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<document ID-CLB-Dataset="6725" ID-DOI="http://dx.doi.org/10.3897/dez.68.59462" ID-GBIF-Dataset="0e6618c3-3d76-400b-8615-4d1c972b9a81" ID-Pensoft-Pub="1860-1324-1-1" ID-Pensoft-UUID="D3A115DA614E53B1BD1AAD6B78A8F38D" ID-ZooBank="458E50692CA2436AABF5BEFE6EBAC3F0" ModsDocID="1860-1324-68-1-1" checkinTime="1621372303991" checkinUser="pensoft" docAuthor="Kaltenbach, Thomas, Garces, Jhoana M. &amp; Gattolliat, Jean-Luc" docDate="2021" docId="46FC58C7148A566E8C27FE7D37840A91" docLanguage="en" docName="DeutEntomolZeit 68(1): 1-20" docOrigin="Deutsche Entomologische Zeitschrift 68 (1)" docPubDate="2021-01-05" docSource="http://dx.doi.org/10.3897/dez.68.59462" docTitle="Philibaetis Kaltenbach &amp; Gattolliat 2021, gen. nov." docType="treatment" docUuid="5C936E67-64CD-43BC-AC21-35709769C2ED" docUuidSource="ZooBank" docVersion="5" id="D3A115DA614E53B1BD1AAD6B78A8F38D" lastPageNumber="1" masterDocId="D3A115DA614E53B1BD1AAD6B78A8F38D" masterDocTitle="Philibaetis gen. nov., a new genus from the Philippines (Ephemeroptera, Baetidae)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="1" updateTime="1732938980440" updateUser="ExternalLinkService">
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<mods:title id="77C83FE2B8E20E88D3FD592609756BCD">Philibaetis gen. nov., a new genus from the Philippines (Ephemeroptera, Baetidae)</mods:title>
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<mods:namePart id="BFF3F7A603DAAFF72F8D42CFCABCEB23">Kaltenbach, Thomas</mods:namePart>
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<mods:affiliation id="140BD8169D67F2EF216952D0953F40CE">Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<mods:namePart id="8ADBE9BBE8C3959B8B266A0AC8F593BC">Garces, Jhoana M.</mods:namePart>
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<mods:affiliation id="36F7DA8239E3466DBCAA78F72235FF62">Department of Biology, School of Science and Engineering, Ateneo de Manila University, Quezon City, 1108 Metro Manila, Philippines</mods:affiliation>
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<mods:namePart id="D200083B4A36337E3F77693EC3B217D4">Gattolliat, Jean-Luc</mods:namePart>
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<mods:affiliation id="D8935AA84CDC4764AB1B6A6ABE0BCBF8">Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<treatment id="46FC58C7148A566E8C27FE7D37840A91" ID-GBIF-Taxon="182223401" LSID="urn:lsid:zoobank.org:act:5C936E67-64CD-43BC-AC21-35709769C2ED" httpUri="http://treatment.plazi.org/id/46FC58C7148A566E8C27FE7D37840A91" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Baetidae" scope_order="Ephemeroptera">
<subSubSection id="B6BE51C0531C2D6858A1DC9EF0120820" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="DB146222690FA60D0B3890360E63E1D3" pageId="0" pageNumber="1">
<taxonomicName id="CCC7EA1189E1D2FE22CC31499C3B5A4F" LSID="http://zoobank.org/5C936E67-64CD-43BC-AC21-35709769C2ED" authority="Kaltenbach &amp; Gattolliat" authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2021" class="Insecta" family="Baetidae" genus="Philibaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philibaetis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus" status="gen. nov.">Philibaetis Kaltenbach &amp; Gattolliat</taxonomicName>
<taxonomicNameLabel id="87B2BBC24AAC03EF09D08C475136B87B" pageId="0" pageNumber="1">gen. nov.</taxonomicNameLabel>
<figureCitation id="F406ADD4F80323CF12B413F69F9DCD72" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Philibaetis luzonensis comb. nov., larva morphology: a. Labrum, dorsal view; b. Labrum, ventral view; c. Seta on anterolateral margin; d. Medial setae; e. Labrum, ventral view, submarginal row of setae; f. Setae of submarginal row; g. Right mandible; h. Right prostheca; i. Left mandible; j. Left prostheca; k. Tuft of setae; l. Right incisor, kinetodontium and prostheca; m. Left incisor, kinetodontium and prostheca." figureDoi="10.3897/dez.68.59462.figure1" httpUri="https://binary.pensoft.net/fig/494102" pageId="0" pageNumber="1">Figures 1</figureCitation>
<figureCitation id="A66A0E651575B87B3DBE414EEA159688" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Philibaetis luzonensis comb. nov., larva morphology: a. Hypopharynx and superlinguae; b. Maxilla; c. Distal and middle denti-setae; d. Labium (left side: ventral view; right side: dorsal view); e. Posterolateral seta on ventral side of paraglossa." figureDoi="10.3897/dez.68.59462.figure2" httpUri="https://binary.pensoft.net/fig/494103" pageId="0" pageNumber="1">, 2</figureCitation>
<figureCitation id="FC80A59D2344F5260A631D766CAF9AB6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Philibaetis luzonensis comb. nov., larva morphology: a. Foreleg; b. Claw; c. Apex of femur (posterior view); d. Basal part of antenna." figureDoi="10.3897/dez.68.59462.figure3" httpUri="https://binary.pensoft.net/fig/494104" pageId="0" pageNumber="1">, 3</figureCitation>
<figureCitation id="5C9AB7F4EB92F37E338123B59112C9B4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Philibaetis luzonensis comb. nov., larva morphology: a. Gill IV; b. Inner margin of gill IV; c. Outer margin of gill IV; d. Gills I-VII; e. Paraproct; f. Patch of notched scales; g. Metanotum; h. Tergite IV." figureDoi="10.3897/dez.68.59462.figure4" httpUri="https://binary.pensoft.net/fig/494105" pageId="0" pageNumber="1">, 4</figureCitation>
<figureCitation id="B33A31276A6025EBB469FF58571E0CF5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Philibaetis luzonensis comb. nov., habitus, larva: a. Dorsal view; b. Lateral view; c. Ventral view." figureDoi="10.3897/dez.68.59462.figure5" httpUri="https://binary.pensoft.net/fig/494106" pageId="0" pageNumber="1">, 5</figureCitation>
<figureCitation id="CD584F097C465E902BDA039E1E4BB26F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Philibaetis realonae comb. nov., larva morphology: a. Labrum; b. Setae of submarginal row (ventral view); c. Right mandible; d. Right prostheca; e. Left mandible; f. Left prostheca; g. Tuft of setae; h. Hypopharynx and superlinguae; i. Maxilla; j. Labium (left side: ventral view; right side: dorsal view); k. Posterolateral seta on ventral side of paraglossa." figureDoi="10.3897/dez.68.59462.figure6" httpUri="https://binary.pensoft.net/fig/494107" pageId="0" pageNumber="1">, 6</figureCitation>
