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<document id="B1D1D17816AD57E95CD2863EA45548B0" ID-CLB-Dataset="31681" ID-DOI="10.11646/zootaxa.4341.2.2" ID-GBIF-Dataset="18949fd5-8f09-44ea-8dfa-c04ba71bcc7b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1039524" ID-ZooBank="05152B19-56AA-4CCD-A3C6-53EA3369A54C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509435537401" checkinUser="plazi" docAuthor="Ünal, Mustafa &amp; Beccaloni, George W." docDate="2017" docId="533F87C4FF92733926E2FDA3FC15CB97" docLanguage="en" docName="zootaxa.4341.2.2.pdf" docOrigin="Zootaxa 4341 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Colossopus redtenbacheri" docType="treatment" docVersion="8" lastPageNumber="220" masterDocId="AF06FFBCFF8B73222675FFCFFF90C85C" masterDocTitle="Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov." pageNumber="218" updateTime="1698470558790" updateUser="ExternalLinkService">
<mods:mods id="46A53E834ABC61C6BE935620F494C4B6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A093BBBEDB6762045A76E5A472191300">Revision of the Madagascan genera Oncodopus Brongniart and Colossopus Saussure (Orthoptera: Tettigoniidae: Conocephalinae; Euconchophorini), with description of Malagasopus gen. nov.</mods:title>
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<mods:namePart id="63F0017481437ADB7B6B79EF5BB1844A">Ünal, Mustafa</mods:namePart>
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<mods:namePart id="06E5AD5FAD60D636EBE7F8CC8B45E998">Beccaloni, George W.</mods:namePart>
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<mods:typeOfResource id="B3631D8D2C7E5DC3B6AE3813C78144BF">text</mods:typeOfResource>
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<mods:title id="621D5E7D95FA3C5CEE4490D527EBF278">Zootaxa</mods:title>
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<mods:date id="3A45302B4879CE59C9C71469EB25F0D1">2017</mods:date>
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<mods:number id="D11D5A5247E96E717153967C3147C0FB">2017-10-31</mods:number>
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<mods:number id="280C73A8B8AF45F466CB8707579D50AD">4341</mods:number>
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<mods:number id="687C8C2ED527BFCF37ACEDF3D4317EAB">2</mods:number>
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<mods:classification id="57C209A2E9EB33373D2EF3418D4D27A4">journal volume</mods:classification>
<mods:identifier id="B59A571970D81F42413F21BF400979B6" type="CLB-Dataset">31681</mods:identifier>
<mods:identifier id="EF2805F87DD8BA919D71220CD3CD85CA" type="DOI">10.11646/zootaxa.4341.2.2</mods:identifier>
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<treatment id="533F87C4FF92733926E2FDA3FC15CB97" ID-DOI="http://doi.org/10.5281/zenodo.5611867" ID-GBIF-Taxon="135489691" ID-Zenodo-Dep="5611867" LSID="urn:lsid:plazi:treatment:533F87C4FF92733926E2FDA3FC15CB97" httpUri="http://treatment.plazi.org/id/533F87C4FF92733926E2FDA3FC15CB97" lastPageId="27" lastPageNumber="220" pageId="25" pageNumber="218">
<subSubSection id="938C6559FF92733B26E2FDA3FE27CAFB" pageId="25" pageNumber="218" type="nomenclature">
<paragraph id="DB2936D2FF92733B26E2FDA3FD2DCADA" blockId="25.[151,1040,620,715]" box="[151,701,620,646]" pageId="25" pageNumber="218">
<heading id="806181BEFF92733B26E2FDA3FD2DCADA" bold="true" box="[151,701,620,646]" fontSize="11" level="1" pageId="25" pageNumber="218" reason="1">
<taxonomicName id="1C964D51FF92733B26E2FDA3FD2DCADA" ID-CoL="X8BW" authority="(Brongniart, 1897)" baseAuthorityName="Brongniart" baseAuthorityYear="1897" box="[151,701,620,646]" class="Insecta" family="Tettigoniidae" genus="Colossopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="218" phylum="Arthropoda" rank="species" species="redtenbacheri">
<emphasis id="E9E2EAC0FF92733B26E2FDA3FD2DCADA" bold="true" box="[151,701,620,646]" pageId="25" pageNumber="218">
<emphasis id="E9E2EAC0FF92733B26E2FDA3FE5ECADA" bold="true" box="[151,462,620,646]" italics="true" pageId="25" pageNumber="218">Colossopus redtenbacheri</emphasis>
(
<bibRefCitation id="BF074B23FF92733B27A8FDA3FD25CADA" author="Brongniart, C." box="[477,693,620,646]" journalOrPublisher="Bulletin de la Societ philomatique de Paris" pageId="25" pageNumber="218" pagination="120 - 212" part="8" refString="Brongniart, C. (1897) Revision des Salomonitae, Locustidae de la Tribu des Conocephalinae. Bulletin de la Societ philomatique de Paris, 8 (8), 120 - 212." title="Revision des Salomonitae, Locustidae de la Tribu des Conocephalinae" type="journal article" year="1897">Brongniart, 1897</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="DB2936D2FF92733B26E2FD41FE27CAFB" blockId="25.[151,1040,620,715]" box="[151,439,654,679]" pageId="25" pageNumber="218">
(
<figureCitation id="43AD2A57FF92733B26EAFD41FF7ECAFB" box="[159,238,654,679]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,264,1902,1924]" captionTargetBox="[190,1390,209,1856]" captionTargetId="figure@4.[176,1420,195,1876]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Fastigium of vertex from above. 1 Oncodopus zonatus, 2 Malagasopus desutterae, 3 Colossopus grandidieri. Figs. 4 – 6. apical half of left fore femur, inner view. 4 Oncodopus zonatus, 5 Malagasopus desutterae, 6 Colossopus redtenbacheri. Figs. 7 – 9. apical part of left hind tibia from above. 7 Oncodopus zonatus, 8 Malagasopus desutterae, 9 Colossopus grandidieri. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039526/files/figure.png" pageId="25" pageNumber="218">Figs. 6</figureCitation>
,
<figureCitation id="43AD2A57FF92733B268FFD40FED3CAFB" box="[250,323,655,679]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">87–97</figureCitation>
,
<figureCitation id="43AD2A57FF92733B273BFD40FEE8CAFB" box="[334,376,655,679]" captionStart="FIGURES 109 – 111" captionStartId="30.[151,264,1176,1198]" captionTargetBox="[153,1434,225,1132]" captionTargetId="figure@30.[151,1436,224,1135]" captionTargetPageId="30" captionText="FIGURES 109 – 111. Maps. 109 map of Madagascar, 110 distributions of Oncodopus spp., 111 distributions of Malagasopus spp. and Colossopus spp." httpUri="https://zenodo.org/record/1039548/files/figure.png" pageId="25" pageNumber="218">111</figureCitation>
,
<figureCitation id="43AD2A57FF92733B27F1FD40FE3FCAFB" box="[388,431,655,679]" captionStart="FIGURES 120 – 129" captionStartId="32.[151,264,1742,1764]" captionTargetBox="[151,1436,213,1703]" captionTargetId="figure@32.[151,1436,213,1704]" captionTargetPageId="32" captionText="FIGURES 120 – 129. Living specimens. 120 Colossopus grandidieri, older (swollen) egg. Captive stock. © G. J. N. González, 121 Colossopus grandidieri, recently hatched 1 st instar nymph. Captive stock. © G. J. N. González, 122 Colossopus grandidieri, late instar male nymph. Captive stock. © G. J. N. González, 123 Colossopus grandidieri, final instar female nymph. Captive stock. © G. J. N. González, 124 Colossopus grandidieri, adult female. Captive stock. © G. J. N. González, 125 Colossopus grandidieri, threat display of adult female, near village of Mangily, Toliara Province. 15.3. 2015. © M. Bader, 126 Colossopus grandidieri, adult male, near village of Mangily, Toliara Province. 19.11. 2004. © G. W. Beccaloni, 127 frontal view of head of the previous individual. © G. W. Beccaloni, 128 Colossopus redtenbacheri, adult female, Berenty Private Reserve, Toliara Province, 4.12. 2004. © G. W. Beccaloni, 129 Spiny forest near village of Mangily, Toliara Province in November 2004, close to where the second author collected Colossopus grandidieri and Oncodopus brongniarti. © G. W. Beccaloni." httpUri="https://zenodo.org/record/1039552/files/figure.png" pageId="25" pageNumber="218">128</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="938C6559FF92733B26E2FD7DFB80CA97" box="[151,1040,690,715]" pageId="25" pageNumber="218" type="description">
<paragraph id="DB2936D2FF92733B26E2FD7DFB80CA97" blockId="25.[151,1040,620,715]" box="[151,1040,690,715]" pageId="25" pageNumber="218">
http://lsid.speciesfile.org/urn:lsid:
<taxonomicName id="1C964D51FF92733B247FFD7DFD1BCA97" box="[522,651,690,715]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="218" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
.speciesfile.org:TaxonName:15117
</paragraph>
</subSubSection>
<subSubSection id="938C6559FF92733B26E2FD35FD7ACB12" pageId="25" pageNumber="218" type="reference_group">
<paragraph id="DB2936D2FF92733B26E2FD35FC49CB4C" blockId="25.[151,985,762,846]" box="[151,985,762,784]" pageId="25" pageNumber="218">
<treatmentCitationGroup id="FB8611FCFF92733B26E2FD35FC49CB4C" box="[151,985,762,784]" pageId="25" pageNumber="218">
<taxonomicName id="1C964D51FF92733B26E2FD35FD18CB4C" ID-CoL="49HSP" authority=": Brongniart 1897: 208" authorityName="Brongniart" authorityPageNumber="208" authorityYear="1897" box="[151,648,762,784]" class="Insecta" family="Tettigoniidae" genus="Oncodopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="218" phylum="Arthropoda" rank="species" species="redtenbacheri">
<emphasis id="E9E2EAC0FF92733B26E2FD35FE0DCB4C" box="[151,413,762,784]" italics="true" pageId="25" pageNumber="218">Oncodopus redtenbacheri</emphasis>
:
<treatmentCitation id="5A3710C3FF92733B27DEFD35FD18CB4C" author="Brongniart, C." box="[427,648,762,784]" page="208" pageId="25" pageNumber="218" year="1897">
<bibRefCitation id="BF074B23FF92733B27DEFD35FDC5CB4C" author="Brongniart, C." box="[427,597,762,784]" journalOrPublisher="Bulletin de la Societ philomatique de Paris" pageId="25" pageNumber="218" pagination="120 - 212" part="8" refString="Brongniart, C. (1897) Revision des Salomonitae, Locustidae de la Tribu des Conocephalinae. Bulletin de la Societ philomatique de Paris, 8 (8), 120 - 212." title="Revision des Salomonitae, Locustidae de la Tribu des Conocephalinae" type="journal article" year="1897">Brongniart 1897</bibRefCitation>
: 208
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="5A3710C3FF92733B24E7FD35FCACCB4C" author="Kirby, W. F." box="[658,828,762,784]" page="270" pageId="25" pageNumber="218" year="1906">
<bibRefCitation id="BF074B23FF92733B24E7FD35FC99CB4C" author="Kirby, W. F." bookContentInfo="562 pp." box="[658,777,762,784]" journalOrPublisher="Trustees of the British Museum, London" pageId="25" pageNumber="218" refString="Kirby, W. F. (1906) A synonymic catalogue of Orthoptera. Orthoptera Saltatoria (Achetidae et Phasgonuridae). Trustees of the British Museum, London, 562 pp." title="A synonymic catalogue of Orthoptera. Orthoptera Saltatoria (Achetidae et Phasgonuridae)" type="book" year="1906">Kirby 1906</bibRefCitation>
: 270
</treatmentCitation>
; Otte 1997: 32.
