482 lines
73 KiB
XML
482 lines
73 KiB
XML
<document id="A3708C4F8FCA6C93310F0A31FA4BB26A" ID-DOI="10.11646/zootaxa.1097.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5051451" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625109780512" checkinUser="felipe" docAuthor="Wesener, Thomas & Sierwald, Petra" docDate="2005" docId="03DF6C009D13BA3ECF6DF923FCD3770B" docLanguage="en" docName="zt01097p060.pdf" docOrigin="Zootaxa 1097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1097.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Zoosphaerium villosum Wesener & Sierwald 2005, sp. nov" docType="treatment" docVersion="4" lastPageNumber="28" masterDocId="FFE614789D01BA22CE65FFB7FE177655" masterDocTitle="New giant pill-millipede species from the littoral forest of Madagascar (Diplopoda, Sphaerotheriida, Zoosphaerium)" masterLastPageNumber="60" masterPageNumber="1" pageNumber="19" updateTime="1698981510058" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="31D862E10CA0F5D803DDBADD5587AA8F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6C81C5004D2E6C0836A4F604B799D1D2">
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<mods:title id="3E91CEE59B86F2B0317142C122E4B656">New giant pill-millipede species from the littoral forest of Madagascar (Diplopoda, Sphaerotheriida, Zoosphaerium)</mods:title>
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</mods:titleInfo>
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<mods:name id="4BD55050DAD1A9549F8FFD47623317E0" type="personal">
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<mods:roleTerm id="2483402D094DA80113DDF0205D4EF205">Author</mods:roleTerm>
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<mods:namePart id="DC1FDA2470042D74B95E95AAF3E5D647">Wesener, Thomas</mods:namePart>
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</mods:name>
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<mods:name id="944D3279A24811BF2F6CDC376977DEF1" type="personal">
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<mods:roleTerm id="3DFE43E9AD0F29BDF000D1A5A068B4EE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6324BA6B13076EE4EE7946101ED85E09">Sierwald, Petra</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="AC84BF8A5D0F0746134C0ACE63FE5FAF">text</mods:typeOfResource>
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<mods:titleInfo id="63B3FBEBC71BF0A149EFF81C2D41FEAC">
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<mods:title id="054F80D2813CFEA88E414E76082E9DF4">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F3C716EFBA8B6F366466F09C68798CFF">
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<mods:date id="03FC77BF50B902BF41550BB846F65CC5">2005</mods:date>
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<mods:detail id="248C813A5B2AF5F5DEF28882CB5F7BC8" type="pubDate">
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<mods:number id="25F037E9BCA1AFE32F4046FEF5256DB2">2005-12-16</mods:number>
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</mods:detail>
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<mods:detail id="8D4A8A4A59003303716FCE92D25DE80E" type="volume">
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<mods:number id="B719D21A3E7FF69078B87507EFBC3C7D">1097</mods:number>
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</mods:detail>
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<mods:detail id="2E5C5E30E2E465AF8823F16851E6552D" type="issue">
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<mods:number id="63FA25FD4EA0A24BA1B049D45585AC71">1</mods:number>
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<mods:extent id="E17625EDB0ED51A0CA800DAA48DE14BF" unit="page">
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<mods:start id="A92BCA28614228D990F1D7AE6EE9DA73">1</mods:start>
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<mods:end id="B5AF3B78C3B3A85014D626166118C515">60</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="71165856DF26F436ED2CB0887257C214">
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<mods:url id="8C2DA260B56301CA8A018682CAEA8F22">https://biotaxa.org/Zootaxa/article/view/zootaxa.1097.1.1</mods:url>
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</mods:location>
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<mods:classification id="D4E66EBA966C60581C796AC7723E8EA8">journal article</mods:classification>
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<mods:identifier id="8A933618B9A7BDABA20412AD60CE0CFF" type="DOI">10.11646/zootaxa.1097.1.1</mods:identifier>
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<mods:identifier id="35A8F77C60C3E5AE6B532A7401A1A165" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="381A3EE3A96265625A974205402101E1" type="Zenodo-Dep">5051451</mods:identifier>
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<mods:identifier id="22A0875EFBF4296820020FC55C36957E" type="ZooBank">4B7F398E-BB84-49D4-94D0-5E03A715CE7A</mods:identifier>
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</mods:mods>
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<treatment id="03DF6C009D13BA3ECF6DF923FCD3770B" ID-DOI="http://doi.org/10.5281/zenodo.5054324" ID-Zenodo-Dep="5054324" LSID="urn:lsid:plazi:treatment:03DF6C009D13BA3ECF6DF923FCD3770B" httpUri="http://treatment.plazi.org/id/03DF6C009D13BA3ECF6DF923FCD3770B" lastPageId="28" lastPageNumber="28" pageId="18" pageNumber="19">
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||
<subSubSection id="C36C8E9D9D13BA30CF6DF923FC9270FB" box="[264,645,1684,1710]" pageId="18" pageNumber="19" type="nomenclature">
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||
<paragraph id="8BC9DD169D13BA30CF6DF923FC9270FB" blockId="18.[264,645,1684,1745]" box="[264,645,1684,1710]" pageId="18" pageNumber="19">
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||
<heading id="D0816A7A9D13BA30CF6DF923FC9270FB" bold="true" box="[264,645,1684,1710]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
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||
<emphasis id="B90201049D13BA30CF6DF923FC9270FB" bold="true" box="[264,645,1684,1710]" pageId="18" pageNumber="19">
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||
<taxonomicName id="4C76A6959D13BA30CF6DF923FC3170FB" ID-CoL="5D7JD" authority="Wesener & Sierwald, 2005" authorityName="Wesener & Sierwald" authorityYear="2005" box="[264,550,1684,1710]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="villosum" status="sp. nov.">
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<emphasis id="B90201049D13BA30CF6DF923FC3170FB" bold="true" box="[264,550,1684,1710]" italics="true" pageId="18" pageNumber="19">Zoosphaerium villosum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A231BC7F9D13BA30CC48F923FC9270FB" box="[557,645,1684,1710]" pageId="18" pageNumber="19" rank="species">sp. nov</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C8E9D9D13BA30CF6DF900FF6C7084" box="[264,379,1719,1745]" pageId="18" pageNumber="19" type="description">
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<paragraph id="8BC9DD169D13BA30CF6DF900FF6C7084" blockId="18.[264,645,1684,1745]" box="[264,379,1719,1745]" pageId="18" pageNumber="19">
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<figureCitation id="134DC1939D13BA30CF6DF900FF6C7084" box="[264,379,1719,1745]" captionStart-0="FIGURES 9" captionStart-1="FIGURES 10" captionStart-2="FIGURES 11" captionStart-3="FIGURES 12" captionStart-4="FIGURE 13" captionStart-5="FIGURES 14" captionStart-6="FIGURES 15" captionStart-7="FIGURES 16" captionStartId-0="20.[264,370,1478,1499]" captionStartId-1="21.[264,370,1067,1088]" captionStartId-2="22.[264,370,1839,1860]" captionStartId-3="23.[264,370,1762,1783]" captionStartId-4="24.[264,357,1481,1502]" captionStartId-5="25.[264,370,1805,1826]" captionStartId-6="26.[264,370,1428,1449]" captionStartId-7="27.[264,370,1080,1101]" captionTargetBox-0="[274,1315,286,1464]" captionTargetBox-1="[278,1265,663,1010]" captionTargetBox-2="[272,1319,556,1825]" captionTargetBox-3="[274,1319,334,1747]" captionTargetBox-4="[368,1213,435,1452]" captionTargetBox-5="[264,1323,409,1793]" captionTargetBox-6="[269,1323,328,1408]" captionTargetBox-7="[264,1323,489,1053]" captionTargetId-0="figure-151@20.[264,1323,286,1478]" captionTargetId-1="figure-413@21.[264,1323,632,1038]" captionTargetId-2="figure-125@22.[264,1323,556,1833]" captionTargetId-3="figure-88@23.[264,1323,324,1753]" captionTargetId-4="figure-189@24.[337,1245,430,1464]" captionTargetId-5="figure-100@25.[264,1323,407,1793]" captionTargetId-6="figure-175@26.[264,1323,324,1408]" captionTargetId-7="figure-322@27.[264,1323,489,1053]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionTargetPageId-3="23" captionTargetPageId-4="24" captionTargetPageId-5="25" captionTargetPageId-6="26" captionTargetPageId-7="27" captionText-0="FIGURES 9 A–D Z. villosum sp. nov., male holotype. a: habitus; b: 7th paratergite, detail; c: left 9th leg, posterior view; d: 5th left pleurite. Scale bars = 1 mm" captionText-1="FIGURES 10 A–B: Z. villosum sp. nov., male holotype. a: coxa of 1st pair of legs with 1st left sternite, posterior view; b: left closing margins of anal shield. 12T = 12th tergite, As = anal shield, PL = pleurite, S = sternite, Scale bars = 1 mm" captionText-2="FIGURES 11 A–B: Z. villosum sp. nov., female paratype. a: subanal plate with washboard; b:coxa of 2nd left leg pair with vulva. EP = exterior plate, IP = inner plate, O = operculum. Scale bars = 1 mm" captionText-3="FIGURES 12 A–F: Z. villosum sp. nov., male holotype. a: anterior telopods, anterior view; b: inner view of 2–3 joint of right anterior telopod; c: detail of tip of process of 2nd joint of right anterior telopod, inner view; d: right anterior telopod, posterior view; e: posterior pair of telopods, anterior view; f: right posterior telopod, posterior view. IH = inner horns. Scale bars = 1 mm" captionText-4="FIGURE 13 a. villosum sp. nov., male holotype. a: left coxa of 2nd pair of legs with male gonopore, posterior view. Scale bars = 1 mm" captionText-5="FIGURES 14 A–D: Z. villosum sp. nov., male paratype, SEM. a: lateral view on antennae; b: lateral view on antennae joints 1–3; c: apical view on sensorial cones of antennae; d: endotergum, (1) = marginal ridge with external row of marginal bristles, (2) = band of flattened nodules, (3) = internal area." captionText-6="FIGURES 15 A–C: Z. villosum sp. nov., male paratype, SEM. a: ventral view of gnathochilarium; b: ventral view of epipharynx; c: right palp of gnathochilarium with sensorial cones, dorsal view, arrow = field of sensorial cones, cP = central pads, LL = lingual lamella, P = palpus." captionText-7="FIGURES 16 A–B: Z. villosum sp. nov., male paratype, SEM. a: left mandible, condylus; b: left mandible, pectinate lamellae." figureDoi-0="http://doi.org/10.5281/zenodo.5051477" figureDoi-1="http://doi.org/10.5281/zenodo.5051479" figureDoi-2="http://doi.org/10.5281/zenodo.5051481" figureDoi-3="http://doi.org/10.5281/zenodo.5051483" figureDoi-4="http://doi.org/10.5281/zenodo.5051487" figureDoi-5="http://doi.org/10.5281/zenodo.5051489" figureDoi-6="http://doi.org/10.5281/zenodo.5051492" figureDoi-7="http://doi.org/10.5281/zenodo.5051496" httpUri-0="https://zenodo.org/record/5051477/files/figure.png" httpUri-1="https://zenodo.org/record/5051479/files/figure.png" httpUri-2="https://zenodo.org/record/5051481/files/figure.png" httpUri-3="https://zenodo.org/record/5051483/files/figure.png" httpUri-4="https://zenodo.org/record/5051487/files/figure.png" httpUri-5="https://zenodo.org/record/5051489/files/figure.png" httpUri-6="https://zenodo.org/record/5051492/files/figure.png" httpUri-7="https://zenodo.org/record/5051496/files/figure.png" pageId="18" pageNumber="19">Figs 9–16</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36C8E9D9D13BA31CF6DF8B1FA8477D3" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="materials_examined">
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||
<paragraph id="8BC9DD169D13BA30CF6DF8B1FCA07124" blockId="18.[264,1323,1798,1905]" pageId="18" pageNumber="19">
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||
<materialsCitation id="3B1ED74B9D13BA30CF6DF8B1FCA07124" collectingDate="1996-04-03" collectingDateMax="1996-04-16" collectingDateMin="1996-04-03" collectorName="S. Goodman" country="Madagascar" elevation="1" latitude="-17.286667" location="Station Forestiere de Tampolo" longLatPrecision="127" longitude="49.408333" municipality="Province" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
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||
<emphasis id="B90201049D13BA30CF6DF8B1FF6D7175" bold="true" box="[264,378,1798,1824]" pageId="18" pageNumber="19">
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||
<typeStatus id="54CD63B49D13BA30CF6DF8B1FF6D7175" box="[264,378,1798,1824]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
:
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<quantity id="4C8E70F39D13BA30CFEEF8B0FFAB7174" box="[395,444,1799,1825]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="19" unit="m" value="1.0">
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||
<elevation id="005B3A259D13BA30CFEEF8B0FFAB7174" box="[395,444,1799,1825]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="19" unit="m" value="1.0">1 m</elevation>
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||
</quantity>
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||
, coll.
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||
<collectingCountry id="F3619D869D13BA30CC6DF8B0FC807174" box="[520,663,1799,1825]" name="Madagascar" pageId="18" pageNumber="19">Madagascar</collectingCountry>
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||
,
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||
<collectingMunicipality id="6BAD476C9D13BA30CCC0F8B0FD1A7174" box="[677,781,1799,1825]" pageId="18" pageNumber="19">Province</collectingMunicipality>
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||
<collectingRegion id="49B213F49D13BA30CD73F8B0FD807174" box="[790,919,1799,1825]" country="Madagascar" name="Toamasina" pageId="18" pageNumber="19">Toamasina</collectingRegion>
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,
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<collectingRegion id="49B213F49D13BA30CDC0F8B0FA317174" box="[933,1062,1799,1825]" country="Madagascar" name="Toamasina" pageId="18" pageNumber="19">Toamasina</collectingRegion>
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,
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<location id="8EA98BCD9D13BA30CA51F8B0FF64711C" LSID="urn:lsid:plazi:treatment:03DF6C009D13BA3ECF6DF923FCD3770B:8EA98BCD9D13BA30CA51F8B0FF64711C" country="Madagascar" latitude="-17.286667" longLatPrecision="127" longitude="49.408333" municipality="Province" name="Station Forestiere de Tampolo" pageId="18" pageNumber="19" stateProvince="Toamasina">Station Forestiére de Tampolo</location>
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;
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<date id="FFC8FBD69D13BA30CF1AF898FC3F711C" box="[383,552,1839,1865]" pageId="18" pageNumber="19" value="1996-04-03" valueMax="1996-04-16" valueMin="1996-04-03">
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||
<collectingDate id="EF8C023E9D13BA30CF1AF898FC3F711C" box="[383,552,1839,1865]" pageId="18" pageNumber="19" value="1996-04-03" valueMax="1996-04-16" valueMin="1996-04-03">3.–16.IV.1996</collectingDate>
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</date>
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; relatively undisturbed littoral forest;
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<geoCoordinate id="EE42BBD19D13BA30CD81F898FA4A711C" box="[996,1117,1839,1865]" degrees="17" direction="south" minutes="17.2" orientation="latitude" pageId="18" pageNumber="19" precision="92" value="-17.286667">17°17.2' S</geoCoordinate>
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||
<geoCoordinate id="EE42BBD19D13BA30CA00F898FAF4711C" box="[1125,1251,1839,1865]" degrees="49" direction="east" minutes="24.5" orientation="longitude" pageId="18" pageNumber="19" precision="92" value="49.408333">49°24.5' E</geoCoordinate>
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||
;
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<quantity id="4C8E70F39D13BA30CA8AF898FB3C711C" box="[1263,1323,1839,1865]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="19" unit="m" value="10.0">
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||
<elevation id="005B3A259D13BA30CA8AF898FB3C711C" box="[1263,1323,1839,1865]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="19" unit="m" value="10.0">10 m</elevation>
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||
</quantity>
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||
NN; leg.
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<collectorName id="2683B8C09D13BA30CF11F8E0FC1D7124" box="[372,522,1879,1905]" pageId="18" pageNumber="19">S. Goodman</collectorName>
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||
; FMMC 3958.
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BC9DD169D12BA31CF5DFEACFAAE7763" blockId="19.[264,1323,283,1842]" box="[312,1209,283,310]" pageId="19" pageNumber="20">
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||
<materialsCitation id="3B1ED74B9D12BA31CF5DFEACFAA27763" box="[312,1205,283,310]" collectionCode="CAS" elevation="1" pageId="19" pageNumber="20" specimenCount="3" typeStatus="paratype">
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||
<emphasis id="B90201049D12BA31CF5DFEACFFA27760" bold="true" box="[312,437,283,309]" pageId="19" pageNumber="20">
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||
<typeStatus id="54CD63B49D12BA31CF5DFEACFFA27760" box="[312,437,283,309]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
|
||
:
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||
<specimenCount id="9D70169F9D12BA31CFA1FEABFFEE7763" box="[452,505,284,310]" pageId="19" pageNumber="20" type="generic">
|
||
<quantity id="4C8E70F39D12BA31CFA1FEABFFE37763" box="[452,500,284,310]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="20" unit="m" value="1.0">
|
||
<elevation id="005B3A259D12BA31CFA1FEABFFE37763" box="[452,500,284,310]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="20" unit="m" value="1.0">1 m</elevation>
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||
</quantity>
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||
.
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||
</specimenCount>
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FMMC 8247;
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<specimenCount id="9D70169F9D12BA31CCCCFEABFCC77763" box="[681,720,284,310]" pageId="19" pageNumber="20" type="generic">1 f.</specimenCount>
|
||
FMMC 8248,
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||
<specimenCount id="9D70169F9D12BA31CDE5FEABFDA37763" box="[896,948,284,310]" pageId="19" pageNumber="20" type="generic">1 m.</specimenCount>
|
||
<collectionCode id="ED6745D39D12BA31CDDEFEABFDE07763" box="[955,1015,284,310]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="19" pageNumber="20" type="Museum">CAS</collectionCode>
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||
, same coll. data
|
||
</materialsCitation>
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||
.
|
||
</paragraph>
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||
<paragraph id="8BC9DD169D12BA31CF5DFEF3FA8477D3" blockId="19.[264,1323,283,1842]" pageId="19" pageNumber="20">
|
||
Additional material:
|
||
<specimenCount id="9D70169F9D12BA31CC55FEF3FC4A770B" box="[560,605,324,350]" pageId="19" pageNumber="20" type="generic">
|
||
<quantity id="4C8E70F39D12BA31CC55FEF3FC4F770B" box="[560,600,324,350]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="20" unit="m" value="1.0">1m</quantity>
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||
.
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||
</specimenCount>
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||
FMMC 7828;
|
||
<materialsCitation id="3B1ED74B9D12BA31CD7BFEF3FA8477D3" collectionCode="MNHN" collectorName="Grandidier" elevation="4" pageId="19" pageNumber="20" specimenCode="CB016" specimenCount="8" stateProvince="West Coast">
|
||
<specimenCount id="9D70169F9D12BA31CD7BFEF3FD42770B" box="[798,853,324,350]" pageId="19" pageNumber="20" type="generic">
|
||
<quantity id="4C8E70F39D12BA31CD7BFEF3FD47770B" box="[798,848,324,350]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="20" unit="m" value="4.0">
|
||
<elevation id="005B3A259D12BA31CD7BFEF3FD47770B" box="[798,848,324,350]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="20" unit="m" value="4.0">4 m</elevation>
|
||
</quantity>
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||
.
