298 lines
35 KiB
XML
298 lines
35 KiB
XML
<document id="656A310EBFDEB9AACCABF3EADC39CB28" ID-CLB-Dataset="22084" ID-DOI="10.5852/ejt.2018.446" ID-GBIF-Dataset="08fb092e-b3df-47c4-a1ad-8ea8ffd2457a" ID-Zenodo-Dep="3829902" ID-ZooBank="56E50F4E-6A7E-4CE6-963E-3B49AA7A03B6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589575450896" checkinUser="valdenar" docAuthor="Schmidt, Joachim & Faille, Arnaud" docDate="2018" docId="03DF87A4FFC3FFA3FD64FD3DFE285D84" docLanguage="en" docName="ejt-446_schmidt_faille.pdf.imf" docOrigin="European Journal of Taxonomy 446" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Trechus nanulus Schmidt & Faille 2018, sp. nov." docType="treatment" docUuid="6C2C58F5-6A6D-4FA3-B32B-486736DFBD91" docUuidSource="ZooBank" docVersion="6" lastPageNumber="15" masterDocId="FFE6FFDCFFCFFFADFFE6F953FFAE5C66" masterDocTitle="Revision of Trechus Clairville, 1806 of the Bale Mountains and adjacent volcanos, Ethiopia (Coleoptera, Carabidae, Trechini)" masterLastPageNumber="82" masterPageNumber="1" pageNumber="13" updateTime="1698839989887" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="7B3BB21CC2F08EF4426AB6D10E24B2B3">
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<mods:title id="3035BC259E8D899964E099E025B4E453">Revision of Trechus Clairville, 1806 of the Bale Mountains and adjacent volcanos, Ethiopia (Coleoptera, Carabidae, Trechini)</mods:title>
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<mods:name id="EF971C8473ADE3AD56DE2532BBC61753" type="personal">
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<mods:role id="DB4508528E2016E4D8D1BACB313BACC6">
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<mods:roleTerm id="E86DDAE4162364AD6B66C9B4FAD43B2B">Author</mods:roleTerm>
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<mods:namePart id="0F85DA0BB21C0FD35D55B4660B956543">Schmidt, Joachim</mods:namePart>
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<mods:affiliation id="3B31FB30A2C0796D69F0A12A6F3E295A">University of Rostock, Institute of Biosciences, General and Systematic Zoology, Universitätsplatz 2, 18055 Rostock, Germany. University of Marburg, Fb. 17 - Biologie, Karl-von-Frisch-Strasse 8, 35043 Marburg, Germany. Institute of Evolutionary Biology (CSIC-Universitat Pompeu Fabra), Passeig Maritim de la Barceloneta 37, 08003 Barcelona, Spain. MECADEV - UMR 7179 MNHN / CNRS, Paris, France.</mods:affiliation>
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<mods:nameIdentifier id="BAAD540F5A49BEA22BA88948450F4DD9" type="email">schmidt@agonum.de</mods:nameIdentifier>
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<mods:roleTerm id="A9146BC8195CC8BD09BE9B4F17B6E910">Author</mods:roleTerm>
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<mods:namePart id="3B635435A7970BE8F2EB0CB51FE8DCD2">Faille, Arnaud</mods:namePart>
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<mods:nameIdentifier id="15A5A7B37166D05D2D1AC71F54852AE0" type="email">arnaud1140@yahoo.fr</mods:nameIdentifier>
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<mods:typeOfResource id="B4D889B65A2318A180A2620604B36479">text</mods:typeOfResource>
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<mods:title id="573D60065051D326A00325BC03E58F6E">European Journal of Taxonomy</mods:title>
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<mods:part id="72D02620EDEA3DA6715E5C33BCAC3D7C">
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<mods:date id="CAEEF06E69DFFC459159AFF7B3C051F1">2018</mods:date>
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<mods:number id="3FB6EC686E635A9A670C694A3A7D15F4">2018-06-21</mods:number>
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<mods:number id="8B826ADA097A22133729A10824C88AD5">446</mods:number>
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<mods:identifier id="A25A47C6D5493E2CD365ED03220F0CAC" type="DOI">10.5852/ejt.2018.446</mods:identifier>
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<treatment id="03DF87A4FFC3FFA3FD64FD3DFE285D84" ID-DOI="http://doi.org/10.5281/zenodo.3848571" ID-GBIF-Taxon="164165564" ID-Zenodo-Dep="3848571" LSID="urn:lsid:zoobank.org:act:6C2C58F5-6A6D-4FA3-B32B-486736DFBD91" httpUri="http://treatment.plazi.org/id/03DF87A4FFC3FFA3FD64FD3DFE285D84" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
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<subSubSection id="C36C6539FFC3FFA1FD64FD3DFB6758CA" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="8BC936B2FFC3FFA1FD64FD3DFC1F58EF" blockId="12.[363,1225,1134,1231]" box="[642,945,1134,1161]" pageId="12" pageNumber="13">
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<heading id="D08181DEFFC3FFA1FD64FD3DFC1F58EF" box="[642,945,1134,1161]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
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<taxonomicName id="4C764D31FFC3FFA1FD64FD3DFCE158EF" ID-CoL="8MJ7S" authority="Schmidt & Faille, 2018" authorityName="Schmidt & Faille" authorityYear="2018" box="[642,847,1134,1161]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nanulus" status="sp. nov.">
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<emphasis id="B902EAA0FFC3FFA1FD64FD3DFCE158EF" bold="true" box="[642,847,1134,1161]" italics="true" pageId="12" pageNumber="13">Trechus nanulus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23157DBFFC3FFA1FCB1FD3CFC1F58EF" box="[855,945,1135,1161]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8BC936B2FFC3FFA1FE8DFDC2FB6758CA" blockId="12.[363,1225,1134,1231]" box="[363,1225,1169,1196]" pageId="12" pageNumber="13">
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<uri id="FFE73AB0FFC3FFA1FE8DFDC2FB6758CA" box="[363,1225,1169,1196]" pageId="12" pageNumber="13">
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urn:lsid:zoobank.org:act:
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<uuid id="FFD00C67FFC3FFA1FD6CFDC2FB6758CA" box="[650,1225,1169,1196]" pageId="12" pageNumber="13">6C2C58F5-6A6D-4FA3-B32B-486736DFBD91</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C6539FFC3FFA1FD43FDE6FC2058A9" box="[677,910,1205,1231]" pageId="12" pageNumber="13" type="description">
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<paragraph id="8BC936B2FFC3FFA1FD43FDE6FC2058A9" blockId="12.[363,1225,1134,1231]" box="[677,910,1205,1231]" pageId="12" pageNumber="13">
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<figureCitation id="134D2A37FFC3FFA1FD43FDE6FD5E58A9" box="[677,752,1205,1231]" captionStart="Figs 1–4" captionStartId="5.[189,241,1965,1991]" captionTargetBox="[279,1312,321,1921]" captionTargetId="figure@5.[276,1312,317,1924]" captionTargetPageId="5" captionText="Figs 1–4. Trechus spp., habitus. 1. T. mattisi sp. nov., paratype, ³. 2. T. transversicollis sp. nov., holotype. 3. T. minitrechus sp. nov., paratype, ³. 4. T. nanulus sp. nov., paratype, ³." figureDoi="http://doi.org/10.5281/zenodo.3829904" httpUri="https://zenodo.org/record/3829904/files/figure.png" pageId="12" pageNumber="13">Figs 4</figureCitation>
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,
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<figureCitation id="134D2A37FFC3FFA1FD1AFDE6FCA258A9" box="[764,780,1205,1231]" captionStart="Figs 5–8" captionStartId="7.[189,241,1965,1991]" captionTargetBox="[271,1316,318,1925]" captionTargetId="figure@7.[271,1316,317,1925]" captionTargetPageId="7" captionText="Figs 5–8. Trechus spp., head, pronotum. 5. T. mattisi sp. nov., paratype, ³. 6. T. transversicollis sp. nov., holotype. 7. T. minitrechus sp. nov., paratype, ³. 8. T. nanulus sp. nov., paratype, ³." figureDoi="http://doi.org/10.5281/zenodo.3829906" httpUri="https://zenodo.org/record/3829906/files/figure.png" pageId="12" pageNumber="13">8</figureCitation>
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,
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<figureCitation id="134D2A37FFC3FFA1FCFFFDE6FC9758A9" box="[793,825,1205,1231]" captionStart="Figs 9–12" captionStartId="9.[189,241,1930,1956]" captionTargetBox="[189,1375,351,1857]" captionTargetId="figure@9.[189,1398,351,1857]" captionTargetPageId="9" captionText="Figs 9–12. Trechus spp., elytra. 9. T. mattisi sp. nov., paratype, ³. 10. T. transversicollis sp. nov., holotype. 11. T. minitrechus sp. nov., paratype, ³. 12. T. nanulus sp. nov., paratype, ³. The arrows point to the insertions of the discal setae, preapical seta (if present), and setae of the umbilical series." figureDoi="http://doi.org/10.5281/zenodo.3829908" httpUri="https://zenodo.org/record/3829908/files/figure.png" pageId="12" pageNumber="13">12</figureCitation>
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,
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<figureCitation id="134D2A37FFC3FFA1FCA3FDE6FC2058A9" box="[837,910,1205,1231]" captionStart="Figs 37–56" captionStartId="28.[189,241,1788,1814]" captionTargetBox="[273,1317,319,1746]" captionTargetId="figure@28.[271,1317,317,1746]" captionTargetPageId="28" captionText="Figs 37–56. Trechus spp., aedeagal median lobe, left lateral view (37, 39, 40, 42, 44–45, 47–48, 50, 52–53, 55–56) and dorsal view (38, 41, 43, 46, 49, 51, 54). 37–38. T. mattisi sp. nov., paratype. 39. T. transversicollis sp. nov., holotype. 40–41. T. minitrechus sp. nov., paratype. 42–43. T. nanulus sp. nov., paratype. 44. T. rira sp. nov., holotype. 45–46. T. iridescens sp. nov., holotype. 47. T. relictus Magrini, Quéinnec & Vigna Taglianti, 2012, specimen from brook valley above Goba. 48–49. T. oppositus sp. nov., paratype. 50–51. T. harryi sp. nov., paratype. 52. T. dodola sp. nov., holotype. 53–54. T. adaba sp. nov., paratype. 55. T. harenna sp. nov., paratype. 56. T. bombi sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829924" httpUri="https://zenodo.org/record/3829924/files/figure.png" pageId="12" pageNumber="13">42–43</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C6539FFC3FFA1FF5BFDA4FB8A5A3B" pageId="12" pageNumber="13" type="diagnosis">