<figureCitation id="F1B6876B8E84206EB023439CF06D221E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Philibaetis realonae comb. nov., larva morphology: a. Foreleg; b. Seta on dorsal margin of femur; c. Seta on dorsal margin of tibia; d. Claw; e. Apex of femur (posterior view); f. Base of antenna; g. Paraproct; h. Patch of notched scales; i. Tergite IV." figureDoi="10.3897/dez.68.59462.figure7" httpUri="https://binary.pensoft.net/fig/494108" pageId="0" pageNumber="1">, 7</figureCitation>
<figureCitation id="6FA3C3AC991781458018434620C3B48F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Philibaetis realonae comb. nov., larva morphology: a. Gill IV; b. Gills I-VII; c. Head." figureDoi="10.3897/dez.68.59462.figure8" httpUri="https://binary.pensoft.net/fig/494109" pageId="0" pageNumber="1">, 8</figureCitation>
<figureCitation id="75EC13AF41E25BFF1C967600CC664DDC" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Philibaetis realonae comb. nov., habitus, larva: a. Dorsal view; b. Lateral view; c. Ventral view." figureDoi="10.3897/dez.68.59462.figure9" httpUri="https://binary.pensoft.net/fig/494110" pageId="0" pageNumber="1">, 9</figureCitation>
<figureCitation id="68E1A9562620F4D23EBC226D8B8595E2" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Protogonostyli developing under cuticle of male last instar larvae: a. Philibaetis luzonensis comb. nov.; b. Philibaetis realonae comb. nov.; c. Labiobaetis type of folding, Labiobaetis pelingeni Kaltenbach, Garces &amp; Gattolliat, 2020; d. Baetis type of folding, Baetis rhodani (Pictet, 1843)." figureDoi="10.3897/dez.68.59462.figure10" httpUri="https://binary.pensoft.net/fig/494111" pageId="0" pageNumber="1">, 10</figureCitation>
<figureCitation id="1AAB44B6EDB3795FCF17CE506D10439B" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Claws and subapical setae, SEM photos: a-c. Philibaetis luzonensis comb. nov., a. Fore claw, anterior view; b. Middle claw, posterior view; c. Hind claw, posterior view; d, e. Philibaetis realonae comb. nov., d. Fore claw, anterior view; e. Middle claw, posterior view." figureDoi="10.3897/dez.68.59462.figure11" httpUri="https://binary.pensoft.net/fig/494112" pageId="0" pageNumber="1">, 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BD5C389EF9FFF87C7A6308C528A7F6E4" pageId="0" pageNumber="1" type="type species">
<paragraph id="2275D631385F8673C0B144224D038921" pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph id="DFC55DA0DE7043D963F27442132B927B" pageId="0" pageNumber="1">
<taxonomicName id="68BF6811228233EC8904706FB8D131D0" authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2021" baseAuthorityName="Mueller-Liebenau" baseAuthorityYear="1982" class="Insecta" family="Baetidae" genus="Philibaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philibaetis luzonensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="luzonensis">
<emphasis id="3DE889569CBEE654EB58510B691E1EB6" italics="true" pageId="0" pageNumber="1">Philibaetis luzonensis</emphasis>
</taxonomicName>
(
<normalizedToken id="9BE385B3BA221B856BFF6BFA779A06AF" originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
, 1982) comb. nov., by present designation.
</paragraph>
</subSubSection>
<subSubSection id="67CEBB508FECEB9E1D9C5ACDD2D2CFDE" pageId="0" pageNumber="1" type="species included">
<paragraph id="BBE1D29DD14B63C20841FF5B942D3328" pageId="0" pageNumber="1">Species included.</paragraph>
<paragraph id="6CD14EA89031AA66350156EEC8932C3B" pageId="0" pageNumber="1">
<taxonomicName id="B006785EC3A48D227DC0024869FF91A0" authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2021" baseAuthorityName="Mueller-Liebenau" baseAuthorityYear="1982" class="Insecta" family="Baetidae" genus="Philibaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philibaetis realonae" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="realonae">
<emphasis id="8CF930E223CE2A212EECC02095A61E61" italics="true" pageId="0" pageNumber="1">Philibaetis realonae</emphasis>
</taxonomicName>
(
<normalizedToken id="A2D44DE4FEF7D052B904F36F75BA96FB" originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
, 1982) comb. nov.
</paragraph>
</subSubSection>
<subSubSection id="57BF53C530E154376B2023FCF18BEAA5" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="9FF9A07EB52C04EE3E5DCF0CC2F202E4" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="907CD1E69DAFE276E7CE3D19881E3C01" pageId="0" pageNumber="1">
<emphasis id="5AFAA00857518CB27B0020CEE912D560" bold="true" pageId="0" pageNumber="1">Larva.</emphasis>
This new genus is distinguished by a combination of the following characters: A) body elongate and slender, head clearly hypognathous (Fig.
<figureCitation id="4C329C128A775C6B7E297756C43A10F4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Philibaetis luzonensis comb. nov., habitus, larva: a. Dorsal view; b. Lateral view; c. Ventral view." figureDoi="10.3897/dez.68.59462.figure5" httpUri="https://binary.pensoft.net/fig/494106" pageId="0" pageNumber="1">5</figureCitation>
); B) bases of antennae not close to each other, without carina between them (Figs
<figureCitation id="BA3B6D77646CCEDAF95E7FAE62571224" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Philibaetis luzonensis comb. nov., habitus, larva: a. Dorsal view; b. Lateral view; c. Ventral view." figureDoi="10.3897/dez.68.59462.figure5" httpUri="https://binary.pensoft.net/fig/494106" pageId="0" pageNumber="1">5c</figureCitation>
,
<figureCitation id="FDF2C0A7EFFB975B23457618E6E9DA25" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Philibaetis realonae comb. nov., larva morphology: a. Gill IV; b. Gills I-VII; c. Head." figureDoi="10.3897/dez.68.59462.figure8" httpUri="https://binary.pensoft.net/fig/494109" pageId="0" pageNumber="1">8c</figureCitation>
); C) labrum rectangular, on dorsal surface with submarginal row of long, simple setae (Fig.
<figureCitation id="D042972AD24955239D11B5FC60322337" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Philibaetis luzonensis comb. nov., larva morphology: a. Labrum, dorsal view; b. Labrum, ventral view; c. Seta on anterolateral margin; d. Medial setae; e. Labrum, ventral view, submarginal row of setae; f. Setae of submarginal row; g. Right mandible; h. Right prostheca; i. Left mandible; j. Left prostheca; k. Tuft of setae; l. Right incisor, kinetodontium and prostheca; m. Left incisor, kinetodontium and prostheca." figureDoi="10.3897/dez.68.59462.figure1" httpUri="https://binary.pensoft.net/fig/494102" pageId="0" pageNumber="1">1a</figureCitation>
); ventrally on lateral margins long, simple, spine-like setae, on anterolateral margins long, feathered setae and medially long, bifid setae and a partial, submarginal row of lanceolate setae (Figs
<figureCitation id="0835181C9F340C18D7D607F65C29D186" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Philibaetis luzonensis comb. nov., larva morphology: a. Labrum, dorsal view; b. Labrum, ventral view; c. Seta on anterolateral margin; d. Medial setae; e. Labrum, ventral view, submarginal row of setae; f. Setae of submarginal row; g. Right mandible; h. Right prostheca; i. Left mandible; j. Left prostheca; k. Tuft of setae; l. Right incisor, kinetodontium and prostheca; m. Left incisor, kinetodontium and prostheca." figureDoi="10.3897/dez.68.59462.figure1" httpUri="https://binary.pensoft.net/fig/494102" pageId="0" pageNumber="1">1b-f</figureCitation>
,
<figureCitation id="A30317655EB1891CAD8B2CF472140DA7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Philibaetis realonae comb. nov., larva morphology: a. Labrum; b. Setae of submarginal row (ventral view); c. Right mandible; d. Right prostheca; e. Left mandible; f. Left prostheca; g. Tuft of setae; h. Hypopharynx and superlinguae; i. Maxilla; j. Labium (left side: ventral view; right side: dorsal view); k. Posterolateral seta on ventral side of paraglossa." figureDoi="10.3897/dez.68.59462.figure6" httpUri="https://binary.pensoft.net/fig/494107" pageId="0" pageNumber="1">6a, b</figureCitation>
); D) right mandible with blade-like incisor, stick-like, apically denticulate prostheca and dorsally mediolateral patch of long, spine-like setae (Fig.
<figureCitation id="338A4923CAEFCB67704B2A8AC299E1EF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Philibaetis luzonensis comb. nov., larva morphology: a. Labrum, dorsal view; b. Labrum, ventral view; c. Seta on anterolateral margin; d. Medial setae; e. Labrum, ventral view, submarginal row of setae; f. Setae of submarginal row; g. Right mandible; h. Right prostheca; i. Left mandible; j. Left prostheca; k. Tuft of setae; l. Right incisor, kinetodontium and prostheca; m. Left incisor, kinetodontium and prostheca." figureDoi="10.3897/dez.68.59462.figure1" httpUri="https://binary.pensoft.net/fig/494102" pageId="0" pageNumber="1">1g, h, l</figureCitation>
); E) left mandible with blade-like incisor, slender prostheca apically denticulate and with comb-shaped structure; tuft of long, partly-branched setae at base of subtriangular process; dorsally mediolateral patch of long, spine-like setae (Fig.
<figureCitation id="52F17E29454365D6B2AB77ED0A726112" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Philibaetis luzonensis comb. nov., larva morphology: a. Labrum, dorsal view; b. Labrum, ventral view; c. Seta on anterolateral margin; d. Medial setae; e. Labrum, ventral view, submarginal row of setae; f. Setae of submarginal row; g. Right mandible; h. Right prostheca; i. Left mandible; j. Left prostheca; k. Tuft of setae; l. Right incisor, kinetodontium and prostheca; m. Left incisor, kinetodontium and prostheca." figureDoi="10.3897/dez.68.59462.figure1" httpUri="https://binary.pensoft.net/fig/494102" pageId="0" pageNumber="1">1i-k, m</figureCitation>
); F) hypopharynx with a medial tuft of stout setae and anterolaterally two smaller tufts of stout setae (Fig.
<figureCitation id="704C557D7E188C32D42AEE3C861BF1D4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Philibaetis luzonensis comb. nov., larva morphology: a. Hypopharynx and superlinguae; b. Maxilla; c. Distal and middle denti-setae; d. Labium (left side: ventral view; right side: dorsal view); e. Posterolateral seta on ventral side of paraglossa." figureDoi="10.3897/dez.68.59462.figure2" httpUri="https://binary.pensoft.net/fig/494103" pageId="0" pageNumber="1">2a</figureCitation>
); G) galea-lacinia with three denti-setae, distal denti-seta tooth-like and directed against canines, middle and proximal denti-setae slender, bifid and pectinate; maxillary palp 2-segmented, apex of segment II constricted (Fig.
<figureCitation id="69978643526C29E3ED2049FD8F3158BD" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Philibaetis luzonensis comb. nov., larva morphology: a. Hypopharynx and superlinguae; b. Maxilla; c. Distal and middle denti-setae; d. Labium (left side: ventral view; right side: dorsal view); e. Posterolateral seta on ventral side of paraglossa." figureDoi="10.3897/dez.68.59462.figure2" httpUri="https://binary.pensoft.net/fig/494103" pageId="0" pageNumber="1">2b</figureCitation>
); H) glossae basally broad, narrowing towards apex, shorter than paraglossae, apically and on both margins with long, spine-like setae; paraglossae truncate in distal part, one stout, bifurcate seta in posterolateral area; labial palps segment II without protuberance (Fig.
<figureCitation id="AC5844816F9CF0910DAF1F008E467E08" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Philibaetis luzonensis comb. nov., larva morphology: a. Hypopharynx and superlinguae; b. Maxilla; c. Distal and middle denti-setae; d. Labium (left side: ventral view; right side: dorsal view); e. Posterolateral seta on ventral side of paraglossa." figureDoi="10.3897/dez.68.59462.figure2" httpUri="https://binary.pensoft.net/fig/494103" pageId="0" pageNumber="1">2d, e</figureCitation>
); I) fore femur apically with stout setae both on anterior and posterior side, femoral patch absent; claw robust and pointed, with one row of denticles and 2-3 long subapical setae on posterior side and one short, fine, subapical seta on anterior side (Figs
<figureCitation id="38B14A92E47EC6A3E38A5B0E966FBB4A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Philibaetis luzonensis comb. nov., larva morphology: a. Foreleg; b. Claw; c. Apex of femur (posterior view); d. Basal part of antenna." figureDoi="10.3897/dez.68.59462.figure3" httpUri="https://binary.pensoft.net/fig/494104" pageId="0" pageNumber="1">3a-c</figureCitation>
,
<figureCitation id="C5C9C05AE5025EA5C2B1D708C1C9B195" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Claws and subapical setae, SEM photos: a-c. Philibaetis luzonensis comb. nov., a. Fore claw, anterior view; b. Middle claw, posterior view; c. Hind claw, posterior view; d, e. Philibaetis realonae comb. nov., d. Fore claw, anterior view; e. Middle claw, posterior view." figureDoi="10.3897/dez.68.59462.figure11" httpUri="https://binary.pensoft.net/fig/494112" pageId="0" pageNumber="1">11</figureCitation>
); J) paraproct with patch of notched scales (Fig.
<figureCitation id="72C59076ABF609F3BD5F016C30B845F7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Philibaetis luzonensis comb. nov., larva morphology: a. Gill IV; b. Inner margin of gill IV; c. Outer margin of gill IV; d. Gills I-VII; e. Paraproct; f. Patch of notched scales; g. Metanotum; h. Tergite IV." figureDoi="10.3897/dez.68.59462.figure4" httpUri="https://binary.pensoft.net/fig/494105" pageId="0" pageNumber="1">4e, f</figureCitation>
); K) protogonostyli under larval cuticle folded in the
<taxonomicName id="664BDC833BDEBF22649784C46199B076" authorityName="Novikova &amp; Kluge" authorityYear="1987" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="E3B9BAF994920A4EEFCD60655FAD67AE" italics="true" pageId="0" pageNumber="1">Labiobaetis</emphasis>
</taxonomicName>
type (Fig.
<figureCitation id="844F4656BB544441A71D6F952799965F" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Protogonostyli developing under cuticle of male last instar larvae: a. Philibaetis luzonensis comb. nov.; b. Philibaetis realonae comb. nov.; c. Labiobaetis type of folding, Labiobaetis pelingeni Kaltenbach, Garces &amp; Gattolliat, 2020; d. Baetis type of folding, Baetis rhodani (Pictet, 1843)." figureDoi="10.3897/dez.68.59462.figure10" httpUri="https://binary.pensoft.net/fig/494111" pageId="0" pageNumber="1">10</figureCitation>
).
</paragraph>
<caption id="1B9940AF493650C6D0FC6419B495D326" doi="10.3897/dez.68.59462.figure1" httpUri="https://binary.pensoft.net/fig/494102" pageId="0" pageNumber="1" start="Figure 1" startId="F1">
<paragraph id="9D0456A5C1E2A6739168D962C94D5335" pageId="0" pageNumber="1">
<emphasis id="1A7ADEAE62D00A27991930B976B3D23F" bold="true" pageId="0" pageNumber="1">Figure 1.</emphasis>
<taxonomicName id="EC3E0342D6244B0C9576E4CC6D8E2178" authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2021" baseAuthorityName="Mueller-Liebenau" baseAuthorityYear="1982" class="Insecta" family="Baetidae" genus="Philibaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philibaetis luzonensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="luzonensis">
<emphasis id="715FCAB5E2F4EA85785A39A1B7861DB7" italics="true" pageId="0" pageNumber="1">Philibaetis luzonensis</emphasis>
</taxonomicName>
comb. nov., larva morphology:
<emphasis id="E7FA38704F768EDC36D101BCC8DD0BAF" bold="true" pageId="0" pageNumber="1">a.</emphasis>
Labrum, dorsal view;
<emphasis id="7FCA3B013476D931B9F45D0ECAE0565C" bold="true" pageId="0" pageNumber="1">b.</emphasis>
Labrum, ventral view;
<emphasis id="52A638C1D334D62C0519CEED85EE1C80" bold="true" pageId="0" pageNumber="1">c.</emphasis>
Seta on anterolateral margin;
<emphasis id="5B81AF6C2F5C1F81A33891F10A85D5DE" bold="true" pageId="0" pageNumber="1">d.</emphasis>
Medial setae;
<emphasis id="DF7214F2B864A25FC3883840B3855A21" bold="true" pageId="0" pageNumber="1">e.</emphasis>
Labrum, ventral view, submarginal row of setae;
<emphasis id="AB1D804F1B7B64EB9BB9A88EC0DE5444" bold="true" pageId="0" pageNumber="1">f.</emphasis>
Setae of submarginal row;
<emphasis id="94E54D626F66DF0752021CA92B1021AD" bold="true" pageId="0" pageNumber="1">g.</emphasis>
Right mandible;
<emphasis id="67E475BCEC9698A09FF0323F83DE96D2" bold="true" pageId="0" pageNumber="1">h.</emphasis>
Right prostheca;
<emphasis id="27D6CFC2AA0028BEAF63C9DB22D0E04B" bold="true" pageId="0" pageNumber="1">i.</emphasis>
Left mandible;
<emphasis id="B26E849CA5660202CB47431F4E6EDC8C" bold="true" pageId="0" pageNumber="1">j.</emphasis>
Left prostheca;
<emphasis id="1EB8D4C7B429B26A5DCBA0CE8D54D7D4" bold="true" pageId="0" pageNumber="1">k.</emphasis>
Tuft of setae;
<emphasis id="DF96B253D7D85D6E5E38422DCC7B88F8" bold="true" pageId="0" pageNumber="1">l.</emphasis>
Right incisor, kinetodontium and prostheca;
<emphasis id="9A66CF0DD7FB40D5C0B92ABC925F8B06" bold="true" pageId="0" pageNumber="1">m.</emphasis>
Left incisor, kinetodontium and prostheca.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="D7677C29F0F105DFB29C446408E75505" pageId="0" pageNumber="1" type="imagines">
<paragraph id="2D732034DDC479966DE6A38D4227DE46" pageId="0" pageNumber="1">Imagines.</paragraph>
<paragraph id="4F10175681540864374D97BFBE65B2F7" pageId="0" pageNumber="1">Unknown.</paragraph>
</subSubSection>
<subSubSection id="6B03892DF940024FA9E13EBDE4033A04" pageId="0" pageNumber="1" type="etymology">
<paragraph id="A6DC1ED0F3683E65CC05633CCC2F526E" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="16238ABEB4F32D428974ABB281763BA4" pageId="0" pageNumber="1">
<taxonomicName id="1AA52B36ACAE303870C99E8A8EA0C208" authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2021" class="Insecta" family="Baetidae" genus="Philibaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philibaetis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="752B83C6602CCBC3168280CB8FA8F182" italics="true" pageId="0" pageNumber="1">Philibaetis</emphasis>
</taxonomicName>
is an arbitrary combination of letters with allusion to the Philippines, where the genus occurs and to the genus
<taxonomicName id="625237D191F7C7E81996B1E3C7F31952" authorityName="Leach" authorityYear="1815" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="B884D65A282C22B8E9546256D11262A1" italics="true" pageId="0" pageNumber="1">Baetis</emphasis>
</taxonomicName>
. The latter is with reference to the superficial similarities between both genera and the fact that the species of
<taxonomicName id="52F0D5E3ADE02DECD930938309131BB5" authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2021" class="Insecta" family="Baetidae" genus="Philibaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philibaetis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="896F9693AE18DB654FC170D7ED643B8C" italics="true" pageId="0" pageNumber="1">Philibaetis</emphasis>
</taxonomicName>
gen. nov. were up to now assigned to
<taxonomicName id="7736EA7B21C189A75B9CDE44B323DFAC" authorityName="Leach" authorityYear="1815" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="B1DF2F2D68EF391C95DA5663FBF689E1" italics="true" pageId="0" pageNumber="1">Baetis</emphasis>
</taxonomicName>
. Gender is masculine.