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||
</specimenCount>
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||
FMMC 7827, same collection data as typematerial.
|
||
<specimenCount id="9D70169F9D12BA31CFD6FEDBFFFE77D3" box="[435,489,364,390]" pageId="19" pageNumber="20" type="generic">
|
||
<quantity id="4C8E70F39D12BA31CFD6FEDBFFF277D3" box="[435,485,364,390]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="20" unit="m" value="1.0">
|
||
<elevation id="005B3A259D12BA31CFD6FEDBFFF277D3" box="[435,485,364,390]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="20" unit="m" value="1.0">1 m</elevation>
|
||
</quantity>
|
||
.
|
||
</specimenCount>
|
||
,
|
||
<specimenCount id="9D70169F9D12BA31CF93FEDBFC0277D3" box="[502,533,364,390]" pageId="19" pageNumber="20" type="generic">3f.</specimenCount>
|
||
coll.
|
||
<collectingRegion id="49B213F49D12BA31CC32FEDBFC9977D3" box="[599,654,364,390]" country="New Zealand" name="West Coast" pageId="19" pageNumber="20">Côte</collectingRegion>
|
||
SudOuest, leg.
|
||
<collectorName id="2683B8C09D12BA31CD36FEDBFDC377D3" box="[851,980,364,390]" pageId="19" pageNumber="20">Grandidier</collectorName>
|
||
,
|
||
<collectionCode id="ED6745D39D12BA31CDBAFEDBFA2E77D3" box="[991,1081,364,390]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="20">MNHN</collectionCode>
|
||
<specimenCode id="DBD0756D9D12BA31CA25FEDBFA8477D3" box="[1088,1171,364,390]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="20">CB016</specimenCode>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C8E9D9D12BA31CF5DFE24FABD74A4" pageId="19" pageNumber="20" type="diagnosis">
|
||
<paragraph id="8BC9DD169D12BA31CF5DFE24FF6B749C" blockId="19.[264,1323,283,1842]" pageId="19" pageNumber="20">
|
||
<emphasis id="B90201049D12BA31CF5DFE24FFAE77F8" bold="true" box="[312,441,403,429]" pageId="19" pageNumber="20">Diagnosis:</emphasis>
|
||
The shape of the posterior telopods easily identifies this species as another member of the
|
||
<taxonomicName id="4C76A6959D12BA31CFA4FE0BFC357780" authorityName="DeSaussure and Zehntner" authorityYear="1897" box="[449,546,444,469]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="alluaudi">
|
||
<emphasis id="B90201049D12BA31CFA4FE0BFC357780" box="[449,546,444,469]" italics="true" pageId="19" pageNumber="20">alluaudi</emphasis>
|
||
</taxonomicName>
|
||
species group. Z
|
||
<emphasis id="B90201049D12BA31CC99FE0BFA167780" box="[764,1025,444,469]" italics="true" pageId="19" pageNumber="20">
|
||
oosphaerium
|
||
<taxonomicName id="4C76A6959D12BA31CDFBFE0BFA167780" authority="Wesener & Sierwald, 2005" authorityName="Wesener & Sierwald" authorityYear="2005" box="[926,1025,444,469]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="villosum" status="sp. nov.">villosum</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="B90201049D12BA31CA6EFE0CFA7C7780" bold="true" box="[1035,1131,443,469]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="A231BC7F9D12BA31CA6EFE0CFA7C7780" box="[1035,1131,443,469]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
features unique characters, which separate this species unambiguously from others in the group. Tergites and anal shield are densely covered with short hairs, each hair inserting in a small pit (
|
||
<figureCitation id="134DC1939D12BA31CA9EFDBBFF3E7404" captionStart="FIGURES 9" captionStartId="20.[264,370,1478,1499]" captionTargetBox="[274,1315,286,1464]" captionTargetId="figure-151@20.[264,1323,286,1478]" captionTargetPageId="20" captionText="FIGURES 9 A–D Z. villosum sp. nov., male holotype. a: habitus; b: 7th paratergite, detail; c: left 9th leg, posterior view; d: 5th left pleurite. Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051477" httpUri="https://zenodo.org/record/5051477/files/figure.png" pageId="19" pageNumber="20">Fig. 9b</figureCitation>
|
||
). Harp on male telopods with two stridulation ridges and the tips of the 3
|
||
<superScript id="7C03705E9D12BA31CADFFD83FAD07417" attach="left" box="[1210,1223,564,578]" fontSize="6" pageId="19" pageNumber="20">rd</superScript>
|
||
joint of anterior telopods rounded (
|
||
<figureCitation id="134DC1939D12BA31CC29FDE8FCCD742C" box="[588,730,607,633]" captionStart="FIGURES 12" captionStartId="23.[264,370,1762,1783]" captionTargetBox="[274,1319,334,1747]" captionTargetId="figure-88@23.[264,1323,324,1753]" captionTargetPageId="23" captionText="FIGURES 12 A–F: Z. villosum sp. nov., male holotype. a: anterior telopods, anterior view; b: inner view of 2–3 joint of right anterior telopod; c: detail of tip of process of 2nd joint of right anterior telopod, inner view; d: right anterior telopod, posterior view; e: posterior pair of telopods, anterior view; f: right posterior telopod, posterior view. IH = inner horns. Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051483" httpUri="https://zenodo.org/record/5051483/files/figure.png" pageId="19" pageNumber="20">Figs. 12a–d</figureCitation>
|
||
). Furthermore, the shape of the first sternite and shape of the male gonopore differs in
|
||
<taxonomicName id="4C76A6959D12BA31CCB0FD30FD4E74F5" authorityName="Wesener & Sierwald" authorityYear="2005" box="[725,857,647,672]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="villosum">
|
||
<emphasis id="B90201049D12BA31CCB0FD30FD4E74F5" box="[725,857,647,672]" italics="true" pageId="19" pageNumber="20">Z. villosum</emphasis>
|
||
</taxonomicName>
|
||
from other species of the genus (
|
||
<figureCitation id="134DC1939D12BA31CA95FD30FF22749C" captionStart="FIGURES 10" captionStartId="21.[264,370,1067,1088]" captionTargetBox="[278,1265,663,1010]" captionTargetId="figure-413@21.[264,1323,632,1038]" captionTargetPageId="21" captionText="FIGURES 10 A–B: Z. villosum sp. nov., male holotype. a: coxa of 1st pair of legs with 1st left sternite, posterior view; b: left closing margins of anal shield. 12T = 12th tergite, As = anal shield, PL = pleurite, S = sternite, Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051479" httpUri="https://zenodo.org/record/5051479/files/figure.png" pageId="19" pageNumber="20">Figs. 10a</figureCitation>
|
||
,
|
||
<figureCitation id="134DC1939D12BA31CF24FD18FF79749C" box="[321,366,687,713]" captionStart="FIGURE 13" captionStartId="24.[264,357,1481,1502]" captionTargetBox="[368,1213,435,1452]" captionTargetId="figure-189@24.[337,1245,430,1464]" captionTargetPageId="24" captionText="FIGURE 13 a. villosum sp. nov., male holotype. a: left coxa of 2nd pair of legs with male gonopore, posterior view. Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051487" httpUri="https://zenodo.org/record/5051487/files/figure.png" pageId="19" pageNumber="20">13a</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC9DD169D12BA31CF5DFD60FABD74A4" blockId="19.[264,1323,283,1842]" box="[312,1194,727,753]" pageId="19" pageNumber="20">
|
||
<emphasis id="B90201049D12BA31CF5DFD60FFEC74A4" bold="true" box="[312,507,727,753]" pageId="19" pageNumber="20">Similar species:</emphasis>
|
||
See
|
||
<tableCitation id="C6F4E8AD9D12BA31CC51FD60FC9D74A4" box="[564,650,727,753]" captionStart="TABLE 1" captionStartId="28.[264,343,403,424]" captionText="TABLE 1: Differences between members of the Zoosphaerium alluaudi speciesgroup. Abbreviations: ant. = anterior; post = posterior; tsh = thoracic shield; SR = stridulation ribs; AS = sensory cones on antennae; aT = anterior telopods; pT = posterior telopods; O = operculum of vulva; St = sternite, md = mandible; gn = gnathochilarium, n°. = number." pageId="19" pageNumber="20">Table 1</tableCitation>
|
||
for a comparison with closely related species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C8E9D9D12BA39CF5DFD48FD0E77FB" lastPageId="27" lastPageNumber="28" pageId="19" pageNumber="20" type="description">
|
||
<paragraph id="8BC9DD169D12BA31CF5DFD48FC1E75C4" blockId="19.[264,1323,283,1842]" pageId="19" pageNumber="20">
|
||
<emphasis id="B90201049D12BA31CF5DFD48FFDE754C" bold="true" box="[312,457,767,793]" pageId="19" pageNumber="20">Description</emphasis>
|
||
: Males (
|
||
<specimenCount id="9D70169F9D12BA31CC24FD48FCF4754C" box="[577,739,767,793]" pageId="19" pageNumber="20" type="generic">4 specimens</specimenCount>
|
||
): length: 43.2–64.6, width of thoracic shield: 21.1–33.6, height of thoracic shield 12.2–17.4. Females (
|
||
<specimenCount id="9D70169F9D12BA31CDCEFC90FA547514" box="[939,1091,807,833]" pageId="19" pageNumber="20" type="generic">4 specimens</specimenCount>
|
||
): length: 43.3–57.9, width: 24.25–30.25, height: 12.2–16.1. Body length of males is not significantly greater than those of females.
|
||
</paragraph>
|
||
<paragraph id="8BC9DD169D12BA31CF5DFC28FC407264" blockId="19.[264,1323,283,1842]" pageId="19" pageNumber="20">
|
||
<emphasis id="B90201049D12BA31CF5DFC28FFD575ED" box="[312,450,927,952]" italics="true" pageId="19" pageNumber="20">Coloration:</emphasis>
|
||
The body is dirty blackish green. Posterior margin of the tergites with a thin darkbrown line. Collum and head dark green. Head, ocelli, antennae and legs green. In alcoholpreserved animals, pieces of the dark green pigment layer break off and brown body tissues become visible.
|
||
</paragraph>
|
||
<paragraph id="8BC9DD169D12BA31CF5DFB88FAD47284" blockId="19.[264,1323,283,1842]" pageId="19" pageNumber="20">
|
||
<emphasis id="B90201049D12BA31CF5DFB88FF60720D" box="[312,375,1087,1112]" italics="true" pageId="19" pageNumber="20">Head</emphasis>
|
||
: with some long hairs and numerous setiferous pits mostly around the clypeus and lateral of the eyes. Some long, isolated hairs around the eyes and more distributed on the entire head. Posterior margin of head towards collum with a patch of very short hairs. Edges of antennal socket each with little crenulated teeth and a single short spine.
|
||
</paragraph>
|
||
<paragraph id="8BC9DD169D12BA31CF5DFB68FAB2732F" blockId="19.[264,1323,283,1842]" pageId="19" pageNumber="20">
|
||
<emphasis id="B90201049D12BA31CF5DFB68FFB172AD" box="[312,422,1247,1272]" italics="true" pageId="19" pageNumber="20">Antennae</emphasis>
|
||
: overall shape as in genus description. Very long, with thin, long, cylindrical joints; length of antennomeres: 1>2>3>4=5<6. 6
|
||
<superScript id="7C03705E9D12BA31CD33FAB0FD757340" attach="left" box="[854,866,1287,1301]" fontSize="6" pageId="19" pageNumber="20">th</superScript>
|
||
antennomere long (
|
||
<figureCitation id="134DC1939D12BA31CA37FABDFAAF7371" box="[1106,1208,1290,1316]" captionStart="FIGURES 14" captionStartId="25.[264,370,1805,1826]" captionTargetBox="[264,1323,409,1793]" captionTargetId="figure-100@25.[264,1323,407,1793]" captionTargetPageId="25" captionText="FIGURES 14 A–D: Z. villosum sp. nov., male paratype, SEM. a: lateral view on antennae; b: lateral view on antennae joints 1–3; c: apical view on sensorial cones of antennae; d: endotergum, (1) = marginal ridge with external row of marginal bristles, (2) = band of flattened nodules, (3) = internal area." figureDoi="http://doi.org/10.5281/zenodo.5051489" httpUri="https://zenodo.org/record/5051489/files/figure.png" pageId="19" pageNumber="20">Fig. 14a</figureCitation>
|
||
), with 35 (25) to 45 sensory cones (
|
||
<figureCitation id="134DC1939D12BA31CC5EFA82FCB6731A" box="[571,673,1333,1359]" captionStart="FIGURES 14" captionStartId="25.[264,370,1805,1826]" captionTargetBox="[264,1323,409,1793]" captionTargetId="figure-100@25.[264,1323,407,1793]" captionTargetPageId="25" captionText="FIGURES 14 A–D: Z. villosum sp. nov., male paratype, SEM. a: lateral view on antennae; b: lateral view on antennae joints 1–3; c: apical view on sensorial cones of antennae; d: endotergum, (1) = marginal ridge with external row of marginal bristles, (2) = band of flattened nodules, (3) = internal area." figureDoi="http://doi.org/10.5281/zenodo.5051489" httpUri="https://zenodo.org/record/5051489/files/figure.png" pageId="19" pageNumber="20">Fig. 14c</figureCitation>
|
||
). 1
|
||
<superScript id="7C03705E9D12BA31CCAAFA85FCCD7315" attach="left" box="[719,730,1330,1344]" fontSize="6" pageId="19" pageNumber="20">st</superScript>
|
||
and 2
|
||
<superScript id="7C03705E9D12BA31CD46FA85FD247315" attach="left" box="[803,819,1330,1344]" fontSize="6" pageId="19" pageNumber="20">nd</superScript>
|
||
joints with small crenulated teeth and one invagination each; 3
|
||
<superScript id="7C03705E9D12BA31CF93FAEAFC14733E" attach="left" box="[502,515,1373,1387]" fontSize="6" pageId="19" pageNumber="20">rd</superScript>
|
||
joint with some small crenulated teeth at base (
|
||
<figureCitation id="134DC1939D12BA31CA57FAD7FA8F732F" box="[1074,1176,1376,1402]" captionStart="FIGURES 14" captionStartId="25.[264,370,1805,1826]" captionTargetBox="[264,1323,409,1793]" captionTargetId="figure-100@25.[264,1323,407,1793]" captionTargetPageId="25" captionText="FIGURES 14 A–D: Z. villosum sp. nov., male paratype, SEM. a: lateral view on antennae; b: lateral view on antennae joints 1–3; c: apical view on sensorial cones of antennae; d: endotergum, (1) = marginal ridge with external row of marginal bristles, (2) = band of flattened nodules, (3) = internal area." figureDoi="http://doi.org/10.5281/zenodo.5051489" httpUri="https://zenodo.org/record/5051489/files/figure.png" pageId="19" pageNumber="20">Fig. 14b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC9DD169D12BA31CF5DFA3FFC0373A7" blockId="19.[264,1323,283,1842]" pageId="19" pageNumber="20">
|
||
<emphasis id="B90201049D12BA31CF5DFA3FFFBF73F4" box="[312,424,1416,1441]" italics="true" pageId="19" pageNumber="20">Mandible</emphasis>
|
||
: molar plate process of mandible with a single, sharp furrow near the apical end (
|
||
<figureCitation id="134DC1939D12BA31CF2BFA07FFA2739F" box="[334,437,1456,1482]" captionStart="FIGURES 16" captionStartId="27.[264,370,1080,1101]" captionTargetBox="[264,1323,489,1053]" captionTargetId="figure-322@27.[264,1323,489,1053]" captionTargetPageId="27" captionText="FIGURES 16 A–B: Z. villosum sp. nov., male paratype, SEM. a: left mandible, condylus; b: left mandible, pectinate lamellae." figureDoi="http://doi.org/10.5281/zenodo.5051496" httpUri="https://zenodo.org/record/5051496/files/figure.png" pageId="19" pageNumber="20">Fig. 16a</figureCitation>
|
||
); 7 pectinate lamellae with long and thin teeth, number of teeth declining proximally (
|
||
<figureCitation id="134DC1939D12BA31CFC5FA6FFC1073A7" box="[416,519,1496,1522]" captionStart="FIGURES 16" captionStartId="27.[264,370,1080,1101]" captionTargetBox="[264,1323,489,1053]" captionTargetId="figure-322@27.[264,1323,489,1053]" captionTargetPageId="27" captionText="FIGURES 16 A–B: Z. villosum sp. nov., male paratype, SEM. a: left mandible, condylus; b: left mandible, pectinate lamellae." figureDoi="http://doi.org/10.5281/zenodo.5051496" httpUri="https://zenodo.org/record/5051496/files/figure.png" pageId="19" pageNumber="20">Fig. 16b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC9DD169D12BA31CF5DF9B7FDF4703F" blockId="19.[264,1323,283,1842]" pageId="19" pageNumber="20">
|
||
<emphasis id="B90201049D12BA31CF5DF9B7FC15704C" box="[312,514,1536,1561]" italics="true" pageId="19" pageNumber="20">Gnathochilarium</emphasis>
|
||
: posterior surface with many hairs but fewer hairs on the lingual lamella (
|
||
<figureCitation id="134DC1939D12BA31CF17F99FFFCD7017" box="[370,474,1576,1602]" captionStart="FIGURES 15" captionStartId="26.[264,370,1428,1449]" captionTargetBox="[269,1323,328,1408]" captionTargetId="figure-175@26.[264,1323,324,1408]" captionTargetPageId="26" captionText="FIGURES 15 A–C: Z. villosum sp. nov., male paratype, SEM. a: ventral view of gnathochilarium; b: ventral view of epipharynx; c: right palp of gnathochilarium with sensorial cones, dorsal view, arrow = field of sensorial cones, cP = central pads, LL = lingual lamella, P = palpus." figureDoi="http://doi.org/10.5281/zenodo.5051492" httpUri="https://zenodo.org/record/5051492/files/figure.png" pageId="19" pageNumber="20">Fig. 15a</figureCitation>
|
||
). Lateral of palpi is a pit with three sensorial cones (
|
||
<figureCitation id="134DC1939D12BA31CA09F99FFAC37017" box="[1132,1236,1576,1602]" captionStart="FIGURES 15" captionStartId="26.[264,370,1428,1449]" captionTargetBox="[269,1323,328,1408]" captionTargetId="figure-175@26.[264,1323,324,1408]" captionTargetPageId="26" captionText="FIGURES 15 A–C: Z. villosum sp. nov., male paratype, SEM. a: ventral view of gnathochilarium; b: ventral view of epipharynx; c: right palp of gnathochilarium with sensorial cones, dorsal view, arrow = field of sensorial cones, cP = central pads, LL = lingual lamella, P = palpus." figureDoi="http://doi.org/10.5281/zenodo.5051492" httpUri="https://zenodo.org/record/5051492/files/figure.png" pageId="19" pageNumber="20">Fig. 15c</figureCitation>
|
||
).
|
||
<emphasis id="B90201049D12BA31CA9DF99FFF7F703C" italics="true" pageId="19" pageNumber="20">Epipharynx</emphasis>
|
||
shaped like in other sphaerotheriid species (
|
||
<figureCitation id="134DC1939D12BA31CD0AF9E7FDC2703F" box="[879,981,1616,1642]" captionStart="FIGURES 15" captionStartId="26.[264,370,1428,1449]" captionTargetBox="[269,1323,328,1408]" captionTargetId="figure-175@26.[264,1323,324,1408]" captionTargetPageId="26" captionText="FIGURES 15 A–C: Z. villosum sp. nov., male paratype, SEM. a: ventral view of gnathochilarium; b: ventral view of epipharynx; c: right palp of gnathochilarium with sensorial cones, dorsal view, arrow = field of sensorial cones, cP = central pads, LL = lingual lamella, P = palpus." figureDoi="http://doi.org/10.5281/zenodo.5051492" httpUri="https://zenodo.org/record/5051492/files/figure.png" pageId="19" pageNumber="20">Fig. 15b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC9DD169D12BA31CF5DF9CFFDD77167" blockId="19.[264,1323,283,1842]" pageId="19" pageNumber="20">
|
||
<emphasis id="B90201049D12BA31CF5DF9CFFF9970C4" box="[312,398,1656,1681]" italics="true" pageId="19" pageNumber="20">Collum</emphasis>
|
||
: anterior margin with three rows of isolated hairs; rest of collum, especially posterior margin with only few isolated hairs.
|
||
<emphasis id="B90201049D12BA31CD4EF917FDFB70EC" box="[811,1004,1696,1721]" italics="true" pageId="19" pageNumber="20">Thoracic shield:</emphasis>
|
||
A few hairs in the concave lateral extension of the thoracic shield, which are remarkably large in this species. Marginal brim very thin, only anteriorly a little broader (
|
||
<figureCitation id="134DC1939D12BA31CDD4F947FA1E715F" box="[945,1033,1776,1802]" captionStart="FIGURES 9" captionStartId="20.[264,370,1478,1499]" captionTargetBox="[274,1315,286,1464]" captionTargetId="figure-151@20.[264,1323,286,1478]" captionTargetPageId="20" captionText="FIGURES 9 A–D Z. villosum sp. nov., male holotype. a: habitus; b: 7th paratergite, detail; c: left 9th leg, posterior view; d: 5th left pleurite. Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051477" httpUri="https://zenodo.org/record/5051477/files/figure.png" pageId="19" pageNumber="20">Fig. 9a</figureCitation>
|
||
). Underside of thoracic shield with two rounded impressions at the posterior edges.
|
||
</paragraph>
|
||
<caption id="DF098D9E9D15BA36CF6DFA71FD1573A9" ID-DOI="http://doi.org/10.5281/zenodo.5051477" ID-Zenodo-Dep="5051477" httpUri="https://zenodo.org/record/5051477/files/figure.png" pageId="20" pageNumber="21" startId="20.[264,370,1478,1499]" targetBox="[274,1315,286,1464]" targetPageId="20">
|
||
<paragraph id="8BC9DD169D15BA36CF6DFA71FD1573A9" blockId="20.[264,1323,1478,1532]" pageId="20" pageNumber="21">
|
||
<emphasis id="B90201049D15BA36CF6DFA71FFA9738E" bold="true" box="[264,446,1478,1499]" pageId="20" pageNumber="21">FIGURES 9 A–D</emphasis>
|
||
<taxonomicName id="4C76A6959D15BA36CFA2FA70FC237389" authority="Wesener & Sierwald, 2005" authorityName="Wesener & Sierwald" authorityYear="2005" box="[455,564,1479,1500]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="villosum" status="sp. nov.">
|
||
<emphasis id="B90201049D15BA36CFA2FA70FC237389" box="[455,564,1479,1500]" italics="true" pageId="20" pageNumber="21">Z. villosum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90201049D15BA36CC59FA71FC9B738E" bold="true" box="[572,652,1478,1499]" pageId="20" pageNumber="21">
|
||
<taxonomicNameLabel id="A231BC7F9D15BA36CC59FA71FC9B738E" box="[572,652,1478,1499]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype. a: habitus; b: 7th paratergite, detail; c: left 9th leg, posterior view; d: 5
|
||
<superScript id="7C03705E9D15BA36CFA6FA53FFDA73A5" attach="left" box="[451,461,1508,1520]" fontSize="5" pageId="20" pageNumber="21">th</superScript>
|
||
left pleurite. Scale bars = 1 mm
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC9DD169D15BA36CF5DF988FAB770AC" blockId="20.[264,1323,1599,1905]" pageId="20" pageNumber="21">
|
||
<emphasis id="B90201049D15BA36CF5DF988FF83700D" box="[312,404,1599,1624]" italics="true" pageId="20" pageNumber="21">Tergites</emphasis>
|
||
: anterior paratergite depression densely covered with hairs and several ridges. Tips of paratergites project posteriorly. Tergites covered with numerous small pits (
|
||
<figureCitation id="134DC1939D15BA36CA9EF9D0FF3F70FC" captionStart="FIGURES 9" captionStartId="20.[264,370,1478,1499]" captionTargetBox="[274,1315,286,1464]" captionTargetId="figure-151@20.[264,1323,286,1478]" captionTargetPageId="20" captionText="FIGURES 9 A–D Z. villosum sp. nov., male holotype. a: habitus; b: 7th paratergite, detail; c: left 9th leg, posterior view; d: 5th left pleurite. Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051477" httpUri="https://zenodo.org/record/5051477/files/figure.png" pageId="20" pageNumber="21">Fig. 9b</figureCitation>
|
||
), some of them with a single hair inserting at the center, giving the body in some specimens a furry look. Posterior margins of tergites with a visible fringe of short hairs, originating from the underside of posterior margin of tergites, the endotergum.
|
||
</paragraph>
|
||
<paragraph id="8BC9DD169D15BA36CF5DF8B0FAA87124" blockId="20.[264,1323,1599,1905]" pageId="20" pageNumber="21">
|
||
<emphasis id="B90201049D15BA36CF5DF8B0FFC67175" box="[312,465,1799,1824]" italics="true" pageId="20" pageNumber="21">Endotergum:</emphasis>
|
||
features two rows of marginal bristles. Marginal ridge regularly rounded; proximal of marginal ridge with a distinct band of cuticular patterns (
|
||
<figureCitation id="134DC1939D15BA36CAD0F898FB09711C" box="[1205,1310,1839,1865]" captionStart="FIGURES 14" captionStartId="25.[264,370,1805,1826]" captionTargetBox="[264,1323,409,1793]" captionTargetId="figure-100@25.[264,1323,407,1793]" captionTargetPageId="25" captionText="FIGURES 14 A–D: Z. villosum sp. nov., male paratype, SEM. a: lateral view on antennae; b: lateral view on antennae joints 1–3; c: apical view on sensorial cones of antennae; d: endotergum, (1) = marginal ridge with external row of marginal bristles, (2) = band of flattened nodules, (3) = internal area." figureDoi="http://doi.org/10.5281/zenodo.5051489" httpUri="https://zenodo.org/record/5051489/files/figure.png" pageId="20" pageNumber="21">Fig. 14d</figureCitation>
|
||
). Inner area covered with numerous hairs and spines. Bristles of endotergum scaly.
|
||
</paragraph>
|
||
<paragraph id="8BC9DD169D14BA37CF5DFEABFF417404" blockId="21.[264,1323,284,593]" pageId="21" pageNumber="22">
|
||
<emphasis id="B90201049D14BA37CF5DFEABFFD47760" box="[312,451,284,309]" italics="true" pageId="21" pageNumber="22">Anal shield</emphasis>
|
||
: rounded, neither bellshaped nor tapered. Anal shield more densely covered with hairs than tergites. Ventral surface carries two black locking carinae on each side, locking carinae sloping weakly towards the posterior end of the anal shield (
|
||
<figureCitation id="134DC1939D14BA37CAA5FEDBFB3177D3" box="[1216,1318,364,390]" captionStart="FIGURES 10" captionStartId="21.[264,370,1067,1088]" captionTargetBox="[278,1265,663,1010]" captionTargetId="figure-413@21.[264,1323,632,1038]" captionTargetPageId="21" captionText="FIGURES 10 A–B: Z. villosum sp. nov., male holotype. a: coxa of 1st pair of legs with 1st left sternite, posterior view; b: left closing margins of anal shield. 12T = 12th tergite, As = anal shield, PL = pleurite, S = sternite, Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051479" httpUri="https://zenodo.org/record/5051479/files/figure.png" pageId="21" pageNumber="22">Fig. 10b</figureCitation>
|
||
: 12T = 12
|
||
<superScript id="7C03705E9D14BA37CF13FE23FF9577F7" attach="left" box="[374,386,404,418]" fontSize="6" pageId="21" pageNumber="22">th</superScript>
|
||
tergite; As = anal shield; PL = pleurite), posterior one up to three times longer than the anterior one. Locking carinae separated from each other by a distance equal to the length of the shorter carina. Distinct suture present between both carinae. Where the suture reaches the margin of the anal shield, a small, distinct triangular invagination is visible
|
||
</paragraph>
|
||
<caption id="DF098D9E9D14BA37CF6DFB9CFC047228" ID-DOI="http://doi.org/10.5281/zenodo.5051479" ID-Zenodo-Dep="5051479" httpUri="https://zenodo.org/record/5051479/files/figure.png" pageId="21" pageNumber="22" startId="21.[264,370,1067,1088]" targetBox="[278,1265,663,1010]" targetPageId="21">
|
||
<paragraph id="8BC9DD169D14BA37CF6DFB9CFC047228" blockId="21.[264,1323,1065,1149]" pageId="21" pageNumber="22">
|
||
<emphasis id="B90201049D14BA37CF6DFB9CFFDB7215" bold="true" box="[264,460,1067,1088]" pageId="21" pageNumber="22">FIGURES 10 A–B</emphasis>
|
||
:
|
||
<taxonomicName id="4C76A6959D14BA37CFB8FB9BFC5C7214" authority="Wesener & Sierwald, 2005" authorityName="Wesener & Sierwald" authorityYear="2005" box="[477,587,1068,1089]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="villosum" status="sp. nov.">
|
||
<emphasis id="B90201049D14BA37CFB8FB9BFC5C7214" box="[477,587,1068,1089]" italics="true" pageId="21" pageNumber="22">Z. villosum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90201049D14BA37CC30FB9CFCB17215" bold="true" box="[597,678,1067,1088]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A231BC7F9D14BA37CC30FB9CFCB17215" box="[597,678,1067,1088]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype. a: coxa of 1
|
||
<superScript id="7C03705E9D14BA37CDAAFB9EFDCF7260" attach="left" box="[975,984,1065,1077]" fontSize="5" pageId="21" pageNumber="22">st</superScript>
|
||
pair of legs with 1
|
||
<superScript id="7C03705E9D14BA37CAC4FB9EFABD7260" attach="left" box="[1185,1194,1065,1077]" fontSize="5" pageId="21" pageNumber="22">st</superScript>
|
||
left sternite, posterior view; b: left closing margins of anal shield. 12T = 12
|
||
<superScript id="7C03705E9D14BA37CD11FBFEFD697200" attach="left" box="[884,894,1097,1109]" fontSize="5" pageId="21" pageNumber="22">th</superScript>
|
||
tergite, As = anal shield, PL = pleurite, S = sternite, Scale bars = 1 mm
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC9DD169D14BA37CF5DFB66FD4773E3" blockId="21.[264,1323,1233,1905]" pageId="21" pageNumber="22">
|
||
<emphasis id="B90201049D14BA37CF5DFB66FF6E72BF" box="[312,377,1233,1258]" italics="true" pageId="21" pageNumber="22">Legs:</emphasis>
|
||
tarsi of first leg pair with only three to six, second pair with only five to eight ventral spines, claws only weakly curved and without an apical spine. Tarsi of the following legs with 8–11 ventral spines, curved claws and one apical spine. Coxae of all legs covered at the inside margin with a patch of long hairs, the following leg joints with some isolated, long hairs as well. 9
|
||
<superScript id="7C03705E9D14BA37CCC8FAC6FCAE732A" attach="left" box="[685,697,1393,1407]" fontSize="6" pageId="21" pageNumber="22">th</superScript>
|
||
pair of legs without coxal lobes and spines. All legs feature a crenulated ridge on the femora (
|
||
<figureCitation id="134DC1939D14BA37CC88FA2BFD5473E3" box="[749,835,1436,1462]" captionStart="FIGURES 9" captionStartId="20.[264,370,1478,1499]" captionTargetBox="[274,1315,286,1464]" captionTargetId="figure-151@20.[264,1323,286,1478]" captionTargetPageId="20" captionText="FIGURES 9 A–D Z. villosum sp. nov., male holotype. a: habitus; b: 7th paratergite, detail; c: left 9th leg, posterior view; d: 5th left pleurite. Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051477" httpUri="https://zenodo.org/record/5051477/files/figure.png" pageId="21" pageNumber="22">Fig. 9c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC9DD169D14BA37CF5DFA73FFDF70F3" blockId="21.[264,1323,1233,1905]" pageId="21" pageNumber="22">
|
||
<emphasis id="B90201049D14BA37CF5DFA73FF8B7388" box="[312,412,1476,1501]" italics="true" pageId="21" pageNumber="22">Sternite:</emphasis>
|
||
First sternite with a sclerotized ledge (
|
||
<figureCitation id="134DC1939D14BA37CDE3FA73FDFA738B" box="[902,1005,1476,1502]" captionStart="FIGURES 10" captionStartId="21.[264,370,1067,1088]" captionTargetBox="[278,1265,663,1010]" captionTargetId="figure-413@21.[264,1323,632,1038]" captionTargetPageId="21" captionText="FIGURES 10 A–B: Z. villosum sp. nov., male holotype. a: coxa of 1st pair of legs with 1st left sternite, posterior view; b: left closing margins of anal shield. 12T = 12th tergite, As = anal shield, PL = pleurite, S = sternite, Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051479" httpUri="https://zenodo.org/record/5051479/files/figure.png" pageId="21" pageNumber="22">Fig. 10a</figureCitation>
|
||
) and long lobe, the latter reaching towards the beginning of prefemur and curved towards the leg pair. Anterior margin irregularly rounded with one small invagination near the tip, covered with long, isolated hairs. Posterior part of the sternite bald (
|
||
<figureCitation id="134DC1939D14BA37CD38F98BFDD27003" box="[861,965,1596,1622]" captionStart="FIGURES 10" captionStartId="21.[264,370,1067,1088]" captionTargetBox="[278,1265,663,1010]" captionTargetId="figure-413@21.[264,1323,632,1038]" captionTargetPageId="21" captionText="FIGURES 10 A–B: Z. villosum sp. nov., male holotype. a: coxa of 1st pair of legs with 1st left sternite, posterior view; b: left closing margins of anal shield. 12T = 12th tergite, As = anal shield, PL = pleurite, S = sternite, Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051479" httpUri="https://zenodo.org/record/5051479/files/figure.png" pageId="21" pageNumber="22">Fig. 10a</figureCitation>
|
||
: S = sternite). Sternites three and beyond with a spine like process which reaches almost the stigma opening of the next anterior sternite.
|
||
</paragraph>
|
||
<paragraph id="8BC9DD169D14BA34CF5DF900FDE677D3" blockId="21.[264,1323,1233,1905]" lastBlockId="22.[264,1323,284,510]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
|
||
<emphasis id="B90201049D14BA37CF5DF900FC637085" box="[312,628,1719,1744]" italics="true" pageId="21" pageNumber="22">Female sexual characters:</emphasis>
|
||
2
|
||
<superScript id="7C03705E9D14BA37CCEEF904FC8C7094" attach="left" box="[651,667,1715,1729]" fontSize="6" pageId="21" pageNumber="22">nd</superScript>
|
||
pair of legs without coxal lobe. Operculum (
|
||
<figureCitation id="134DC1939D14BA37CADAF900FB317084" box="[1215,1318,1719,1745]" captionStart="FIGURES 11" captionStartId="22.[264,370,1839,1860]" captionTargetBox="[272,1319,556,1825]" captionTargetId="figure-125@22.[264,1323,556,1833]" captionTargetPageId="22" captionText="FIGURES 11 A–B: Z. villosum sp. nov., female paratype. a: subanal plate with washboard; b:coxa of 2nd left leg pair with vulva. EP = exterior plate, IP = inner plate, O = operculum. Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051481" httpUri="https://zenodo.org/record/5051481/files/figure.png" pageId="21" pageNumber="22">Fig. 11b</figureCitation>
|
||
: O) of vulvae small, not reaching upper margin of coxa. Apical margin constricted in the middle (= subreniform), lower margin straight. Exterior and interior plate (
|
||
<figureCitation id="134DC1939D14BA37CAEAF8B0FAEF7174" box="[1167,1272,1799,1825]" captionStart="FIGURES 11" captionStartId="22.[264,370,1839,1860]" captionTargetBox="[272,1319,556,1825]" captionTargetId="figure-125@22.[264,1323,556,1833]" captionTargetPageId="22" captionText="FIGURES 11 A–B: Z. villosum sp. nov., female paratype. a: subanal plate with washboard; b:coxa of 2nd left leg pair with vulva. EP = exterior plate, IP = inner plate, O = operculum. Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051481" httpUri="https://zenodo.org/record/5051481/files/figure.png" pageId="21" pageNumber="22">Fig. 11b</figureCitation>
|
||
: EP, IP) of vulvae long and broad, its anterior margin reaching around the base of the operculum; interior plate about 2/3 the size of the operculum, exterior plate only 1/3 the size of the operculum. Cyphopod sclerites consisting of two triangular apical sclerites and a much larger third sclerite shaped like a tuningfork; all visible as dark structures near the suture of the vulva between inner and exterior plates (
|
||
<figureCitation id="134DC1939D17BA34CD1AFEDBFDF377D3" box="[895,996,364,390]" captionStart="FIGURES 11" captionStartId="22.[264,370,1839,1860]" captionTargetBox="[272,1319,556,1825]" captionTargetId="figure-125@22.[264,1323,556,1833]" captionTargetPageId="22" captionText="FIGURES 11 A–B: Z. villosum sp. nov., female paratype. a: subanal plate with washboard; b:coxa of 2nd left leg pair with vulva. EP = exterior plate, IP = inner plate, O = operculum. Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051481" httpUri="https://zenodo.org/record/5051481/files/figure.png" pageId="22" pageNumber="23">Fig. 11b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC9DD169D17BA34CF5DFE23FC2977AB" blockId="22.[264,1323,284,510]" pageId="22" pageNumber="23">
|
||
<emphasis id="B90201049D17BA34CF5DFE23FFCC77F8" box="[312,475,404,429]" italics="true" pageId="22" pageNumber="23">Subanal plate</emphasis>
|
||
rounded, center of anterior margin slightly excavated. The washboard with two small, weakly developed, symmetrical stridulation ribs, ending well in front of anterior margin (
|
||
<figureCitation id="134DC1939D17BA34CFA8FE53FC2777AB" box="[461,560,484,510]" captionStart="FIGURES 11" captionStartId="22.[264,370,1839,1860]" captionTargetBox="[272,1319,556,1825]" captionTargetId="figure-125@22.[264,1323,556,1833]" captionTargetPageId="22" captionText="FIGURES 11 A–B: Z. villosum sp. nov., female paratype. a: subanal plate with washboard; b:coxa of 2nd left leg pair with vulva. EP = exterior plate, IP = inner plate, O = operculum. Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051481" httpUri="https://zenodo.org/record/5051481/files/figure.png" pageId="22" pageNumber="23">Fig. 11a</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF098D9E9D17BA34CF6DF898FA977137" ID-DOI="http://doi.org/10.5281/zenodo.5051481" ID-Zenodo-Dep="5051481" httpUri="https://zenodo.org/record/5051481/files/figure.png" pageId="22" pageNumber="23" startId="22.[264,370,1839,1860]" targetBox="[272,1319,556,1825]" targetPageId="22">
|
||
<paragraph id="8BC9DD169D17BA34CF6DF898FA977137" blockId="22.[264,1323,1837,1890]" pageId="22" pageNumber="23">
|
||
<emphasis id="B90201049D17BA34CF6DF898FFD47111" bold="true" box="[264,451,1839,1860]" pageId="22" pageNumber="23">FIGURES 11 A–B</emphasis>
|
||
:
|
||
<taxonomicName id="4C76A6959D17BA34CFB5F887FC2C7110" authority="Wesener & Sierwald, 2005" authorityName="Wesener & Sierwald" authorityYear="2005" box="[464,571,1840,1861]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="villosum" status="sp. nov.">
|
||
<emphasis id="B90201049D17BA34CFB5F887FC2C7110" box="[464,571,1840,1861]" italics="true" pageId="22" pageNumber="23">Z. villosum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90201049D17BA34CC27F898FC987111" bold="true" box="[578,655,1839,1860]" pageId="22" pageNumber="23">
|
||
<taxonomicNameLabel id="A231BC7F9D17BA34CC27F898FC987111" box="[578,655,1839,1860]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female paratype. a: subanal plate with washboard; b:coxa of 2
|
||
<superScript id="7C03705E9D17BA34CA92F89AFB13716C" attach="left" box="[1271,1284,1837,1849]" fontSize="5" pageId="22" pageNumber="23">nd</superScript>
|
||
left leg pair with vulva. EP = exterior plate, IP = inner plate, O = operculum. Scale bars = 1 mm
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF098D9E9D16BA35CF6DF955FFAD7125" ID-DOI="http://doi.org/10.5281/zenodo.5051483" ID-Zenodo-Dep="5051483" httpUri="https://zenodo.org/record/5051483/files/figure.png" pageId="23" pageNumber="24" startId="23.[264,370,1762,1783]" targetBox="[274,1319,334,1747]" targetPageId="23">
|
||
<paragraph id="8BC9DD169D16BA35CF6DF955FC5F7106" blockId="23.[264,1325,1762,1904]" pageId="23" pageNumber="24">
|
||
<emphasis id="B90201049D16BA35CF6DF955FFD370A2" bold="true" box="[264,452,1762,1783]" pageId="23" pageNumber="24">FIGURES 12 A–F</emphasis>
|
||
:
|
||
<taxonomicName id="4C76A6959D16BA35CFB6F954FC2970AD" authority="Wesener & Sierwald, 2005" authorityName="Wesener & Sierwald" authorityYear="2005" box="[467,574,1763,1784]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="villosum" status="sp. nov.">
|
||
<emphasis id="B90201049D16BA35CFB6F954FC2970AD" box="[467,574,1763,1784]" italics="true" pageId="23" pageNumber="24">Z. villosum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90201049D16BA35CC23F955FC8470A2" bold="true" box="[582,659,1762,1783]" pageId="23" pageNumber="24">
|
||
<taxonomicNameLabel id="A231BC7F9D16BA35CC23F955FC8470A2" box="[582,659,1762,1783]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype. a: anterior telopods, anterior view; b: inner view of 2–3 joint of right anterior telopod; c: detail of tip of process of 2
|
||
<superScript id="7C03705E9D16BA35CD15F8B7FD6A7159" attach="left" box="[880,893,1792,1804]" fontSize="5" pageId="23" pageNumber="24">nd</superScript>
|
||
joint of right anterior telopod, inner view; d: right anterior telopod, posterior view; e: posterior pair of telopods, anterior view; f: right posterior telopod, posterior view. IH = inner horns.
|
||
</paragraph>
|
||
<paragraph id="8BC9DD169D16BA35CF6DF8ECFFAD7125" blockId="23.[264,1325,1762,1904]" box="[264,442,1883,1904]" pageId="23" pageNumber="24">Scale bars = 1 mm</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC9DD169D19BA3ACF5DFEABFF9477D3" blockId="24.[264,1323,284,390]" pageId="24" pageNumber="25">
|
||
<emphasis id="B90201049D19BA3ACF5DFEABFC4C7760" box="[312,603,284,309]" italics="true" pageId="24" pageNumber="25">Male sexual characters</emphasis>
|
||
: male gonopore covered by a single large, broad plate. Sclerotized part of plate with a few long hairs. Second pair of legs without coxal lobe (
|
||
<figureCitation id="134DC1939D19BA3ACF74FEDBFF6277D3" box="[273,373,364,390]" captionStart="FIGURE 13" captionStartId="24.[264,357,1481,1502]" captionTargetBox="[368,1213,435,1452]" captionTargetId="figure-189@24.[337,1245,430,1464]" captionTargetPageId="24" captionText="FIGURE 13 a. villosum sp. nov., male holotype. a: left coxa of 2nd pair of legs with male gonopore, posterior view. Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051487" httpUri="https://zenodo.org/record/5051487/files/figure.png" pageId="24" pageNumber="25">Fig. 13a</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF098D9E9D19BA3ACF6DFA7EFFE373A9" ID-DOI="http://doi.org/10.5281/zenodo.5051487" ID-Zenodo-Dep="5051487" httpUri="https://zenodo.org/record/5051487/files/figure.png" pageId="24" pageNumber="25" startId="24.[264,357,1481,1502]" targetBox="[368,1213,435,1452]" targetPageId="24">
|
||
<paragraph id="8BC9DD169D19BA3ACF6DFA7EFFE373A9" blockId="24.[264,1322,1479,1532]" pageId="24" pageNumber="25">
|
||
<emphasis id="B90201049D19BA3ACF6DFA7EFF93738B" bold="true" box="[264,388,1481,1502]" pageId="24" pageNumber="25">FIGURE 13</emphasis>
|
||
a.
|
||
<taxonomicName id="4C76A6959D19BA3ACFC7FA7EFFE4738B" authority="Wesener & Sierwald, 2005" authorityName="Wesener & Sierwald" authorityYear="2005" box="[418,499,1481,1502]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="villosum" status="sp. nov.">
|
||
<emphasis id="B90201049D19BA3ACFC7FA7EFFE4738B" box="[418,499,1481,1502]" italics="true" pageId="24" pageNumber="25">villosum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90201049D19BA3ACF9CFA7EFC51738B" bold="true" box="[505,582,1481,1502]" pageId="24" pageNumber="25">
|
||
<taxonomicNameLabel id="A231BC7F9D19BA3ACF9CFA7EFC51738B" box="[505,582,1481,1502]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype. a: left coxa of 2
|
||
<superScript id="7C03705E9D19BA3ACDE4FA70FD997386" attach="left" box="[897,910,1479,1491]" fontSize="5" pageId="24" pageNumber="25">nd</superScript>
|
||
pair of legs with male gonopore, posterior view. Scale bars = 1 mm
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC9DD169D19BA3BCF5DF996FADA770B" blockId="24.[264,1323,1569,1875]" lastBlockId="25.[264,1322,284,350]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
|
||
<emphasis id="B90201049D19BA3ACF5DF996FC1E706F" box="[312,521,1569,1594]" italics="true" pageId="24" pageNumber="25">Anterior telopods</emphasis>
|
||
: with three joints distally of syncoxite, inner margin of first joint with ‘harp’ consisting of one long and one short stridulation ridge, long ridge very strong and regular, curved centrally towards middle of the body, beginning on the posterior margin and ending just in front of the anterior margin of first joint (
|
||
<figureCitation id="134DC1939D19BA3ACA5CF92EFAB670E6" box="[1081,1185,1689,1715]" captionStart="FIGURES 12" captionStartId="23.[264,370,1762,1783]" captionTargetBox="[274,1319,334,1747]" captionTargetId="figure-88@23.[264,1323,324,1753]" captionTargetPageId="23" captionText="FIGURES 12 A–F: Z. villosum sp. nov., male holotype. a: anterior telopods, anterior view; b: inner view of 2–3 joint of right anterior telopod; c: detail of tip of process of 2nd joint of right anterior telopod, inner view; d: right anterior telopod, posterior view; e: posterior pair of telopods, anterior view; f: right posterior telopod, posterior view. IH = inner horns. Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051483" httpUri="https://zenodo.org/record/5051483/files/figure.png" pageId="24" pageNumber="25">Fig. 12a</figureCitation>
|
||
). Posterior side of second telopod joint at the inside with a projection, which reaches up to half the length of the third joint (
|
||
<figureCitation id="134DC1939D19BA3ACC4AF95EFC8F7156" box="[559,664,1769,1795]" captionStart="FIGURES 12" captionStartId="23.[264,370,1762,1783]" captionTargetBox="[274,1319,334,1747]" captionTargetId="figure-88@23.[264,1323,324,1753]" captionTargetPageId="23" captionText="FIGURES 12 A–F: Z. villosum sp. nov., male holotype. a: anterior telopods, anterior view; b: inner view of 2–3 joint of right anterior telopod; c: detail of tip of process of 2nd joint of right anterior telopod, inner view; d: right anterior telopod, posterior view; e: posterior pair of telopods, anterior view; f: right posterior telopod, posterior view. IH = inner horns. Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051483" httpUri="https://zenodo.org/record/5051483/files/figure.png" pageId="24" pageNumber="25">Fig. 12b</figureCitation>
|
||
). The apical part features round sclerotized spots (
|
||
<figureCitation id="134DC1939D19BA3ACA9EF95EFF21717E" captionStart="FIGURES 12" captionStartId="23.[264,370,1762,1783]" captionTargetBox="[274,1319,334,1747]" captionTargetId="figure-88@23.[264,1323,324,1753]" captionTargetPageId="23" captionText="FIGURES 12 A–F: Z. villosum sp. nov., male holotype. a: anterior telopods, anterior view; b: inner view of 2–3 joint of right anterior telopod; c: detail of tip of process of 2nd joint of right anterior telopod, inner view; d: right anterior telopod, posterior view; e: posterior pair of telopods, anterior view; f: right posterior telopod, posterior view. IH = inner horns. Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051483" httpUri="https://zenodo.org/record/5051483/files/figure.png" pageId="24" pageNumber="25">Fig. 12c</figureCitation>
|
||
). The third joint long and wellrounded, tapering distally, posterior side with large membranous invagination covered with few sclerotized spines, one of them extraordinarily big. Border of invagination with up to 11 sclerotized black teeth positioned juxtaposed the round sclerotized spots of the second joint (
|
||
<figureCitation id="134DC1939D18BA3BCA5CFEF3FAD7770B" box="[1081,1216,324,350]" captionStart="FIGURES 12" captionStartId="23.[264,370,1762,1783]" captionTargetBox="[274,1319,334,1747]" captionTargetId="figure-88@23.[264,1323,324,1753]" captionTargetPageId="23" captionText="FIGURES 12 A–F: Z. villosum sp. nov., male holotype. a: anterior telopods, anterior view; b: inner view of 2–3 joint of right anterior telopod; c: detail of tip of process of 2nd joint of right anterior telopod, inner view; d: right anterior telopod, posterior view; e: posterior pair of telopods, anterior view; f: right posterior telopod, posterior view. IH = inner horns. Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051483" httpUri="https://zenodo.org/record/5051483/files/figure.png" pageId="25" pageNumber="26">Figs 12b–d</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF098D9E9D18BA3BCF6DF8BAFA2C7135" ID-DOI="http://doi.org/10.5281/zenodo.5051489" ID-Zenodo-Dep="5051489" httpUri="https://zenodo.org/record/5051489/files/figure.png" pageId="25" pageNumber="26" startId="25.[264,370,1805,1826]" targetBox="[264,1323,409,1793]" targetPageId="25">
|
||
<paragraph id="8BC9DD169D18BA3BCF6DF8BAFA2C7135" blockId="25.[264,1322,1805,1888]" pageId="25" pageNumber="26">
|
||
<emphasis id="B90201049D18BA3BCF6DF8BAFFD97177" bold="true" box="[264,462,1805,1826]" pageId="25" pageNumber="26">FIGURES 14 A–D:</emphasis>
|
||
<taxonomicName id="4C76A6959D18BA3BCFB0F8B9FC577176" authority="Wesener & Sierwald, 2005" authorityName="Wesener & Sierwald" authorityYear="2005" box="[469,576,1806,1827]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="villosum" status="sp. nov.">
|
||
<emphasis id="B90201049D18BA3BCFB0F8B9FC577176" box="[469,576,1806,1827]" italics="true" pageId="25" pageNumber="26">Z. villosum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90201049D18BA3BCC23F8BAFC847177" bold="true" box="[582,659,1805,1826]" pageId="25" pageNumber="26">
|
||
<taxonomicNameLabel id="A231BC7F9D18BA3BCC23F8BAFC847177" box="[582,659,1805,1826]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male paratype, SEM. a: lateral view on antennae; b: lateral view on antennae joints 1–3; c: apical view on sensorial cones of antennae; d: endotergum, (1) = marginal ridge with external row of marginal bristles, (2) = band of flattened nodules, (3) = internal area.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF098D9E9D1BBA38CF6DFA23FD8273B1" ID-DOI="http://doi.org/10.5281/zenodo.5051492" ID-Zenodo-Dep="5051492" httpUri="https://zenodo.org/record/5051492/files/figure.png" pageId="26" pageNumber="27" startId="26.[264,370,1428,1449]" targetBox="[269,1323,328,1408]" targetPageId="26">
|
||
<paragraph id="8BC9DD169D1BBA38CF6DFA23FD8273B1" blockId="26.[264,1322,1428,1508]" pageId="26" pageNumber="27">
|
||
<emphasis id="B90201049D1BBA38CF6DFA23FFD073FC" bold="true" box="[264,455,1428,1449]" pageId="26" pageNumber="27">FIGURES 15 A–C</emphasis>
|
||
:
|
||
<taxonomicName id="4C76A6959D1BBA38CFB1FA22FC2873FF" authority="Wesener & Sierwald, 2005" authorityName="Wesener & Sierwald" authorityYear="2005" box="[468,575,1429,1450]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="villosum" status="sp. nov.">
|
||
<emphasis id="B90201049D1BBA38CFB1FA22FC2873FF" box="[468,575,1429,1450]" italics="true" pageId="26" pageNumber="27">Z. villosum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90201049D1BBA38CC23FA23FC8473FC" bold="true" box="[582,659,1428,1449]" pageId="26" pageNumber="27">
|
||
<taxonomicNameLabel id="A231BC7F9D1BBA38CC23FA23FC8473FC" box="[582,659,1428,1449]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male paratype, SEM. a: ventral view of gnathochilarium; b: ventral view of epipharynx; c: right palp of gnathochilarium with sensorial cones, dorsal view, arrow = field of sensorial cones, cP = central pads, LL = lingual lamella, P = palpus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC9DD169D1BBA39CF5DF985FD0E77FB" blockId="26.[264,1323,1586,1892]" lastBlockId="27.[264,1323,284,430]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
|
||
<emphasis id="B90201049D1BBA38CF5DF985FC0C701E" box="[312,539,1586,1611]" italics="true" pageId="26" pageNumber="27">Posterior telopods</emphasis>
|
||
: chela of very specific shape. Third joint very broad, the membranous concavity extending over only a part of the anterior margin. Four nonsclerotized spines, basal spine big, two smaller spines in the middle and a fourth spine on the distal end of the membranous invagination. Posterior side of third joint with oblique sclerotized teeth restricted to the membranous invagination. Immovable finger of second joint thin, but as long as movable finger. Inside of immovable finger invaginated, with round sclerotized knobs juxtaposed the tip of the membranous field of the movable finger (third joint). Remarkable is one short spine on the anterior side near the base of the immovable digit. Base of second and third joint densely covered with hairs, apically almost hairless. Syncoxite and first joint almost hairless. Tips of inner horns of telopod coxa with terminal portion of inner lobe bent posteriorly more than 25 degree. Subanal lobe densely covered with hairs (
|
||
<figureCitation id="134DC1939D1ABA39CCEEFE23FD1C77FB" box="[651,779,404,430]" captionStart="FIGURES 12" captionStartId="23.[264,370,1762,1783]" captionTargetBox="[274,1319,334,1747]" captionTargetId="figure-88@23.[264,1323,324,1753]" captionTargetPageId="23" captionText="FIGURES 12 A–F: Z. villosum sp. nov., male holotype. a: anterior telopods, anterior view; b: inner view of 2–3 joint of right anterior telopod; c: detail of tip of process of 2nd joint of right anterior telopod, inner view; d: right anterior telopod, posterior view; e: posterior pair of telopods, anterior view; f: right posterior telopod, posterior view. IH = inner horns. Scale bars = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.5051483" httpUri="https://zenodo.org/record/5051483/files/figure.png" pageId="27" pageNumber="28">Figs 12e–f</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF098D9E9D1ABA39CF6DFB8FFFAA7238" ID-DOI="http://doi.org/10.5281/zenodo.5051496" ID-Zenodo-Dep="5051496" httpUri="https://zenodo.org/record/5051496/files/figure.png" pageId="27" pageNumber="28" startId="27.[264,370,1080,1101]" targetBox="[264,1323,489,1053]" targetPageId="27">
|
||
<paragraph id="8BC9DD169D1ABA39CF6DFB8FFFAA7238" blockId="27.[264,1323,1080,1133]" pageId="27" pageNumber="28">
|
||
<emphasis id="B90201049D1ABA39CF6DFB8FFFD07218" bold="true" box="[264,455,1080,1101]" pageId="27" pageNumber="28">FIGURES 16 A–B</emphasis>
|
||
:
|
||
<taxonomicName id="4C76A6959D1ABA39CFB3FB8FFC557218" authority="Wesener & Sierwald, 2005" authorityName="Wesener & Sierwald" authorityYear="2005" box="[470,578,1080,1101]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="villosum" status="sp. nov.">
|
||
<emphasis id="B90201049D1ABA39CFB3FB8FFC557218" box="[470,578,1080,1101]" italics="true" pageId="27" pageNumber="28">Z. villosum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90201049D1ABA39CC2FFB8FFC8F7218" bold="true" box="[586,664,1080,1101]" pageId="27" pageNumber="28">
|
||
<taxonomicNameLabel id="A231BC7F9D1ABA39CC2FFB8FFC8F7218" box="[586,664,1080,1101]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male paratype, SEM. a: left mandible, condylus; b: left mandible, pectinate lamellae.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C36C8E9D9D1ABA3ECF5DFB17FCD3770B" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="distribution">
|
||
<paragraph id="8BC9DD169D1ABA39CF5DFB17FAD27386" blockId="27.[264,1323,1184,1891]" pageId="27" pageNumber="28">
|
||
<emphasis id="B90201049D1ABA39CF5DFB17FC7372EF" bold="true" box="[312,612,1184,1210]" pageId="27" pageNumber="28">Distribution & Ecology:</emphasis>
|
||
This species, belonging to the six largest sphaerotheriid species of the world, is only known from the small littoral forest of
|
||
<collectingRegion id="49B213F49D1ABA39CA7DFB7EFA8E72B6" box="[1048,1177,1225,1251]" country="Madagascar" name="Toamasina" pageId="27" pageNumber="28">Toamasina</collectingRegion>
|
||
, and may be endemic in this area. Ecological data, not yet available, may shed light on the function of the dense hair coverage on the tergites of this species. The hairs may have sensorial functions or may keep dirt particles away from the body of the animal. Such suggestions may indicate euedaphic and burrowing habits of this species. Behavioral observations can elucidate the function of the extraordinarily big movable finger of the posterior telopods. The dissected female, collected in the middle of
|
||
<date id="FFC8FBD69D1ABA39CD27FA0EFDD37386" box="[834,964,1465,1491]" pageId="27" pageNumber="28" value="1996-04">April 1996</date>
|
||
, did not contain eggs.
|
||
</paragraph>
|
||
<paragraph id="8BC9DD169D1ABA39CF5DFA57FD637146" blockId="27.[264,1323,1184,1891]" pageId="27" pageNumber="28">
|
||
<emphasis id="B90201049D1ABA39CF5DFA57FFFE73AF" bold="true" box="[312,489,1504,1530]" pageId="27" pageNumber="28">Conservation:</emphasis>
|
||
The fate of this species is dependent on the future of the littoral forest fragment near
|
||
<collectingRegion id="49B213F49D1ABA39CFD2F9BEFC2E7076" box="[439,569,1545,1571]" country="Madagascar" name="Toamasina" pageId="27" pageNumber="28">Toamasina</collectingRegion>
|
||
, where the specimens were collected. No other collecting sites have been recorded for this species thus far. All recently published records indicate a very fast destruction of the scattered remaining patches of littoral forest on the east coast of
|
||
<collectingCountry id="F3619D869D1ABA39CF6DF936FF8270CE" box="[264,405,1665,1691]" name="Madagascar" pageId="27" pageNumber="28">Madagascar</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFE7A0E79D1ABA39CFD5F936FC4870CE" author="Dumetz, N." box="[432,607,1665,1691]" pageId="27" pageNumber="28" pagination="273 - 315" refId="ref17844" refString="Dumetz, N. (1999) High plant diversity of lowland rainforest vestiges in eastern Madagascar. Biodiversity and Conservation 8, 273 - 315." type="journal article" year="1999">Dumetz 1999</bibRefCitation>
|
||
, deGouvenain & Srilander 2003). These largebodied millipedes play an important, albeit understudied role in the decomposition cycle (Ashwini & Sridhar 2003) and millipede populations will have to be carefully monitored if planned reforestation with endemic plants is to be successful.
|
||
</paragraph>
|
||
<paragraph id="8BC9DD169D1ABA3ECF5DF897FCD3770B" blockId="27.[264,1323,1184,1891]" lastBlockId="28.[264,1322,284,350]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
|
||
<emphasis id="B90201049D1ABA39CF5DF897FC4F716F" bold="true" box="[312,600,1824,1850]" pageId="27" pageNumber="28">Intraspecific variation:</emphasis>
|
||
The lack of material precludes examination of intraspecific variability. The
|
||
<specimenCount id="9D70169F9D1ABA39CFAEF8FEFC417136" box="[459,598,1865,1891]" pageId="27" pageNumber="28" type="male">three males</specimenCount>
|
||
show identical formed telopods. The
|
||
<typeStatus id="54CD63B49D1ABA39CA48F8FEFA857136" box="[1069,1170,1865,1891]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
|
||
selected for SEM shows a unique feature in sphaerotheriids, his antennae has seven joints before the small disc carrying the sensory cones.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |