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<mods:title id="038175E75C2A0258E16869465DE16F9C">Review of the Nearctic species of Drusilla Leach (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<heading id="446881ACFF846638FF1F3339A6C2F896" bold="true" box="[151,655,1818,1844]" fontSize="11" level="1" pageId="1" pageNumber="46" reason="1">
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<emphasis id="2DEBEAD2FF846638FF1F3339A6C2F896" bold="true" box="[151,655,1818,1844]" pageId="1" pageNumber="46">
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1.
<taxonomicName id="D89F4D43FF846638FF3C3339A6C2F896" ID-CoL="37TBP" authority="Fabricius, 1787" authorityName="Fabricius" authorityYear="1787" box="[180,655,1818,1844]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="canaliculata">
<emphasis id="2DEBEAD2FF846638FF3C3339A5F9F896" bold="true" box="[180,436,1818,1844]" italics="true" pageId="1" pageNumber="46">Drusilla canaliculata</emphasis>
(
<bibRefCitation id="7B0E4B31FF846638FE4B3339A6CAF896" author="Fabricius" box="[451,647,1818,1844]" pageId="1" pageNumber="46" refString="Fabricius, J. C. (1787) Mantissa insectorum sistens eorum species nuper detectas adiectis characteribus genericis, differentiis specificis, emendationibus, observationibus. Tom. 1. Christ. Gottl. Proft., Hafnia [Copenhagen], xx + 348 pp." type="book" year="1787">Fabricius, 1787</bibRefCitation>
)
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</emphasis>
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</paragraph>
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<subSubSection id="5785654BFF846638FF1F3342A722F871" pageId="1" pageNumber="46" type="reference_group">
<paragraph id="1F2036C0FF846638FF1F3342A634F8D5" blockId="1.[151,879,1889,2004]" box="[151,633,1889,1911]" pageId="1" pageNumber="46">
<treatmentCitationGroup id="3F8F11EEFF846638FF1F3342A634F8D5" box="[151,633,1889,1911]" pageId="1" pageNumber="46">
<taxonomicName id="D89F4D43FF846638FF1F3342A638F8D5" authority="Fabricius, 1787: 221" authorityName="Fabricius" authorityPageNumber="221" authorityYear="1787" box="[151,629,1889,1911]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="2DEBEAD2FF846638FF1F3342A5D6F8D5" box="[151,411,1889,1911]" italics="true" pageId="1" pageNumber="46">Staphylinus canaliculatus</emphasis>
<treatmentCitation id="9E3E10D1FF846638FE293342A638F8D5" author="Fabricius" box="[417,629,1889,1911]" page="221" pageId="1" pageNumber="46" year="1787">
<bibRefCitation id="7B0E4B31FF846638FE293342A60EF8D5" author="Fabricius" box="[417,579,1889,1911]" pageId="1" pageNumber="46" refString="Fabricius, J. C. (1787) Mantissa insectorum sistens eorum species nuper detectas adiectis characteribus genericis, differentiis specificis, emendationibus, observationibus. Tom. 1. Christ. Gottl. Proft., Hafnia [Copenhagen], xx + 348 pp." type="book" year="1787">Fabricius, 1787</bibRefCitation>
: 221
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</taxonomicName>
.
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</paragraph>
<paragraph id="1F2036C0FF846638FF1F33A3A642F816" blockId="1.[151,879,1889,2004]" pageId="1" pageNumber="46">
<treatmentCitationGroup id="3F8F11EEFF846638FF1F33A3A642F816" pageId="1" pageNumber="46">
<taxonomicName id="D89F4D43FF846638FF1F33A3A5F6F834" ID-CoL="37TBP" authority="Leach" authorityName="Leach" box="[151,443,1920,1942]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="canaliculata">
<emphasis id="2DEBEAD2FF846638FF1F33A3A53DF834" box="[151,368,1920,1942]" italics="true" pageId="1" pageNumber="46">Drusilla canaliculata</emphasis>
: Leach
</taxonomicName>
<emphasis id="2DEBEAD2FF846638FE4A33A2A59BF834" box="[450,470,1921,1942]" italics="true" pageId="1" pageNumber="46">in</emphasis>
Samuelle, 1819: 177.
<taxonomicName id="D89F4D43FF846638FF1F33BCA646F816" ID-CoL="6DLZH" authority="Casey, 1906: 322" authorityName="Casey" authorityPageNumber="322" authorityYear="1906" box="[151,523,1950,1973]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="cavicollis">
<emphasis id="2DEBEAD2FF846638FF1F33BCA51EF817" box="[151,339,1951,1973]" italics="true" pageId="1" pageNumber="46">Drusilla cavicollis</emphasis>
<treatmentCitation id="9E3E10D1FF846638FED133BCA646F816" author="Casey" box="[345,523,1950,1972]" page="322" pageId="1" pageNumber="46" year="1906">
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: 322
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.
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</paragraph>
<paragraph id="1F2036C0FF846638FF1F339DA722F871" blockId="1.[151,879,1889,2004]" box="[151,879,1981,2004]" pageId="1" pageNumber="46">
See
<bibRefCitation id="7B0E4B31FF846638FF4A339DA52CF876" author="Gusarov" box="[194,353,1982,2004]" pageId="1" pageNumber="46" refString="Gusarov, V. I. (2003) Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa, 353, 1 - 134." type="book chapter" year="2003">Gusarov (2003)</bibRefCitation>
and
<bibRefCitation id="7B0E4B31FF846638FE1B339EA66AF871" author="Assing" box="[403,551,1981,2003]" pageId="1" pageNumber="46" refString="Assing, V. (2005 a) A revision of the Middle Asian species of Drusilla Leach (Coleoptera, Staphylinidae, Aleocharinae). Entomologische Blatter, 101, 43 - 56." type="journal article" year="2005" yearSuffix="a">Assing (2005a</bibRefCitation>
,
<bibRefCitation id="7B0E4B31FF846638FDBA339EA63EF871" author="Assing" box="[562,627,1981,2003]" pageId="1" pageNumber="46" refString="Assing, V. (2005 b) On the western Palaearctic species of Drusilla Leach, with special reference to the species of the eastern Mediterranean (Coleoptera: Staphylinidae, Aleocharinae). Koleopterologische Rundschau, 75, 111 - 149." type="journal article" year="2005" yearSuffix="b">2005b</bibRefCitation>
) for further synonymies.
</paragraph>
</subSubSection>
<subSubSection id="5785654BFF87663BFF1F34B4A573FE25" pageId="2" pageNumber="47" type="diagnosis">
<paragraph id="1F2036C0FF87663BFF1F34B4A111FF76" blockId="2.[151,1436,151,716]" pageId="2" pageNumber="47">
<emphasis id="2DEBEAD2FF87663BFF1F34B4A55CFF12" bold="true" box="[151,273,151,176]" pageId="2" pageNumber="47">Diagnosis.</emphasis>
<taxonomicName id="D89F4D43FF87663BFE9234BBA640FF12" box="[282,525,152,176]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="canaliculata">
<emphasis id="2DEBEAD2FF87663BFE9234BBA640FF12" box="[282,525,152,176]" italics="true" pageId="2" pageNumber="47">Drusilla canaliculata</emphasis>
</taxonomicName>
can be easily separated from
<taxonomicName id="D89F4D43FF87663BFCE134BAA7BAFF12" box="[873,1015,153,176]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="nearctica">
<emphasis id="2DEBEAD2FF87663BFCE134BAA7BAFF12" box="[873,1015,153,176]" italics="true" pageId="2" pageNumber="47">D. nearctica</emphasis>
</taxonomicName>
and
<taxonomicName id="D89F4D43FF87663BFBBA34BAA0DCFF12" box="[1074,1169,152,176]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="ashei">
<emphasis id="2DEBEAD2FF87663BFBBA34BAA0DCFF12" box="[1074,1169,152,176]" italics="true" pageId="2" pageNumber="47">D. ashei</emphasis>
</taxonomicName>
by its larger body size (significantly greater than
<quantity id="D8679B25FF87663BFE37349FA65BFF76" box="[447,534,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="47" unit="mm" value="3.6">3.6 mm</quantity>
) and the broadly emarginate apex of the eighth tergite with serrate margin.
</paragraph>
<paragraph id="1F2036C0FF87663BFF4F34FCA573FE25" blockId="2.[151,1436,151,716]" pageId="2" pageNumber="47">
See:
<bibRefCitation id="7B0E4B31FF87663BFE8834FCA59EFF55" author="Casey" box="[256,467,223,247]" pageId="2" pageNumber="47" refString="Casey, T. L. (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis, 16, 125 - 434." type="journal article" year="1906">Casey (1906: 322)</bibRefCitation>
and
<bibRefCitation id="7B0E4B31FF87663BFD8634FCA754FF55" author="Lohse" box="[526,793,223,247]" pageId="2" pageNumber="47" refString="Lohse, G. A. 1974. 23. Familie Staphylinidae II (Hypocyphtinae und Aleocharinae). Tribus 15 - 19 (Schistogenini - Aleocharinae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Ka? fer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 221 - 292." type="book chapter" year="1974">Lohse (1974: 222223)</bibRefCitation>
for descriptions;
<bibRefCitation id="7B0E4B31FF87663BFC6C34FCA08DFF55" author="Assing" box="[996,1216,223,247]" pageId="2" pageNumber="47" refString="Assing, V. (2006) On the Italian species of Drusilla Leach, 1819, with a note of D. taygetana Assing. Beitrage zur Entomologie, 56 (2), 281 - 296." type="journal article" year="2006">Assing (2006: 283)</bibRefCitation>
for habitus images (both sexes), apex of the eighth tergite (both sexes; females have a slightly more shallow emargination and lateral apices of emargination less produced), lateral view of the median lobe and spermatheca;
<bibRefCitation id="7B0E4B31FF87663BFB013504A13EFE9D" author="Gusarov" box="[1161,1395,295,319]" pageId="2" pageNumber="47" refString="Gusarov, V. I. (2003) Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa, 353, 1 - 134." type="book chapter" year="2003">Gusarov (2003: 112)</bibRefCitation>
for an illustration of the internal sac;
<bibRefCitation id="7B0E4B31FF87663BFD9F356EA6BAFEC6" author="Werner" box="[535,759,332,356]" pageId="2" pageNumber="47" refString="Werner, M., Gack, C., Speck, T. &amp; Peschke, K. (2007) Queue up, please! Spermathecal filling in the rove beetle Drusilla canaliculata (Coleoptera, Staphylinidae). Naturwissenschaften, 94, 837 - 841." type="journal article" year="2007">
Werner
<emphasis id="2DEBEAD2FF87663BFDF9356EA6E7FEC6" box="[625,682,332,356]" italics="true" pageId="2" pageNumber="47">et al.</emphasis>
(2007
</bibRefCitation>
: 838) for a SEM of the copulatory piece and mechanism of sperm transfer.
</paragraph>
</subSubSection>
<subSubSection id="5785654BFF87663BFF4F35B0A642FDBA" pageId="2" pageNumber="47" type="discussion">
<paragraph id="1F2036C0FF87663BFF4F35B0A642FDBA" blockId="2.[151,1436,151,716]" pageId="2" pageNumber="47">
<emphasis id="2DEBEAD2FF87663BFF4F35B0A501FE0E" bold="true" box="[199,332,403,428]" pageId="2" pageNumber="47">Discussion.</emphasis>
The
<typeStatus id="C0248862FF87663BFE0235B6A5F7FE0F" box="[394,442,405,429]" pageId="2" pageNumber="47">type</typeStatus>
material of
<taxonomicName id="D89F4D43FF87663BFDC235B7A064FE0E" authority="Fabricius, 1787" authorityName="Fabricius" authorityYear="1787" box="[586,1065,404,428]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="canaliculatus">
<emphasis id="2DEBEAD2FF87663BFDC235B7A721FE0E" box="[586,876,404,428]" italics="true" pageId="2" pageNumber="47">Staphylinus canaliculatus</emphasis>
<bibRefCitation id="7B0E4B31FF87663BFCFD35B7A064FE0E" author="Fabricius" box="[885,1065,404,428]" pageId="2" pageNumber="47" refString="Fabricius, J. C. (1787) Mantissa insectorum sistens eorum species nuper detectas adiectis characteribus genericis, differentiis specificis, emendationibus, observationibus. Tom. 1. Christ. Gottl. Proft., Hafnia [Copenhagen], xx + 348 pp." type="book" year="1787">Fabricius, 1787</bibRefCitation>
</taxonomicName>
, was not examined in this study because this is a widespread, well-documented species. Possibly due to a combination of wide distribution and aptery (
<bibRefCitation id="7B0E4B31FF87663BFF6335FFA5C1FE56" author="Assing" box="[235,396,476,500]" pageId="2" pageNumber="47" refString="Assing, V. (2005 b) On the western Palaearctic species of Drusilla Leach, with special reference to the species of the eastern Mediterranean (Coleoptera: Staphylinidae, Aleocharinae). Koleopterologische Rundschau, 75, 111 - 149." type="journal article" year="2005" yearSuffix="b">Assing 2005b</bibRefCitation>
), this species shows a range of subtle morphological variation and this may have contributed to the large number of synonyms.
</paragraph>
</subSubSection>
<subSubSection id="5785654BFF87663BFF4F3600A667FD6E" pageId="2" pageNumber="47" type="distribution">
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<bibRef id="C63FA236FF87663BFF4F3600A667FD6E" pageId="2" pageNumber="47" type="journal article">
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<title id="6D9BDDF6FF87663BFF4F3600A511FD9E" box="[199,348,547,572]" jopScore="0.002739726" pageId="2" pageNumber="47">Distribution</title>
.
</emphasis>
<author id="C722DE25FF87663BFEE23606A629FD9E" box="[362,612,548,572]" fns="I" npo="LF" pageId="2" pageNumber="47">
In
<editor id="D903DEC3FF87663BFE043607A629FD9E" box="[396,612,548,572]" pageId="2" pageNumber="47">
North
<collectingCountry id="67887650FF87663BFE5F3607A677FD9E" box="[471,570,548,572]" name="United States of America" pageId="2" pageNumber="47">America</collectingCountry>
,
<emphasis id="2DEBEAD2FF87663BFDC13606A629FD9E" box="[585,612,549,572]" italics="true" pageId="2" pageNumber="47">D.</emphasis>
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canaliculata is known mainly from the northeastern
<collectingCountry id="67887650FF87663BFB593607A124FD9E" box="[1233,1385,548,572]" name="United States of America" pageId="2" pageNumber="47">United States</collectingCountry>
and provinces of
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, including: Alaska (
<bibRefCitation id="7B0E4B31FF87663BFDE43664A6BBFDFD" author="Casey" box="[620,758,583,607]" pageId="2" pageNumber="47" refString="Casey, T. L. (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis, 16, 125 - 434." type="journal article" year="1906">Casey 1906</bibRefCitation>
; but most likely not established,
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), Kentucky (
<bibRefCitation id="7B0E4B31FF87663BFF16364FA50EFD26" author="Gusarov" box="[158,323,620,644]" pageId="2" pageNumber="47" refString="Gusarov, V. I. (2003) Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa, 353, 1 - 134." type="book chapter" year="2003">Gusarov 2003</bibRefCitation>
), Massachusetts (FMNH), New York (
<bibRefCitation id="7B0E4B31FF87663BFD71364FA7D9FD26" author="Seevers" box="[761,916,620,644]" pageId="2" pageNumber="47" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp." type="book" year="1978">Seevers 1978</bibRefCitation>
,
<bibRefCitation id="7B0E4B31FF87663BFC29364EA079FD26" author="Muona" box="[929,1076,620,644]" pageId="2" pageNumber="47" refString="Muona, J. (1984) Review of Palearctic Aleocharinae also occurring in North America (Coleoptera: Staphylinidae). Entomologica Scandinavica, 15, 227 - 231." type="journal article" year="1984">Muona 1984</bibRefCitation>
,
<bibRefCitation id="7B0E4B31FF87663BFBC8364FA0AEFD26" author="Gusarov" box="[1088,1251,620,644]" pageId="2" pageNumber="47" refString="Gusarov, V. I. (2003) Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa, 353, 1 - 134." type="book chapter" year="2003">Gusarov 2003</bibRefCitation>
; personal observations)
</volumeTitle>
, Pennsylvania (
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Seevers
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), Ontario (
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Gusarov
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),
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Prince Edward Island (
<bibRefCitation id="7B0E4B31FF87663BFB0236ACA1D1FD05" author="Majka" box="[1162,1436,655,679]" pageId="2" pageNumber="47" refString="Majka, C. G. &amp; Klimaszewski, J. (2008) Introduced Staphylinidae (Coleoptera) in the maritime provinces of Canada. Canadian Entomologist, 140, 48 - 72." type="journal article" year="2008">Majka &amp; Klimaszewski</bibRefCitation>
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2008), and Quebec (
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Gusarov
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).
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</paragraph>
</subSubSection>
</treatment>
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