</paragraph>
<caption id="70CC0DB9C02DE090F3E6EC6899C3569D" doi="10.3897/dez.68.59462.figure2" httpUri="https://binary.pensoft.net/fig/494103" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
<paragraph id="2D6E91E400F28BBF374D04A8B55D3E94" pageId="0" pageNumber="1">
<emphasis id="272EA3AE063820D58140D86D409A7189" bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
<taxonomicName id="EAD36AA8331429F71A604D5E52BE3558" authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2021" baseAuthorityName="Mueller-Liebenau" baseAuthorityYear="1982" class="Insecta" family="Baetidae" genus="Philibaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philibaetis luzonensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="luzonensis">
<emphasis id="E119A7F43EE92E4C5312135DB73ED053" italics="true" pageId="0" pageNumber="1">Philibaetis luzonensis</emphasis>
</taxonomicName>
comb. nov., larva morphology:
<emphasis id="EC2BA51451599DA3EA2044D97E60C744" bold="true" pageId="0" pageNumber="1">a.</emphasis>
<taxonomicName id="7386540E000E4DB569D4B506E1F06EF9" genus="Hypopharynx" lsidName="Hypopharynx" pageId="0" pageNumber="1" rank="genus">Hypopharynx</taxonomicName>
and superlinguae;
<emphasis id="140CF204FDEF5C916CB8DCC62DE8A36A" bold="true" pageId="0" pageNumber="1">b.</emphasis>
Maxilla;
<emphasis id="6CD21A6AC7FA6E3C046482D1479A5D8F" bold="true" pageId="0" pageNumber="1">c.</emphasis>
Distal and middle denti-setae;
<emphasis id="66B86B6259488C3EF7484892F25344EE" bold="true" pageId="0" pageNumber="1">d.</emphasis>
Labium (left side: ventral view; right side: dorsal view);
<emphasis id="DFD82EC938DC647E1B7566E67FCADFEA" bold="true" pageId="0" pageNumber="1">e.</emphasis>
Posterolateral seta on ventral side of paraglossa.
</paragraph>
</caption>
<caption id="28B3ACF0FE004BEAEEA54904F31B097E" doi="10.3897/dez.68.59462.figure3" httpUri="https://binary.pensoft.net/fig/494104" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph id="A6A068527F6236BCC55B6E15BD670C58" pageId="0" pageNumber="1">
<emphasis id="855859D5F0A9480FEDAF162E93B63D8E" bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
<taxonomicName id="11272ACFD90D2DBF8A0CF4352B2F14CD" authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2021" baseAuthorityName="Mueller-Liebenau" baseAuthorityYear="1982" class="Insecta" family="Baetidae" genus="Philibaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philibaetis luzonensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="luzonensis">
<emphasis id="7DB11E444E82040768F6745C17659FB1" italics="true" pageId="0" pageNumber="1">Philibaetis luzonensis</emphasis>
</taxonomicName>
comb. nov., larva morphology:
<emphasis id="0889E2F961258CA93AED4E92E9901237" bold="true" pageId="0" pageNumber="1">a.</emphasis>
Foreleg;
<emphasis id="9219B63FE296590C1FC869CA01374855" bold="true" pageId="0" pageNumber="1">b.</emphasis>
Claw;
<emphasis id="5C21592E92F7A8467581D07484DCA5DC" bold="true" pageId="0" pageNumber="1">c.</emphasis>
Apex of femur (posterior view);
<emphasis id="D77EA50583F930CB1CC176D0DFB03368" bold="true" pageId="0" pageNumber="1">d.</emphasis>
Basal part of antenna.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="9A19BDBFBA76F609512A830D058A670B" pageId="0" pageNumber="1" type="description">
<paragraph id="B30A3DE0CD46100E871165D9305A9D30" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="2437F4B692FC7D61141E7C87F1D7187C" pageId="0" pageNumber="1">
<emphasis id="12595A32C5BAC20710E3467B72C6237F" bold="true" pageId="0" pageNumber="1">Larva.</emphasis>
(Figs
<figureCitation id="A89E728A6AB4A031D1C402CBC37CCC15" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Philibaetis luzonensis comb. nov., larva morphology: a. Labrum, dorsal view; b. Labrum, ventral view; c. Seta on anterolateral margin; d. Medial setae; e. Labrum, ventral view, submarginal row of setae; f. Setae of submarginal row; g. Right mandible; h. Right prostheca; i. Left mandible; j. Left prostheca; k. Tuft of setae; l. Right incisor, kinetodontium and prostheca; m. Left incisor, kinetodontium and prostheca." figureDoi="10.3897/dez.68.59462.figure1" httpUri="https://binary.pensoft.net/fig/494102" pageId="0" pageNumber="1">1</figureCitation>
-
<figureCitation id="615C1B37390AA80B0DBF7E343379F5C7" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Claws and subapical setae, SEM photos: a-c. Philibaetis luzonensis comb. nov., a. Fore claw, anterior view; b. Middle claw, posterior view; c. Hind claw, posterior view; d, e. Philibaetis realonae comb. nov., d. Fore claw, anterior view; e. Middle claw, posterior view." figureDoi="10.3897/dez.68.59462.figure11" httpUri="https://binary.pensoft.net/fig/494112" pageId="0" pageNumber="1">11</figureCitation>
).
</paragraph>
<paragraph id="617FA2203213B5A049E4F4D3C04F2D95" pageId="0" pageNumber="1">
<emphasis id="D8AF16F21B4AB4E2688C2E54CE1CBEED" bold="true" pageId="0" pageNumber="1">
<emphasis id="E452FDF4906A2D314BAD666B865206CB" italics="true" pageId="0" pageNumber="1">Body</emphasis>
.
</emphasis>
Elongate and slender, head hypognathous (Fig.
<figureCitation id="47EEEADDF27526F651CE26DD6B9EE72C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Philibaetis luzonensis comb. nov., habitus, larva: a. Dorsal view; b. Lateral view; c. Ventral view." figureDoi="10.3897/dez.68.59462.figure5" httpUri="https://binary.pensoft.net/fig/494106" pageId="0" pageNumber="1">5</figureCitation>
).
</paragraph>
<paragraph id="698B117E8EA7F73B1577F36AEC1C8A69" pageId="0" pageNumber="1">
<emphasis id="65120940F69F5E4A3F2C8B52FD2E5B13" bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
<emphasis id="E9D0D645D9C36B4E15368E7A351F448F" italics="true" pageId="0" pageNumber="1">Antenna</emphasis>
(Figs
<figureCitation id="0DEF2C3759F648355615D69CE58F3F03" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Philibaetis luzonensis comb. nov., larva morphology: a. Foreleg; b. Claw; c. Apex of femur (posterior view); d. Basal part of antenna." figureDoi="10.3897/dez.68.59462.figure3" httpUri="https://binary.pensoft.net/fig/494104" pageId="0" pageNumber="1">3d</figureCitation>
,
<figureCitation id="07FD69B9683C33849475EA1FBEB16310" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Philibaetis luzonensis comb. nov., habitus, larva: a. Dorsal view; b. Lateral view; c. Ventral view." figureDoi="10.3897/dez.68.59462.figure5" httpUri="https://binary.pensoft.net/fig/494106" pageId="0" pageNumber="1">5c</figureCitation>
,
<figureCitation id="DDBA428C6B7F2E91D9E925F916A02335" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Philibaetis realonae comb. nov., larva morphology: a. Gill IV; b. Gills I-VII; c. Head." figureDoi="10.3897/dez.68.59462.figure8" httpUri="https://binary.pensoft.net/fig/494109" pageId="0" pageNumber="1">8c</figureCitation>
). More than 2
<normalizedToken id="BB5973A683313B1AD8E815A8374BD152" originalValue="×">x</normalizedToken>
longer than head length, bases of antennae not close to each other, without carina between them (Figs
<figureCitation id="FDF58A8772DB48F762B8F7B635BCEFBB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Philibaetis luzonensis comb. nov., habitus, larva: a. Dorsal view; b. Lateral view; c. Ventral view." figureDoi="10.3897/dez.68.59462.figure5" httpUri="https://binary.pensoft.net/fig/494106" pageId="0" pageNumber="1">5c</figureCitation>
,
<figureCitation id="FC3A5B14A64E2C72F8A0FDA8C1DA6BEB" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Philibaetis realonae comb. nov., larva morphology: a. Gill IV; b. Gills I-VII; c. Head." figureDoi="10.3897/dez.68.59462.figure8" httpUri="https://binary.pensoft.net/fig/494109" pageId="0" pageNumber="1">8c</figureCitation>
).
</paragraph>
<paragraph id="D408B8E3B3ABD18949AD8F049F3B3AB7" pageId="0" pageNumber="1">
<emphasis id="B4C7280DA94C7D82DD8674CB0ED0EEAD" italics="true" pageId="0" pageNumber="1">Labrum</emphasis>
(Figs
<figureCitation id="8BE65C2B9E4F1B1483B30FB063C56E8C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Philibaetis luzonensis comb. nov., larva morphology: a. Labrum, dorsal view; b. Labrum, ventral view; c. Seta on anterolateral margin; d. Medial setae; e. Labrum, ventral view, submarginal row of setae; f. Setae of submarginal row; g. Right mandible; h. Right prostheca; i. Left mandible; j. Left prostheca; k. Tuft of setae; l. Right incisor, kinetodontium and prostheca; m. Left incisor, kinetodontium and prostheca." figureDoi="10.3897/dez.68.59462.figure1" httpUri="https://binary.pensoft.net/fig/494102" pageId="0" pageNumber="1">1a-f</figureCitation>
,
<figureCitation id="FEE39944FE54D8D315B5C25E446C4CE7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Philibaetis realonae comb. nov., larva morphology: a. Labrum; b. Setae of submarginal row (ventral view); c. Right mandible; d. Right prostheca; e. Left mandible; f. Left prostheca; g. Tuft of setae; h. Hypopharynx and superlinguae; i. Maxilla; j. Labium (left side: ventral view; right side: dorsal view); k. Posterolateral seta on ventral side of paraglossa." figureDoi="10.3897/dez.68.59462.figure6" httpUri="https://binary.pensoft.net/fig/494107" pageId="0" pageNumber="1">6a, b</figureCitation>
). Rectangular, wider than long; on dorsal surface with submarginal row of long, simple setae; ventrally on lateral margin long, simple, spine-like setae, on anterolateral margin long, feathered setae and medially long, bifid setae and a partial, submarginal row of lanceolate setae.
</paragraph>
<paragraph id="4B18697B4D07B95C2FFF3AD00CE0268F" pageId="0" pageNumber="1">
<emphasis id="8D9502E3BBF36A9E152D209C941918C7" italics="true" pageId="0" pageNumber="1">Right mandible</emphasis>
(Fig.
<figureCitation id="497117F4FB5F4ADE46F5C164463CC56F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Philibaetis luzonensis comb. nov., larva morphology: a. Labrum, dorsal view; b. Labrum, ventral view; c. Seta on anterolateral margin; d. Medial setae; e. Labrum, ventral view, submarginal row of setae; f. Setae of submarginal row; g. Right mandible; h. Right prostheca; i. Left mandible; j. Left prostheca; k. Tuft of setae; l. Right incisor, kinetodontium and prostheca; m. Left incisor, kinetodontium and prostheca." figureDoi="10.3897/dez.68.59462.figure1" httpUri="https://binary.pensoft.net/fig/494102" pageId="0" pageNumber="1">1g, h, l</figureCitation>
). Incisor and kinetodontium fused, incisor blade-like, kinetodontium with denticles; inner margin of innermost denticle of kinetodontium without row of thin setae; prostheca stick-like, apically denticulate; dorsally with mediolateral patch of long, spine-like setae.
</paragraph>
<paragraph id="9F3CD46E4541FB5280BC8FD1415328AD" pageId="0" pageNumber="1">
<emphasis id="8C9177BABF794166A7D0ACC6E3D5B8C9" italics="true" pageId="0" pageNumber="1">Left mandible</emphasis>
(Fig.
<figureCitation id="B6F05502F8E34102F5A23DAD0693E62D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Philibaetis luzonensis comb. nov., larva morphology: a. Labrum, dorsal view; b. Labrum, ventral view; c. Seta on anterolateral margin; d. Medial setae; e. Labrum, ventral view, submarginal row of setae; f. Setae of submarginal row; g. Right mandible; h. Right prostheca; i. Left mandible; j. Left prostheca; k. Tuft of setae; l. Right incisor, kinetodontium and prostheca; m. Left incisor, kinetodontium and prostheca." figureDoi="10.3897/dez.68.59462.figure1" httpUri="https://binary.pensoft.net/fig/494102" pageId="0" pageNumber="1">1i-k, m</figureCitation>
). Incisor and kinetodontium fused, incisor blade-like, kinetodontium with denticles; prostheca apically denticulate and with comb-shaped structure; with tuft of long, partly-branched setae at base of subtriangular process; dorsally with mediolateral patch of long, spine-like setae.
</paragraph>
<paragraph id="B95D00A572FE4B83188C0613EDF416D8" pageId="0" pageNumber="1">
<taxonomicName id="848A6DA2FCBA3BCD510F6183F4144338" genus="Hypopharynx" lsidName="Hypopharynx" pageId="0" pageNumber="1" rank="genus">
<emphasis id="A86419B6E02B981947FC95FBD8A5A5A6" italics="true" pageId="0" pageNumber="1">Hypopharynx</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="8B1D79D16BD08AFD6A9B32B227B487BC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Philibaetis luzonensis comb. nov., larva morphology: a. Hypopharynx and superlinguae; b. Maxilla; c. Distal and middle denti-setae; d. Labium (left side: ventral view; right side: dorsal view); e. Posterolateral seta on ventral side of paraglossa." figureDoi="10.3897/dez.68.59462.figure2" httpUri="https://binary.pensoft.net/fig/494103" pageId="0" pageNumber="1">2a</figureCitation>
). With medial tuft of stout setae and anterolaterally two smaller tufts of stout setae.
</paragraph>
<paragraph id="0B3A0FC6A83F7B9DA5A99A6A0AD9039F" pageId="0" pageNumber="1">
<emphasis id="9BFD544A959AD1D64C840EEAFFFDCE53" italics="true" pageId="0" pageNumber="1">Maxilla</emphasis>
(Fig.
<figureCitation id="99292DE12F5DA9CF4FFF53D2D3B0314D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Philibaetis luzonensis comb. nov., larva morphology: a. Hypopharynx and superlinguae; b. Maxilla; c. Distal and middle denti-setae; d. Labium (left side: ventral view; right side: dorsal view); e. Posterolateral seta on ventral side of paraglossa." figureDoi="10.3897/dez.68.59462.figure2" httpUri="https://binary.pensoft.net/fig/494103" pageId="0" pageNumber="1">2b, c</figureCitation>
). Apically with three stout canines and three denti-setae; distal denti-seta tooth-like, with opposite direction to canines, other denti-setae slender, bifid and pectinate; maxillary palp with two segments.
</paragraph>
<paragraph id="BB7FDFA2272CD7ECEF27424E675BE90C" pageId="0" pageNumber="1">
<emphasis id="A34F3DDB2609C1B7E1FB65E8262EBA8B" italics="true" pageId="0" pageNumber="1">Labium</emphasis>
(Fig.
<figureCitation id="69FB12215B7A14358E285D0DB3F51525" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Philibaetis luzonensis comb. nov., larva morphology: a. Hypopharynx and superlinguae; b. Maxilla; c. Distal and middle denti-setae; d. Labium (left side: ventral view; right side: dorsal view); e. Posterolateral seta on ventral side of paraglossa." figureDoi="10.3897/dez.68.59462.figure2" httpUri="https://binary.pensoft.net/fig/494103" pageId="0" pageNumber="1">2d, e</figureCitation>
). Glossae basally broad, narrowing towards apex, shorter than paraglossae; apically and on both margins with long, spine-like setae; paraglossae truncate in distal part, on apex with three rows of long, robust, distally-pectinate setae and one stout, bifurcate seta in posterolateral area; labial palps segment II without protuberance, dorsally with row of spine-like setae near outer lateral margin.
</paragraph>
<paragraph id="C0A3BE1D590F80F3BD187727FA6DE5D9" pageId="0" pageNumber="1">
<emphasis id="F6610BA4C721BC7122EF9D79EC01A630" bold="true" pageId="0" pageNumber="1">
<emphasis id="5CD49FCF95B4F1C1A63C4F298D0EFE37" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
.
</emphasis>
<emphasis id="E5DBB63FEAD5C06CC0735C1A170C064F" italics="true" pageId="0" pageNumber="1">Hind protoptera</emphasis>
(Fig.
<figureCitation id="81C6A0B135DDA3087F881591534C52C7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Philibaetis luzonensis comb. nov., larva morphology: a. Gill IV; b. Inner margin of gill IV; c. Outer margin of gill IV; d. Gills I-VII; e. Paraproct; f. Patch of notched scales; g. Metanotum; h. Tergite IV." figureDoi="10.3897/dez.68.59462.figure4" httpUri="https://binary.pensoft.net/fig/494105" pageId="0" pageNumber="1">4g</figureCitation>
). Absent.
</paragraph>
<caption id="3B930722344C2FAE40DC626E556DA706" doi="10.3897/dez.68.59462.figure4" httpUri="https://binary.pensoft.net/fig/494105" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph id="B962068C7D754E09A93BDEB59D18787E" pageId="0" pageNumber="1">
<emphasis id="9A6E30AE9EC0BB2838AF3741F4A049FC" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
<taxonomicName id="B793670863ABF262F8AA13DA208D5FB8" authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2021" baseAuthorityName="Mueller-Liebenau" baseAuthorityYear="1982" class="Insecta" family="Baetidae" genus="Philibaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philibaetis luzonensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="luzonensis">
<emphasis id="4011B5621A0C92496829650B509C609B" italics="true" pageId="0" pageNumber="1">Philibaetis luzonensis</emphasis>
</taxonomicName>
comb. nov., larva morphology:
<emphasis id="9EE13B8BCF84568D6EF42B02362AAEE7" bold="true" pageId="0" pageNumber="1">a.</emphasis>
Gill IV;
<emphasis id="FE2ADD837C01993CD90EFC87A9EE5F82" bold="true" pageId="0" pageNumber="1">b.</emphasis>
Inner margin of gill IV;
<emphasis id="3E763958C55C2D919C1FFCE98F78F7ED" bold="true" pageId="0" pageNumber="1">c.</emphasis>
Outer margin of gill IV;
<emphasis id="F4841A0A2D9DC3B6D72A922B6AF25033" bold="true" pageId="0" pageNumber="1">d.</emphasis>
Gills I-VII;
<emphasis id="EAE823A5499452E801965C7C0722E986" bold="true" pageId="0" pageNumber="1">e.</emphasis>
Paraproct;
<emphasis id="5A346CF25B71EB3E3232C583028A81DF" bold="true" pageId="0" pageNumber="1">f.</emphasis>
Patch of notched scales;
<emphasis id="6C9772D1EBA32453C600237527363C46" bold="true" pageId="0" pageNumber="1">g.</emphasis>
Metanotum;
<emphasis id="6BCD3E54C54991CE82F88F9AC494FE9F" bold="true" pageId="0" pageNumber="1">h.</emphasis>
Tergite IV.
</paragraph>
</caption>
<paragraph id="2499EDD461B1F03F03001B420CF342D3" pageId="0" pageNumber="1">
<emphasis id="7F46D09EFCB9EDD2ABB7C1304A909EA8" italics="true" pageId="0" pageNumber="1">Foreleg</emphasis>
(Fig.
<figureCitation id="08D1EBE941F58BC3B7DF4D079A5AB2BA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Philibaetis luzonensis comb. nov., larva morphology: a. Foreleg; b. Claw; c. Apex of femur (posterior view); d. Basal part of antenna." figureDoi="10.3897/dez.68.59462.figure3" httpUri="https://binary.pensoft.net/fig/494104" pageId="0" pageNumber="1">3a-c</figureCitation>
).
<emphasis id="2CB90F3BC0F916A9DF7D7552BCBF064A" italics="true" pageId="0" pageNumber="1">Femur</emphasis>
with dense row of curved, spine-like setae on dorsal margin, on apex short, stout setae on anterior and posterior side; femoral patch absent;
<emphasis id="2B9F956D1D3D9CE9FBFA900E772F92E2" italics="true" pageId="0" pageNumber="1">tibia</emphasis>
with patellotibial suture, dorsal margin with row of medium, clavate setae;
<emphasis id="6F308EEC3C589BFF9DDE9CEDF335680A" italics="true" pageId="0" pageNumber="1">claw</emphasis>
robust and pointed, with one row of denticles, 2-3 long subapical setae on posterior side and one short, thin subapical seta on anterior side.
</paragraph>
<paragraph id="B80458F28B9E5657E9F5DBB0824DDC84" pageId="0" pageNumber="1">
<emphasis id="C66A94D0D45FE4FB3BE9C84A2FC637C7" bold="true" pageId="0" pageNumber="1">
<emphasis id="8B4F0983FC42A142FCB221B0946F9BC1" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
.
</emphasis>
<emphasis id="A2B0392A7D91F441D35E938671D63BFD" italics="true" pageId="0" pageNumber="1">Gills</emphasis>
(Figs
<figureCitation id="D5525B21595BF2BAD70CB60BB54F9CCD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Philibaetis luzonensis comb. nov., larva morphology: a. Gill IV; b. Inner margin of gill IV; c. Outer margin of gill IV; d. Gills I-VII; e. Paraproct; f. Patch of notched scales; g. Metanotum; h. Tergite IV." figureDoi="10.3897/dez.68.59462.figure4" httpUri="https://binary.pensoft.net/fig/494105" pageId="0" pageNumber="1">4a-d</figureCitation>
). Seven pairs of gills on segments I-VII, dorsally orientated. Margin with minute denticles intercalating fine, simple setae.
</paragraph>
<paragraph id="A3A6FAD0C3D16DC0B12D5AFBA515AA12" pageId="0" pageNumber="1">
<emphasis id="6815B270D13C4949AF54893B2434F2A9" italics="true" pageId="0" pageNumber="1">Paraproct</emphasis>
(Figs
<figureCitation id="73225857F86DFB669C351C9B2C173ABF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Philibaetis luzonensis comb. nov., larva morphology: a. Gill IV; b. Inner margin of gill IV; c. Outer margin of gill IV; d. Gills I-VII; e. Paraproct; f. Patch of notched scales; g. Metanotum; h. Tergite IV." figureDoi="10.3897/dez.68.59462.figure4" httpUri="https://binary.pensoft.net/fig/494105" pageId="0" pageNumber="1">4e, f</figureCitation>
). Without prolongation at posterior margin, with stout, marginal spines; surface with patch of notched scales.
</paragraph>
<paragraph id="A0107C9855950FE6D795B9EED4DBE471" pageId="0" pageNumber="1">
<emphasis id="450943EE36CC6D908D4AAB4687D26388" italics="true" pageId="0" pageNumber="1">Caudal filaments</emphasis>
(Fig.
<figureCitation id="0DEA845DA1D3E33A1B98432F41CA0E40" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Philibaetis luzonensis comb. nov., habitus, larva: a. Dorsal view; b. Lateral view; c. Ventral view." figureDoi="10.3897/dez.68.59462.figure5" httpUri="https://binary.pensoft.net/fig/494106" pageId="0" pageNumber="1">5a, d</figureCitation>
). Inner margin of cerci with long, thin setae; paracercus bilaterally with long, thin setae.
</paragraph>
</subSubSection>
<subSubSection id="453186EE30BF3466B17A990B1D06224D" pageId="0" pageNumber="1" type="imago">
<paragraph id="4ABAF96CD9A6763DEDB2CC0BB505C2AB" pageId="0" pageNumber="1">Imago.</paragraph>
<paragraph id="7C3E6A1EF3780EF9CFA17D63587E07E6" pageId="0" pageNumber="1">Unknown.</paragraph>
</subSubSection>
<subSubSection id="175639087660C90C35109B8E8771A46F" pageId="0" pageNumber="1" type="distribution">
<paragraph id="F48E37AE16240D3E6A307C5DB3406EDE" pageId="0" pageNumber="1">Distribution</paragraph>
<paragraph id="6AE0D5B44C1B75A5024F04A51CE15ADD" pageId="0" pageNumber="1">
<emphasis id="28266BDF032741DC364833C457980B3C" bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation id="68167937CFFDFB8B8007A3650ED32D72" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Philibaetis gen. nov., distribution, Philippines." figureDoi="10.3897/dez.68.59462.figure12" httpUri="https://binary.pensoft.net/fig/494113" pageId="0" pageNumber="1">12</figureCitation>
).
</emphasis>
Philippines: Luzon.
</paragraph>
</subSubSection>
<subSubSection id="009DA9664D03C0CAC114732C680E405F" pageId="0" pageNumber="1" type="redescription">
<paragraph id="4A1EC092C7F768A2133179305FDE3F3F" pageId="0" pageNumber="1">Redescription</paragraph>
</subSubSection>
</treatment>
</document>