</treatmentCitationGroup>
</paragraph>
<paragraph id="DB2936D2FF92733B26E2FCD6FC94CB73" blockId="25.[151,985,762,846]" box="[151,772,793,815]" pageId="25" pageNumber="218">
<treatmentCitationGroup id="FB8611FCFF92733B26E2FCD6FC94CB73" box="[151,772,793,815]" pageId="25" pageNumber="218">
<taxonomicName id="1C964D51FF92733B26E2FCD6FC90CB73" ID-CoL="X8BW" authority=": Saussure 1899: 628" authorityName="Saussure" authorityPageNumber="628" authorityYear="1899" box="[151,768,793,815]" class="Insecta" family="Tettigoniidae" genus="Colossopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="218" phylum="Arthropoda" rank="species" species="redtenbacheri" subGenus="Oncodopus">
<emphasis id="E9E2EAC0FF92733B26E2FCD6FDB9CB73" box="[151,553,793,815]" italics="true" pageId="25" pageNumber="218">Colossopus (Oncodopus) redtenbacheri</emphasis>
:
<treatmentCitation id="5A3710C3FF92733B2442FCD6FC90CB73" author="Saussure, H." box="[567,768,793,815]" page="628" pageId="25" pageNumber="218" year="1899">
<bibRefCitation id="BF074B23FF92733B2442FCD6FD5DCB73" author="Saussure, H." box="[567,717,793,815]" journalOrPublisher="in Wissenschaftliche ergebnisse der reisen in Madagascar und Ostafrika in den jahren 1889 - 95 von Dr. A. Voeltzkow. Abhandlungen herausgegeben von der Senckenbergischen Naturforschenden Gesellschaft" pageId="25" pageNumber="218" pagination="567 - 664" part="21" refString="Saussure, H. (1899) Orthoptera, in Wissenschaftliche ergebnisse der reisen in Madagascar und Ostafrika in den jahren 1889 - 95 von Dr. A. Voeltzkow. Abhandlungen herausgegeben von der Senckenbergischen Naturforschenden Gesellschaft, 21, 567 - 664." title="Orthoptera" type="journal article" year="1899">Saussure 1899</bibRefCitation>
: 628
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="DB2936D2FF92733B26E2FCF7FD7ACB12" blockId="25.[151,985,762,846]" box="[151,746,824,846]" pageId="25" pageNumber="218">
<treatmentCitationGroup id="FB8611FCFF92733B26E2FCF7FD7ACB12" box="[151,746,824,846]" pageId="25" pageNumber="218">
<taxonomicName id="1C964D51FF92733B26E2FCF7FDDECB12" ID-CoL="X8BW" authority=": Karny 1912: 39" authorityName=": Karny" authorityPageNumber="39" authorityYear="1912" box="[151,590,824,846]" class="Insecta" family="Tettigoniidae" genus="Colossopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="25" pageNumber="218" phylum="Arthropoda" rank="species" species="redtenbacheri">
<emphasis id="E9E2EAC0FF92733B26E2FCF7FE0FCB12" box="[151,415,824,846]" italics="true" pageId="25" pageNumber="218">Colossopus redtenbacheri</emphasis>
:
<treatmentCitation id="5A3710C3FF92733B27D9FCF6FDDECB12" author="Karny, H. H." box="[428,590,824,846]" page="39" pageId="25" pageNumber="218" year="1912">
<bibRefCitation id="BF074B23FF92733B27D9FCF6FDB8CB12" author="Karny, H. H." box="[428,552,824,846]" editor="Wystman, P." journalOrPublisher="Bruxelles" pageId="25" pageNumber="218" pagination="1 - 47" refString="Karny, H. H. (1912). Orthoptera Fam. Locustidae Subfam. Agraeciinae. In: Wystman, P. (Ed.), Genera Insectorum. V. Verteneuil &amp; L. Desmet, Bruxelles, pp. 1 - 47." title="Orthoptera Fam. Locustidae Subfam. Agraeciinae" type="book chapter" volumeTitle="Genera Insectorum. V. Verteneuil &amp; L. Desmet" year="1912">Karny 1912</bibRefCitation>
: 39
</treatmentCitation>
</taxonomicName>
; Otte 1997: 27.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="938C6559FF92733B26E2FCB5FBD0CBCF" box="[151,1088,890,915]" pageId="25" pageNumber="218" type="materials_examined">
<paragraph id="DB2936D2FF92733B26E2FCB5FBD0CBCF" blockId="25.[151,1437,890,2031]" box="[151,1088,890,915]" pageId="25" pageNumber="218">
<emphasis id="E9E2EAC0FF92733B26E2FCB5FEA5CBCF" bold="true" box="[151,309,890,915]" pageId="25" pageNumber="218">Type locality.</emphasis>
Madagascar.
<emphasis id="E9E2EAC0FF92733B27A5FCB5FDD6CBCF" bold="true" box="[464,582,890,915]" pageId="25" pageNumber="218">Lectotype</emphasis>
male (in alcohol) (MNHN), here designated.
</paragraph>
</subSubSection>
<subSubSection id="938C6559FF92733B26B2FC51FE05CFB3" pageId="25" pageNumber="218" type="description">
<paragraph id="DB2936D2FF92733B26B2FC51FC38CE63" blockId="25.[151,1437,890,2031]" pageId="25" pageNumber="218">
<emphasis id="E9E2EAC0FF92733B26B2FC51FEE4CBEB" bold="true" box="[199,372,926,951]" pageId="25" pageNumber="218">Redescription.</emphasis>
Body size is intermediate between the other two members of the genus (
<tableCitation id="96140369FF92733B22D7FC51FB6ACBEB" box="[1186,1274,926,951]" captionStart="TABLE 4" captionStartId="29.[151,239,1445,1468]" captionTargetBox="[159,1426,1496,1980]" captionText="TABLE 4. Measurements (mm) of the body parts of Colossopus spp. M male, F female." httpUri="http://table.plazi.org/id/8FE9665AFF96733F26E2FA6AFBD5CDE0" pageId="25" pageNumber="218" tableUuid="8FE9665AFF96733F26E2FA6AFBD5CDE0">Table 4</tableCitation>
). Fastigium of vertex (
<figureCitation id="43AD2A57FF92733B2699FC0DFEF6CB87" box="[236,358,962,987]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">Fig. 87, 88</figureCitation>
) with three-pronged horn; upper 2 arms short and equal, slightly upcurved; lower one longer and situated between the upper arms; 1.4 times narrower than diameter of eye; its basal part 1.1 times wider than antennal scape. Fastigium of frons (
<figureCitation id="43AD2A57FF92733B244AFBC5FD05CC7F" box="[575,661,1034,1059]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">Fig. 88</figureCitation>
) with a large spiniform tooth. Face almost flattened with distinct tubercles some of which are like short spines. Pronotum (
<figureCitation id="43AD2A57FF92733B255DFBE1FC37CC1B" box="[808,935,1070,1095]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">Fig. 87, 88</figureCitation>
) slightly longer than wide, 3.4 times longer than its height; anterior margin convex, posterior margin with a deep angular median incision; prozona 1.5 times longer than metazona. Tegmina (
<figureCitation id="43AD2A57FF92733B2463FBB9FD09CCD3" box="[534,665,1142,1167]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">Fig. 87, 88</figureCitation>
) wavy, clearly visible from above, reaching 4/5 of the way down metanotum. Prothoracic auditory spiracle (
<figureCitation id="43AD2A57FF92733B24FBFB55FD75CCEF" box="[654,741,1178,1203]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">Fig. 90</figureCitation>
) almost rounded, almost fully covered by lateral lobes of pronotum. Mesothoracic auricle (
<figureCitation id="43AD2A57FF92733B2467FB71FDF5CC8B" box="[530,613,1214,1239]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">Fig. 90</figureCitation>
) oval, visible in lateral view, longer than wide, without ventral lobe; its inner gap long and elliptical. Prosternum with two, V-shaped spines, their apices not reaching to level of ventral margin of fore coxa. Fore coxa with a short dorsal spine. Fore femur (
<figureCitation id="43AD2A57FF92733B25A5FAC9FBB3CD43" box="[976,1059,1286,1311]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,264,1902,1924]" captionTargetBox="[190,1390,209,1856]" captionTargetId="figure@4.[176,1420,195,1876]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Fastigium of vertex from above. 1 Oncodopus zonatus, 2 Malagasopus desutterae, 3 Colossopus grandidieri. Figs. 4 – 6. apical half of left fore femur, inner view. 4 Oncodopus zonatus, 5 Malagasopus desutterae, 6 Colossopus redtenbacheri. Figs. 7 – 9. apical part of left hind tibia from above. 7 Oncodopus zonatus, 8 Malagasopus desutterae, 9 Colossopus grandidieri. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039526/files/figure.png" pageId="25" pageNumber="218">Figs. 6</figureCitation>
,
<figureCitation id="43AD2A57FF92733B2247FAC8FBDFCD43" box="[1074,1103,1287,1311]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">87</figureCitation>
) with a large inner spine on anterio-lateral margin; with a large inner and a large outer apical spine ventrally; with 5–6 additional ventral spines on inner side. Fore tibia (
<figureCitation id="43AD2A57FF92733B27C4FA81FD93CD3B" box="[433,515,1358,1383]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">Fig. 87</figureCitation>
) with 5 inner and 4–7 outer spines ventrally, inner ones larger. Mid femur with 2– 4 and hind femur with 1–3 outer spines ventrally. Hind tibia with 5–7 dorsal spines on inner side, with a single inner apical spur dorsally; with 4 apical spurs ventrally, inner two slightly shorter than outer ones. Last abdominal tergite (
<figureCitation id="43AD2A57FF92733B269BFA75FED1CD8F" box="[238,321,1466,1491]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">Fig. 91</figureCitation>
) twice as wide as its length, posterior margin with a deep narrow incision. Cercus (
<figureCitation id="43AD2A57FF92733B228FFA75FADDCD8F" box="[1274,1357,1466,1491]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">Fig. 92</figureCitation>
) short, basal part broad, end of basal part with a distinct projection on inner margin; apical part narrower and slightly incurved, apex blunt, slightly rounded, with a small apical tooth. Subgenital plate (
<figureCitation id="43AD2A57FF92733B223BF9CDFB31CE47" box="[1102,1185,1538,1563]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">Fig. 93</figureCitation>
) almost as long as its width, its styli slightly longer than depth of posterior incision of plate.
</paragraph>
<paragraph id="DB2936D2FF92733B26B2F985FE05CFB3" blockId="25.[151,1437,890,2031]" pageId="25" pageNumber="218">
Female: Head as in male. Fastigium of vertex (
<figureCitation id="43AD2A57FF92733B24A1F985FCCACE3F" box="[724,858,1610,1635]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">Figs. 94, 95</figureCitation>
) 1.4 times narrower than diameter of eye, 1.3 times wider than antennal scape. Pronotum (
<figureCitation id="43AD2A57FF92733B243DF9A1FD40CEDB" box="[584,720,1646,1671]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">Figs. 94, 95</figureCitation>
) with long, narrow prozona and wide, short metazona; prozona 1.4 times narrower and 4.3 times longer than metazona; anterior margin convex, posterior margin strongly concave. Tegmina strongly reduced to very small scale-like projections. Prothoracic auditory spiracle almost concealed by lateral lobe of pronotum. Mesothoracic auricle smaller than that of male, its inner gap narrow elliptical, not covered by pronotum. Fore tibia (
<figureCitation id="43AD2A57FF92733B27DBF931FE6ECF4B" box="[430,510,1790,1815]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">Fig. 94</figureCitation>
) with 5–6 ventral spines on outer side; mid femur with 1–4 and hind femur with 1– 4 outer spines ventrally; hind tibia with 6–7 dorsal spines on inner side, with an inner apical spur dorsally and 4 apical spurs ventrally. Last abdominal tergite broad and narrow, posterior margin with a small distinct median incision. Cercus short, conical, its apical part slightly incurved, pointed at apex. Subgenital plate (
<figureCitation id="43AD2A57FF92733B22A3F8A5FAB7CFDF" box="[1238,1319,1898,1923]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">Fig. 97</figureCitation>
) 3.5 times broader than long, with a broad rectangular median carina; lateral lobes with many transverse ridges. Ovipositor (
<figureCitation id="43AD2A57FF92733B26EAF87DFF63CF97" box="[159,243,1970,1995]" captionStart="FIGURES 87 – 97" captionStartId="26.[151,264,1905,1927]" captionTargetBox="[192,1404,197,1865]" captionTargetId="figure@26.[170,1426,193,1884]" captionTargetPageId="26" captionText="FIGURES 87 – 97. Colossopus redtenbacheri. Male: 87 forelegs, head and pronotum, dorsal view, 88 head and pronotum, lateral view, 89 habitus, dorso-lateral view, 90 prothoracic auditory spiracle and mesothoracic auricle, 91 last abdominal tergite, 92 cercus, 93 subgenital plate. Female: 94 forelegs, head and pronotum, dorsal view, 95 head and pronotum, lateral view, 96 habitus, dorso-lateral view, 97 subgenital plate. Scale bars 1 mm." httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="25" pageNumber="218">Fig. 96</figureCitation>
) straight, 1.5–1.6 times longer than hind femur and slightly longer than abdomen; apical part of ventral valve with small teeth.
</paragraph>
</subSubSection>
<caption id="8FE9665AFF91733826E2F8BEFCBACFBF" httpUri="https://zenodo.org/record/1039544/files/figure.png" pageId="26" pageNumber="219" targetBox="[192,1404,197,1865]" targetPageId="26">
<paragraph id="DB2936D2FF91733826E2F8BEFCBACFBF" blockId="26.[151,1437,1905,2019]" pageId="26" pageNumber="219">
<emphasis id="E9E2EAC0FF91733826E2F8BEFEC7CFDB" bold="true" box="[151,343,1905,1927]" pageId="26" pageNumber="219">FIGURES 87–97.</emphasis>
<taxonomicName id="1C964D51FF9173382714F8BEFDFCCFDB" authorityName=": Karny" authorityYear="1912" box="[353,620,1905,1927]" class="Insecta" family="Tettigoniidae" genus="Colossopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="26" pageNumber="219" phylum="Arthropoda" rank="species" species="redtenbacheri">
<emphasis id="E9E2EAC0FF9173382714F8BEFDFCCFDB" box="[353,620,1905,1927]" italics="true" pageId="26" pageNumber="219">Colossopus redtenbacheri</emphasis>
</taxonomicName>
. Male:
<emphasis id="E9E2EAC0FF91733824B4F8BEFD4ACFDB" bold="true" box="[705,730,1905,1927]" pageId="26" pageNumber="219">87</emphasis>
forelegs, head and pronotum, dorsal view,
<emphasis id="E9E2EAC0FF91733822DDF8BEFB52CFDB" bold="true" box="[1192,1218,1905,1927]" pageId="26" pageNumber="219">88</emphasis>
head and pronotum, lateral view,
<emphasis id="E9E2EAC0FF9173382762F840FEA0CFF9" bold="true" box="[279,304,1935,1957]" pageId="26" pageNumber="219">89</emphasis>
habitus, dorso-lateral view,
<emphasis id="E9E2EAC0FF9173382438F840FDF7CFF9" bold="true" box="[589,615,1935,1957]" pageId="26" pageNumber="219">90</emphasis>
prothoracic auditory spiracle and mesothoracic auricle,
<emphasis id="E9E2EAC0FF91733822E9F840FB26CFF9" bold="true" box="[1180,1206,1935,1957]" pageId="26" pageNumber="219">91</emphasis>
last abdominal tergite,
<emphasis id="E9E2EAC0FF91733826E2F861FF21CF98" bold="true" box="[151,177,1966,1988]" pageId="26" pageNumber="219">92</emphasis>
cercus,
<emphasis id="E9E2EAC0FF9173382773F861FE8FCF98" bold="true" box="[262,287,1966,1988]" pageId="26" pageNumber="219">93</emphasis>
subgenital plate. Female:
<emphasis id="E9E2EAC0FF9173382445F861FDDACF98" bold="true" box="[560,586,1966,1988]" pageId="26" pageNumber="219">94</emphasis>
forelegs, head and pronotum, dorsal view,
<emphasis id="E9E2EAC0FF917338227DF861FBB2CF98" bold="true" box="[1032,1058,1966,1988]" pageId="26" pageNumber="219">95</emphasis>
head and pronotum, lateral view,
<emphasis id="E9E2EAC0FF91733823F6F861FA0DCF98" bold="true" box="[1411,1437,1966,1988]" pageId="26" pageNumber="219">96</emphasis>
habitus, dorso-lateral view,
<emphasis id="E9E2EAC0FF91733827DAF802FE58CFBF" bold="true" box="[431,456,1997,2019]" pageId="26" pageNumber="219">97</emphasis>
subgenital plate. Scale bars 1 mm.
</paragraph>
</caption>
<subSubSection id="938C6559FF90733926B2FF58FAE6C91C" pageId="27" pageNumber="220" type="materials_examined">
<paragraph id="DB2936D2FF90733926B2FF58FAE6C91C" blockId="27.[151,1437,151,971]" pageId="27" pageNumber="220">
<emphasis id="E9E2EAC0FF90733926B2FF58FE7AC8EC" bold="true" box="[199,490,151,176]" pageId="27" pageNumber="220">Colour (dry specimens).</emphasis>
Body with different shades of brown, mostly milky and light brown. Face darker, reddish brown; clypeus and labrum yellowish brown. Metazona of pronotum darker with reddish brown, prozona light brown. Fore tibia, anterior part of fore femur, knees of mid and hind legs and tarsi reddish brown, remaining parts of legs milky brown. Male tegmina yellowish brown. Abdominal tergites reddish brown, with a very narrow black stripe on hind margin in female. Basal part of ovipositor milky brown, its apical part light reddish brown.
</paragraph>
</subSubSection>
<subSubSection id="938C6559FF90733926B2FE83FEEFC9A9" pageId="27" pageNumber="220" type="diagnosis">
<paragraph id="DB2936D2FF90733926B2FE83FEEFC9A9" blockId="27.[151,1437,151,971]" pageId="27" pageNumber="220">
<emphasis id="E9E2EAC0FF90733926B2FE83FED1C939" bold="true" box="[199,321,332,357]" pageId="27" pageNumber="220">Diagnosis.</emphasis>
This species differs from
<taxonomicName id="1C964D51FF907339240FFE82FC8EC939" authorityName=": Saussure" authorityYear="1899" box="[634,798,332,357]" class="Insecta" family="Tettigoniidae" genus="Colossopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="27" pageNumber="220" phylum="Arthropoda" rank="species" species="grandidieri">
<emphasis id="E9E2EAC0FF907339240FFE82FC8EC939" box="[634,798,332,357]" italics="true" pageId="27" pageNumber="220">C. grandidieri</emphasis>
</taxonomicName>
in the structure of the pronotum, male cercus, shorter ovipositor, the ratios of ovipositor/hind femur and ovipositor/abdomen, clearly lighter body colour and smaller size. It differs from
<taxonomicName id="1C964D51FF9073392701FE5AFD9FC9F0" box="[372,527,405,428]" class="Insecta" family="Tettigoniidae" genus="Colossopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="27" pageNumber="220" phylum="Arthropoda" rank="species" species="parvicavus" status="sp. nov.">
<emphasis id="E9E2EAC0FF9073392701FE5AFD9FC9F0" box="[372,527,405,428]" italics="true" pageId="27" pageNumber="220">C. parvicavus</emphasis>
</taxonomicName>
<emphasis id="E9E2EAC0FF9073392463FE5BFDFFC9F1" bold="true" box="[534,623,404,429]" pageId="27" pageNumber="220">
<taxonomicNameLabel id="F2D157BBFF9073392463FE5BFDFFC9F1" box="[534,623,404,429]" pageId="27" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the shape of the male pronotum, prothoracic auditory spiracle (which has a wide opening), the presence of ventral spines on the hind femur, and the ratios of ovipositor/hind femur and ovipositor/abdomen.
</paragraph>
</subSubSection>
<subSubSection id="938C6559FF90733926B2FE30FD7DCA93" pageId="27" pageNumber="220" type="materials_examined">
<paragraph id="DB2936D2FF90733926B2FE30FD7DCA93" blockId="27.[151,1437,151,971]" pageId="27" pageNumber="220">
<materialsCitation id="6BFE3C8FFF90733926B2FE30FCEFCA44" ID-GBIF-Occurrence="1676482555" box="[199,895,511,536]" country="Madagascar" location="Madagascar" pageId="27" pageNumber="220" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis id="E9E2EAC0FF90733926B2FE30FE24CA44" bold="true" box="[199,436,511,536]" pageId="27" pageNumber="220">Material examined.</emphasis>
<collectingCountry id="A3817642FF90733927CBFE30FDD7CA44" box="[446,583,511,536]" name="Madagascar" pageId="27" pageNumber="220">Madagascar</collectingCountry>
, 1867,
<specimenCount id="CD90FD5BFF90733924EAFDCFFD7ECA44" box="[671,750,511,536]" pageId="27" pageNumber="220" type="male">1 male</specimenCount>
(
<typeStatus id="042D8870FF9073392574FDCFFCE7CA44" box="[769,887,512,536]" pageId="27" pageNumber="220" type="lectotype">Lectotype</typeStatus>
)
</materialsCitation>
,
<materialsCitation id="6BFE3C8FFF90733925F8FDCFFEA7CA3D" ID-GBIF-Occurrence="1676482568" collectingDate="2004-01-12" collectionCode="MNHN" collectorName="M. Grandidier" country="Madagascar" latitude="-24.93" location="Tsimelahy" municipality="Andohahela National Park" pageId="27" pageNumber="220" specimenCount="1" specimenCount-female="1" stateProvince="Toliara Province" typeStatus="paralectotype">
<specimenCount id="CD90FD5BFF90733925F8FDCFFC62CA44" box="[909,1010,511,536]" pageId="27" pageNumber="220" type="female">1 female</specimenCount>
(
<typeStatus id="042D8870FF9073392271FE30FB30CA44" box="[1028,1184,511,536]" pageId="27" pageNumber="220" type="paralectotype">Paralectotype</typeStatus>
) (leg.
<collectorName id="76635304FF907339229AFDCFFA04CA44" box="[1263,1428,511,536]" pageId="27" pageNumber="220">M. Grandidier</collectorName>
) (
<collectionCode id="BD87AE17FF90733926EAFDEAFF6BCA61" box="[159,251,549,573]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="220">MNHN</collectionCode>
).
<collectingRegion id="1952F830FF9073392778FDEBFE5BCA61" box="[269,459,548,573]" country="Madagascar" name="Toliara" pageId="27" pageNumber="220">Toliara Province</collectingRegion>
,
<collectingMunicipality id="3B4DACA8FF90733927A3FDEBFC95CA61" box="[470,773,548,573]" pageId="27" pageNumber="220">Andohahela National Park</collectingMunicipality>
:
<location id="DE496009FF9073392565FDEBFC14CA61" LSID="urn:lsid:plazi:treatment:533F87C4FF92733926E2FDA3FC15CB97:DE496009FF9073392565FDEBFC14CA61" box="[784,900,548,573]" country="Madagascar" latitude="-24.93" municipality="Andohahela National Park" name="Tsimelahy" pageId="27" pageNumber="220" stateProvince="Toliara Province">Tsimelahy</location>
parcel, (-24.93, 46.65) [
<geoCoordinate id="BEA25015FF90733922E3FDEAFAB0CA61" box="[1174,1312,547,573]" direction="south" orientation="latitude" pageId="27" pageNumber="220" precision="15" value="-24.93">
24
<emphasis id="E9E2EAC0FF90733922C1FDECFB2FCA61" box="[1204,1215,547,573]" italics="true" pageId="27" pageNumber="220">°</emphasis>
55'48'' S
</geoCoordinate>
, 46
<emphasis id="E9E2EAC0FF907339233CFDECFAC4CA61" box="[1353,1364,547,573]" italics="true" pageId="27" pageNumber="220">°</emphasis>
38'60'' E],
<date id="AF281012FF90733926B1FD86FEA7CA3D" box="[196,311,585,609]" pageId="27" pageNumber="220" value="2004-01-12">
<collectingDate id="BF6CE9FAFF90733926B1FD86FEA7CA3D" box="[196,311,585,609]" pageId="27" pageNumber="220" value="2004-01-12">1.12.2004</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="6BFE3C8FFF907339273DFD86FA84CADA" ID-GBIF-Occurrence="1676482560" collectingDate="1961-01" collectionCode="NHMUK" collectorName="Coll." country="Madagascar" location="Tolagnaro" municipality="Tsimelahy" pageId="27" pageNumber="220" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Toliara" typeStatus="paralectotype">
<collectorName id="76635304FF907339273DFD86FEEFCA3E" box="[328,383,585,610]" pageId="27" pageNumber="220">Coll.</collectorName>
leg. G.W. Beccaloni,
<specimenCount id="CD90FD5BFF90733924F2FD86FD4ACA3E" box="[647,730,585,610]" pageId="27" pageNumber="220" type="male">1 male</specimenCount>
,
<specimenCount id="CD90FD5BFF907339249EFD86FCC1CA3E" box="[747,849,585,610]" pageId="27" pageNumber="220" type="female">1 female</specimenCount>
(
<collectionCode id="BD87AE17FF9073392510FD86FC46CA3D" box="[869,982,585,609]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="27" pageNumber="220">NHMUK</collectionCode>
).
<collectingCountry id="A3817642FF9073392598FD86FBE6CA3E" box="[1005,1142,585,610]" name="Madagascar" pageId="27" pageNumber="220">Madagascar</collectingCountry>
,
<collectingRegion id="1952F830FF90733922F3FD86FB45CA3E" box="[1158,1237,585,610]" country="Madagascar" name="Toliara" pageId="27" pageNumber="220">Toliara</collectingRegion>
Prov., Parc Nat. d’Andohahela, Forét d’Ambohibory,
<quantity id="1C6E9B37FF907339243DFDA1FD0BCADA" box="[584,667,621,646]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="27" pageNumber="220" unit="km" value="1.7">1.7 km</quantity>
<date id="AF281012FF90733924DDFDA2FC96CAD9" box="[680,774,621,646]" pageId="27" pageNumber="220" value="1961-01">
<collectingDate id="BF6CE9FAFF90733924DDFDA2FC96CAD9" box="[680,774,621,646]" pageId="27" pageNumber="220" value="1961-01">61 ENE</collectingDate>
</date>
<collectingMunicipality id="3B4DACA8FF9073392567FDA2FC17CADA" box="[786,903,621,646]" pageId="27" pageNumber="220">Tsimelahy</collectingMunicipality>
,
<quantity id="1C6E9B37FF90733925EDFDA2FC69CADA" box="[920,1017,621,646]" metricMagnitude="4" metricUnit="m" metricValue="3.61" pageId="27" pageNumber="220" unit="km" value="36.1">36.1 km</quantity>
308 NW
<location id="DE496009FF9073392202FDA2FB7BCADA" LSID="urn:lsid:plazi:treatment:533F87C4FF92733926E2FDA3FC15CB97:DE496009FF9073392202FDA2FB7BCADA" box="[1143,1259,621,646]" country="Madagascar" municipality="Tsimelahy" name="Tolagnaro" pageId="27" pageNumber="220" stateProvince="Toliara">Tolagnaro</location>
, 16
</materialsCitation>
–
<materialsCitation id="6BFE3C8FFF9073392356FDA2FD7DCA93" ID-GBIF-Occurrence="1676482566" collectingDate="2002-01-20" collectionCode="CAS" country="Madagascar" latitude="-24.93" location="Castlot" longLatPrecision="21" longitude="46.645557" pageId="27" pageNumber="220" specimenCount="10" specimenCount-female="4" specimenCount-male="6" stateProvince="Toliara" typeStatus="paralectotype">
<date id="AF281012FF9073392356FDA2FA08CADA" box="[1315,1432,621,646]" pageId="27" pageNumber="220" value="2002-01-20">
<collectingDate id="BF6CE9FAFF9073392356FDA2FA08CADA" box="[1315,1432,621,646]" pageId="27" pageNumber="220" value="2002-01-20">20.1.2002</collectingDate>
</date>
,
<geoCoordinate id="BEA25015FF90733926E2FD5DFEB0CAF6" box="[151,288,656,683]" direction="south" orientation="latitude" pageId="27" pageNumber="220" precision="15" value="-24.93">
24
<emphasis id="E9E2EAC0FF90733926C6FD5FFF2ECAF6" box="[179,190,656,682]" italics="true" pageId="27" pageNumber="220">°</emphasis>
55'48&quot; S
</geoCoordinate>
,
<geoCoordinate id="BEA25015FF9073392758FD5DFE28CAF6" box="[301,440,656,683]" direction="east" orientation="longitude" pageId="27" pageNumber="220" precision="15" value="46.645557">
46
<emphasis id="E9E2EAC0FF907339273CFD5FFEC4CAF6" box="[329,340,656,682]" italics="true" pageId="27" pageNumber="220">°</emphasis>
38'44&quot; E
</geoCoordinate>
,
<specimenCount id="CD90FD5BFF90733927B0FD5DFD8FCAF7" box="[453,543,658,683]" pageId="27" pageNumber="220" type="male">2 males</specimenCount>
,
<specimenCount id="CD90FD5BFF907339245EFD5DFD1CCAF7" box="[555,652,658,683]" pageId="27" pageNumber="220" type="female">1 female</specimenCount>
(Fisher, Griswold
<emphasis id="E9E2EAC0FF9073392514FD5CFC03CAF7" box="[865,915,658,683]" italics="true" pageId="27" pageNumber="220">et al</emphasis>
.), tropical dry forest;
<location id="DE496009FF90733922F9FD5DFB4BCAF7" LSID="urn:lsid:plazi:treatment:533F87C4FF92733926E2FDA3FC15CB97:DE496009FF90733922F9FD5DFB4BCAF7" box="[1164,1243,658,683]" country="Madagascar" latitude="-24.93" longLatPrecision="21" longitude="46.645557" name="Castlot" pageId="27" pageNumber="220" stateProvince="Toliara">Castlot</location>
0 0 6432 [locality unknown],
<specimenCount id="CD90FD5BFF9073392763FD78FEFFCA93" box="[278,367,694,719]" pageId="27" pageNumber="220" type="male">3 males</specimenCount>
,
<specimenCount id="CD90FD5BFF907339270EFD78FE7ACA93" box="[379,490,694,719]" pageId="27" pageNumber="220" type="female">3 females</specimenCount>
,
<specimenCount id="CD90FD5BFF9073392780FD78FDD0CA93" box="[501,576,694,719]" pageId="27" pageNumber="220" type="male">1 male</specimenCount>
nymph (
<collectionCode id="BD87AE17FF90733924D1FD78FD70CA93" box="[676,736,695,719]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="27" pageNumber="220">CAS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="938C6559FF90733926B2FD15FCFACB67" pageId="27" pageNumber="220" type="description">
<paragraph id="DB2936D2FF90733926B2FD15FCFACB67" blockId="27.[151,1437,151,971]" pageId="27" pageNumber="220">
<emphasis id="E9E2EAC0FF90733926B2FD15FE55CAAF" bold="true" box="[199,453,730,755]" pageId="27" pageNumber="220">Measurements (mm).</emphasis>
<typeStatus id="042D8870FF90733927B9FD14FDACCAAF" box="[460,572,731,755]" pageId="27" pageNumber="220" type="lectotype">Lectotype</typeStatus>
(male): Body length: 40; pronotum length: 10.6; width of pronotum: 10; fore femur length: 12.9; width of fore femur: 4.6; fore tibia length: 10.3; hind femur length: 14.4. See
<tableCitation id="96140369FF9073392281FD31FAD8CB4B" box="[1268,1352,766,791]" captionStart="TABLE 4" captionStartId="29.[151,239,1445,1468]" captionTargetBox="[159,1426,1496,1980]" captionText="TABLE 4. Measurements (mm) of the body parts of Colossopus spp. M male, F female." httpUri="http://table.plazi.org/id/8FE9665AFF96733F26E2FA6AFBD5CDE0" pageId="27" pageNumber="220" tableUuid="8FE9665AFF96733F26E2FA6AFBD5CDE0">Table 4</tableCitation>
for the range of measurement of the other specimens including females.
</paragraph>
</subSubSection>
<subSubSection id="938C6559FF90733926B2FC89FDF9CBFB" pageId="27" pageNumber="220" type="distribution">
<paragraph id="DB2936D2FF90733926B2FC89FB56CB03" blockId="27.[151,1437,151,971]" box="[199,1222,838,863]" pageId="27" pageNumber="220">
<emphasis id="E9E2EAC0FF90733926B2FC89FEF0CB03" bold="true" box="[199,352,838,863]" pageId="27" pageNumber="220">Distribution.</emphasis>
South-eastern
<collectingCountry id="A3817642FF907339247DFC89FD1FCB03" box="[520,655,838,863]" name="Madagascar" pageId="27" pageNumber="220">Madagascar</collectingCountry>
(
<figureCitation id="43AD2A57FF90733924EBFC89FD6FCB03" box="[670,767,838,863]" captionStart="FIGURES 109 – 111" captionStartId="30.[151,264,1176,1198]" captionTargetBox="[153,1434,225,1132]" captionTargetId="figure@30.[151,1436,224,1135]" captionTargetPageId="30" captionText="FIGURES 109 – 111. Maps. 109 map of Madagascar, 110 distributions of Oncodopus spp., 111 distributions of Malagasopus spp. and Colossopus spp." httpUri="https://zenodo.org/record/1039548/files/figure.png" pageId="27" pageNumber="220">Fig. 111</figureCitation>
).
<collectingRegion id="1952F830FF9073392567FC89FC40CB03" box="[786,976,838,863]" country="Madagascar" name="Toliara" pageId="27" pageNumber="220">Toliara Province</collectingRegion>
:
<collectingRegion id="1952F830FF90733925AFFC89FBBCCB03" box="[986,1068,838,863]" country="Madagascar" name="Toliara" pageId="27" pageNumber="220">Toliara</collectingRegion>
, Andohahela.
</paragraph>
<paragraph id="DB2936D2FF90733926B2FCA5FDF9CBFB" blockId="27.[151,1437,151,971]" pageId="27" pageNumber="220">
<emphasis id="E9E2EAC0FF90733926B2FCA5FEBACBDF" bold="true" box="[199,298,874,899]" pageId="27" pageNumber="220">Habitat.</emphasis>
This species has been found in transitional tropical dry forest. The specimens collected by the second author were found under loose dead bark.
</paragraph>
</subSubSection>
<subSubSection id="938C6559FF90733926B2FC7DFC15CB97" box="[199,901,946,971]" pageId="27" pageNumber="220" type="description">
<paragraph id="DB2936D2FF90733926B2FC7DFC15CB97" blockId="27.[151,1437,151,971]" box="[199,901,946,971]" pageId="27" pageNumber="220">
<emphasis id="E9E2EAC0FF90733926B2FC7DFED9CB97" bold="true" box="[199,329,946,971]" pageId="27" pageNumber="220">Phenology.</emphasis>
Adults have been found in December and January.
</paragraph>
</subSubSection>
</treatment>
</document>
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