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112 lines
16 KiB
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<document ID-DOI="10.1093/zoolinnean/zlac062" ID-ISSN="0024-4082" ID-Zenodo-Dep="7381299" approvalRequired="2" approvalRequired_for_taxonomicNames="2" checkinTime="1669808489868" checkinUser="plazi" docAuthor="Norman, David B, Baron, Matthew G, Garcia, Mauricio S & Müller, Rodrigo Temp" docDate="2022" docId="03DF87C3D22A0742AF9FFACEFD10F8AE" docLanguage="en" docName="zlac062.pdf" docOrigin="Zoological Journal of the Linnean Society 196 (4)" docSource="https://academic.oup.com/zoolinnean/article/196/4/1273/6680019" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-.journal_article" docStyleVersion="7" docTitle="Ornithischia" docType="treatment" docVersion="3" lastPageNumber="1286" masterDocId="FFE6FFBBD227074FAF3CFF8BFF8CFFFB" masterDocTitle="Taxonomic, palaeobiological and evolutionary implications of a phylogenetic hypothesis for Ornithischia (Archosauria: Dinosauria)" masterLastPageNumber="1309" masterPageNumber="1273" pageNumber="1286" updateTime="1669992736928" updateUser="diego">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic, palaeobiological and evolutionary implications of a phylogenetic hypothesis for Ornithischia (Archosauria: Dinosauria)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Norman, David B</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Baron, Matthew G</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Garcia, Mauricio S</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Müller, Rodrigo Temp</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>196</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1273</mods:start>
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<mods:end>1309</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>https://academic.oup.com/zoolinnean/article/196/4/1273/6680019</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1093/zoolinnean/zlac062</mods:identifier>
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<mods:identifier type="ISSN">0024-4082</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7381299</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03DF87C3D22A0742AF9FFACEFD10F8AE" httpUri="http://treatment.plazi.org/id/03DF87C3D22A0742AF9FFACEFD10F8AE" lastPageNumber="1286" pageId="13" pageNumber="1286">
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<subSubSection box="[163,607,1349,1373]" pageId="13" pageNumber="1286" type="nomenclature">
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<paragraph blockId="13.[163,607,1349,1373]" box="[163,607,1349,1373]" pageId="13" pageNumber="1286">
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<heading box="[163,607,1349,1373]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="1286" reason="2">
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<emphasis box="[163,607,1349,1373]" italics="true" pageId="13" pageNumber="1286">
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<taxonomicName baseAuthorityName="Seeley" baseAuthorityYear="1888" box="[163,319,1349,1373]" class="Reptilia" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="1286" phylum="Chordata" rank="order">Ornithischia</taxonomicName>
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(
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<figureCitation box="[335,403,1349,1373]" captionStart="Figure 6" captionStartId="10.[145,223,1518,1540]" captionTargetBox="[146,1426,197,1477]" captionTargetId="figure-160@10.[145,1428,195,1478]" captionTargetPageId="10" captionText="Figure 6. Strict consensus treefrom the unconstrained analysis depicting nodenumbers mentioned in thetext and bootstrap values equal to or higher than 40%. Node numbers: 1, Dinosauria; 2, Saurischia; 3, Ornithischia; 4, Sulcimentisauria; 5, Parapredentata; 6, Unnamed; 7, Prionodontia; 8, Thyreophora; 9, Neornithischia; 10, Heterodontosauridae. Saltopus elginensis was removed from the strict consensus tree after an iterPCR analysis (Pol & Escapa, 2009). Silhouettes based on the artwork by Márcio L. Castro, Gabriel Lio, Rodrigo T. Müller, Maurício S. Garcia and John Sibbick." figureDoi="http://doi.org/10.5281/zenodo.7381323" httpUri="https://zenodo.org/record/7381323/files/figure.png" pageId="13" pageNumber="1286">Fig. 6</figureCitation>
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: stem/branch 3)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="1286" type="synonymic_list">
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<paragraph blockId="13.[163,778,1389,1503]" pageId="13" pageNumber="1286">
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<emphasis box="[163,398,1389,1410]" italics="true" pageId="13" pageNumber="1286">Definition adopted:</emphasis>
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The most inclusive clade that includes
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<taxonomicName baseAuthorityName="Boulenger" baseAuthorityYear="1881" box="[280,619,1420,1441]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="1286" phylum="Chordata" rank="species" species="bernissartensis">
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<emphasis box="[280,619,1420,1441]" italics="true" pageId="13" pageNumber="1286">Iguanodon bernissartensis</emphasis>
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</taxonomicName>
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but neither
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<taxonomicName baseAuthorityName="Mantell" baseAuthorityYear="1827" box="[163,474,1450,1471]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="13" pageNumber="1286" phylum="Chordata" rank="species" species="bucklandii">
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<emphasis box="[163,474,1450,1471]" italics="true" pageId="13" pageNumber="1286">Megalosaurus bucklandii</emphasis>
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</taxonomicName>
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nor
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<taxonomicName baseAuthorityName="Hatcher" baseAuthorityYear="1901" box="[535,778,1450,1472]" class="Reptilia" family="Diplodocidae" genus="Diplodocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="13" pageNumber="1286" phylum="Chordata" rank="species" species="carnegii">
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<emphasis box="[535,778,1450,1472]" italics="true" pageId="13" pageNumber="1286">Diplodocus carnegii</emphasis>
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</taxonomicName>
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(after
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<bibRefCitation author="Baron MG & Norman DB & Barrett PM" box="[233,446,1481,1503]" pageId="13" pageNumber="1286" pagination="501 - 506" refId="ref23261" refString="Baron MG, Norman DB, Barrett PM. 2017 a. A new hypothesis of dinosaur relationships and early dinosaur evolution. Nature 543: 501 - 506." type="journal article" year="2017">
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Baron
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<emphasis box="[309,365,1481,1503]" italics="true" pageId="13" pageNumber="1286">et al.</emphasis>
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, 2017a
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="1286" type="description">
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<paragraph blockId="13.[163,779,1549,1878]" pageId="13" pageNumber="1286">
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<emphasis box="[163,410,1549,1571]" italics="true" pageId="13" pageNumber="1286">Characters in support:</emphasis>
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Horizontal or only gently arched premaxillary palate (Char. 17: 0 → 1); strongly curved and hook-shaped jugal process of the ectopterygoid (Char. 57: 0 → 1;
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<figureCitation box="[374,459,1641,1663]" captionStart="Figure 5" captionStartId="9.[165,245,1678,1700]" captionTargetBox="[199,1406,197,1635]" captionTargetId="figure-91@9.[196,1410,195,1638]" captionTargetPageId="9" captionText="Figure 5. Third analysis forcing Chilesaurus diegosuarezi in Theropoda. Abbreviations: Aphan, Aphanosauria; Herrer, Herrerasauridae. Silhouettes are based on artwork by Márcio L. Castro, Gabriel Lio, Rodrigo T. Müller, Maurício S. Garcia, John Sibbick and Douglas M. Heman." figureDoi="http://doi.org/10.5281/zenodo.7381321" httpUri="https://zenodo.org/record/7381321/files/figure.png" pageId="13" pageNumber="1286">Fig. 5E</figureCitation>
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); ventrally inclined dorsal surface of the rostrum of the dentary (Char. 74: 0 → 1); retroarticular is slightly upturned at its distal end (Char. 82: 0 → 1;
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<figureCitation box="[309,401,1733,1755]" captionStart="Figure 7" captionStartId="14.[145,225,1678,1700]" captionTargetBox="[191,1375,197,1633]" captionTargetId="figure-24@14.[190,1380,195,1638]" captionTargetPageId="14" captionText="Figure 7. Body plan evolution within Ornithischia and character states with phylogenetic importance for the present hypothesis. A, skeletal reconstruction of Silesaurus opolensis, a parapredentatan (stem lineage) ornithischian (after Dzik, 2003). B, skeletal reconstruction of Laquintasaura venezuelae, a parapredentatan ornithischian (after Barrett et al., 2014). C, skeletal reconstruction of Eocursor parvus, a prionodontian ornithischian (after Butler et al., 2010). D, left dentary of Kwanasaurus williamparkeri (DMNH EPV.63136) in lateral view (modified from Martz & Small, 2019). E, left ectopterygoid of Asilisaurus kongwe (NMT RB159) in posterior (modified from Nesbitt et al., 2019). F, posterior portion of the right hemimandible of Asilisaurus kongwe (NMT RB159) in lateral (modified from Nesbitt et al., 2019). G, partial left dentary of Asilisaurus kongwe (NMT RB159) in occlusal view (modified from Nesbitt et al., 2019). H, left dentary of Kwanasaurus williamparkeri (DMNH EPV.63136) in medial view (modified from Martz & Small, 2019). I, isolated tooth of Kwanasaurus williamparkeri (DMNH EPV.63843) in labial view (modified from Martz & Small, 2019). J, proximal portion of the right femur of Sacisaurus agudoensis (MCN PV100014) in caudomedial view. K, right maxilla of Echinodon becklesii (NHMUK OR48211) in lateral view (modified from Sereno, 2012). L, right tibia of Eocursor parvus (SAM-PK-K8025) in proximal view. M, left pelvic elements of Heterodontosaurus tucki (SAM-PK-K1332) in lateral view. N, skull of Heterodontosaurus tucki (SAM-PK-K1332) in right lateral view. O, cervical 6 and 7 of Heterodontosaurus tucki (SAM-PK-K1332) in left lateral view. P, proximal portion of the right tibia of Eocursor parvus (SAM-PK-K8025) in lateral view. Q, left ischium of Lesothosaurus diagnosticus (SAM-PK-K1105) in medial view (modified from Baron et al., 2017a). R, proximal portion of the left femur of Lesothosaurus diagnosticus (BP/1/6582) in lateral view. Elements are not to scale." figureDoi="http://doi.org/10.5281/zenodo.7381325" httpUri="https://zenodo.org/record/7381325/files/figure.png" pageId="13" pageNumber="1286">Fig. 7F</figureCitation>
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); premaxilla with four teeth (Char. 84: 0 → 1); teeth ankylosed into the alveoli (Char. 100: 0 → 1;
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<figureCitation box="[374,457,1794,1816]" captionStart="Figure 7" captionStartId="14.[145,225,1678,1700]" captionTargetBox="[191,1375,197,1633]" captionTargetId="figure-24@14.[190,1380,195,1638]" captionTargetPageId="14" captionText="Figure 7. Body plan evolution within Ornithischia and character states with phylogenetic importance for the present hypothesis. A, skeletal reconstruction of Silesaurus opolensis, a parapredentatan (stem lineage) ornithischian (after Dzik, 2003). B, skeletal reconstruction of Laquintasaura venezuelae, a parapredentatan ornithischian (after Barrett et al., 2014). C, skeletal reconstruction of Eocursor parvus, a prionodontian ornithischian (after Butler et al., 2010). D, left dentary of Kwanasaurus williamparkeri (DMNH EPV.63136) in lateral view (modified from Martz & Small, 2019). E, left ectopterygoid of Asilisaurus kongwe (NMT RB159) in posterior (modified from Nesbitt et al., 2019). F, posterior portion of the right hemimandible of Asilisaurus kongwe (NMT RB159) in lateral (modified from Nesbitt et al., 2019). G, partial left dentary of Asilisaurus kongwe (NMT RB159) in occlusal view (modified from Nesbitt et al., 2019). H, left dentary of Kwanasaurus williamparkeri (DMNH EPV.63136) in medial view (modified from Martz & Small, 2019). I, isolated tooth of Kwanasaurus williamparkeri (DMNH EPV.63843) in labial view (modified from Martz & Small, 2019). J, proximal portion of the right femur of Sacisaurus agudoensis (MCN PV100014) in caudomedial view. K, right maxilla of Echinodon becklesii (NHMUK OR48211) in lateral view (modified from Sereno, 2012). L, right tibia of Eocursor parvus (SAM-PK-K8025) in proximal view. M, left pelvic elements of Heterodontosaurus tucki (SAM-PK-K1332) in lateral view. N, skull of Heterodontosaurus tucki (SAM-PK-K1332) in right lateral view. O, cervical 6 and 7 of Heterodontosaurus tucki (SAM-PK-K1332) in left lateral view. P, proximal portion of the right tibia of Eocursor parvus (SAM-PK-K8025) in lateral view. Q, left ischium of Lesothosaurus diagnosticus (SAM-PK-K1105) in medial view (modified from Baron et al., 2017a). R, proximal portion of the left femur of Lesothosaurus diagnosticus (BP/1/6582) in lateral view. Elements are not to scale." figureDoi="http://doi.org/10.5281/zenodo.7381325" httpUri="https://zenodo.org/record/7381325/files/figure.png" pageId="13" pageNumber="1286">Fig. 7G</figureCitation>
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); and medial articular facet of the proximal portion of the femur is straight in caudomedial view (Char. 213: 0 → 1;
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<figureCitation box="[577,653,1856,1878]" captionStart="Figure 7" captionStartId="14.[145,225,1678,1700]" captionTargetBox="[191,1375,197,1633]" captionTargetId="figure-24@14.[190,1380,195,1638]" captionTargetPageId="14" captionText="Figure 7. Body plan evolution within Ornithischia and character states with phylogenetic importance for the present hypothesis. A, skeletal reconstruction of Silesaurus opolensis, a parapredentatan (stem lineage) ornithischian (after Dzik, 2003). B, skeletal reconstruction of Laquintasaura venezuelae, a parapredentatan ornithischian (after Barrett et al., 2014). C, skeletal reconstruction of Eocursor parvus, a prionodontian ornithischian (after Butler et al., 2010). D, left dentary of Kwanasaurus williamparkeri (DMNH EPV.63136) in lateral view (modified from Martz & Small, 2019). E, left ectopterygoid of Asilisaurus kongwe (NMT RB159) in posterior (modified from Nesbitt et al., 2019). F, posterior portion of the right hemimandible of Asilisaurus kongwe (NMT RB159) in lateral (modified from Nesbitt et al., 2019). G, partial left dentary of Asilisaurus kongwe (NMT RB159) in occlusal view (modified from Nesbitt et al., 2019). H, left dentary of Kwanasaurus williamparkeri (DMNH EPV.63136) in medial view (modified from Martz & Small, 2019). I, isolated tooth of Kwanasaurus williamparkeri (DMNH EPV.63843) in labial view (modified from Martz & Small, 2019). J, proximal portion of the right femur of Sacisaurus agudoensis (MCN PV100014) in caudomedial view. K, right maxilla of Echinodon becklesii (NHMUK OR48211) in lateral view (modified from Sereno, 2012). L, right tibia of Eocursor parvus (SAM-PK-K8025) in proximal view. M, left pelvic elements of Heterodontosaurus tucki (SAM-PK-K1332) in lateral view. N, skull of Heterodontosaurus tucki (SAM-PK-K1332) in right lateral view. O, cervical 6 and 7 of Heterodontosaurus tucki (SAM-PK-K1332) in left lateral view. P, proximal portion of the right tibia of Eocursor parvus (SAM-PK-K8025) in lateral view. Q, left ischium of Lesothosaurus diagnosticus (SAM-PK-K1105) in medial view (modified from Baron et al., 2017a). R, proximal portion of the left femur of Lesothosaurus diagnosticus (BP/1/6582) in lateral view. Elements are not to scale." figureDoi="http://doi.org/10.5281/zenodo.7381325" httpUri="https://zenodo.org/record/7381325/files/figure.png" pageId="13" pageNumber="1286">Fig. 7J</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |