</figureCitation>
.
<emphasis id="B902EAD6FF8BAB129D83118A427D39EA" bold="true" box="[988,1056,431,457]" pageId="103" pageNumber="104">Wing</emphasis>
(Fig. 68M): similar to male but with additional somewhat darkened scales in basal band.
<emphasis id="B902EAD6FF8BAB129D1511F245D839D2" bold="true" box="[842,901,471,497]" pageId="103" pageNumber="104">Legs</emphasis>
: Claws of each leg equal to those of others; equal on each leg, simple (without inner teeth).
<emphasis id="B902EAD6FF8BAB129CB811DA453C3A3A" bold="true" box="[743,865,511,537]" pageId="103" pageNumber="104">Abdomen</emphasis>
: Light brown with sternites 12, segments 89 dark brown. Cercus pale.
</paragraph>
<paragraph id="8BC936C4FF8BAB129E99127547433AC1" blockId="103.[151,1437,152,1538]" pageId="103" pageNumber="104">
<emphasis id="B902EAD6FF8BAB129E99127547593A4A" box="[198,260,592,617]" italics="true" pageId="103" pageNumber="104">Pupa</emphasis>
. Mostly undescribed.
<emphasis id="B902EAD6FF8BAB129C76126A44DB3A4A" bold="true" box="[553,646,591,617]" pageId="103" pageNumber="104">Thorax</emphasis>
: Scutum, metathorax each with spherical sensory pit (as in Fig. 100A). Respiratory organ (Fig. 101I): Tubular.
<emphasis id="B902EAD6FF8BAB129C881252450C3AB2" bold="true" box="[727,849,631,657]" pageId="103" pageNumber="104">Abdomen</emphasis>
(Fig. 105D): Segments 37 somewhat expanded laterally. Paddle only moderately elongate; apicodorsal thick spine articulating; apicoventral seta longer than thick spine.
</paragraph>
<paragraph id="8BC936C4FF8BAB129E9912D547E33B29" blockId="103.[151,1437,152,1538]" box="[198,446,752,778]" pageId="103" pageNumber="104">
<emphasis id="B902EAD6FF8BAB129E9912D547513B2A" box="[198,268,752,777]" italics="true" pageId="103" pageNumber="104">Larva</emphasis>
. Undescribed.
</paragraph>
<paragraph id="8BC936C4FF8BAB129E99133D47DA3B11" blockId="103.[151,1437,152,1538]" box="[198,391,792,818]" pageId="103" pageNumber="104">
<emphasis id="B902EAD6FF8BAB129E99133D46A83B12" box="[198,245,792,817]" italics="true" pageId="103" pageNumber="104">Egg</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF8BAB129E99136547953C51" pageId="103" pageNumber="104" type="distribution">
<paragraph id="8BC936C4FF8BAB129E99136544AF3BF1" blockId="103.[151,1437,152,1538]" pageId="103" pageNumber="104">
DISTRIBUTION AND BIONOMICS:
<taxonomicName id="4C764D47FF8BAB129CF1136545F73B7A" authorityName="Borkent" authorityYear="2008" box="[686,938,832,857]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="briannae">
<emphasis id="B902EAD6FF8BAB129CF1136545F73B7A" box="[686,938,832,857]" italics="true" pageId="103" pageNumber="104">Corethrella briannae</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="54CD8866FF8BAB129ABB1365434B3B79" box="[1252,1302,832,858]" pageId="103" pageNumber="104">type</typeStatus>
locality in
<collectingCountry id="F3617654FF8BAB129EC8134D46A43BA1" box="[151,249,872,898]" name="Ecuador" pageId="103" pageNumber="104">Ecuador</collectingCountry>
(Fig. 123) at an altitude of
<specimenCount id="9D70FD4DFF8BAB129C1C134D44CB3BA1" box="[579,662,872,898]" pageId="103" pageNumber="104" type="generic">
<quantity id="4C8E9B21FF8BAB129C1C134D44CC3BA1" box="[579,657,872,898]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="103" pageNumber="104" unit="m" value="280.0">280 m</quantity>
.
</specimenCount>
Specimens have been reared from immatures collected from the leaf axils of
<taxonomicName id="4C764D47FF8BAB129F7913B547C43B8A" box="[294,409,912,937]" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Animalia" order="Zingiberales" pageId="103" pageNumber="104" phylum="Tracheophyta" rank="genus">
<emphasis id="B902EAD6FF8BAB129F7913B547C43B8A" box="[294,409,912,937]" italics="true" pageId="103" pageNumber="104">Heliconia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C764D47FF8BAB129F8D13B544203B8A" box="[466,637,912,937]" class="Liliopsida" family="Araceae" genus="Montrichardia" kingdom="Plantae" order="Alismatales" pageId="103" pageNumber="104" phylum="Tracheophyta" rank="genus">
<emphasis id="B902EAD6FF8BAB129F8D13B544203B8A" box="[466,637,912,937]" italics="true" pageId="103" pageNumber="104">Montrichardia</emphasis>
</taxonomicName>
(a terrestrial aroid). The specimens from the Belkin “Mosquitoes of Middle America” project are identified in
<tableCitation id="C6F4037FFF8BAB129CDA139D44B33BF1" box="[645,750,952,978]" captionStart="TABLE 12" captionStartId="295.[151,241,151,175]" captionText="TABLE 12. The identity of specimens of species of Corethrella previously studied by Belkin and his associates for the “Mosquitoes of Middle America” project included in the current study. Abbreviations and collection numbers are provided in (Belkin and Heinemann 1973, 1975a, 1975b, 1976a, 1976b, 1976c; Heinemann and Belkin 1977a, 1977b, 1977c, 1978a, 1978b, 1978c, 1979; Belkin et al. 1970; Heinemann et al. 1980)." pageId="103" pageNumber="104">Table 12</tableCitation>
.
</paragraph>
<paragraph id="8BC936C4FF8BAB129E9913C547953C51" blockId="103.[151,1437,152,1538]" pageId="103" pageNumber="104">
TAXONOMIC DISCUSSION: Males and females were associated through the shared presence of a distinctive pigmentation pattern and were reared together from the same locality. One male
<typeStatus id="54CD8866FF8BAB129AF1142D434E3C01" box="[1198,1299,1032,1058]" pageId="103" pageNumber="104" type="paratype">paratype</typeStatus>
(ECU 164) had deformed gonostyli with these being compressed and more curved apically and one with two subbasal setae (one of these small).
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF8BAB129E9914A5420F3E21" pageId="103" pageNumber="104" type="description">
<paragraph id="8BC936C4FF8BAB129E9914A546A03DA9" blockId="103.[151,1437,152,1538]" pageId="103" pageNumber="104">
TYPES:
<materialsCitation id="3B1E3C99FF8BAB129F6414A547833D31" collectingDate="1966-02-13" collectionCode="USNM" collectorName="J. N. Belkin" country="Ecuador" elevation="280" location="Cochancay" pageId="103" pageNumber="104" specimenCount="2" stateProvince="Canar" typeStatus="holotype">
<typeStatus id="54CD8866FF8BAB129F6414A547F63CB9" box="[315,427,1152,1178]" pageId="103" pageNumber="104" type="holotype">Holotype</typeStatus>
, male adult on microscope slide, larval and pupal exuviae on separate slide, labeled &quot;
<typeStatus id="54CD8866FF8BAB129EC0148D47633CE1" box="[159,318,1192,1218]" pageId="103" pageNumber="104" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C764D47FF8BAB129F19148D44F73CE1" authority="Borkent" authorityName="Borkent" authorityYear="2008" box="[326,682,1192,1218]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="briannae">
<emphasis id="B902EAD6FF8BAB129F19148D441D3CE2" box="[326,576,1192,1217]" italics="true" pageId="103" pageNumber="104">Corethrella briannae</emphasis>
Borkent
</taxonomicName>
,
<locationDeviation id="9A7B2FE6FF8BAB129CE7148D45AF3CE1" box="[696,1010,1192,1218]" location="Guayaquil on Tr." pageId="103" pageNumber="104">
<quantity id="4C8E9B21FF8BAB129CE7148D455E3CE1" box="[696,771,1192,1218]" metricMagnitude="4" metricUnit="m" metricValue="8.6" pageId="103" pageNumber="104" unit="km" value="86.0">86 km</quantity>
E Guayaquil on Tr.
</locationDeviation>
8,
<location id="8EA9601FFF8BAB129A46148D42C13CE1" LSID="urn:lsid:plazi:treatment:03DF87D2FF8AAB129EC811F2420F3E21:8EA9601FFF8BAB129A46148D42C13CE1" box="[1049,1180,1192,1218]" country="Ecuador" name="Cochancay" pageId="103" pageNumber="104" stateProvince="Canar">Cochancay</location>
,
<collectingRegion id="49B2F826FF8BAB129AF5148D42AF3CE1" box="[1194,1266,1192,1218]" country="Ecuador" name="Canar" pageId="103" pageNumber="104">Canar</collectingRegion>
,
<collectingCountry id="F3617654FF8BAB129AA0148D433F3CE1" box="[1279,1378,1192,1218]" name="Ecuador" pageId="103" pageNumber="104">Ecuador</collectingCountry>
,
<quantity id="4C8E9B21FF8BAB129B2F148D46ED3CC9" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="103" pageNumber="104" unit="m" value="280.0">
<elevation id="005BD1F7FF8BAB129B2F148D46ED3CC9" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="103" pageNumber="104" unit="m" value="280.0">280 m</elevation>
</quantity>
,
<date id="FFC81004FF8BAB129E9F14F5471D3CC9" box="[192,320,1232,1258]" pageId="103" pageNumber="104" value="1966-02-13">
<collectingDate id="EF8CE9ECFF8BAB129E9F14F5471D3CC9" box="[192,320,1232,1258]" pageId="103" pageNumber="104" value="1966-02-13">13-II-1966</collectingDate>
</date>
,
<collectorName id="26835312FF8BAB129F0F14F547863CC9" box="[336,475,1232,1258]" pageId="103" pageNumber="104">J.N. Belkin</collectorName>
, Leaf axils of
<taxonomicName id="4C764D47FF8BAB129CD114F5455B3CC9" box="[654,774,1232,1258]" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Animalia" order="Zingiberales" pageId="103" pageNumber="104" phylum="Tracheophyta" rank="genus">Heliconia</taxonomicName>
&quot;, “
<taxonomicName id="4C764D47FF8BAB129D7514F545813CC9" box="[810,988,1232,1258]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B902EAD6FF8BAB129D7514F545EC3CCA" box="[810,945,1232,1257]" italics="true" pageId="103" pageNumber="104">Corethrella</emphasis>
sp.
</taxonomicName>
14”, “ECU 165-20,
<collectingCountry id="F3617654FF8BAB129A8614F543613CC9" box="[1241,1340,1232,1258]" name="Ecuador" pageId="103" pageNumber="104">Ecuador</collectingCountry>
, 6566, Mosq Mid Amer” (
<collectionCode id="ED67AE01FF8BAB129F2514DD47883D31" box="[378,469,1272,1298]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="103" pageNumber="104" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<typeStatus id="54CD8866FF8BAB129FB414DD44113D31" box="[491,588,1272,1298]" pageId="103" pageNumber="104" type="allotype">allotype</typeStatus>
, female adult on microscope slide, larval and pupal exuviae on separate slide, labeled as for
<materialsCitation id="3B1E3C99FF8BAB129FDD150544063D19" box="[386,603,1312,1338]" collectionCode="USNM" country="Ecuador" pageId="103" pageNumber="104" specimenCount="1" stateProvince="Canar" typeStatus="holotype">
<typeStatus id="54CD8866FF8BAB129FDD150547B53D19" box="[386,488,1312,1338]" pageId="103" pageNumber="104" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED67AE01FF8BAB129FA81505440F3D19" box="[503,594,1312,1338]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="103" pageNumber="104" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B1E3C99FF8BAB129C39150546A43DA9" collectionCode="CNCI, USNM" country="Ecuador" pageId="103" pageNumber="104" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Canar" typeStatus="paratype">
<typeStatus id="54CD8866FF8BAB129C39150544813D19" box="[614,732,1312,1338]" pageId="103" pageNumber="104" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D70FD4DFF8BAB129CB8150545503D10" box="[743,781,1312,1338]" pageId="103" pageNumber="104" type="male">2 ♂</specimenCount>
each with pupal exuviae,
<specimenCount id="9D70FD4DFF8BAB129A61150542393D10" box="[1086,1124,1312,1338]" pageId="103" pageNumber="104" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D70FD4DFF8BAB129A2C150542C43D10" box="[1139,1177,1312,1338]" pageId="103" pageNumber="104" type="female">1 ♀</specimenCount>
with pupal exuviae, 1 pupal exuviae, 5 fourth instar larvae, all from type locality (
<specimenCount id="9D70FD4DFF8BAB129D31156D45CA3D78" box="[878,919,1352,1378]" pageId="103" pageNumber="104" type="male">1 ♂</specimenCount>
with pupal exuviae to
<collectionCode id="ED67AE01FF8BAB129AF2156D42AA3D41" box="[1197,1271,1352,1378]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="103" pageNumber="104">CNCI</collectionCode>
; remainder to
<collectionCode id="ED67AE01FF8BAB129EC8155546AC3DA9" box="[151,241,1392,1418]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="103" pageNumber="104" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC936C4FF8BAB129E9915BD420F3E21" blockId="103.[151,1437,152,1538]" pageId="103" pageNumber="104">
DERIVATION OF SPECIFIC EPITHET: The name
<taxonomicName id="4C764D47FF8BAB129D6915BD45FD3D92" authorityName="Borkent" authorityYear="2008" box="[822,928,1432,1457]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="briannae">
<emphasis id="B902EAD6FF8BAB129D6915BD45FD3D92" box="[822,928,1432,1457]" italics="true" pageId="103" pageNumber="104">briannae</emphasis>
</taxonomicName>
is proposed for my wonderful daughter-inlaw Brianna Borkent, not only because I am so proud to have her in my life and to experience her love but also in recognition of her lively spirit and talented writing (as Brianna Brash-Nyberg).
</paragraph>
</subSubSection>
</treatment>
</document>