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<paragraph id="8BC936B2FFC3FFA1FF5BFDA4FE985977" blockId="12.[189,310,1271,1297]" box="[189,310,1271,1297]" pageId="12" pageNumber="13">
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<heading id="D08181DEFFC3FFA1FF5BFDA4FE985977" bold="true" box="[189,310,1271,1297]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
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<emphasis id="B902EAA0FFC3FFA1FF5BFDA4FE985977" bold="true" box="[189,310,1271,1297]" pageId="12" pageNumber="13">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC936B2FFC3FFA1FF5BFC75FB8A5A3B" blockId="12.[189,1399,1318,1629]" pageId="12" pageNumber="13">
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Habitus and male genital characters very similar to
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<taxonomicName id="4C764D31FFC3FFA1FCDDFC74FC1C5926" authorityName="Vigna Taglianti & Magrini" authorityYear="2010" box="[827,946,1319,1344]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="gypaeti">
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<emphasis id="B902EAA0FFC3FFA1FCDDFC74FC1C5926" box="[827,946,1319,1344]" italics="true" pageId="12" pageNumber="13">T. gypaeti</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C764D31FFC3FFA1FC12FC74FB305926" authority="Schmidt & Faille, 2018" authorityName="Schmidt & Faille" authorityYear="2018" box="[1012,1182,1318,1344]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="minitrechus" status="sp. nov.">
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<emphasis id="B902EAA0FFC3FFA1FC12FC74FB305926" box="[1012,1182,1318,1344]" italics="true" pageId="12" pageNumber="13">T. minitrechus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23157DBFFC3FFA1FB4CFC75FAAB5926" box="[1194,1285,1318,1344]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
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from Mt Enkuolo, but slightly smaller, colour of body distinctly darker brown and shinier with elytra iridescent, pronotum and elytra with meshes of microsculpture less deeply engraved, eyes larger and more protruded, pronotum more transverse, elytra broader and shorter with preapical seta developed, and aedeagal median lobe stouter.
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<taxonomicName id="4C764D31FFC3FFA1FDFBFCE7FD7059A8" authority="Schmidt & Faille, 2018" authorityName="Schmidt & Faille" authorityYear="2018" box="[541,734,1460,1486]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nanulus" status="sp. nov.">
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<emphasis id="B902EAA0FFC3FFA1FDFBFCE7FD7059A8" box="[541,734,1460,1486]" italics="true" pageId="12" pageNumber="13">Trechus nanulus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23157DBFFC3FFA1FD03FCE6FC9159A9" box="[741,831,1461,1487]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
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is also very similar to
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<taxonomicName id="4C764D31FFC3FFA1FBA1FCE6FB3F59A8" authority="Schmidt & Faille, 2018" authorityName="Schmidt & Faille" authorityYear="2018" box="[1095,1169,1461,1486]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="rira" status="sp. nov.">
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<emphasis id="B902EAA0FFC3FFA1FBA1FCE6FB3F59A8" box="[1095,1169,1461,1486]" italics="true" pageId="12" pageNumber="13">T. rira</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23157DBFFC3FFA1FB7EFCE6FB5C59A9" box="[1176,1266,1461,1487]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
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but readily distinguished by larger eyes and stouter aedeagal median lobe (see below for details). Compared to all other
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<taxonomicName id="4C764D31FFC3FFA1FECBFCA8FE245A73" authorityName="Clairville" authorityYear="1806" box="[301,394,1531,1557]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="B902EAA0FFC3FFA1FECBFCA8FE245A73" box="[301,394,1531,1557]" italics="true" pageId="12" pageNumber="13">Trechus</emphasis>
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</taxonomicName>
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species of the Ethiopian fauna, the new species differs by the combination of the following characters: body length below
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<quantity id="4C8E9B57FFC3FFA1FD42FF4CFCAC5A5C" box="[676,770,1567,1594]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="13" unit="mm" value="2.5">2.5 mm</quantity>
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, only one protarsomere of male dilated, elytral striae indistinctly impressed on disc; aedeagal median lobe apex simply tapered.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C6539FFC3FFA1FF5BFFD6FC3F5A97" pageId="12" pageNumber="13" type="etymology">
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<paragraph id="8BC936B2FFC3FFA1FF5BFFD6FEEA5AF9" blockId="12.[189,324,1669,1695]" box="[189,324,1669,1695]" pageId="12" pageNumber="13">
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<heading id="D08181DEFFC3FFA1FF5BFFD6FEEA5AF9" bold="true" box="[189,324,1669,1695]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
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<emphasis id="B902EAA0FFC3FFA1FF5BFFD6FEEA5AF9" bold="true" box="[189,324,1669,1695]" pageId="12" pageNumber="13">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC936B2FFC3FFA1FF5BFFE0FC3F5A97" blockId="12.[189,1398,1715,1777]" pageId="12" pageNumber="13">
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The specific epithet refers to the very small body length of the species and particularly to the fact that it represents the smallest
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<taxonomicName id="4C764D31FFC3FFA1FE28FF84FD855A97" authorityName="Clairville" authorityYear="1806" box="[462,555,1751,1777]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="B902EAA0FFC3FFA1FE28FF84FD855A97" box="[462,555,1751,1777]" italics="true" pageId="12" pageNumber="13">Trechus</emphasis>
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</taxonomicName>
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species known from
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<collectingCountry id="F3617622FFC3FFA1FCC0FF84FC235A97" box="[806,909,1751,1777]" name="Ethiopia" pageId="12" pageNumber="13">Ethiopia</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C6539FFC3FFA0FF5BFE4AFB735DD4" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="materials_examined">
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<paragraph id="8BC936B2FFC3FFA1FF5BFE4AFE045B55" blockId="12.[189,426,1817,1843]" box="[189,426,1817,1843]" pageId="12" pageNumber="13">
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<heading id="D08181DEFFC3FFA1FF5BFE4AFE045B55" bold="true" box="[189,426,1817,1843]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
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<emphasis id="B902EAA0FFC3FFA1FF5BFE4AFE045B55" bold="true" box="[189,426,1817,1843]" pageId="12" pageNumber="13">Material examined</emphasis>
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</heading>
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</paragraph>
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<materialsCitation id="3B1E3CEFFFC3FFA1FF0CFE1BFCEB5BCF" ID-GBIF-Occurrence="2620317305" collectingDate="2006-12" collectionCode="CSCHM, ZSM" country="Ethiopia" elevation="2380" latitude="6.721389" location="Rira" longLatPrecision="21" longitude="39.720554" municipality="Bale Mts" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Oromiya" typeStatus="holotype">
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<paragraph id="8BC936B2FFC3FFA1FF0CFE1BFEF25B04" blockId="12.[189,1399,1864,1961]" box="[234,348,1864,1890]" pageId="12" pageNumber="13">
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<emphasis id="B902EAA0FFC3FFA1FF0CFE1BFEF25B04" bold="true" box="[234,348,1864,1890]" pageId="12" pageNumber="13">
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<typeStatus id="54CD8810FFC3FFA1FF0CFE1BFEF25B04" box="[234,348,1864,1890]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC936B2FFC3FFA1FF5BFE38FCEB5BCF" blockId="12.[189,1399,1864,1961]" pageId="12" pageNumber="13">
|
||
<collectingCountry id="F3617622FFC3FFA1FF5BFE38FEE75BE3" box="[189,329,1899,1925]" name="Ethiopia" pageId="12" pageNumber="13">ETHIOPIA</collectingCountry>
|
||
: ³,
|
||
<collectingRegion id="49B2F850FFC3FFA1FE90FE38FE7D5BE3" box="[374,467,1899,1925]" country="Ethiopia" name="Oromiya" pageId="12" pageNumber="13">Oromia</collectingRegion>
|
||
,
|
||
<collectionCode id="ED67AE77FFC3FFA1FE3BFE38FE435BE3" box="[477,493,1899,1925]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="12" pageNumber="13" type="Herbarium">S</collectionCode>
|
||
slope
|
||
<collectingMunicipality id="6BADACC8FFC3FFA1FDD1FE38FD0F5BE0" box="[567,673,1899,1926]" pageId="12" pageNumber="13">Bale Mts</collectingMunicipality>
|
||
below
|
||
<location id="8EA96069FFC3FFA1FD13FE38FC845BE3" LSID="urn:lsid:plazi:treatment:03DF87A4FFC3FFA3FD64FD3DFE285D84:8EA96069FFC3FFA1FD13FE38FC845BE3" box="[757,810,1899,1925]" country="Ethiopia" latitude="6.721389" longLatPrecision="21" longitude="39.720554" municipality="Bale Mts" name="Rira" pageId="12" pageNumber="13" stateProvince="Oromiya">Rira</location>
|
||
, alt.
|
||
<elevation id="005BD181FFC3FFA1FCB9FE38FC175BE0" box="[863,953,1899,1926]" metricMagnitude="3" metricUnit="m" metricValue="2.38" pageId="12" pageNumber="13" unit="m" value="2380.0">
|
||
<quantity id="4C8E9B57FFC3FFA1FCB9FE38FC175BE0" box="[863,953,1899,1926]" metricMagnitude="3" metricUnit="m" metricValue="2.38" pageId="12" pageNumber="13" unit="m" value="2380.0">2380 m</quantity>
|
||
</elevation>
|
||
,
|
||
<geoCoordinate id="EE425075FFC3FFA1FC22FE3AFBF75BE0" box="[964,1113,1897,1926]" degrees="06" direction="north" minutes="43" orientation="latitude" pageId="12" pageNumber="13" precision="15" seconds="17" value="6.721389">
|
||
06°43′17″
|
||
<collectionCode id="ED67AE77FFC3FFA1FBA7FE3FFBF75BE0" box="[1089,1113,1900,1926]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="12" pageNumber="13" type="Herbarium">N</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE425075FFC3FFA1FB85FE3AFB5A5BE0" box="[1123,1268,1897,1926]" degrees="39" direction="east" minutes="43" orientation="longitude" pageId="12" pageNumber="13" precision="15" seconds="14" value="39.720554">
|
||
39°43′14″
|
||
<collectionCode id="ED67AE77FFC3FFA1FB06FE3FFB5A5BE0" box="[1248,1268,1900,1926]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="12" pageNumber="13">E</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<date id="FFC81072FFC3FFA1FB19FE3FFAD95BE3" box="[1279,1399,1899,1926]" pageId="12" pageNumber="13" value="2006-12">
|
||
<collectingDate id="EF8CE99AFFC3FFA1FB19FE3FFAD95BE3" box="[1279,1399,1899,1926]" pageId="12" pageNumber="13" value="2006-12">Dec. 2006</collectingDate>
|
||
</date>
|
||
(
|
||
<collectionCode id="ED67AE77FFC3FFA1FF23FEDCFE9E5BCF" box="[197,304,1935,1961]" pageId="12" pageNumber="13">CSCHM</collectionCode>
|
||
, registration number
|
||
<collectionCode id="ED67AE77FFC3FFA1FDCCFEDCFDC65BCF" box="[554,616,1935,1961]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="12" pageNumber="13" type="Museum">ZSM</collectionCode>
|
||
_COL_2018_004).
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="8BC936B2FFC2FFA0FF0CF858FEC95D43" blockId="13.[189,1399,267,435]" box="[234,359,267,293]" pageId="13" pageNumber="14">
|
||
<emphasis id="B902EAA0FFC2FFA0FF0CF858FEC95D43" bold="true" box="[234,359,267,293]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54CD8810FFC2FFA0FF0CF858FEC95D43" box="[234,359,267,293]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC936B2FFC2FFA0FF5BF87DFB735DD4" blockId="13.[189,1399,267,435]" pageId="13" pageNumber="14">
|
||
<collectingCountry id="F3617622FFC2FFA0FF5BF87DFEE75D2E" box="[189,329,302,328]" name="Ethiopia" pageId="13" pageNumber="14">ETHIOPIA</collectingCountry>
|
||
: 3 ³³,
|
||
<specimenCount id="9D70FD3BFFC2FFA0FE43F87DFE465D2F" box="[421,488,302,329]" pageId="13" pageNumber="14" type="female">3 ♀♀</specimenCount>
|
||
, same data as for
|
||
<typeStatus id="54CD8810FFC2FFA0FD5DF87DFC8C5D2E" box="[699,802,302,328]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="ED67AE77FFC2FFA0FCD7F87DFCC25D2E" box="[817,876,302,328]" pageId="13" pageNumber="14">CAF</collectionCode>
|
||
,
|
||
<collectionCode id="ED67AE77FFC2FFA0FC9EF87DFC4A5D2E" box="[888,996,302,328]" pageId="13" pageNumber="14">CSCHM</collectionCode>
|
||
). 7 ³³,
|
||
<specimenCount id="9D70FD3BFFC2FFA0FBAFF87DFB225D2F" box="[1097,1164,302,329]" pageId="13" pageNumber="14" type="female">6 ♀♀</specimenCount>
|
||
, Bale Mts, Harenna Forest, E Rira, alt.
|
||
<quantity id="4C8E9B57FFC2FFA0FE71F802FE5F5D0A" box="[407,497,337,364]" metricMagnitude="3" metricUnit="m" metricValue="2.69" pageId="13" pageNumber="14" unit="m" value="2690.0">2690 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE425075FFC2FFA0FE1BF803FD3F5D0A" box="[509,657,336,364]" degrees="06" direction="north" minutes="43" orientation="latitude" pageId="13" pageNumber="14" precision="15" seconds="25" value="6.7236114">
|
||
06°43′25″
|
||
<collectionCode id="ED67AE77FFC2FFA0FD9FF801FD3F5D0A" box="[633,657,338,364]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="13" pageNumber="14" type="Herbarium">N</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE425075FFC2FFA0FD7AF803FC835D0A" box="[668,813,336,364]" degrees="39" direction="east" minutes="45" orientation="longitude" pageId="13" pageNumber="14" precision="15" seconds="25" value="39.756943">
|
||
39°45′25″
|
||
<collectionCode id="ED67AE77FFC2FFA0FCFFF801FC835D0A" box="[793,813,338,364]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="13" pageNumber="14">E</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<date id="FFC81072FFC2FFA0FCDEF801FC035D0A" box="[824,941,338,364]" pageId="13" pageNumber="14" value="2007-02">Feb. 2007</date>
|
||
(
|
||
<collectionCode id="ED67AE77FFC2FFA0FC5DF801FC585D0A" box="[955,1014,338,364]" pageId="13" pageNumber="14">CAF</collectionCode>
|
||
,
|
||
<collectionCode id="ED67AE77FFC2FFA0FBE7F801FBC35D0A" box="[1025,1133,338,364]" pageId="13" pageNumber="14">CSCHM</collectionCode>
|
||
); 37 ³³,
|
||
<specimenCount id="9D70FD3BFFC2FFA0FB3BF801FA805D0A" box="[1245,1326,338,364]" pageId="13" pageNumber="14" type="female">
|
||
29
|
||
<specimenCount id="9D70FD3BFFC2FFA0FAE7F801FA805D0A" box="[1281,1326,338,364]" pageId="13" pageNumber="14">♀♀</specimenCount>
|
||
</specimenCount>
|
||
, same data as for preceding but
|
||
<date id="FFC81072FFC2FFA0FE06F826FDD55DE9" box="[480,635,373,400]" pageId="13" pageNumber="14" value="2017-12-13">13 Dec. 2017</date>
|
||
(
|
||
<collectionCode id="ED67AE77FFC2FFA0FD6FF826FD535DE9" box="[649,765,373,399]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="13" pageNumber="14" type="Museum">NHMUK</collectionCode>
|
||
,
|
||
<collectionCode id="ED67AE77FFC2FFA0FCE1F826FCEF5DE9" box="[775,833,373,399]" pageId="13" pageNumber="14">CAF</collectionCode>
|
||
,
|
||
<collectionCode id="ED67AE77FFC2FFA0FCADF826FC225DE9" box="[843,908,373,399]" pageId="13" pageNumber="14">CDH</collectionCode>
|
||
,
|
||
<collectionCode id="ED67AE77FFC2FFA0FC71F826FBAF5DE9" box="[919,1025,373,399]" pageId="13" pageNumber="14">CSCHM</collectionCode>
|
||
,
|
||
<collectionCode id="ED67AE77FFC2FFA0FBEDF825FBC65DF6" box="[1035,1128,374,400]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="14">MNHN</collectionCode>
|
||
,
|
||
<collectionCode id="ED67AE77FFC2FFA0FB94F826FB605DE9" box="[1138,1230,373,399]" pageId="13" pageNumber="14">ZMAA</collectionCode>
|
||
); 7 ³³,
|
||
<specimenCount id="9D70FD3BFFC2FFA0FAC8F826FADE5DF6" box="[1326,1392,373,400]" pageId="13" pageNumber="14" type="female">
|
||
8
|
||
<specimenCount id="9D70FD3BFFC2FFA0FAA5F825FADE5DF6" box="[1347,1392,374,400]" pageId="13" pageNumber="14">♀♀</specimenCount>
|
||
</specimenCount>
|
||
, Harenna Forest,
|
||
<collectionCode id="ED67AE77FFC2FFA0FE99F8CAFE345DD5" box="[383,410,409,435]" country="Austria" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="13" pageNumber="14" type="Herbarium">W</collectionCode>
|
||
Rira, alt.
|
||
<quantity id="4C8E9B57FFC2FFA0FDE9F8CBFDC45DD5" box="[527,618,408,435]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="14" unit="m" value="3000.0">3000 m</quantity>
|
||
,
|
||
<geoCoordinate id="EE425075FFC2FFA0FD90F8C4FCA25DD5" box="[630,780,407,435]" degrees="06" direction="north" minutes="45" orientation="latitude" pageId="13" pageNumber="14" precision="15" seconds="57" value="6.7658334">
|
||
06°45′57″
|
||
<collectionCode id="ED67AE77FFC2FFA0FD12F8CAFCA25DD5" box="[756,780,409,435]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="13" pageNumber="14" type="Herbarium">N</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE425075FFC2FFA0FCFEF8C4FC045DD5" box="[792,938,407,435]" degrees="39" direction="east" minutes="41" orientation="longitude" pageId="13" pageNumber="14" precision="15" seconds="47" value="39.69639">
|
||
39°41′47″
|
||
<collectionCode id="ED67AE77FFC2FFA0FC70F8CAFC045DD5" box="[918,938,409,435]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="13" pageNumber="14">E</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<date id="FFC81072FFC2FFA0FC50F8CBFBFA5DD4" box="[950,1108,408,435]" pageId="13" pageNumber="14" value="2017-12-14">14 Dec. 2017</date>
|
||
(
|
||
<collectionCode id="ED67AE77FFC2FFA0FB85F8CBFB7F5DD4" box="[1123,1233,408,434]" pageId="13" pageNumber="14">CSCHM</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C6539FFC2FFA3FF5BF889FAE25D2C" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
|
||
<paragraph id="8BC936B2FFC2FFA0FF5BF889FEE05D92" blockId="13.[189,334,474,500]" box="[189,334,474,500]" pageId="13" pageNumber="14">
|
||
<heading id="D08181DEFFC2FFA0FF5BF889FEE05D92" bold="true" box="[189,334,474,500]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
|
||
<emphasis id="B902EAA0FFC2FFA0FF5BF889FEE05D92" bold="true" box="[189,334,474,500]" pageId="13" pageNumber="14">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC936B2FFC2FFA0FF5BFB5AFCA55E45" blockId="13.[189,779,519,547]" box="[189,779,519,547]" pageId="13" pageNumber="14">
|
||
BODY LENGTH.
|
||
<quantity id="4C8E9B57FFC2FFA0FE83FB5AFE5C5E45" box="[357,498,521,547]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="13" pageNumber="14" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.2–2.4 mm</quantity>
|
||
(Ø =
|
||
<quantity id="4C8E9B57FFC2FFA0FDDEFB5AFD0D5E45" box="[568,675,521,547]" metricMagnitude="-3" metricUnit="m" metricValue="2.29" pageId="13" pageNumber="14" unit="mm" value="2.29">2.29 mm</quantity>
|
||
, n = 20).
|
||
</paragraph>
|
||
<paragraph id="8BC936B2FFC2FFA0FF5BFB18FB6A5EEE" blockId="13.[189,1399,584,649]" pageId="13" pageNumber="14">PROPORTIONS (n = 10): PW/HW = 1.35–1.42 (Ø = 1.38); PW/PL = 1.50–1.60 (Ø = 1.56); PW/PBW = 1.22–1.30 (Ø = 1.25); EW/PW = 1.41–1.51 (Ø = 1.46); EL/EW = 1.24–1.31 (Ø = 1.28).</paragraph>
|
||
<paragraph id="8BC936B2FFC2FFA0FF5BFBFCFEF25F77" blockId="13.[189,1398,687,785]" pageId="13" pageNumber="14">COLOUR. Head, pronotum and elytra dark brown, markedly shiny, elytra slightly iridescent. Clypeus, labrum, palpi, antennal base and legs yellowish; antennae distally darkened beginning from third antennomere.</paragraph>
|
||
<paragraph id="8BC936B2FFC2FFA0FF5BFA6BFB0E5FFF" blockId="13.[189,1398,824,921]" pageId="13" pageNumber="14">MICROSCULPTURE. Same in males and females. Head with comparatively large, almost isodiametric meshes on disc and supraorbital area, and smaller meshes on clypeus. Pronotum and elytra with slightly engraved narrow transverse meshes, which are narrower on elytra than on pronotum.</paragraph>
|
||
<paragraph id="8BC936B2FFC2FFA0FF5BFA92FAD95801" blockId="13.[189,1399,960,1128]" pageId="13" pageNumber="14">HEAD. Comparatively robust. Mandibles short. Labrum with apical margin widely emarginated. Eyes moderately small, distinctly convexly protruded. Tempora convex, markedly wrinkled to the neck, 0.9 times as long as eyes, very finely and sparsely pubescent. Frons and supraorbital area strongly convex, with supraorbital furrows almost uniformly bent on disc, deep throughout. Antennae short, proportions of the first four antennomeres as follows: 1/1/0.9/0.7; antennomeres 7–9 only slightly longer than broad.</paragraph>
|
||
<paragraph id="8BC936B2FFC2FFA0FF5BFDC3FEB459C7" blockId="13.[189,1399,1167,1441]" pageId="13" pageNumber="14">PROTHORAX. Pronotum large, markedly transverse, broadest portion slightly before middle, base as wide as apical margin. Disc markedly convex. Anterior margin slightly concave with anterior angles slightly protruded. Sides almost evenly rounded throughout. Laterobasal angles very obtuse, blunt at apex. Marginal gutter very narrow, not widened near laterobasal angles. Base straight in middle, distinctly bent anteriorly at outer quarter. Median longitudinal impression fine but distinct, not deepened near base, disappearing at apex; anterior transverse impression very shallow, smooth; posterior transverse impression shallow, laterobasal foveae very small, notch-like, smooth. Pronotum with laterobasal setae present.</paragraph>
|
||
<paragraph id="8BC936B2FFC2FFA0FF5BFC9AFDF25B47" blockId="13.[189,1399,1480,1825]" pageId="13" pageNumber="14">
|
||
PTEROTHORAX. Elytra convex on disc, in dorsal view markedly short and broad, broadest in mid-length, with shoulders fully rounded; broadly rounded at apex. Striae impunctate, parascutellar stria and first stria absent, latter sometimes finely impressed near apex, striae 2–5 indistinct but present, striae 6–7 usually absent, stria 8 impressed only at levels of the middle and subapical groups of the marginal umbilicate pores. All intervals flat. Recurrent preapical stria deep, curved in front, directed to the fifth stria. Third stria with two setiferous dorsal pores, the anterior one slightly behind the anterior elytral quarter, the posterior one slightly behind middle (except in one specimen without posterior discal seta!), and with preapical seta which is situated at the apical anastomosis of second and third striae slightly closer to the elytral apex than to the suture. Number and positions of the setae of the marginal umbilicate series as in
|
||
<taxonomicName id="4C764D31FFC2FFA0FEA3FE54FE0C5B47" authorityName="Clairville" authorityYear="1806" box="[325,418,1799,1825]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B902EAA0FFC2FFA0FEA3FE54FE0C5B47" box="[325,418,1799,1825]" italics="true" pageId="13" pageNumber="14">Trechus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23157DBFFC2FFA0FE4FFE54FE495B47" box="[425,487,1799,1825]" pageId="13" pageNumber="14" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="134D2A37FFC2FFA0FE10FE54FDE05B47" box="[502,590,1799,1825]" captionStart="Figs 9–12" captionStartId="9.[189,241,1930,1956]" captionTargetBox="[189,1375,351,1857]" captionTargetId="figure@9.[189,1398,351,1857]" captionTargetPageId="9" captionText="Figs 9–12. Trechus spp., elytra. 9. T. mattisi sp. nov., paratype, ³. 10. T. transversicollis sp. nov., holotype. 11. T. minitrechus sp. nov., paratype, ³. 12. T. nanulus sp. nov., paratype, ³. The arrows point to the insertions of the discal setae, preapical seta (if present), and setae of the umbilical series." figureDoi="http://doi.org/10.5281/zenodo.3829908" httpUri="https://zenodo.org/record/3829908/files/figure.png" pageId="13" pageNumber="14">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC936B2FFC2FFA0FF5BFE1AFB365BE0" blockId="13.[189,1398,1864,1926]" pageId="13" pageNumber="14">LEGS. Short and comparatively thin; protibia slightly dilated towards apex, hardly bowed, finely but distinctly grooved on external surface. Only one basal protarsomere of male dilated.</paragraph>
|
||
<paragraph id="8BC936B2FFC2FFA0FF5BFEFDFAD85B8D" blockId="13.[189,1398,1963,2027]" pageId="13" pageNumber="14">MALE GENITALIA. EL/AL = 4.00–4.28 (Ø = 4.13, n = 6). Aedeagal median lobe small and stout, in lateral view with ventral margin slightly bent near base and apex, straight in middle; apical lamella moderately</paragraph>
|
||
<paragraph id="8BC936B2FFC1FFA3FF5BF858FAE25D2C" blockId="14.[189,1398,267,331]" pageId="14" pageNumber="15">short with apex simply tapered; basal bulb moderately large, sagittal aileron average. Endophallus with copulatory piece bag-like, triangular in lateral view, arcuate towards apex, very slightly sclerotized.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C6539FFC1FFA3FF5BF820FE285D84" pageId="14" pageNumber="15" type="distribution">
|
||
<paragraph id="8BC936B2FFC1FFA3FF5BF820FEF85DEB" blockId="14.[189,342,371,397]" box="[189,342,371,397]" pageId="14" pageNumber="15">
|
||
<heading id="D08181DEFFC1FFA3FF5BF820FEF85DEB" bold="true" box="[189,342,371,397]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
|
||
<emphasis id="B902EAA0FFC1FFA3FF5BF820FEF85DEB" bold="true" box="[189,342,371,397]" pageId="14" pageNumber="15">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC936B2FFC1FFA3FF5BF8F7FE285D84" blockId="14.[189,1398,419,482]" pageId="14" pageNumber="15">
|
||
Known only from Harenna Forest on the southern slope of the Bale Mts, below Rira village, at altitudes of
|
||
<quantity id="4C8E9B57FFC1FFA3FF3BF89BFE2C5D84" box="[221,386,456,482]" metricMagnitude="3" metricUnit="m" metricValue="2.535" metricValueMax="2.69" metricValueMin="2.38" pageId="14" pageNumber="15" unit="m" value="2535.0" valueMax="2690.0" valueMin="2380.0">2380–2690 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |